215 lines
20 KiB
XML
215 lines
20 KiB
XML
<document ID-DOI="10.1080/00222933.2018.1524032" ID-GBIF-Dataset="034ebcd9-efa7-416a-822c-b6aee0e57a1c" ID-ISSN="1464-5262" ID-Zenodo-Dep="3670229" checkinTime="1581941790231" checkinUser="valdenar" docAuthor="DeVries, Thomas J." docDate="2019" docId="03BB2338E369E219FE49A5445E88FA5A" docLanguage="en" docName="JNATHIST.53.25–26.1533–1584.pdf.imf" docOrigin="Journal of Natural History 53 (25)" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Nodifaunus gainesi DeVries 2019, sp. nov." docType="treatment" docVersion="21" lastPageNumber="1566" masterDocId="FF825B40E348E23BFFA1A1435A0CFFCC" masterDocTitle="Early Paleogene brackish-water molluscs from the Caballas Formation of the East Pisco Basin (Southern Peru)" masterLastPageNumber="1584" masterPageNumber="1533" pageNumber="1565" updateTime="1636522412784" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Early Paleogene brackish-water molluscs from the Caballas Formation of the East Pisco Basin (Southern Peru)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>DeVries, Thomas J.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Journal of Natural History</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2018-12-17</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>53</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>25</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1533</mods:start>
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<mods:end>1584</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1080/00222933.2018.1524032</mods:identifier>
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<mods:identifier type="GBIF-Dataset">034ebcd9-efa7-416a-822c-b6aee0e57a1c</mods:identifier>
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<mods:identifier type="ISSN">1464-5262</mods:identifier>
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<mods:identifier type="Zenodo-Dep">3670229</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3671262" ID-GBIF-Taxon="162146488" ID-Zenodo-Dep="3671262" LSID="urn:lsid:plazi:treatment:03BB2338E369E219FE49A5445E88FA5A" httpUri="http://treatment.plazi.org/id/03BB2338E369E219FE49A5445E88FA5A" lastPageId="34" lastPageNumber="1566" pageId="33" pageNumber="1565">
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<subSubSection box="[488,828,1031,1056]" pageId="33" pageNumber="1565" type="nomenclature">
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<paragraph blockId="33.[488,828,1031,1091]" box="[488,828,1031,1056]" pageId="33" pageNumber="1565">
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<heading box="[488,828,1031,1056]" centered="true" fontSize="11" level="2" pageId="33" pageNumber="1565" reason="2">
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<taxonomicName authority="DeVries, 2019" authorityName="DeVries" authorityYear="2019" box="[488,725,1031,1056]" class="Gastropoda" family="Pachychilidae" genus="Nodifaunus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="33" pageNumber="1565" phylum="Mollusca" rank="species" species="gainesi" status="sp. nov.">
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<emphasis bold="true" box="[488,725,1031,1056]" italics="true" pageId="33" pageNumber="1565">Nodifaunus gainesi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[735,828,1031,1056]" pageId="33" pageNumber="1565" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[563,752,1066,1091]" pageId="33" pageNumber="1565" type="description">
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<paragraph blockId="33.[488,828,1031,1091]" box="[563,752,1066,1091]" pageId="33" pageNumber="1565">
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<heading box="[563,752,1066,1091]" centered="true" fontSize="11" level="2" pageId="33" pageNumber="1565" reason="5">
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(
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<figureCitation box="[571,670,1066,1091]" captionStart="Figure 4" captionStartId="13.[160,225,936,959]" captionTargetId="figure@14.[162,1153,142,1037]" captionTargetPageId="14" captionText="Figure 4. Gastropods from the Caballas Formation. (a–e) Nerita squatina sp. nov., B8771. (a) UWBM 107577, holotype, W 5.7 mm, spire. (b) UWBM 107648, paratype, W 8.5 mm, shoulder of last whorl. (c) UWBM 107649,paratype, W 10.1 mm, anterior of outer lip.(d) UWBM 107647,paratype,W 7.8 mm,inner surface of outer lip. (e) UWBM 107646, paratype, W 6.2 mm, columnar teeth. (f) Calyptraea cf. C. aperta Solander, 1766, UWBM 107578, B8772,W (diameter) 9.9 mm, external mould. (g) Ampullina ortoni Gabb, 1870, UWBM 107,627, B8772, L 36.8 mm, abapertural view, last whorl mostly missing; left margin annotated. (h–l) Melanella indicaformis sp. nov. (h) UWBM 107620, paratype, B8772, L 8.2 mm, apertural view. (i) UWBM 107619, paratype, B8769, L 9.4 mm, apertural/lateral view. (j) UWBM 107623, paratype, B8769,L 8.2 mm, apertural view. (k) UWBM 107618, paratype, B8772, L 10.6, abapertural view. (l) UWBM 107617, holotype, B8769, L 11.1 mm, abapertural view. (m–q) Potamides henryi sp. nov. (m) UWBM 107582, holotype, B8772, L 17.8 mm, lateral view. (n) UWBM 107582, W 6.9 mm, basal view. (o) UWBM 107586, paratype, B8771, L 11.2 mm, lateral view. (p) UWBM 107587, paratype, B8769, L 9.5 mm, abapertural view. (q) UWBM 107650, paratype, B8770, L 15.4 mm, lateral view, shell compressed. (r–t) Potamides janeae sp. nov., B8771.(r) UWBM 107579, holotype, L 9.0 mm, lateral view. (s) MUSM INV 249, L 8.8 mm, lateral and columellar view. (t) UWBM 107580, L 9.5 mm, layeral and columellar view. (u, v, x) Papposilenus utriculus gen.nov., sp. nov. (u) UWBM 107592,paratype, B8772, L 18.7 mm, oblique view of base. (v) UWBM 107589, holotype, B8772, L 25.5 mm, lateral view. (x) UWBM 107590, B8770, L 17.6 mm, lateral view.(w, y,z,jj) Rhinotamides everriculum sp.nov. (w) UWBM 107598,paratype, B8772,L 23.5 mm, abapertural view, juvenile. (y) UWBM 107593, holotype, B8769, L 21.1 mm, abapertural view. (z) UWBM 107594, paratype, B8772, L 19.4 mm, apertural view, posterior and anterior missing. (jj) UWBM 107593, apertural view.(aa,bb) Terebralia pauli gen. nov.sp.nov., UWBM 107599,holotype,B8771, L 9.2 mm.(aa) Abapertural view. (bb) Apertural view, outer lip missing. (cc) Terebralia marki gen. nov. sp. nov., UWBM 107600, holotype, B8771, L 9.8, lateral view. (dd–ii). Nodifaunus gainesi sp. nov. (dd) UWBM 107609, holotype, B8769, L 42.4 mm, apertural view. (ee) MUSM INV 264, paratype, B8772, L 31.0 mm, apertural view, anterior missing. (ff) UWBM 107609, abapertural view. (gg) UWBM 107612, paratype, B8770, L 14.2 mm, abapertural view, juvenile. (hh) UWBM 107612, apertural view. (ii) UWBM 107610, paratype, B8769, L 25.9 mm, apertural view, anterior missing." figureDoi="http://doi.org/10.5281/zenodo.3686966" httpUri="https://zenodo.org/record/3686966/files/figure.png" pageId="33" pageNumber="1565">Figure 4</figureCitation>
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(dd
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<emphasis box="[709,723,1066,1091]" italics="true" pageId="33" pageNumber="1565">–</emphasis>
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ii))
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="33" pageNumber="1565" type="diagnosis">
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<paragraph blockId="33.[160,1155,1141,1270]" box="[160,276,1141,1167]" pageId="33" pageNumber="1565">
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<emphasis bold="true" box="[160,276,1141,1167]" italics="true" pageId="33" pageNumber="1565">Diagnosis</emphasis>
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</paragraph>
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<paragraph blockId="33.[160,1155,1141,1270]" pageId="33" pageNumber="1565">Early teleoconch whorls smooth; later whorls with tubercles or tuberculate axial ribs situated near posterior suture. Base smooth, separated from side of whorl by single spiral cord. Anterior canal short.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="34" lastPageNumber="1566" pageId="33" pageNumber="1565" type="description">
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<paragraph blockId="33.[160,1156,1320,1727]" box="[160,294,1320,1346]" pageId="33" pageNumber="1565">
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<emphasis bold="true" box="[160,294,1320,1346]" italics="true" pageId="33" pageNumber="1565">Description</emphasis>
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</paragraph>
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<paragraph blockId="33.[160,1156,1320,1727]" lastBlockId="34.[160,1156,142,305]" lastPageId="34" lastPageNumber="1566" pageId="33" pageNumber="1565">Shell length to about 45 mm, high-spired, cerithiform, spire angle about 35° for early whorls, 15° for later whorls. Protoconch unknown. Teleoconch of nine or 10 slightly convex whorls; sutures impressed; sutures of early teleoconch whorls weakly canaliculate to turriculate. Spiral sculpture on earliest five or six whorls smooth; later whorls with spiral row of tubercles along posterior margin, present as blunt spines or developed as coarse prosocline, orthocline, or opisthocline axial ribs (variable orientation between and within individual shells) with blunt erect spines rising above posterior suture and extending partly or entirely to anterior suture; as many as 12 tubercles/axial ribs per whorl at first appearance, as few as eight on last whorl. No varices or ventrolateral varix. Faint spiral lines may be diagenetic artefacts. Growth lines orthocline to opisthocyrt. Base poorly preserved; steeply rounded, smooth, separated from side of whorl by spiral cord. Aperture rhombohedral; parietal area with broad callus, raised above body whorl; columella straight, thickened, without folds; anterior canal very short, slightly recurved abaxially. Outer lip mostly missing, posterior portion adjacent to suture extended abaxially with extension aligned with last subsutural tubercle; shallow posterior canal present.</paragraph>
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</subSubSection>
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<subSubSection pageId="34" pageNumber="1566" type="discussion">
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<paragraph blockId="34.[160,1155,370,811]" box="[160,262,370,396]" pageId="34" pageNumber="1566">
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<emphasis bold="true" box="[160,262,370,396]" italics="true" pageId="34" pageNumber="1566">Remarks</emphasis>
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</paragraph>
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<paragraph blockId="34.[160,1155,370,811]" pageId="34" pageNumber="1566">
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<taxonomicName authority="DeVries, 2019" authorityName="DeVries" authorityYear="2019" box="[160,374,406,430]" class="Gastropoda" family="Pachychilidae" genus="Nodifaunus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="34" pageNumber="1566" phylum="Mollusca" rank="species" species="gainesi" status="sp. nov.">
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<emphasis box="[160,374,406,430]" italics="true" pageId="34" pageNumber="1566">Nodifaunus gainesi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[387,483,406,430]" pageId="34" pageNumber="1566" rank="species">sp. nov.</taxonomicNameLabel>
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, with one spiral row of tubercles, differs from the early Eocene species,
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<taxonomicName authorityName="Woodsi" authorityYear="1922" box="[353,512,441,465]" class="Gastropoda" family="Thiaridae" genus="Melanatria" kingdom="Animalia" pageId="34" pageNumber="1566" phylum="Mollusca" rank="species" species="dimorphica">
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<emphasis box="[353,512,441,465]" italics="true" pageId="34" pageNumber="1566">N. dimorphica</emphasis>
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</taxonomicName>
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, from the Talara Basin of northern Peru, which has a second row of nodes at the angled boundary of the base.
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<taxonomicName authorityName="DeVries" authorityYear="2019" box="[811,1018,475,499]" class="Gastropoda" family="Pachychilidae" genus="Nodifaunus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="34" pageNumber="1566" phylum="Mollusca" rank="species" species="gainesi">
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<emphasis box="[811,1018,475,499]" italics="true" pageId="34" pageNumber="1566">Nodifaunus gainesi</emphasis>
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</taxonomicName>
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differs from
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<taxonomicName authorityName="Woodsi" authorityYear="1922" box="[160,306,510,534]" class="Gastropoda" family="Thiaridae" genus="Melanatria" kingdom="Animalia" pageId="34" pageNumber="1566" phylum="Mollusca" rank="species" species="propinqua">
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<emphasis box="[160,306,510,534]" italics="true" pageId="34" pageNumber="1566">N. propinqua</emphasis>
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</taxonomicName>
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and
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<taxonomicName authorityName="Woodsi" authorityYear="1922" box="[369,487,510,534]" class="Gastropoda" family="Thiaridae" genus="Melanatria" kingdom="Animalia" pageId="34" pageNumber="1566" phylum="Mollusca" rank="species" species="venusta">
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<emphasis box="[369,487,510,534]" italics="true" pageId="34" pageNumber="1566">N. venusta</emphasis>
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</taxonomicName>
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, both also from the Talara Basin, which have one row of tubercles, but medially located. All three northern Peruvian species also have spiral cords or threads on the base and on the early whorls. The posterior suture-hugging single row of tubercles on specimens of
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<taxonomicName authorityName="DeVries" authorityYear="2019" box="[504,612,614,638]" class="Gastropoda" family="Pachychilidae" genus="Nodifaunus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="34" pageNumber="1566" phylum="Mollusca" rank="species" species="gainesi">
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<emphasis box="[504,612,614,638]" italics="true" pageId="34" pageNumber="1566">N. gainesi</emphasis>
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</taxonomicName>
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most resembles the sculpture of specimens of Late Cretaceous
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<taxonomicName authorityName="Olsson" authorityYear="1944" box="[347,467,649,673]" class="Gastropoda" family="Pachychilidae" genus="Nodifaunus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="34" pageNumber="1566" phylum="Mollusca" rank="species" species="nodosus">
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<emphasis box="[347,467,649,673]" italics="true" pageId="34" pageNumber="1566">N. nodosus</emphasis>
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</taxonomicName>
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, which also lack spiral cords, but the number of tubercles on the earliest nodose teleoconch whorls of
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<taxonomicName authorityName="DeVries" authorityYear="2019" box="[631,738,683,707]" class="Gastropoda" family="Pachychilidae" genus="Nodifaunus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="34" pageNumber="1566" phylum="Mollusca" rank="species" species="gainesi">
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<emphasis box="[631,738,683,707]" italics="true" pageId="34" pageNumber="1566">N. gainesi</emphasis>
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</taxonomicName>
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(about 12) is double the number on the Late Cretaceous species.
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</paragraph>
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<paragraph blockId="34.[160,1155,370,811]" pageId="34" pageNumber="1566">
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Juvenile specimens of
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<taxonomicName authorityName="DeVries" authorityYear="2019" box="[468,683,753,777]" class="Gastropoda" family="Pachychilidae" genus="Nodifaunus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="34" pageNumber="1566" phylum="Mollusca" rank="species" species="gainesi">
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<emphasis box="[468,683,753,777]" italics="true" pageId="34" pageNumber="1566">Nodifaunus gainesi</emphasis>
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</taxonomicName>
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have a widely flaring aperture with a thickened outer lip. Both a twisted anterior canal and posterior canal are present.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="34" pageNumber="1566" type="etymology">
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<paragraph blockId="34.[160,1155,876,1006]" box="[160,285,876,902]" pageId="34" pageNumber="1566">
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<emphasis bold="true" box="[160,285,876,902]" italics="true" pageId="34" pageNumber="1566">Etymology</emphasis>
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</paragraph>
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<paragraph blockId="34.[160,1155,876,1006]" pageId="34" pageNumber="1566">
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<emphasis box="[160,166,913,937]" italics="true" pageId="34" pageNumber="1566">‘</emphasis>
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Gainesi
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<emphasis box="[247,253,913,937]" italics="true" pageId="34" pageNumber="1566">’</emphasis>
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, in recognition of Arthur G. Gaines, Oceanographer Emeritus, of the Woods Hole Oceanographic Institution, Woods Hole, MA, USA, and ocean sciences mentor to the author in the early 1970s.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="34" pageNumber="1566" type="materials_examined">
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<paragraph blockId="34.[160,1157,1071,1305]" box="[160,259,1071,1097]" pageId="34" pageNumber="1566">
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<emphasis bold="true" box="[160,259,1071,1097]" italics="true" pageId="34" pageNumber="1566">Material</emphasis>
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</paragraph>
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<paragraph blockId="34.[160,1157,1071,1305]" pageId="34" pageNumber="1566">
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<materialsCitation ID-GBIF-Occurrence="2572183373" box="[160,869,1107,1131]" collectionCode="UWBM" pageId="34" pageNumber="1566" specimenCode="UWBM 107609" specimenCount="1" typeStatus="holotype">
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<specimenCode box="[160,326,1107,1131]" pageId="34" pageNumber="1566">UWBM 107609</specimenCode>
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,
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<typeStatus box="[336,437,1107,1131]" pageId="34" pageNumber="1566">holotype</typeStatus>
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, B8769 (type locality), L (42.4), W (19.1)
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</materialsCitation>
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;
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<materialsCitation ID-GBIF-Occurrence="2572183363" collectionCode="UWBM" pageId="34" pageNumber="1566" specimenCode="UWBM 107591" specimenCount="1" typeStatus="paratype">
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remainder are
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<typeStatus box="[1042,1153,1107,1131]" pageId="34" pageNumber="1566">paratypes</typeStatus>
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:
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<specimenCode box="[160,327,1142,1166]" pageId="34" pageNumber="1566">UWBM 107591</specimenCode>
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, B8769, L (31.8), W (19.9)
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</materialsCitation>
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;
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<materialsCitation ID-GBIF-Occurrence="2572183374" box="[620,1068,1142,1166]" collectionCode="UWBM" pageId="34" pageNumber="1566" specimenCode="UWBM 107610" specimenCount="1">
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<specimenCode box="[620,787,1142,1166]" pageId="34" pageNumber="1566">UWBM 107610</specimenCode>
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, B8769, L (25.9), W (17.0)
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</materialsCitation>
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;
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<materialsCitation ID-GBIF-Occurrence="2572183356" collectionCode="UWBM" pageId="34" pageNumber="1566" specimenCode="UWBM 107611" specimenCount="1">
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<specimenCode pageId="34" pageNumber="1566">UWBM 107611</specimenCode>
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, B8770, L (36.9), W (18.2)
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</materialsCitation>
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;
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<materialsCitation ID-GBIF-Occurrence="2572183359" box="[550,970,1177,1201]" collectionCode="UWBM" pageId="34" pageNumber="1566" specimenCode="UWBM 107612" specimenCount="1">
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<specimenCode box="[550,720,1177,1201]" pageId="34" pageNumber="1566">UWBM 107612</specimenCode>
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, B8770, L 14.2, W 7.4
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</materialsCitation>
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;
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<materialsCitation ID-GBIF-Occurrence="2572183357" collectionCode="UWBM" pageId="34" pageNumber="1566" specimenCode="UWBM 107613" specimenCount="1">
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<specimenCode box="[983,1153,1177,1201]" pageId="34" pageNumber="1566">UWBM 107613</specimenCode>
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, B8769, L 18.0, W 8.0
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</materialsCitation>
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;
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<materialsCitation ID-GBIF-Occurrence="2572183365" box="[392,805,1211,1235]" collectionCode="UWBM" pageId="34" pageNumber="1566" specimenCode="UWBM 107614" specimenCount="1">
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<specimenCode box="[392,558,1211,1235]" pageId="34" pageNumber="1566">UWBM 107614</specimenCode>
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, B8772, L (11.9), W 7.4
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</materialsCitation>
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;
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<materialsCitation ID-GBIF-Occurrence="2572183368" collectionCode="UWBM" pageId="34" pageNumber="1566" specimenCode="UWBM 107615" specimenCount="1">
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<specimenCode box="[814,980,1211,1235]" pageId="34" pageNumber="1566">UWBM 107615</specimenCode>
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, B8769, L (22.0), W (13.7)
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</materialsCitation>
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;
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<materialsCitation ID-GBIF-Occurrence="2572183358" box="[264,694,1246,1270]" collectionCode="UWBM" pageId="34" pageNumber="1566" specimenCode="UWBM 107616" specimenCount="1">
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<specimenCode box="[264,430,1246,1270]" pageId="34" pageNumber="1566">UWBM 107616</specimenCode>
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, B8769, L (11.9), W (6.0)
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</materialsCitation>
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;
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<materialsCitation ID-GBIF-Occurrence="2572183364" box="[705,1152,1246,1270]" collectionCode="MUSM" pageId="34" pageNumber="1566" specimenCode="INV 263" specimenCount="1">
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<collectionCode box="[705,778,1246,1270]" pageId="34" pageNumber="1566">MUSM</collectionCode>
|
||
<specimenCode box="[785,874,1246,1270]" pageId="34" pageNumber="1566">INV 263</specimenCode>
|
||
, B8769, L (33.0), W (15.8)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation ID-GBIF-Occurrence="2572183361" box="[160,607,1281,1305]" collectionCode="MUSM" pageId="34" pageNumber="1566" specimenCode="INV 264" specimenCount="1">
|
||
<collectionCode box="[160,233,1281,1305]" pageId="34" pageNumber="1566">MUSM</collectionCode>
|
||
<specimenCode box="[240,329,1281,1305]" pageId="34" pageNumber="1566">INV 264</specimenCode>
|
||
, B8772, L (31.0), W (16.5)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation ID-GBIF-Occurrence="2565890729" box="[618,1026,1281,1305]" collectionCode="MUSM" pageId="34" pageNumber="1566" specimenCode="INV 265" specimenCount="1">
|
||
<collectionCode box="[618,691,1281,1305]" pageId="34" pageNumber="1566">MUSM</collectionCode>
|
||
<specimenCode box="[698,787,1281,1305]" pageId="34" pageNumber="1566">INV 265</specimenCode>
|
||
, B8769, L 15.9, W 8.1.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph blockId="34.[160,1156,1370,1430]" box="[160,292,1370,1396]" pageId="34" pageNumber="1566">
|
||
<emphasis bold="true" box="[160,292,1370,1396]" italics="true" pageId="34" pageNumber="1566">Occurrence</emphasis>
|
||
</paragraph>
|
||
<paragraph blockId="34.[160,1156,1370,1430]" box="[160,1156,1406,1430]" pageId="34" pageNumber="1566">
|
||
Lower Paleogene, Cuenca Member, Caballas Formation, East Pisco Basin, southern
|
||
<collectingCountry box="[1099,1153,1406,1430]" name="Peru" pageId="34" pageNumber="1566">Peru</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |