248 lines
42 KiB
XML
248 lines
42 KiB
XML
<document id="AFB87517A5F3C6624494262AA710D39A" ID-CLB-Dataset="26919" ID-DOI="10.11646/zootaxa.4598.1.1" ID-GBIF-Dataset="475fd53e-c5c6-4a99-bb2e-cc2382f0171f" ID-ISSN="1175-5326" ID-Zenodo-Dep="2668063" ID-ZooBank="6BA5F086-1014-46B5-B6D5-77FE0CD07AEC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1557127411994" checkinUser="plazi" docAuthor="Seldon, David S. & Buckley, Thomas R." docDate="2019" docId="03BBD0260241B74890FBB693EFABFA3B" docLanguage="en" docName="zootaxa.4598.1.1.pdf.imf" docOrigin="Zootaxa 4598 (1)" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Mecodema manaia Seldon & Leschen 2011" docType="treatment" docVersion="12" lastPageNumber="71" masterDocId="FF82A85E0205B70E906CB267ED0BFFA4" masterDocTitle="The genus Mecodema Blanchard 1853 (Coleoptera: Carabidae: Broscini) from the North Island, New Zealand" masterLastPageNumber="148" masterPageNumber="1" pageNumber="69" updateTime="1698726038790" updateUser="ExternalLinkService">
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<mods:titleInfo id="C3265D016E7AD1B2F027ED8990791E6C">
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<mods:title id="FADC1DD773BD5AC437113B439A9F372D">The genus Mecodema Blanchard 1853 (Coleoptera: Carabidae: Broscini) from the North Island, New Zealand</mods:title>
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<mods:name id="8870D795546A22FB34605CBC8714D7E3" type="personal">
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<mods:namePart id="125E4A8DBEB73B143D067A78F63EB5CE">Seldon, David S.</mods:namePart>
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<mods:roleTerm id="CB55883493315ED683DCB089783A511F">Author</mods:roleTerm>
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<mods:namePart id="43743AFFB730EC4192C09E88B1DE1EA1">Buckley, Thomas R.</mods:namePart>
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<mods:typeOfResource id="AAD7F8699D772E8938549A01F251EBFC">text</mods:typeOfResource>
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<mods:title id="DDE104E06E8F6FB34A6C97C2BDCE76F4">Zootaxa</mods:title>
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<mods:part id="5B778399702EBA930E54C78A99520547">
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<mods:date id="5AB9F306ED404FCD08511DF835BE6A48">2019</mods:date>
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<mods:number id="CA57DAF0ACCEC4399F20E34C75DE584B">2019-05-06</mods:number>
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<mods:number id="8908F0261437501D7702924DE4E893EE">4598</mods:number>
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<mods:number id="9DE98D220AF7C7FE94E6AB60A975BF78">1</mods:number>
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<mods:classification id="AA6DC1BAD29BFB7B65C73095B8A8CCA9">journal article</mods:classification>
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<mods:identifier id="DF9D33CB53AC11B9DE4FDFB0235B5EBB" type="CLB-Dataset">26919</mods:identifier>
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<mods:identifier id="EC28B897AEBC62F1626CB556F82BD130" type="DOI">10.11646/zootaxa.4598.1.1</mods:identifier>
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<mods:identifier id="8BEB6770224BA5683BD3FC1A465BF47E" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03BBD0260241B74890FBB693EFABFA3B" ID-DOI="http://doi.org/10.5281/zenodo.5678398" ID-GBIF-Taxon="157062679" ID-Zenodo-Dep="5678398" LSID="urn:lsid:plazi:treatment:03BBD0260241B74890FBB693EFABFA3B" httpUri="http://treatment.plazi.org/id/03BBD0260241B74890FBB693EFABFA3B" lastPageId="70" lastPageNumber="71" pageId="68" pageNumber="69">
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<subSubSection id="C30832BB0241B74A90FBB693EFBDFAAA" box="[151,694,1268,1294]" pageId="68" pageNumber="69" type="nomenclature">
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<paragraph id="8BAD61300241B74A90FBB693EFBDFAAA" blockId="68.[151,694,1268,1327]" box="[151,694,1268,1294]" pageId="68" pageNumber="69">
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<heading id="D0E5D65C0241B74A90FBB693EFBDFAAA" bold="true" box="[151,694,1268,1294]" fontSize="11" level="1" pageId="68" pageNumber="69" reason="1">
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<emphasis id="B966BD220241B74A90FBB693EFBDFAAA" bold="true" box="[151,694,1268,1294]" pageId="68" pageNumber="69">
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<taxonomicName id="4C121AB30241B74A90FBB693EFBAFAAA" ID-CoL="3YKTG" authority="Seldon & Leschen, 2011" authorityName="Seldon & Leschen" authorityYear="2011" box="[151,689,1268,1294]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="manaia">
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<emphasis id="B966BD220241B74A90FBB693EC76FAAA" bold="true" box="[151,381,1268,1294]" italics="true" pageId="68" pageNumber="69">Mecodema manaia</emphasis>
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<bibRefCitation id="EF831CC10241B74A91E8B693EFBAFAAA" author="Seldon, D. S. & Leschen, R. A. B." box="[388,689,1268,1294]" pageId="68" pageNumber="69" pagination="1 - 45" refId="ref93265" refString="Seldon, D. S. & Leschen, R. A. B. (2011) Revision of the Mecodema curvidens species group (Coleoptera: Carabidae: Broscini). Zootaxa, 2829, 1 - 45. https: // doi. org / 10.11646 / zootaxa. 2829.1.1" type="journal article" year="2011">Seldon & Leschen, 2011</bibRefCitation>
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</taxonomicName>
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.
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30832BB0241B74A90FBB771ECFEFA8B" box="[151,501,1302,1327]" pageId="68" pageNumber="69" type="description">
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<paragraph id="8BAD61300241B74A90FBB771ECFEFA8B" blockId="68.[151,694,1268,1327]" box="[151,501,1302,1327]" pageId="68" pageNumber="69">
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<figureCitation id="13297DB50241B74A90FBB771EC2FFA8B" box="[151,292,1302,1327]" captionStart="FIGURE 12" captionStartId="18.[151,250,1905,1927]" captionTargetBox="[311,1244,697,1878]" captionTargetId="figure@18.[310,1250,689,1889]" captionTargetPageId="18" captionText="FIGURE 12. Ventral view of the mentum with a number of different character states. Illustrations A–D are the different forms of the indentation in the apex of the median process: (A) distinctly indentate (M. manaia); (B) moderately indentate (M. pluto); (C) slightly indentate (M. infimate); (D) indentation notched (M. parataiko). Illustrations E–G are the three general forms (i.e., hashed lines) of the mentum lobes: (E) triangular; (F) rounded; (G) squared." httpUri="https://zenodo.org/record/2668087/files/figure.png" pageId="68" pageNumber="69">Figures 12A</figureCitation>
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,
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<figureCitation id="13297DB50241B74A915CB771EC6AFA8B" box="[304,353,1302,1327]" captionStart="FIGURE 15" captionStartId="20.[151,250,1943,1965]" captionTargetBox="[162,1424,1039,1914]" captionTargetId="figure@20.[162,1424,1038,1921]" captionTargetPageId="20" captionText="FIGURE 15. The penis lobe in ventral view showing the relative deflection to right of vertical: (A) distinctly deflected (M. regulus); (B) moderately deflected (M. haunoho); (C) slightly deflected (M. oregoides). Scale bars = 1 mm. Structures of penis lobe indicated are: base (B); lateral edge (LE); apical portion (AP); ventral edge (ƑE)." httpUri="https://zenodo.org/record/2668093/files/figure.png" pageId="68" pageNumber="69">15B</figureCitation>
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,
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<figureCitation id="13297DB50241B74A9101B771EC90FA8B" box="[365,411,1302,1327]" captionStart="FIGURE 16" captionStartId="21.[151,250,897,919]" captionTargetBox="[151,1436,181,869]" captionTargetId="figure@21.[151,1436,181,876]" captionTargetPageId="21" captionText="FIGURE 16. Relative strength of the ventral curve over the length of the penis lobe (lateral view): (A) slightly curved (M. oblongum); (B) moderately curved (M. ponaiti); (C) distinctly curved (M. manaia). Scale bar = 1 mm." httpUri="https://zenodo.org/record/2668095/files/figure.png" pageId="68" pageNumber="69">16C</figureCitation>
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and
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<figureCitation id="13297DB50241B74A91BEB770ECFAFA8B" box="[466,497,1303,1327]" captionStart="FIGURE 37" captionStartId="69.[151,250,1852,1874]" captionTargetBox="[258,1330,199,1821]" captionTargetId="figure@69.[252,1336,193,1831]" captionTargetPageId="69" captionText="FIGURE 37. Mecodema manaia plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668137/files/figure.png" pageId="68" pageNumber="69">37</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30832BB0241B74A90FBB739E97AFA47" pageId="68" pageNumber="69" type="diagnosis">
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<paragraph id="8BAD61300241B74A90FBB739E97AFA47" blockId="68.[151,1437,1374,2011]" pageId="68" pageNumber="69">
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<emphasis id="B966BD220241B74A90FBB739EC18FAD3" bold="true" box="[151,275,1374,1399]" pageId="68" pageNumber="69">Diagnosis:</emphasis>
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Distinguished from other North Island
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<taxonomicName id="4C121AB30241B74A92B7B739EE5DFAD2" authorityName="Blanchard" authorityYear="1853" box="[731,854,1374,1398]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="genus">
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<emphasis id="B966BD220241B74A92B7B739EE5DFAD2" box="[731,854,1374,1398]" italics="true" pageId="68" pageNumber="69">Mecodema</emphasis>
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</taxonomicName>
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species by having:
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<emphasis id="B966BD220241B74A9456B739E943FAD3" bold="true" box="[1082,1096,1374,1399]" pageId="68" pageNumber="69">1</emphasis>
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, the pronotal carina with 3–4 setae along each side (
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<taxonomicName id="4C121AB30241B74A91FDB7E4ECF5FA3F" authorityName="Broun" authorityYear="1915" box="[401,510,1411,1435]" class="Insecta" family="Carabidae" genus="Metaglymma" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="curvidens">
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<emphasis id="B966BD220241B74A91FDB7E4ECF5FA3F" box="[401,510,1411,1435]" italics="true" pageId="68" pageNumber="69">curvidens</emphasis>
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</taxonomicName>
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species group);
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<emphasis id="B966BD220241B74A92D8B7E5EFC9FA3F" bold="true" box="[692,706,1410,1435]" pageId="68" pageNumber="69">2</emphasis>
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, pronotum overall shape squared, midline poorly defined, medial impressions and microsculpture absent;
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<emphasis id="B966BD220241B74A9236B7C1EF63FA1B" bold="true" box="[602,616,1446,1471]" pageId="68" pageNumber="69">3</emphasis>
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, elytral interval 7 curved inwardly to parallel humeral angle;
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<emphasis id="B966BD220241B74A9573B7C1E826FA1B" bold="true" box="[1311,1325,1446,1471]" pageId="68" pageNumber="69">4</emphasis>
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, ventrites 3 and 4 without setose punctures, ventrite 5 with 1 setose puncture each side of midline.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30832BB0241B74890ABB789EDE7FB06" lastPageId="70" lastPageNumber="71" pageId="68" pageNumber="69" type="description">
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<paragraph id="8BAD61300241B74A90ABB789ECC0F98F" blockId="68.[151,1437,1374,2011]" pageId="68" pageNumber="69">
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<emphasis id="B966BD220241B74A90ABB789EC50F9A3" bold="true" box="[199,347,1518,1543]" pageId="68" pageNumber="69">Description:</emphasis>
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Length
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<quantity id="4CEACCD50241B74A91D7B789EF69F9A2" box="[443,610,1518,1543]" metricMagnitude="-2" metricUnit="m" metricValue="2.66" metricValueMax="2.87" metricValueMin="2.45" pageId="68" pageNumber="69" unit="mm" value="26.6" valueMax="28.7" valueMin="24.5">24.5–28.7 mm</quantity>
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, pronotal width
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<quantity id="4CEACCD50241B74A9372B789EEA3F9A2" box="[798,936,1518,1543]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" metricValueMax="7.7" metricValueMin="6.5" pageId="68" pageNumber="69" unit="mm" value="7.1" valueMax="7.7" valueMin="6.5">6.5–7.7 mm</quantity>
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, elytral width
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<quantity id="4CEACCD50241B74A9422B788E9D1F9A2" box="[1102,1242,1519,1543]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="9.2" metricValueMin="7.8" pageId="68" pageNumber="69" unit="mm" value="8.5" valueMax="9.2" valueMin="7.8">7.8–9.2 mm</quantity>
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. Colour of entire body matte to glossy black.
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</paragraph>
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<paragraph id="8BAD61300241B74A90ABB451E94CF8CA" blockId="68.[151,1437,1374,2011]" pageId="68" pageNumber="69">
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<emphasis id="B966BD220241B74A90ABB451EC06F9EA" box="[199,269,1590,1614]" italics="true" pageId="68" pageNumber="69">Head:</emphasis>
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Broad and convex (
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<figureCitation id="13297DB50241B74A9194B451EF6BF9EB" box="[504,608,1590,1615]" captionStart="FIGURE 14" captionStartId="20.[151,250,936,958]" captionTargetBox="[171,1412,204,911]" captionTargetId="figure@20.[160,1412,193,921]" captionTargetPageId="20" captionText="FIGURE 14. Diagrammatic cross sections that represent the width and convexity of head or deflection of pronotum (frontal view): (A) broad and flat; (B) broad and convex; (C) narrow and flat; (D) narrow and convex." httpUri="https://zenodo.org/record/2668091/files/figure.png" pageId="68" pageNumber="69">Fig. 14B</figureCitation>
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). Vertex smooth; vertexal groove narrow, well-defined the entire length, curved posterad at midpoint; small supraorbital puncture bearing 3 setae; 3 moderately impressed supraorbital grooves; frons with a large and shallow depression each side of midline; frontoclypeal suture (
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<figureCitation id="13297DB50241B74A94D9B419E9F3F933" box="[1205,1272,1662,1687]" captionStart="FIGURE 9" captionStartId="15.[151,250,1706,1728]" captionTargetBox="[157,1433,759,1683]" captionTargetId="figure@15.[151,1436,757,1683]" captionTargetPageId="15" captionText="FIGURE 9. Dorsal view of head with detail of the morphological structures used in the species descriptions. Ƒ = vertex; SPS = supraorbital puncture setae; SP = supraorbital puncture; F = frons area; FCS = frontoclypeal suture; C = clypeus; M = mandible; LS = labrum setae; L = labrum; ACM = anterior clypeal microsculpture; CSP = clypeal setose puncture; TP = tentorial pit; SG = supraorbital grooves; E = eye; ƑG = vertexal groove." httpUri="https://zenodo.org/record/2668081/files/figure.png" pageId="68" pageNumber="69">Fig. 9</figureCitation>
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) well-defined, tentorial pits large; anterior area of clypeus with broad and shallow grooves, 1 setose puncture each side bearing 2 setae (
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<figureCitation id="13297DB50241B74A90B3B4A1EC3FF97B" box="[223,308,1734,1759]" captionStart="FIGURE 37" captionStartId="69.[151,250,1852,1874]" captionTargetBox="[258,1330,199,1821]" captionTargetId="figure@69.[252,1336,193,1831]" captionTargetPageId="69" captionText="FIGURE 37. Mecodema manaia plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668137/files/figure.png" pageId="68" pageNumber="69">Fig. 37</figureCitation>
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). Labrum rounded, anterior edge outwardly curved with 2 proximate central setae, 2 setae each side evenly spaced. Mentum lobes rounded (
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<figureCitation id="13297DB50241B74A9234B48DEFB0F8A7" box="[600,699,1770,1795]" captionStart="FIGURE 12" captionStartId="18.[151,250,1905,1927]" captionTargetBox="[311,1244,697,1878]" captionTargetId="figure@18.[310,1250,689,1889]" captionTargetPageId="18" captionText="FIGURE 12. Ventral view of the mentum with a number of different character states. Illustrations A–D are the different forms of the indentation in the apex of the median process: (A) distinctly indentate (M. manaia); (B) moderately indentate (M. pluto); (C) slightly indentate (M. infimate); (D) indentation notched (M. parataiko). Illustrations E–G are the three general forms (i.e., hashed lines) of the mentum lobes: (E) triangular; (F) rounded; (G) squared." httpUri="https://zenodo.org/record/2668087/files/figure.png" pageId="68" pageNumber="69">Fig. 12F</figureCitation>
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), median process narrow and short, slightly angled upward (15°), moderately indentate (
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<figureCitation id="13297DB50241B74A91F9B569ECF7F883" box="[405,508,1806,1831]" captionStart="FIGURE 12" captionStartId="18.[151,250,1905,1927]" captionTargetBox="[311,1244,697,1878]" captionTargetId="figure@18.[310,1250,689,1889]" captionTargetPageId="18" captionText="FIGURE 12. Ventral view of the mentum with a number of different character states. Illustrations A–D are the different forms of the indentation in the apex of the median process: (A) distinctly indentate (M. manaia); (B) moderately indentate (M. pluto); (C) slightly indentate (M. infimate); (D) indentation notched (M. parataiko). Illustrations E–G are the three general forms (i.e., hashed lines) of the mentum lobes: (E) triangular; (F) rounded; (G) squared." httpUri="https://zenodo.org/record/2668087/files/figure.png" pageId="68" pageNumber="69">Fig. 12B</figureCitation>
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); mentum setae present. Submentum sclerite constriction narrow with 4–6 evenly spaced setae. Stipes with 2 basal setae. Gula pits large, suture indistinct, gula flat with transverse lines present. Gena (
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<figureCitation id="13297DB50241B74A908DB531EC28F8CB" box="[225,291,1878,1903]" captionStart="FIGURE 8" captionStartId="13.[151,250,1097,1119]" captionTargetBox="[155,1432,309,1072]" captionTargetId="figure@13.[151,1436,303,1075]" captionTargetPageId="13" captionText="FIGURE 8. Detail of the ventral head with the taxonomic structures used in the species descriptions. SBMS = submentum setae; SBM = submentum sclerite; ML = mentum lobe; MP = mentum process (with indentation); MPW = mentum process width; SBS = stipes basal setae; MPL = mentum process length; MPS = mentum process setae; SBMC = submentum sclerite constriction; GP = gula pits; GS = gula suture." httpUri="https://zenodo.org/record/2668079/files/figure.png" pageId="68" pageNumber="69">Fig. 8</figureCitation>
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) with very fine lines forming an isodiametric pattern across entire area.
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</paragraph>
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<paragraph id="8BAD61300241B74890ABB51CEF9AFF5C" blockId="68.[151,1437,1374,2011]" lastBlockId="70.[151,1437,151,1439]" lastPageId="70" lastPageNumber="71" pageId="68" pageNumber="69">
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<emphasis id="B966BD220241B74A90ABB51CEC49F837" box="[199,322,1915,1939]" italics="true" pageId="68" pageNumber="69">Prothorax:</emphasis>
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Prothoracic carina narrow the entire length, slightly crenulated with 3–4 setae each side (
|
||
<figureCitation id="13297DB50241B74A9557B51DE884F837" box="[1339,1423,1914,1939]" captionStart="FIGURE 37" captionStartId="69.[151,250,1852,1874]" captionTargetBox="[258,1330,199,1821]" captionTargetId="figure@69.[252,1336,193,1831]" captionTargetPageId="69" captionText="FIGURE 37. Mecodema manaia plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668137/files/figure.png" pageId="68" pageNumber="69">Fig. 37</figureCitation>
|
||
), extended to anterior angle; posterior lateral sinuation indistinctly carinate, parallel; pronotum broad and laterally deflected, overall shape squared; midline poorly defined, medial impressions and microsculpture absent; pronotal foveae deep and narrow; anterior edge (
|
||
<figureCitation id="13297DB50243B748920BB2F0EFB2FF14" box="[615,697,151,176]" captionStart="FIGURE 11" captionStartId="17.[151,250,949,971]" captionTargetBox="[221,1358,181,923]" captionTargetId="figure@17.[221,1359,179,929]" captionTargetPageId="17" captionText="FIGURE 11. Detail of pronotum showing the taxonomic structures used in the species descriptions. PPC = posterior pronotum constriction; PS = pronotal setae; PAA = pronotum anterior angle; AE = anterior edge; PC = pronotal carina; ML = midline; PF= pronotal foveae; PPS = pronotum posterior sinuation; PE = posterior edge." httpUri="https://zenodo.org/record/2668085/files/figure.png" pageId="70" pageNumber="71">Fig. 11</figureCitation>
|
||
) inwardly curved, posterior edge curved, emarginated medially. Prosternum flat with fine transverse lines; proepisternum without microsculpture. Procoxal setae absent; protibia distally expanded and shovel-like (
|
||
<figureCitation id="13297DB50243B7489272B2B8EF8FFF5C" box="[542,644,223,248]" captionStart="FIGURE 10" captionStartId="16.[151,250,1442,1464]" captionTargetBox="[152,1428,191,1421]" captionTargetId="figure@16.[151,1435,183,1427]" captionTargetPageId="16" captionText="FIGURE 10. The protibia form is a character used mainly to determine between genera, except for one species in Mecodema where the difference between (A) greatly distally expanded (M. antarctica) and (B) expanded distally (all other Mecodema species) is used." httpUri="https://zenodo.org/record/2668083/files/figure.png" pageId="70" pageNumber="71">Fig. 10B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF6D31B80240B74B90FBB55BE808F834" httpUri="https://zenodo.org/record/2668137/files/figure.png" pageId="69" pageNumber="70" startId="69.[151,250,1852,1874]" targetBox="[258,1330,199,1821]" targetPageId="69">
|
||
<paragraph id="8BAD61300240B74B90FBB55BE808F834" blockId="69.[151,1436,1852,1936]" pageId="69" pageNumber="70">
|
||
<emphasis id="B966BD220240B74B90FBB55BEC29F8F6" bold="true" box="[151,290,1852,1874]" pageId="69" pageNumber="70">FIGURE 37.</emphasis>
|
||
<taxonomicName id="4C121AB30240B74B9146B55BECE5F8F7" authorityName="Seldon & Leschen" authorityYear="2011" box="[298,494,1852,1875]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="manaia">
|
||
<emphasis id="B966BD220240B74B9146B55BECE5F8F7" box="[298,494,1852,1875]" italics="true" pageId="69" pageNumber="70">Mecodema manaia</emphasis>
|
||
</taxonomicName>
|
||
plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BAD61300243B74890ABB363EDDDFE53" blockId="70.[151,1437,151,1439]" pageId="70" pageNumber="71">
|
||
<emphasis id="B966BD220243B74890ABB363EC1DFEB8" box="[199,278,260,284]" italics="true" pageId="70" pageNumber="71">Elytra:</emphasis>
|
||
Broad and moderately deflected; humeral angle evenly convex (
|
||
<figureCitation id="13297DB50243B7489460B363E963FEB9" box="[1036,1128,260,285]" captionStart="FIGURE 6" captionStartId="10.[151,250,519,541]" captionTargetBox="[156,1432,193,498]" captionTargetId="figure@10.[151,1436,193,498]" captionTargetPageId="10" captionText="FIGURE 6. Detail of the three different forms of the elytral humeral angle: (A) = anteriorly convergent (M. parataiko), (B) = evenly convex (M. oconnori), (C) = subangulate (M. atuanui)." httpUri="https://zenodo.org/record/2668075/files/figure.png" pageId="70" pageNumber="71">Fig. 6B</figureCitation>
|
||
); basal margin moderately curved and bevelled to base, interval 1 extended to margin; lateral carina narrow the entire length, extended to humeral angle; humerus with 2–3 setose punctures (
|
||
<figureCitation id="13297DB50243B7489288B32BEE33FEC0" box="[740,824,332,357]" captionStart="FIGURE 37" captionStartId="69.[151,250,1852,1874]" captionTargetBox="[258,1330,199,1821]" captionTargetId="figure@69.[252,1336,193,1831]" captionTargetPageId="69" captionText="FIGURE 37. Mecodema manaia plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668137/files/figure.png" pageId="70" pageNumber="71">Fig. 37</figureCitation>
|
||
); suture poorly impressed; striae with small, regularly distributed, asetose punctures that slightly increased in size laterally; intervals 1–4 weakly convex, intervals 5–9 moderately convex, interval 7 curved inwardly to parallel humeral angle; interval microsculpture absent; 7
|
||
<superScript id="7C67CC780243B748953CB3F4E857FE05" attach="left" box="[1360,1372,403,417]" fontSize="6" pageId="70" pageNumber="71">th</superScript>
|
||
strial setal pattern with 2 setose punctures in anterior ½, 5 setose punctures in posterior ½ (
|
||
<figureCitation id="13297DB50243B7489419B3DDE9C0FE77" box="[1141,1227,442,467]" captionStart="FIGURE 37" captionStartId="69.[151,250,1852,1874]" captionTargetBox="[258,1330,199,1821]" captionTargetId="figure@69.[252,1336,193,1831]" captionTargetPageId="69" captionText="FIGURE 37. Mecodema manaia plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668137/files/figure.png" pageId="70" pageNumber="71">Fig. 37</figureCitation>
|
||
), setose punctures large.
|
||
</paragraph>
|
||
<paragraph id="8BAD61300243B74890ABB064EDEEFD57" blockId="70.[151,1437,151,1439]" pageId="70" pageNumber="71">
|
||
<emphasis id="B966BD220243B74890ABB064EC70FDBF" box="[199,379,515,539]" italics="true" pageId="70" pageNumber="71">Ventral surface:</emphasis>
|
||
Mesepisternum and metepisternum (
|
||
<figureCitation id="13297DB50243B748934CB065EE6FFDBF" box="[800,868,514,539]" captionStart="FIGURE 2" captionStartId="6.[151,250,800,822]" captionTargetBox="[159,1432,197,773]" captionTargetId="figure@6.[151,1436,193,779]" captionTargetPageId="6" captionText="FIGURE 2. Ventral view of a Mecodema specimen showing specific morphological structures, excluding taxonomic structures indicated in detail figures, used in the species descriptions. ƑLF = ventrite lateral foveae; ƑSP = ventrite setose punctures; MTC = metacoxa; MTƑP = metaventrite process; MSC = mesocoxa; PC = procoxa; PS = prosternum; G = gena; PES = proepisternum; MSE = mesepisternum; MTE = metepisternum; Ƒ1–Ƒ6 = ventrites 1-6 (ventrites 1-3 may be fused); M = midline (dashed line, not a taxonomic structure)." httpUri="https://zenodo.org/record/2668067/files/figure.png" pageId="70" pageNumber="71">Fig. 2</figureCitation>
|
||
) without microsculpture; setose punctures present on mesocoxa (1) and metacoxa (1). All abdominal ventrites finely lineate laterally, ventrites 3 and 4 without setose punctures, ventrite 5 (
|
||
<figureCitation id="13297DB50243B74891E1B02DECDBFDC7" box="[397,464,586,611]" captionStart="FIGURE 2" captionStartId="6.[151,250,800,822]" captionTargetBox="[159,1432,197,773]" captionTargetId="figure@6.[151,1436,193,779]" captionTargetPageId="6" captionText="FIGURE 2. Ventral view of a Mecodema specimen showing specific morphological structures, excluding taxonomic structures indicated in detail figures, used in the species descriptions. ƑLF = ventrite lateral foveae; ƑSP = ventrite setose punctures; MTC = metacoxa; MTƑP = metaventrite process; MSC = mesocoxa; PC = procoxa; PS = prosternum; G = gena; PES = proepisternum; MSE = mesepisternum; MTE = metepisternum; Ƒ1–Ƒ6 = ventrites 1-6 (ventrites 1-3 may be fused); M = midline (dashed line, not a taxonomic structure)." httpUri="https://zenodo.org/record/2668067/files/figure.png" pageId="70" pageNumber="71">Fig. 2</figureCitation>
|
||
) with 1 setose puncture each side of midline; ventrite 6 setae present:
|
||
<specimenCount id="9D14AAB90243B748948BB02CE9F7FDC6" box="[1255,1276,587,610]" pageId="70" pageNumber="71" type="male">♂</specimenCount>
|
||
with 2 evenly spaced setose punctures each side, most lateral pair at junction of straight apical edge and curve anterad,
|
||
<specimenCount id="9D14AAB90243B7489520B017E86AFD23" box="[1356,1377,624,647]" pageId="70" pageNumber="71" type="female">♀</specimenCount>
|
||
with 2–3 setose punctures proximate, but distantly separated on bluntly rounded apical edge; lateral foveae absent. Anterior metaventrite process (
|
||
<figureCitation id="13297DB50243B7489193B0D1EF4DFD6B" box="[511,582,694,719]" captionStart="FIGURE 2" captionStartId="6.[151,250,800,822]" captionTargetBox="[159,1432,197,773]" captionTargetId="figure@6.[151,1436,193,779]" captionTargetPageId="6" captionText="FIGURE 2. Ventral view of a Mecodema specimen showing specific morphological structures, excluding taxonomic structures indicated in detail figures, used in the species descriptions. ƑLF = ventrite lateral foveae; ƑSP = ventrite setose punctures; MTC = metacoxa; MTƑP = metaventrite process; MSC = mesocoxa; PC = procoxa; PS = prosternum; G = gena; PES = proepisternum; MSE = mesepisternum; MTE = metepisternum; Ƒ1–Ƒ6 = ventrites 1-6 (ventrites 1-3 may be fused); M = midline (dashed line, not a taxonomic structure)." httpUri="https://zenodo.org/record/2668067/files/figure.png" pageId="70" pageNumber="71">Fig. 2</figureCitation>
|
||
) a bluntly rounded triangle with a well-defined, narrow carina the entire length.
|
||
</paragraph>
|
||
<paragraph id="8BAD61300243B74890ABB098E9C0FB93" blockId="70.[151,1437,151,1439]" pageId="70" pageNumber="71">
|
||
<emphasis id="B966BD220243B74890ABB098EC7EFCB3" box="[199,373,767,791]" italics="true" pageId="70" pageNumber="71">Male genitalia:</emphasis>
|
||
Apical portion of penis lobe symmetrically rounded (
|
||
<figureCitation id="13297DB50243B74893B2B099E924FCB3" box="[990,1071,766,791]" captionStart="FIGURE 37" captionStartId="69.[151,250,1852,1874]" captionTargetBox="[258,1330,199,1821]" captionTargetId="figure@69.[252,1336,193,1831]" captionTargetPageId="69" captionText="FIGURE 37. Mecodema manaia plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668137/files/figure.png" pageId="70" pageNumber="71">Fig. 37</figureCitation>
|
||
PL) with a distinct deflection to right of vertical axis (VV) (
|
||
<figureCitation id="13297DB50243B74891A4B145EF24FC9F" box="[456,559,802,827]" captionStart="FIGURE 15" captionStartId="20.[151,250,1943,1965]" captionTargetBox="[162,1424,1039,1914]" captionTargetId="figure@20.[162,1424,1038,1921]" captionTargetPageId="20" captionText="FIGURE 15. The penis lobe in ventral view showing the relative deflection to right of vertical: (A) distinctly deflected (M. regulus); (B) moderately deflected (M. haunoho); (C) slightly deflected (M. oregoides). Scale bars = 1 mm. Structures of penis lobe indicated are: base (B); lateral edge (LE); apical portion (AP); ventral edge (ƑE)." httpUri="https://zenodo.org/record/2668093/files/figure.png" pageId="70" pageNumber="71">Fig. 15A</figureCitation>
|
||
); shaft of penis lobe straight and relatively broad entire (VV), overall lenght of penis lobe without curve ventrally (LV). Structures of the endophallus (
|
||
<figureCitation id="13297DB50243B74893B4B121E92AFCFB" box="[984,1057,838,863]" captionStart="FIGURE 4" captionStartId="7.[151,250,973,995]" captionTargetBox="[354,1232,193,952]" captionTargetId="figure@7.[354,1232,193,952]" captionTargetPageId="7" captionText="FIGURE 4. An Environmental Scanning Electron Microscope (ESEM) photograph of the four main structures of the apical plate of the endophallus within the penis lobe. This is an example from Mecodema haunoho showing the central spicule (c.s), left (l.s.f) and the right setose flange (r.s.f), and microtrichial field (m.f). These structures are variable among species but within a species, they are very consistent, even over distant geographical ranges. The microtrichial field can be covered in flattened scales as in photo above, or with elongated setae-like scales." httpUri="https://zenodo.org/record/2668071/files/figure.png" pageId="70" pageNumber="71">Fig. 4</figureCitation>
|
||
): lateral form of the apex of the central spicule rounded; dorsal form of the apex of the central spicule short and narrow; relative extent of setae (scales) of the apical plate sparse (1–25%); size of left setose flange small; size of right setose flange large. Left paramere basal lobe narrowly rectangular, short slope to arm; arm long and relatively broad, gradually narrowed to terminal lobe; terminal lobe indistinct from arm, apical tuft of long setae; medium-lengthed setae continued along apical ¾ of ventral edge (
|
||
<figureCitation id="13297DB50243B74891D9B19DEF0CFBB7" box="[437,519,1018,1043]" captionStart="FIGURE 37" captionStartId="69.[151,250,1852,1874]" captionTargetBox="[258,1330,199,1821]" captionTargetId="figure@69.[252,1336,193,1831]" captionTargetPageId="69" captionText="FIGURE 37. Mecodema manaia plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668137/files/figure.png" pageId="70" pageNumber="71">Fig. 37</figureCitation>
|
||
LP); ventral edge slightly curved. Right paramere narrowly rectangular, gradually narrowed apically with medium to long setae extended along apical ¾ ventral edge (
|
||
<figureCitation id="13297DB50243B7489424B679E992FB93" box="[1096,1177,1054,1079]" captionStart="FIGURE 37" captionStartId="69.[151,250,1852,1874]" captionTargetBox="[258,1330,199,1821]" captionTargetId="figure@69.[252,1336,193,1831]" captionTargetPageId="69" captionText="FIGURE 37. Mecodema manaia plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668137/files/figure.png" pageId="70" pageNumber="71">Fig. 37</figureCitation>
|
||
RP).
|
||
</paragraph>
|
||
<paragraph id="8BAD61300243B74890ABB624EDE7FB06" blockId="70.[151,1437,151,1439]" pageId="70" pageNumber="71">
|
||
<emphasis id="B966BD220243B74890ABB624EC9BFBFF" box="[199,400,1091,1115]" italics="true" pageId="70" pageNumber="71">Female genitalia:</emphasis>
|
||
Basal gonocoxite 1 (
|
||
<figureCitation id="13297DB50243B74892EFB625EFCCFBFF" box="[643,711,1090,1115]" captionStart="FIGURE 5" captionStartId="9.[151,250,1714,1736]" captionTargetBox="[167,1436,790,1674]" captionTargetId="figure@9.[151,1436,790,1693]" captionTargetPageId="9" captionText="FIGURE 5. The entire female reproductive tract (ventral view) including taxonomic structures (for all other abbreviated structure names see Seldon et al. 2012). However, for this revision only the following structures are important: HS = helminthoid sclerite; G1 = gonocoxite 1; G2 = gonocoxite 2; TDC = transverse dorsal carina; R = ramus." httpUri="https://zenodo.org/record/2668073/files/figure.png" pageId="70" pageNumber="71">Fig. 5</figureCitation>
|
||
) long and broad, flattened apically, internal dorso-lateral carina with 2 relatively long setae, ventral surface smooth. Gonocoxite 2 bluntly rounded and triangular. Ramus short and narrow.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30832BB0243B74890ABB6C9EFD3FAAB" pageId="70" pageNumber="71" type="discussion">
|
||
<paragraph id="8BAD61300243B74890ABB6C9EFD3FAAB" blockId="70.[151,1437,151,1439]" pageId="70" pageNumber="71">
|
||
<emphasis id="B966BD220243B74890ABB6C9EC59FB63" bold="true" box="[199,338,1198,1223]" pageId="70" pageNumber="71">Comments:</emphasis>
|
||
<taxonomicName id="4C121AB30243B7489134B6C8EF25FB63" authorityName="Seldon & Leschen" authorityYear="2011" box="[344,558,1199,1223]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="manaia">
|
||
<emphasis id="B966BD220243B7489134B6C8EF25FB63" box="[344,558,1199,1223]" italics="true" pageId="70" pageNumber="71">Mecodema manaia</emphasis>
|
||
</taxonomicName>
|
||
shares similar morphological characters with
|
||
<taxonomicName id="4C121AB30243B748945FB6D7E84CFB63" authority="(Cape Reinga)" baseAuthorityName="Cape Reinga" box="[1075,1351,1198,1223]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="tenaki">
|
||
<emphasis id="B966BD220243B748945FB6D7E995FB63" box="[1075,1182,1199,1223]" italics="true" pageId="70" pageNumber="71">M. tenaki</emphasis>
|
||
(Cape Reinga)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C121AB30243B7489511B6D7ECE2FB4F" authority="(Three Kings Islands)" baseAuthorityName="Three Kings Islands" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="regulus">
|
||
<emphasis id="B966BD220243B7489511B6D7EDE1FB4F" italics="true" pageId="70" pageNumber="71">M. regulus</emphasis>
|
||
(Three Kings Islands)
|
||
</taxonomicName>
|
||
, and although there is suitable forest habitat along the eastern coastline to the north, no specimens have been collected from these areas.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30832BB0243B74890ABB77DE93EFA96" box="[199,1077,1306,1331]" pageId="70" pageNumber="71" type="distribution">
|
||
<paragraph id="8BAD61300243B74890ABB77DE93EFA96" blockId="70.[151,1437,151,1439]" box="[199,1077,1306,1331]" pageId="70" pageNumber="71">
|
||
<emphasis id="B966BD220243B74890ABB77DEC69FA97" bold="true" box="[199,354,1306,1331]" pageId="70" pageNumber="71">Distribution:</emphasis>
|
||
<collectingCountry id="F30521A00243B7489105B77CEF09FA97" box="[361,514,1306,1331]" name="New Zealand" pageId="70" pageNumber="71">New Zealand</collectingCountry>
|
||
, North Island, Northland, Whangarei Head’s area.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30832BB0243B74890ABB759EFABFA3B" pageId="70" pageNumber="71" type="materials_examined">
|
||
<paragraph id="8BAD61300243B74890ABB759E956FADF" blockId="70.[151,1437,151,1439]" pageId="70" pageNumber="71">
|
||
<materialsCitation id="3B7A6B6D0243B74890ABB759E88FFAF3" ID-GBIF-Occurrence="2237851760" box="[199,1412,1342,1367]" collectionCode="NZAC" collectorName="ND Whangarei Heads Mt Lion & F. Brook" country="New Zealand" location="NEW ZEALAND" pageId="70" pageNumber="71" specimenCount="1" typeStatus="holotype">
|
||
<emphasis id="B966BD220243B74890ABB759EC36FAF3" bold="true" box="[199,317,1342,1367]" pageId="70" pageNumber="71">
|
||
<typeStatus id="54A9DF920243B74890ABB759EC33FAF3" box="[199,312,1342,1367]" pageId="70" pageNumber="71" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<collectionCode id="ED03F9F50243B7489121B758EC92FAF3" box="[333,409,1343,1367]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="70" pageNumber="71" type="Other institutional type">NZAC</collectionCode>
|
||
male labelled.
|
||
<collectingCountry id="F30521A00243B7489233B727EE38FAF3" box="[607,819,1343,1367]" name="New Zealand" pageId="70" pageNumber="71">NEW ZEALAND</collectingCountry>
|
||
<collectorName id="26E704E60243B748932FB727E9C9FAF3" box="[835,1218,1342,1367]" pageId="70" pageNumber="71">ND Whangarei Heads Mt Lion</collectorName>
|
||
2005
|
||
<collectorName id="26E704E60243B7489576B727E88FFAF3" box="[1306,1412,1342,1367]" pageId="70" pageNumber="71">F. Brook</collectorName>
|
||
</materialsCitation>
|
||
/
|
||
<typeStatus id="54A9DF920243B74890FBB704EC20FADF" box="[151,299,1379,1403]" pageId="70" pageNumber="71" type="holotype">HOLOTYPE</typeStatus>
|
||
<taxonomicName id="4C121AB30243B748915EB705EF0DFADF" authority="Seldon & Buckley, 2019" authorityName="Seldon & Buckley" authorityYear="2019" box="[306,518,1378,1403]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="manaia" status="sp. nov.">Mecodema manaia</taxonomicName>
|
||
<emphasis id="B966BD220243B7489261B705EF40FADF" bold="true" box="[525,587,1378,1403]" pageId="70" pageNumber="71">
|
||
<taxonomicNameLabel id="A25500590243B7489261B705EF40FADF" box="[525,587,1378,1403]" pageId="70" pageNumber="71" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
design. by Seldon & Leschen 2010 [red label].
|
||
</paragraph>
|
||
<paragraph id="8BAD61300243B74890ABB7E1EFABFA3B" blockId="70.[151,1437,151,1439]" box="[199,672,1414,1439]" pageId="70" pageNumber="71">
|
||
<emphasis id="B966BD220243B74890ABB7E1EC43FA3B" bold="true" box="[199,328,1414,1439]" pageId="70" pageNumber="71">
|
||
<typeStatus id="54A9DF920243B74890ABB7E1EC48FA3B" box="[199,323,1414,1439]" pageId="70" pageNumber="71" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
</emphasis>
|
||
(see
|
||
<bibRefCitation id="EF831CC10243B74891EFB7E1EF9FFA3B" author="Seldon, D. S. & Leschen, R. A. B." box="[387,660,1414,1439]" pageId="70" pageNumber="71" pagination="1 - 45" refId="ref93265" refString="Seldon, D. S. & Leschen, R. A. B. (2011) Revision of the Mecodema curvidens species group (Coleoptera: Carabidae: Broscini). Zootaxa, 2829, 1 - 45. https: // doi. org / 10.11646 / zootaxa. 2829.1.1" type="journal article" year="2011">Seldon & Leschen 2011</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |