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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.901.39432" ID-GBIF-Dataset="654566e8-947b-4dfe-8fbc-baea3d254fa8" ID-PMC="PMC6965333" ID-Pensoft-Pub="1313-2970-901-1" ID-Pensoft-UUID="D8C38C1C3844549F8BEC7C1D884CDF8E" ID-PubMed="31975782" ModsDocID="1313-2970-901-1" checkinTime="1578615027095" checkinUser="pensoft" docAuthor="Balkenohl, Michael" docDate="2020" docId="9A02A9FC7D495695BB7DB3E4F23798EE" docLanguage="en" docName="ZooKeys 901: 1-81" docOrigin="ZooKeys 901" docSource="http://dx.doi.org/10.3897/zookeys.901.39432" docTitle="Salcedia coquilhati Alluaud 1932" docType="treatment" docVersion="4" id="D8C38C1C3844549F8BEC7C1D884CDF8E" lastPageNumber="1" masterDocId="D8C38C1C3844549F8BEC7C1D884CDF8E" masterDocTitle="A genus in disguise. Revision of the genus Salcedia Fairmaire, 1899 with descriptions of nine new species (Coleoptera, Carabidae, Scaritinae, Salcediini)" masterLastPageNumber="81" masterPageNumber="1" pageNumber="1" updateTime="1668168267548" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>A genus in disguise. Revision of the genus Salcedia Fairmaire, 1899 with descriptions of nine new species (Coleoptera, Carabidae, Scaritinae, Salcediini)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Balkenohl, Michael</mods:namePart>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>901</mods:number>
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<mods:start>1</mods:start>
<mods:end>81</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.901.39432</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.901.39432</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-901-1</mods:identifier>
<mods:identifier type="Pensoft-UUID">D8C38C1C3844549F8BEC7C1D884CDF8E</mods:identifier>
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<treatment ID-GBIF-Taxon="161671753" LSID="urn:lsid:plazi:treatment:9A02A9FC7D495695BB7DB3E4F23798EE" httpUri="http://treatment.plazi.org/id/9A02A9FC7D495695BB7DB3E4F23798EE" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="9A02A9FC-7D49-5695-BB7D-B3E4F23798EE" authority="Alluaud, 1932" authorityName="Alluaud" authorityYear="1932" class="Insecta" family="Carabidae" genus="Salcedia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salcedia coquilhati" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="coquilhati">Salcedia coquilhati Alluaud, 1932</taxonomicName>
<figureCitation captionStart="Figures 813" captionStartId="F7" captionText="Figures 8 - 13. Salcedia species, habitus, dorsal view, holotype 8 S. perrieri Fairmaire 9 S. unifoveata sp. nov. 10 S. faillei sp. nov. 11 S. coquilhati Alluaud 12 S. elongata Alluaud 13 S. africana (Britton)." figureDoi="10.3897/zookeys.901.39432.figures8-13" httpUri="https://binary.pensoft.net/fig/370608" pageId="0" pageNumber="1">Figs 11</figureCitation>
<figureCitation captionStart="Figures 2635" captionStartId="F10" captionText="Figures 26 - 35. Salcedia species, pronotum, dorsal view 26 S. perrieri Fairmaire 27 S. unifoveata sp. nov. 28 S. faillei sp. nov. 29 S. coquilhati Alluaud 30 S. elongata Alluaud 31 S. africana (Britton) 32 S. schoutedeni Alluaud 33 S. nigeriensis Alluaud 34 S. putzeysi (Oberthuer) 35 S. matsumotoi sp. nov." figureDoi="10.3897/zookeys.901.39432.figures26-35" httpUri="https://binary.pensoft.net/fig/370611" pageId="0" pageNumber="1">, 29</figureCitation>
<figureCitation captionStart="Figures 5257" captionStartId="F13" captionText="Figures 52 - 57. Salcedia species, male genitalia, dorsal view of aedeagus (a) and parameres (b, c), lateral view (d). Small sketches in black symbolising the apex in cross section 52 S. perrieri Fairmaire 53 S. unifoveata sp. nov. 54 S. faillei sp. nov. 55 S. coquilhati Alluaud 56 S. elongata Alluaud 57 S. africana (Britton)." figureDoi="10.3897/zookeys.901.39432.figures52-57" httpUri="https://binary.pensoft.net/fig/370614" pageId="0" pageNumber="1">, 55</figureCitation>
<figureCitation captionStart="Figures 6876" captionStartId="F16" captionText="Figures 68 - 76. Salcedia species, female coxostyli (gonopod IX) and laterotergite IX (epipleutite IX) 68 S. perrieri Fairmaire 69 S. unifoveata sp. nov. 70 S. coquilhati Alluaud 71 S. elongata Alluaud 72 S. africana (Britton) 73 S. schoutedeni Alluaud 74 S. nigeriensis Alluaud 75 S. putzeysi (Oberthuer) 76 S. matsumotoi sp. nov." figureDoi="10.3897/zookeys.901.39432.figures68-76" httpUri="https://binary.pensoft.net/fig/370617" pageId="0" pageNumber="1">, 70</figureCitation>
<figureCitation captionStart="Figure 84" captionStartId="F18" captionText="Figure 84. Map of tropical Africa providing an overview of the occurrence of all African Salcedia species described to date (recorded localities plotted). 1 (black S. perrieri Fairmaire; 2 (white) S. unifoveata sp. nov.; 3 (black circle) S. faillei sp. nov.; 4 (red) S. coquilhati Alluaud; 5 (yellow) S. elongata Alluaud; 6 (blue) S. africana (Britton); 7 (orange) S. utetea sp. nov.; 8 (grey) S. procera sp. nov.; 9 (light green) S. robusta sp. nov.; 10 (turquoise) S. schoutedeni Alluaud; 11 (purple) S. lukulua sp. nov.; 12 (dark green) S. tuberculata sp. nov.; 13 (brown) S. baroensis sp. nov.; 14 (light blue) S. nigeriensis Alluaud; 16 (dark blue) S. putzeysi (Oberthuer); 17 (black) S. matsumotoi sp. nov.;? in a circle (twice): collection locality of a single female Salcedia specimen of uncertain species. Basic schematic map of Africa taken from SimpleMappr. net." figureDoi="10.3897/zookeys.901.39432.figure84" httpUri="https://binary.pensoft.net/fig/370619" pageId="0" pageNumber="1">, 84</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Alluaud" authorityYear="1932" class="Insecta" family="Carabidae" genus="Salcedia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salcedia coquilhati" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="coquilhati">Salcedia coquilhati</taxonomicName>
Alluaud, 1932: 3;
<bibRefCitation author="Csiki, E" journalOrPublisher="Entomologist's monthly magazine, May" pageId="0" pageNumber="1" refId="B21" refString="Csiki, E, 1933. Coleopterorum Catalogus auspiciis et auxilio. W. Junk editus a S. Schenkling. 127: Carabidae: Carabinae III, Corrigenda et Addenda. W. Junk, Berlin, 623-641." title="Coleopterorum Catalogus auspiciis et auxilio. W. Junk editus a S. Schenkling. 127: Carabidae: Carabinae III, Corrigenda et Addenda. W. Junk, Berlin, 623 - 641." year="1933">Csiki 1933</bibRefCitation>
: 641;
<bibRefCitation author="Alluaud, C" journalOrPublisher="Afra" pageId="0" pageNumber="1" pagination="19 - 20" refId="B4" refString="Alluaud, C, 1935. Note diverses. Localites nouvelles. Afra 9: 19 - 20" title="Note diverses. Localites nouvelles." volume="9" year="1935">Alluaud 1935</bibRefCitation>
: 19;
<bibRefCitation author="Burgeon, L" journalOrPublisher="Entomologist's monthly magazine, May" pageId="0" pageNumber="1" refId="B19" refString="Burgeon, L, 1935. Catalogues raisonnes de la faune entomologique du Congo Belge. Coleopteres - Carabides. Annales du Musee du Congo Belge, Zoologie, Serie III. Section II. Tome II: 135-257." title="Catalogues raisonnes de la faune entomologique du Congo Belge. Coleopteres - Carabides. Annales du Musee du Congo Belge, Zoologie, Serie III. Section II. Tome II: 135 - 257." year="1935">Burgeon 1935</bibRefCitation>
: 159;
<bibRefCitation author="Reichardt, H" journalOrPublisher="Revista Brasileira Entomologia" pageId="0" pageNumber="1" pagination="99 - 110" refId="B38" refString="Reichardt, H, 1975. The genera of Salcediina (Coleoptera, Carabidae, Scaritini). Revista Brasileira Entomologia 19: 99 - 110" title="The genera of Salcediina (Coleoptera, Carabidae, Scaritini)." volume="19" year="1975">Reichardt 1975</bibRefCitation>
: 103; 1932,
<bibRefCitation author="Dostal, A" journalOrPublisher="Zeitschrift der Arbeitsgemeinschaft Oesterreichischer Entomologen" pageId="0" pageNumber="1" pagination="117 - 121" refId="B24" refString="Dostal, A, 1993. Beschreibung einer neuen Gattung aus Vietnam: Androzelma gen. nov. (Coleoptera, Carabidae, Scaritinae: Salcediini). Zeitschrift der Arbeitsgemeinschaft Oesterreichischer Entomologen 45: 117 - 121" title="Beschreibung einer neuen Gattung aus Vietnam: Androzelma gen. nov. (Coleoptera, Carabidae, Scaritinae: Salcediini)." volume="45" year="1993">Dostal 1993</bibRefCitation>
: 121;
<bibRefCitation author="Lorenz, W" journalOrPublisher="Printed by the author, Tutzing" pageId="0" pageNumber="1" refId="B34" refString="Lorenz, W, 2005. Systematic list of extant Ground Beetles of the World (InsectaColeoptera &quot;Geadephaga&quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodidae), Second Edition. Printed by the author, Tutzing" title="Systematic list of extant Ground Beetles of the World (InsectaColeoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodidae), Second Edition." year="2005">Lorenz 2005</bibRefCitation>
: 155.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype:</emphasis>
♂, with labels and data: white, printed and handwritten
<normalizedToken originalValue="“MUSÉE">&quot;MUSEE</normalizedToken>
DU CONGO Coquilhatville
<normalizedToken originalValue="XII">-XII-</normalizedToken>
1924 DrH.Schouteden&quot; / red, glued
<normalizedToken originalValue="“TYPE”">&quot;TYPE&quot;</normalizedToken>
/ white printed and handwritten &quot;R.
<normalizedToken originalValue="DÉT">DET</normalizedToken>
. 2019 f&quot; / white, black framed, handwritten and printed &quot;
<emphasis italics="true" pageId="0" pageNumber="1">Salcedia coquilhati</emphasis>
All. Alluaud det. 19&quot; / red, black double framed
<normalizedToken originalValue="“TYPE”">&quot;TYPE&quot;</normalizedToken>
/ white, black printed &quot;RMCA ENT 000019377&quot; and square barcode (MRACT).
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratype</emphasis>
: 1 ♀, with labels and data: white, printed and handwritten
<normalizedToken originalValue="“MUSÉE">&quot;MUSEE</normalizedToken>
DU CONGO Coquilhatville -XII- 1924 DrH.Schouteden&quot; / white, handwritten &quot;
<emphasis italics="true" pageId="0" pageNumber="1">Salcedia coquilhati</emphasis>
n.sp. Ch. Alluaud det.&quot; / red, black double framed
<normalizedToken originalValue="“PARATYPE”">&quot;PARATYPE&quot;</normalizedToken>
(MRACT).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remark.</paragraph>
<paragraph pageId="0" pageNumber="1">In the holotype the left anterior leg is missing and the hind body shows some slight damages. In the paratype the left hind leg is missing.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Additional material.</paragraph>
<paragraph pageId="0" pageNumber="1">
1 ♂, 1 ♀, 1 spec., Eala I-1935 and XI-1934 J.
<normalizedToken originalValue="Ghesquière">Ghesquiere</normalizedToken>
<normalizedToken originalValue="MUSÉE">MUSEE</normalizedToken>
DU CONGO; 1 ♂, 1 ♀, 2 specs., Irebu 30-XII-1920 DrH.Schouteden
<normalizedToken originalValue="MUSÉE">MUSEE</normalizedToken>
DU CONGO (MRACT, CBB); 10 specs., COLL. MUS. Congo Ubangi: Nouvelle-Anvers 9-XII-1952 P. Basilewsky / A la
<normalizedToken originalValue="lumière">lumiere</normalizedToken>
(MRACT, CBB); 1 spec., I.R.S.A.-MUS.R.A.C. Equateur: terr. Bikoro, Mabali IX.1959 N. Leleup /
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
<normalizedToken originalValue="inondée">inondee</normalizedToken>
<normalizedToken originalValue="à">a</normalizedToken>
la
<normalizedToken originalValue="lumière">lumiere</normalizedToken>
(MRACT); 1 spec., Coquilhatville - a table le soir - G Juillet 1909 /
<normalizedToken originalValue="MUSÉE">MUSEE</normalizedToken>
DU CONGO Voyage de S.A.R. le Prince Albert 1909 (MRACT); 1 spec.,
<normalizedToken originalValue="MUSÉE">MUSEE</normalizedToken>
DU CONGO Stanleyville a Coquilhatville Dr.
<normalizedToken originalValue="Géranrd">Geranrd</normalizedToken>
. (MRACT); 1 spec.,
<normalizedToken originalValue="MUSÉE">MUSEE</normalizedToken>
DU CONGO Kai Bumba DrH.Schouteden 11-X-1920 (MRACT); 1 spec., COLL. MUS. CONGO Tshuapa: Bamania XII-1952 R. P. Hulstaert (MRACT); 1 spec., Congo Belge, P.N.G. Miss. H. De Saeger Mabanga 1949-1952
<normalizedToken originalValue="Réc">Rec</normalizedToken>
. H. De Saeger (MRACT); 1 spec., Congo belge, P.N.G. Miss. H. De Saeger I/b/3
<normalizedToken originalValue="">''</normalizedToken>
, Mabanga 14-IV-1950
<normalizedToken originalValue="Réc">Rec</normalizedToken>
. H. De Saeger.458 /
<taxonomicName authorityName="Fairmaire" authorityYear="1899" class="Insecta" family="Carabidae" genus="Salcedia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salcedia" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Salcedia</taxonomicName>
(MRACT); 1 spec., MUSEUM PARIS Afrique
<normalizedToken originalValue="Équatoriale">Equatoriale</normalizedToken>
Fr. BRAZZAVILLE
<normalizedToken originalValue="ANDRÉ">ANDRE</normalizedToken>
GIDE 1925 / AOUT /
<taxonomicName authorityName="Alluaud" authorityYear="1932" class="Insecta" family="Carabidae" genus="Salcedia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salcedia coquilhati" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="coquilhati">Salcedia coquilhati</taxonomicName>
Alluaud det. 1934 (MNHN).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">A small sized species, with long-ovoid outline of the elytra with maximum width at posterior third. The pseudohumerus is obtuse angled and not dentate. The antennomeres are elongate. Distinguished from all other species of the genus by the extraordinary transverse pronotum which shows at the lateral margin five large transverse pits instead of six pits present in all other species, and the head exhibits a reniform outline in dorsal view.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="redescription">
<paragraph pageId="0" pageNumber="1">Redescription.</paragraph>
<paragraph pageId="0" pageNumber="1">
Measurements in Table
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Measurements and ratios of Salcedia species. N = number of specimens measured; x ̄ = arithmetic mean; &quot; term used &quot; refers to the shape of the elytra." httpUri="http://table.plazi.org/id/FDDA58A1B347C9F0593A738CF707345F" pageId="0" pageNumber="1" tableUuid="FDDA58A1B347C9F0593A738CF707345F">1</tableCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Colour and surface:</emphasis>
Melaneous, opaque; top of carinae griseous; mandibles and legs fuscous, antennae and palpi hinnuleous.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Head:</emphasis>
Two-thirds of width of pronotum. Outline reniform. Clypeus wide, slightly but distinctly convex anteriorly, fused with clypeal wings, separated from supra-antennal plates by obtuse notches, frons with two flattened paramedian carinae, joining anteriorly into a convex bulge not clearly separated from frons, with two small glossy teeth bilaterally anterior to central bulge, with two parallel carinae paralaterally near base; frons and clypeus separated from supra-antennal and supraorbital plates by deep broad furrows, each furrow with moderately deep circular pit between supra-antennal plate and clypeus; supra-antennal and supraorbital plates distinctly margined, supra-antennal plates slightly vaulted, margin of supraorbital plates conspicuously raised. Base emarginated at middle, laterally rounded-off and constricted to neck without angle. Eyes large, convex, genae slightly convex, both of them clearly visible from above, with nearly circular shape in lateral view. Antenna with segments five to ten elongate (L/W 1.26), densely pubescent, segments two to four scarcely pubescent, scapus with minute longitudinal reticulation. Labrum straight anteriorly. Mandible moderately short, wide, slightly arcuate at apex. Mentum small, epilobes wide, projecting and nearly rectangular anteriorly, completely margined, surface coriaciate-like with flat impressions.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Pronotum</emphasis>
(Fig.
<figureCitation captionStart="Figures 2635" captionStartId="F10" captionText="Figures 26 - 35. Salcedia species, pronotum, dorsal view 26 S. perrieri Fairmaire 27 S. unifoveata sp. nov. 28 S. faillei sp. nov. 29 S. coquilhati Alluaud 30 S. elongata Alluaud 31 S. africana (Britton) 32 S. schoutedeni Alluaud 33 S. nigeriensis Alluaud 34 S. putzeysi (Oberthuer) 35 S. matsumotoi sp. nov." figureDoi="10.3897/zookeys.901.39432.figures26-35" httpUri="https://binary.pensoft.net/fig/370611" pageId="0" pageNumber="1">29</figureCitation>
): Outline rectangular, conspicuously transverse, nearly half times wider than long. Lateral margin straight but converging anteriorly, maximum width at posterior third, slightly rounded anteriorly and to posterior angles. Lateral margin distinctly crenulated, with nine tubercles, with distinct emargination at posterior angles. Base with wide tubercle laterally. Disc with two raised paramedian irregularly sinuate carinae parallel to median line, joining at anterior margin, with median line indistinct, with two additional shorter carinae bilaterally, the paralateral one joining with the paramedian carinae at base and forming tooth-like tubercle pointing posteriorly, extended anteriorly as less raised paralateral carina. With one distinctly long outer lateral carina running up to anterior margin. Lateral margin and space between lateral margin and lateral carina wing-like bent up, with five large transverse pits.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Elytron:</emphasis>
Slightly convex in anterior half (lateral view), convex in frontal view. Outline in posterior five sixths long-ovoid, maximum width at middle. Pseudohumerus sub-rectangular, with small tubercle-like tooth. Apex acute at suture. Disc with interneur six crenulated, interneur two running up to apex as distinctly convex line, conspicuously raised; interneur four running in parallel to interneur six, reaching base, not reaching apex. Interneur four and six with two to three indistinct rows of pits becoming transverse apically.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Hind wings:</emphasis>
Fully developed.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Lower surface:</emphasis>
Pronotum with surface of antennal channel with irregular reticulation. Pseudoepipleura with two rows of serial pits, lateral margin of elytron smooth. Metepisternum elongate, with broad longitudinal groove. Metasternum and last five abdominal sternites with numerous pits, regularly arranged at carinae and margins. Sternum four to six distinctly sulcate.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Legs:</emphasis>
Profemora with dorsal surface indistinctly reticulated. Protibia with terminal spine arcuate at apex, laterally with three teeth, dorsally and ventrally with two carinae. Movable spur arcuate, acute, length nearly as half as first tarsomere. First tarsomeres distinctly elongated, almost as long as tarsomeres two to five together.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">External sexual dimorphism:</emphasis>
Not observed.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Male genitalia</emphasis>
(Fig.
<figureCitation captionStart="Figures 5257" captionStartId="F13" captionText="Figures 52 - 57. Salcedia species, male genitalia, dorsal view of aedeagus (a) and parameres (b, c), lateral view (d). Small sketches in black symbolising the apex in cross section 52 S. perrieri Fairmaire 53 S. unifoveata sp. nov. 54 S. faillei sp. nov. 55 S. coquilhati Alluaud 56 S. elongata Alluaud 57 S. africana (Britton)." figureDoi="10.3897/zookeys.901.39432.figures52-57" httpUri="https://binary.pensoft.net/fig/370614" pageId="0" pageNumber="1">55</figureCitation>
): Median lobe short, stout, in dorsal view straight, in lateral view straight, with a few fine pili in apical third, oroficium small, apex circular in cross-section, slightly turned dorsally. Endophallus with two elongated groups of microtrichia, groups as long as one fifth of median lobe. Dorsal paramere sinuate in apical half, lateral and basal apophysis slender; both parameres slightly distorted.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Female genitalia</emphasis>
(Fig.
<figureCitation captionStart="Figures 6876" captionStartId="F16" captionText="Figures 68 - 76. Salcedia species, female coxostyli (gonopod IX) and laterotergite IX (epipleutite IX) 68 S. perrieri Fairmaire 69 S. unifoveata sp. nov. 70 S. coquilhati Alluaud 71 S. elongata Alluaud 72 S. africana (Britton) 73 S. schoutedeni Alluaud 74 S. nigeriensis Alluaud 75 S. putzeysi (Oberthuer) 76 S. matsumotoi sp. nov." figureDoi="10.3897/zookeys.901.39432.figures68-76" httpUri="https://binary.pensoft.net/fig/370617" pageId="0" pageNumber="1">70</figureCitation>
): Coxostylus less sclerotised, broadened just anterior base, distinctly curved dorsally and slightly medially, slender in apical third, at end of basal third with one shorter strong and three slender nematiform setae laterally, with another slender seta at middle, with one long strong nematiform seta ventrally.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Variation:</emphasis>
On the elytron between the interneurs four and six, the rows of pits vary in the number of pits and rows. In some of the specimens the labrum is indistinctly convex.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution</paragraph>
<paragraph pageId="0" pageNumber="1">
(Fig.
<figureCitation captionStart="Figure 84" captionStartId="F18" captionText="Figure 84. Map of tropical Africa providing an overview of the occurrence of all African Salcedia species described to date (recorded localities plotted). 1 (black S. perrieri Fairmaire; 2 (white) S. unifoveata sp. nov.; 3 (black circle) S. faillei sp. nov.; 4 (red) S. coquilhati Alluaud; 5 (yellow) S. elongata Alluaud; 6 (blue) S. africana (Britton); 7 (orange) S. utetea sp. nov.; 8 (grey) S. procera sp. nov.; 9 (light green) S. robusta sp. nov.; 10 (turquoise) S. schoutedeni Alluaud; 11 (purple) S. lukulua sp. nov.; 12 (dark green) S. tuberculata sp. nov.; 13 (brown) S. baroensis sp. nov.; 14 (light blue) S. nigeriensis Alluaud; 16 (dark blue) S. putzeysi (Oberthuer); 17 (black) S. matsumotoi sp. nov.;? in a circle (twice): collection locality of a single female Salcedia specimen of uncertain species. Basic schematic map of Africa taken from SimpleMappr. net." figureDoi="10.3897/zookeys.901.39432.figure84" httpUri="https://binary.pensoft.net/fig/370619" pageId="0" pageNumber="1">84</figureCitation>
). Known from several localities along the course of the Congo River and at the mouth of the river Tshuapa. Some of the specimens were collected at light. The historic name Coquilhatville refers to the city today called Mbandaka, that of Stanleyville to Kisangani, and Brazzaville to Kinshasa.
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