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<mods:title id="630F1A63B521ACF90D5FF89FD14812BE">Re-appraisal of thylacocephalans (Euarthropoda, Thylacocephala) from the Jurassic La Voulte-sur-Rhône Lagerstätte</mods:title>
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<mods:namePart id="DEEE5E9F6A980E8E886C86001EAE6A3E">Laville, Thomas</mods:namePart>
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<mods:affiliation id="94B3C4AE486FA7F7CFF0CA188A46A8A5">Muséum national dHistoire naturelle, Centre de Recherche en Paléontologie - Paris (CR 2 P, UMR 7207), Sorbonne Université-MNHN-CNRS, 57 rue Cuvier, 75005 Paris, France. &amp; Biogéosciences, UMR 6282 CNRS, Université de Bourgogne, 6 boulevard Gabriel, 2100 Dijon, France.</mods:affiliation>
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<mods:namePart id="A5C6FC29904BC34675E7AEA05F0B0900">Forel, Marie-Béatrice</mods:namePart>
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<mods:affiliation id="844788DAA6120F6EF4261F3A0A6364A1">Muséum national dHistoire naturelle, Centre de Recherche en Paléontologie - Paris (CR 2 P, UMR 7207), Sorbonne Université-MNHN-CNRS, 57 rue Cuvier, 75005 Paris, France.</mods:affiliation>
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<mods:affiliation id="1940F2505E1F5D3DB1EEF44A3B1658B4">Muséum national dHistoire naturelle, Centre de Recherche en Paléontologie - Paris (CR 2 P, UMR 7207), Sorbonne Université-MNHN-CNRS, 57 rue Cuvier, 75005 Paris, France.</mods:affiliation>
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<taxonomicName id="8E864D47997CFFD82D33FEC5FA7AF739" authority="Laville &amp; Forel &amp; Charbonnier, 2023" authorityName="Laville &amp; Forel &amp; Charbonnier" authorityYear="2023" box="[603,887,265,291]" family="Austriocarididae" genus="Austriocaris" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="secretanae" status="sp. nov.">
<emphasis id="7BF2EAD6997CFFD82D33FEC5FA7AF739" bold="true" box="[603,887,265,291]" italics="true" pageId="10" pageNumber="11">Austriocaris secretanae</emphasis>
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<subSubSection id="019C654F997CFFD82D8FFE9CFA40F771" box="[743,845,336,363]" pageId="10" pageNumber="11" type="description">
<paragraph id="493936C4997CFFD82D8FFE9CFA40F771" blockId="10.[356,1231,265,363]" box="[743,845,336,363]" pageId="10" pageNumber="11">
<figureCitation id="D1BD2A41997CFFD82D8FFE9CFA40F771" box="[743,845,336,363]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="12.[189,231,1716,1742]" captionStartId-1="13.[187,230,1704,1730]" captionTargetBox-0="[269,1317,265,1679]" captionTargetBox-1="[189,1398,268,1649]" captionTargetId-0="figure-12@12.[269,1318,265,1679]" captionTargetId-1="figure-11@13.[189,1398,268,1649]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionText-0="Fig. 3. Austriocaris secretanae sp. nov. AF. Holotype MNHN.F.R50963. A. General view (part). B. General view (counter-part). C. Drawing (part). D. Drawing (counter-part). E. Close-up of pores. F. Close-up of muscles of raptorial appendages. G. Paratype MNHN.F.A29257. HI. Paratype MNHN.F.R06203. H. General view. I. Drawing. Abbreviations: am = anterior margin; ce = compound eye; dlc = dorso-lateral carina; dm = dorsal midline; g = gills; mlc = medio-lateral carina; os = oval structure; p15 = podomeres of raptorial appendages; pm = posterior margin; po = pores; ra 13 = raptorial appendages; sp = spines; st = stalk; t = posterior trunk; ta 8 = posterior trunk appendage 8; tp 111 = pleural parts of posterior trunk; vlg = ventro-lateral groove; vm = ventral margin. Scale bars: AD, GI = 10 mm; E = 1 mm; F = 5 mm. Photos: AB, G: L. Cazes; EF: T. Laville. H: C. Haug &amp; Joachim T. Haug." captionText-1="Fig. 4. Austriocaris secretanae sp. nov. A, C, F. Paratype MNHN.F.A29286. A. General view. C. Closeup of ornamentation. F. Close-up of compound eye. B. Paratype MNHN.F.R50971. D, G. Paratype MNHN.F.A29249. D. General view. G. Drawing. EH. Paratype MNHN.F.A84029. E. General view. H. Close-up of posterior trunk appendages. IJ. Paratype MSNM i7591. I. General view. J. Close-up of raptorial appendages. Abbreviations: cs = circular structure; dm = dorsal midline; os = oval structure; pm = posterior margin; t = posterior trunk; tp111 = pleural parts of posterior trunk appendages. Scale bars: AB, DE, G, I = 10 mm; C = 1 mm; F = 400 µm; H, J = 5 mm. Photos: A, CF, HJ: T. Laville; B: L. Cazes." figureDoi-0="http://doi.org/10.5281/zenodo.8413508" figureDoi-1="http://doi.org/10.5281/zenodo.8413512" httpUri-0="https://zenodo.org/record/8413508/files/figure.png" httpUri-1="https://zenodo.org/record/8413512/files/figure.png" pageId="10" pageNumber="11">Figs 34</figureCitation>
</paragraph>
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<paragraph id="493936C4997CFFD82FD5FE5EF87BF7CA" blockId="10.[189,1399,402,465]" pageId="10" pageNumber="11">
<treatmentCitationGroup id="699611EA997CFFD82FD5FE5EF87BF7CA" pageId="10" pageNumber="11">
<taxonomicName id="8E864D47997CFFD82FD5FE5EF882F7B6" authorityName="Van Straelen" authorityYear="1923" box="[189,399,402,428]" family="Dollocarididae" genus="Dollocaris" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="ingens">
<emphasis id="7BF2EAD6997CFFD82FD5FE5EF882F7B6" box="[189,399,402,428]" italics="true" pageId="10" pageNumber="11">Dollocaris ingens</emphasis>
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<treatmentCitation id="C82710D5997CFFD82EC6FE5FFBC2F7B6" author="Secretan S." box="[430,719,402,429]" page="381" pageId="10" pageNumber="11" year="1985">
<bibRefCitation id="2D174B35997CFFD82EC6FE5FFBC2F7B6" author="Secretan S." box="[430,719,402,429]" pageId="10" pageNumber="11" pagination="381 - 389" refId="ref29069" refString="Secretan S. 1985. Conchyliocarida, a class of fossil crustaceans: relationships to Malacostraca and postulated behaviour. Transactions of the Royal Society of Edinburgh: Earth Sciences 76 (2 - 3): 381 - 389. https: // doi. org / 10.1017 / S 0263593300010592" type="journal article" year="1985">Secrétan 1985: 381386</bibRefCitation>
</treatmentCitation>
, fig. 5ab. —
<treatmentCitation id="C82710D5997CFFD82C1EFE5EFDC1F7B7" author="Charbonnier S. &amp; Vannier J. &amp; Hantzpergue P. &amp; Gaillard C." box="[886,1228,402,429]" page="119" pageId="10" pageNumber="11" year="2010">
<bibRefCitation id="2D174B35997CFFD82C1EFE5EFDC1F7B7" author="Charbonnier S. &amp; Vannier J. &amp; Hantzpergue P. &amp; Gaillard C." box="[886,1228,402,429]" pageId="10" pageNumber="11" pagination="111 - 132" refId="ref24994" refString="Charbonnier S., Vannier J., Hantzpergue P. &amp; Gaillard C. 2010. Ecological significance of the arthropod fauna from the Jurassic (Callovian) La Voulte Lagerstatte. Acta Palaeontologica Polonica 55 (1): 111 - 132. https: // doi. org / 10.4202 / app. 2009.0036" type="journal article" year="2010">
Charbonnier
<emphasis id="7BF2EAD6997CFFD82B78FE5FFD49F7B6" box="[1040,1092,402,428]" italics="true" pageId="10" pageNumber="11">et al</emphasis>
. 2010: 119
</bibRefCitation>
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, fig. 6d; 2014: 375, fig. 4a.
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</paragraph>
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<paragraph id="493936C4997CFFD82FD5FE34F83BF408" blockId="10.[189,310,504,530]" box="[189,310,504,530]" pageId="10" pageNumber="11">
<heading id="127181A8997CFFD82FD5FE34F83BF408" bold="true" box="[189,310,504,530]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="7BF2EAD6997CFFD82FD5FE34F83BF408" bold="true" box="[189,310,504,530]" pageId="10" pageNumber="11">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="493936C4997CFFD82FD5FDE4FBFEF493" blockId="10.[189,1399,551,649]" pageId="10" pageNumber="11">
<taxonomicName id="8E864D47997CFFD82FD5FDE4F843F458" authorityName="Glaessner" authorityYear="1931" box="[189,334,552,578]" family="Austriocarididae" genus="Austriocaris" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="7BF2EAD6997CFFD82FD5FDE4F843F458" box="[189,334,552,578]" italics="true" pageId="10" pageNumber="11">Austriocaris</emphasis>
</taxonomicName>
with stout and long raptorial appendages, the third pair having spines on the second and third podomeres of the distal part, and a posterior trunk consisting of 12 segments bearing rod-like appendages. Absence of ventro-lateral structure.
</paragraph>
</subSubSection>
<subSubSection id="019C654F997CFFD82FD5FD7DFB1DF507" pageId="10" pageNumber="11" type="etymology">
<paragraph id="493936C4997CFFD82FD5FD7DF849F4D1" blockId="10.[189,324,689,715]" box="[189,324,689,715]" pageId="10" pageNumber="11">
<heading id="127181A8997CFFD82FD5FD7DF849F4D1" bold="true" box="[189,324,689,715]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="7BF2EAD6997CFFD82FD5FD7DF849F4D1" bold="true" box="[189,324,689,715]" pageId="10" pageNumber="11">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="493936C4997CFFD82FD5FD2CFB1DF507" blockId="10.[189,1399,736,797]" pageId="10" pageNumber="11">In reference to Sylvie Secrétan (CNRS, Paris), for her important contribution to the study of La Voulte-sur-Rhône thylacocephalans.</paragraph>
</subSubSection>
<subSubSection id="019C654F997CFFD82FD5FC89FC28F221" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="493936C4997CFFD82FD5FC89F8A7F545" blockId="10.[189,426,837,863]" box="[189,426,837,863]" pageId="10" pageNumber="11">
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<emphasis id="7BF2EAD6997CFFD82FD5FC89F8A7F545" bold="true" box="[189,426,837,863]" pageId="10" pageNumber="11">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="493936C4997CFFD82F82FCB9FC7AF5A9" blockId="10.[189,1399,885,947]" pageId="10" pageNumber="11">
<materialsCitation id="F9EE3C99997CFFD82F82FCB9FC7AF5A9" collectionCode="MNHN" collectorName="La Boissine &amp; La Voulte-sur-Rhone" country="France" location="Ardeche" pageId="10" pageNumber="11" specimenCode="R50963" specimenCount="1" typeStatus="holotype">
<emphasis id="7BF2EAD6997CFFD82F82FCB9F851F595" bold="true" box="[234,348,885,911]" pageId="10" pageNumber="11">
<typeStatus id="963D8866997CFFD82F82FCB9F851F595" box="[234,348,885,911]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="31917654997CFFD82FD5FC54F83CF5A8" box="[189,305,920,946]" name="France" pageId="10" pageNumber="11">FRANCE</collectingCountry>
<collectorName id="E4735312997CFFD82E20FC55F8DBF5A8" box="[328,470,920,947]" pageId="10" pageNumber="11">La Boissine</collectorName>
,
<collectorName id="E4735312997CFFD82E89FC55FBD4F5A8" box="[481,729,920,947]" pageId="10" pageNumber="11">La Voulte-sur-Rhône</collectorName>
,
<location id="4C59601F997CFFD82D8AFC54FA44F5A8" LSID="urn:lsid:plazi:treatment:C12F87D2997CFFDC2D33FEC5FA8CF418:4C59601F997CFFD82D8AFC54FA44F5A8" box="[738,841,920,946]" country="France" name="Ardeche" pageId="10" pageNumber="11">Ardèche</location>
;
<geologicalTimeScale id="3BA6B66F997CFFD82C3DFC54FD9CF5A8" box="[853,1169,920,946]" pageId="10" pageNumber="11">Callovian, Middle Jurassic</geologicalTimeScale>
;
<collectionCode id="2F97AE01997CFFD82BF4FC54FDF6F5A9" box="[1180,1275,920,947]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="10" pageNumber="11">MNHN</collectionCode>
.F.
<specimenCode id="19209EBF997CFFD82A7BFC54FC7EF5A9" box="[1299,1395,920,947]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="10" pageNumber="11">R50963</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="493936C4997CFFD82F82FC16FC28F221" blockId="10.[189,1398,986,1084]" pageId="10" pageNumber="11">
<materialsCitation id="F9EE3C99997CFFD82F82FC16F888F226" collectionCode="MNHN" collectorName="La Boissine &amp; La Voulte-sur-Rhone" country="France" location="FRANCE" pageId="10" pageNumber="11" specimenCode="A29249, A29257, A29286, A84029, R06203, R50971" specimenCount="6" typeStatus="paratype">
<emphasis id="7BF2EAD6997CFFD82F82FC16F86AF5EE" bold="true" box="[234,359,986,1012]" pageId="10" pageNumber="11">
<typeStatus id="963D8866997CFFD82F82FC16F86AF5EE" box="[234,359,986,1012]" pageId="10" pageNumber="11" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="31917654997CFFD82FD5FC32F83CF202" box="[189,305,1022,1048]" name="France" pageId="10" pageNumber="11">FRANCE</collectingCountry>
<specimenCount id="5F80FD4D997CFFD82E20FC32F8ADF202" box="[328,416,1022,1048]" pageId="10" pageNumber="11">6 specs</specimenCount>
; same collection data as for holotype;
<collectionCode id="2F97AE01997CFFD82C33FC32FAB4F202" box="[859,953,1022,1048]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="10" pageNumber="11">MNHN</collectionCode>
.F.
<specimenCode id="19209EBF997CFFD82CB9FC32FD3FF202" box="[977,1074,1022,1048]" pageId="10" pageNumber="11">A29249</specimenCode>
,
<specimenCode id="19209EBF997CFFD82B53FC32FD90F203" box="[1083,1181,1022,1049]" pageId="10" pageNumber="11">A29257</specimenCode>
,
<specimenCode id="19209EBF997CFFD82BCEFC32FC05F202" box="[1190,1288,1022,1048]" pageId="10" pageNumber="11">A29286</specimenCode>
,
<specimenCode id="19209EBF997CFFD82A79FC32FC7FF202" box="[1297,1394,1022,1048]" pageId="10" pageNumber="11">A84029</specimenCode>
,
<specimenCode id="19209EBF997CFFD82FD5FBEDF810F221" box="[189,285,1057,1083]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="10" pageNumber="11">R06203</specimenCode>
,
<specimenCode id="19209EBF997CFFD82E41FBEDF888F226" box="[297,389,1057,1084]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="10" pageNumber="11">R50971</specimenCode>
</materialsCitation>
<materialsCitation id="F9EE3C99997CFFD82EF6FBEEFC2CF221" box="[414,1313,1057,1084]" collectionCode="MSNM" collectorName="La Boissine &amp; La Voulte-sur-Rhone" country="France" location="FRANCE" pageId="10" pageNumber="11" specimenCode="i7591, i13276, i13603" specimenCount="3" typeStatus="paratype">
<specimenCount id="5F80FD4D997CFFD82EF6FBEEF8FAF226" box="[414,503,1058,1084]" pageId="10" pageNumber="11">3 specs</specimenCount>
; same collection data as for holotype;
<collectionCode id="2F97AE01997CFFD82CD2FBEEFD19F226" box="[954,1044,1058,1084]" collectionName="Italy, Milano, Museo Civico di Storia Naturale" pageId="10" pageNumber="11">MSNM</collectionCode>
<specimenCode id="19209EBF997CFFD82B73FBEDFD6DF226" box="[1051,1120,1057,1084]" pageId="10" pageNumber="11">i7591</specimenCode>
,
<specimenCode id="19209EBF997CFFD82B04FBEDFDCCF221" box="[1132,1217,1057,1083]" pageId="10" pageNumber="11">i13276</specimenCode>
,
<specimenCode id="19209EBF997CFFD82BA4FBEDFC2CF221" box="[1228,1313,1057,1083]" pageId="10" pageNumber="11">i13603</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="019C654F997CFFD92FD5FBA8F846F168" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="description">
<paragraph id="493936C4997CFFD82FD5FBA8F878F264" blockId="10.[189,373,1124,1150]" box="[189,373,1124,1150]" pageId="10" pageNumber="11">
<heading id="127181A8997CFFD82FD5FBA8F878F264" bold="true" box="[189,373,1124,1150]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="7BF2EAD6997CFFD82FD5FBA8F878F264" bold="true" box="[189,373,1124,1150]" pageId="10" pageNumber="11">Measurements</emphasis>
</heading>
</paragraph>
<paragraph id="493936C4997CFFD82FD5FB5FF841F2B7" blockId="10.[189,332,1171,1197]" box="[189,332,1171,1197]" pageId="10" pageNumber="11">
See
<tableCitation id="0404037F997CFFD82F86FB5FF845F2B7" box="[238,328,1171,1197]" captionStart="Table 2" captionStartId="7.[185,255,546,574]" captionTargetPageId="7" captionText="Table 2 (continued on nex page). Descriptive statistics on the shield measurements. Abbreviations: Aad = anterodorsal angle; Aav = anteroventral angle; Apd = posterodorsal angle; Apv = posteroventral angle; Ha = anterior shield height; Hmax = maximum shield height; Hp = posterior shield height; Inf = unknown data; Lr = length of the rostrum; Ls = length of the shield; Lw = length of the shield without the rostrum; N = number of measured specimens; NA = non-applicable; SD = standard deviation." httpUri="http://table.plazi.org/id/1DF9664C9971FFD52FD1FDEEFAE8F4BD" pageId="10" pageNumber="11" tableUuid="1DF9664C9971FFD52FD1FDEEFAE8F4BD">Table 2</tableCitation>
.
</paragraph>
<paragraph id="493936C4997CFFD82FD5FB18F843F2F4" blockId="10.[189,334,1236,1262]" box="[189,334,1236,1262]" pageId="10" pageNumber="11">
<heading id="127181A8997CFFD82FD5FB18F843F2F4" bold="true" box="[189,334,1236,1262]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="7BF2EAD6997CFFD82FD5FB18F843F2F4" bold="true" box="[189,334,1236,1262]" pageId="10" pageNumber="11">Description</emphasis>
</heading>
</paragraph>
<paragraph id="493936C4997CFFD82F82FACFF8D5F304" blockId="10.[187,1400,1283,1560]" box="[234,472,1283,1310]" pageId="10" pageNumber="11">
<emphasis id="7BF2EAD6997CFFD82F82FACFF8D5F304" bold="true" box="[234,472,1283,1310]" pageId="10" pageNumber="11">Shield morphology</emphasis>
</paragraph>
<paragraph id="493936C4997CFFD82FD5FAEBF868F002" blockId="10.[187,1400,1283,1560]" pageId="10" pageNumber="11">
Sub-oval shield in lateral view with a short, sinuous anterior margin, a sharp antero-dorsal corner, a straight and horizontal dorsal midline, endings posteriorly in a sharp postero-dorsal corner, a posterior margin divided into a very short and straight dorsal part antero-ventrally oriented and a straight, sub-vertical ventral part, a rounded postero-ventral corner, a ventral margin divided into a posterior half, steeply descending antero-ventrally and a horizontal anterior part merging approximately at the posterior third of the length, a rounded antero-ventral corner (
<figureCitation id="D1BD2A41997CFFD82C44FA16FABCF3EE" box="[812,945,1498,1524]" captionStart="Fig" captionStartId="12.[189,231,1716,1742]" captionTargetBox="[269,1317,265,1679]" captionTargetId="figure-12@12.[269,1318,265,1679]" captionTargetPageId="12" captionText="Fig. 3. Austriocaris secretanae sp. nov. AF. Holotype MNHN.F.R50963. A. General view (part). B. General view (counter-part). C. Drawing (part). D. Drawing (counter-part). E. Close-up of pores. F. Close-up of muscles of raptorial appendages. G. Paratype MNHN.F.A29257. HI. Paratype MNHN.F.R06203. H. General view. I. Drawing. Abbreviations: am = anterior margin; ce = compound eye; dlc = dorso-lateral carina; dm = dorsal midline; g = gills; mlc = medio-lateral carina; os = oval structure; p15 = podomeres of raptorial appendages; pm = posterior margin; po = pores; ra 13 = raptorial appendages; sp = spines; st = stalk; t = posterior trunk; ta 8 = posterior trunk appendage 8; tp 111 = pleural parts of posterior trunk; vlg = ventro-lateral groove; vm = ventral margin. Scale bars: AD, GI = 10 mm; E = 1 mm; F = 5 mm. Photos: AB, G: L. Cazes; EF: T. Laville. H: C. Haug &amp; Joachim T. Haug." figureDoi="http://doi.org/10.5281/zenodo.8413508" httpUri="https://zenodo.org/record/8413508/files/figure.png" pageId="10" pageNumber="11">Fig. 3AD</figureCitation>
). Rim of all free margins delimiting a marginal fold.
</paragraph>
<paragraph id="493936C4997CFFD82F82F9F3FB5EF040" blockId="10.[189,1400,1599,1805]" box="[234,595,1599,1626]" pageId="10" pageNumber="11">
<emphasis id="7BF2EAD6997CFFD82F82F9F3FB5EF040" bold="true" box="[234,595,1599,1626]" pageId="10" pageNumber="11">Shield macro-ornamentation</emphasis>
</paragraph>
<paragraph id="493936C4997CFFD82FD5F9AFFC0EF117" blockId="10.[189,1400,1599,1805]" pageId="10" pageNumber="11">
Shield with a dorso-lateral carina located on its anterior area and a medio-lateral carina protruding from the postero-ventral angle (
<figureCitation id="D1BD2A41997CFFD82E9DF94BFB30F0BB" box="[501,573,1671,1697]" captionStart="Fig" captionStartId="12.[189,231,1716,1742]" captionTargetBox="[269,1317,265,1679]" captionTargetId="figure-12@12.[269,1318,265,1679]" captionTargetPageId="12" captionText="Fig. 3. Austriocaris secretanae sp. nov. AF. Holotype MNHN.F.R50963. A. General view (part). B. General view (counter-part). C. Drawing (part). D. Drawing (counter-part). E. Close-up of pores. F. Close-up of muscles of raptorial appendages. G. Paratype MNHN.F.A29257. HI. Paratype MNHN.F.R06203. H. General view. I. Drawing. Abbreviations: am = anterior margin; ce = compound eye; dlc = dorso-lateral carina; dm = dorsal midline; g = gills; mlc = medio-lateral carina; os = oval structure; p15 = podomeres of raptorial appendages; pm = posterior margin; po = pores; ra 13 = raptorial appendages; sp = spines; st = stalk; t = posterior trunk; ta 8 = posterior trunk appendage 8; tp 111 = pleural parts of posterior trunk; vlg = ventro-lateral groove; vm = ventral margin. Scale bars: AD, GI = 10 mm; E = 1 mm; F = 5 mm. Photos: AB, G: L. Cazes; EF: T. Laville. H: C. Haug &amp; Joachim T. Haug." figureDoi="http://doi.org/10.5281/zenodo.8413508" httpUri="https://zenodo.org/record/8413508/files/figure.png" pageId="10" pageNumber="11">Fig. 3</figureCitation>
). Dorso-lateral carina horizontal until mid-length of the shield where it folds down on itself. Presence of a row of 33 pores on the top of the carina (
<figureCitation id="D1BD2A41997CFFD82B0EF967FDCCF0DF" box="[1126,1217,1707,1733]" captionStart="Fig" captionStartId="12.[189,231,1716,1742]" captionTargetBox="[269,1317,265,1679]" captionTargetId="figure-12@12.[269,1318,265,1679]" captionTargetPageId="12" captionText="Fig. 3. Austriocaris secretanae sp. nov. AF. Holotype MNHN.F.R50963. A. General view (part). B. General view (counter-part). C. Drawing (part). D. Drawing (counter-part). E. Close-up of pores. F. Close-up of muscles of raptorial appendages. G. Paratype MNHN.F.A29257. HI. Paratype MNHN.F.R06203. H. General view. I. Drawing. Abbreviations: am = anterior margin; ce = compound eye; dlc = dorso-lateral carina; dm = dorsal midline; g = gills; mlc = medio-lateral carina; os = oval structure; p15 = podomeres of raptorial appendages; pm = posterior margin; po = pores; ra 13 = raptorial appendages; sp = spines; st = stalk; t = posterior trunk; ta 8 = posterior trunk appendage 8; tp 111 = pleural parts of posterior trunk; vlg = ventro-lateral groove; vm = ventral margin. Scale bars: AD, GI = 10 mm; E = 1 mm; F = 5 mm. Photos: AB, G: L. Cazes; EF: T. Laville. H: C. Haug &amp; Joachim T. Haug." figureDoi="http://doi.org/10.5281/zenodo.8413508" httpUri="https://zenodo.org/record/8413508/files/figure.png" pageId="10" pageNumber="11">Fig. 3E</figureCitation>
). Medio-lateral carina horizontal until mid-length then becoming antero-ventrally oriented. Rim of all free margins delimiting the marginal fold. Shield also ornamented with horseshoe-like ridges (
<figureCitation id="D1BD2A41997CFFD82B1DF93EFDFBF117" box="[1141,1270,1778,1805]" captionStart="Fig" captionStartId="13.[187,230,1704,1730]" captionTargetBox="[189,1398,268,1649]" captionTargetId="figure-11@13.[189,1398,268,1649]" captionTargetPageId="13" captionText="Fig. 4. Austriocaris secretanae sp. nov. A, C, F. Paratype MNHN.F.A29286. A. General view. C. Closeup of ornamentation. F. Close-up of compound eye. B. Paratype MNHN.F.R50971. D, G. Paratype MNHN.F.A29249. D. General view. G. Drawing. EH. Paratype MNHN.F.A84029. E. General view. H. Close-up of posterior trunk appendages. IJ. Paratype MSNM i7591. I. General view. J. Close-up of raptorial appendages. Abbreviations: cs = circular structure; dm = dorsal midline; os = oval structure; pm = posterior margin; t = posterior trunk; tp111 = pleural parts of posterior trunk appendages. Scale bars: AB, DE, G, I = 10 mm; C = 1 mm; F = 400 µm; H, J = 5 mm. Photos: A, CF, HJ: T. Laville; B: L. Cazes." figureDoi="http://doi.org/10.5281/zenodo.8413512" httpUri="https://zenodo.org/record/8413512/files/figure.png" pageId="10" pageNumber="11">Fig. 4A, C</figureCitation>
).
</paragraph>
<paragraph id="493936C4997CFFD82F82F8F8F8D9F155" blockId="10.[187,1399,1844,1980]" box="[234,468,1844,1871]" pageId="10" pageNumber="11">
<emphasis id="7BF2EAD6997CFFD82F82F8F8F8D9F155" bold="true" box="[234,468,1844,1871]" pageId="10" pageNumber="11">Sensory structures</emphasis>
</paragraph>
<paragraph id="493936C4997CFFD82FD5F895FC7BF18C" blockId="10.[187,1399,1844,1980]" pageId="10" pageNumber="11">
Large, oval, stalked compound eyes protruding from the anterior margin and occupying the entire margin (
<figureCitation id="D1BD2A41997CFFD82FACF8B0F830F18C" box="[196,317,1916,1942]" captionStart="Fig" captionStartId="12.[189,231,1716,1742]" captionTargetBox="[269,1317,265,1679]" captionTargetId="figure-12@12.[269,1318,265,1679]" captionTargetPageId="12" captionText="Fig. 3. Austriocaris secretanae sp. nov. AF. Holotype MNHN.F.R50963. A. General view (part). B. General view (counter-part). C. Drawing (part). D. Drawing (counter-part). E. Close-up of pores. F. Close-up of muscles of raptorial appendages. G. Paratype MNHN.F.A29257. HI. Paratype MNHN.F.R06203. H. General view. I. Drawing. Abbreviations: am = anterior margin; ce = compound eye; dlc = dorso-lateral carina; dm = dorsal midline; g = gills; mlc = medio-lateral carina; os = oval structure; p15 = podomeres of raptorial appendages; pm = posterior margin; po = pores; ra 13 = raptorial appendages; sp = spines; st = stalk; t = posterior trunk; ta 8 = posterior trunk appendage 8; tp 111 = pleural parts of posterior trunk; vlg = ventro-lateral groove; vm = ventral margin. Scale bars: AD, GI = 10 mm; E = 1 mm; F = 5 mm. Photos: AB, G: L. Cazes; EF: T. Laville. H: C. Haug &amp; Joachim T. Haug." figureDoi="http://doi.org/10.5281/zenodo.8413508" httpUri="https://zenodo.org/record/8413508/files/figure.png" pageId="10" pageNumber="11">Fig. 3HI</figureCitation>
). Eyes tilted in the same orientation as the anterior margin, formed by hexagonal ommatidia
</paragraph>
<paragraph id="493936C4997CFFD82FD5F86CFA7CF1A0" blockId="10.[187,1399,1844,1980]" box="[189,881,1951,1980]" pageId="10" pageNumber="11">
(w = ~65 µm; l = ~70 µm, about 196 per mm²;
<figureCitation id="D1BD2A41997CFFD82D80F86CFA69F1A0" box="[744,868,1952,1978]" captionStart="Fig" captionStartId="13.[187,230,1704,1730]" captionTargetBox="[189,1398,268,1649]" captionTargetId="figure-11@13.[189,1398,268,1649]" captionTargetPageId="13" captionText="Fig. 4. Austriocaris secretanae sp. nov. A, C, F. Paratype MNHN.F.A29286. A. General view. C. Closeup of ornamentation. F. Close-up of compound eye. B. Paratype MNHN.F.R50971. D, G. Paratype MNHN.F.A29249. D. General view. G. Drawing. EH. Paratype MNHN.F.A84029. E. General view. H. Close-up of posterior trunk appendages. IJ. Paratype MSNM i7591. I. General view. J. Close-up of raptorial appendages. Abbreviations: cs = circular structure; dm = dorsal midline; os = oval structure; pm = posterior margin; t = posterior trunk; tp111 = pleural parts of posterior trunk appendages. Scale bars: AB, DE, G, I = 10 mm; C = 1 mm; F = 400 µm; H, J = 5 mm. Photos: A, CF, HJ: T. Laville; B: L. Cazes." figureDoi="http://doi.org/10.5281/zenodo.8413512" httpUri="https://zenodo.org/record/8413512/files/figure.png" pageId="10" pageNumber="11">Fig. 4A, F</figureCitation>
).
</paragraph>
<paragraph id="493936C4997DFFD92F82FEC4F8FEF739" blockId="11.[189,612,264,327]" box="[234,499,264,291]" pageId="11" pageNumber="12">
<emphasis id="7BF2EAD6997DFFD92F82FEC4F8FEF739" bold="true" box="[234,499,264,291]" pageId="11" pageNumber="12">Cephalic appendages</emphasis>
</paragraph>
<paragraph id="493936C4997DFFD92FD5FEE0FB69F75C" blockId="11.[189,612,264,327]" box="[189,612,300,327]" pageId="11" pageNumber="12">Cephalic appendages not preserved.</paragraph>
<paragraph id="493936C4997DFFD92F82FEA1F8F0F792" blockId="11.[189,1399,365,533]" box="[234,509,365,392]" pageId="11" pageNumber="12">
<emphasis id="7BF2EAD6997DFFD92F82FEA1F8F0F792" bold="true" box="[234,509,365,392]" pageId="11" pageNumber="12">Raptorial appendages</emphasis>
</paragraph>
<paragraph id="493936C4997DFFD92FD5FE5DFB1CF40E" blockId="11.[189,1399,365,533]" pageId="11" pageNumber="12">
Presence of three pairs of stout raptorial appendages recognised by preservation of muscles, protruding from the anterior part of the ventral margin (
<figureCitation id="D1BD2A41997DFFD92DA1FE78FA47F7D4" box="[713,842,436,462]" captionStart="Fig" captionStartId="12.[189,231,1716,1742]" captionTargetBox="[269,1317,265,1679]" captionTargetId="figure-12@12.[269,1318,265,1679]" captionTargetPageId="12" captionText="Fig. 3. Austriocaris secretanae sp. nov. AF. Holotype MNHN.F.R50963. A. General view (part). B. General view (counter-part). C. Drawing (part). D. Drawing (counter-part). E. Close-up of pores. F. Close-up of muscles of raptorial appendages. G. Paratype MNHN.F.A29257. HI. Paratype MNHN.F.R06203. H. General view. I. Drawing. Abbreviations: am = anterior margin; ce = compound eye; dlc = dorso-lateral carina; dm = dorsal midline; g = gills; mlc = medio-lateral carina; os = oval structure; p15 = podomeres of raptorial appendages; pm = posterior margin; po = pores; ra 13 = raptorial appendages; sp = spines; st = stalk; t = posterior trunk; ta 8 = posterior trunk appendage 8; tp 111 = pleural parts of posterior trunk; vlg = ventro-lateral groove; vm = ventral margin. Scale bars: AD, GI = 10 mm; E = 1 mm; F = 5 mm. Photos: AB, G: L. Cazes; EF: T. Laville. H: C. Haug &amp; Joachim T. Haug." figureDoi="http://doi.org/10.5281/zenodo.8413508" httpUri="https://zenodo.org/record/8413508/files/figure.png" pageId="11" pageNumber="12">Fig. 3A, C</figureCitation>
). Raptorial appendages consisting in two parts: a proximal postero-ventrally oriented and a distal one, antero-ventrally oriented. Articulation between both parts forming an elbow.
</paragraph>
<paragraph id="493936C4997DFFD92FD5FDF0FC7AF4F9" blockId="11.[189,1401,572,739]" pageId="11" pageNumber="12">
First pair of raptorial appendages (
<figureCitation id="D1BD2A41997DFFD92D01FDF0FBE8F44C" box="[617,741,572,598]" captionStart="Fig" captionStartId="12.[189,231,1716,1742]" captionTargetBox="[269,1317,265,1679]" captionTargetId="figure-12@12.[269,1318,265,1679]" captionTargetPageId="12" captionText="Fig. 3. Austriocaris secretanae sp. nov. AF. Holotype MNHN.F.R50963. A. General view (part). B. General view (counter-part). C. Drawing (part). D. Drawing (counter-part). E. Close-up of pores. F. Close-up of muscles of raptorial appendages. G. Paratype MNHN.F.A29257. HI. Paratype MNHN.F.R06203. H. General view. I. Drawing. Abbreviations: am = anterior margin; ce = compound eye; dlc = dorso-lateral carina; dm = dorsal midline; g = gills; mlc = medio-lateral carina; os = oval structure; p15 = podomeres of raptorial appendages; pm = posterior margin; po = pores; ra 13 = raptorial appendages; sp = spines; st = stalk; t = posterior trunk; ta 8 = posterior trunk appendage 8; tp 111 = pleural parts of posterior trunk; vlg = ventro-lateral groove; vm = ventral margin. Scale bars: AD, GI = 10 mm; E = 1 mm; F = 5 mm. Photos: AB, G: L. Cazes; EF: T. Laville. H: C. Haug &amp; Joachim T. Haug." figureDoi="http://doi.org/10.5281/zenodo.8413508" httpUri="https://zenodo.org/record/8413508/files/figure.png" pageId="11" pageNumber="12">Fig. 3HI</figureCitation>
), the shortest, consisting of a proximal part with two rectangular podomeres: podomere 1 being 4.2 times as long as wide; podomere 2 being 4.7 times as long as wide. Distal part of the appendages formed by three rectangular podomeres: podomere 3 being 3.4 times as long as wide, bearing six spines on its medial margin; podomere 4 being 2.3 times as long as wide; podomere 5, the shortest, being 1.4 times as long as wide and ending in a cluster of three spines.
</paragraph>
<paragraph id="493936C4997DFFD92FD5FCC6FB45F5AB" blockId="11.[188,1399,778,945]" pageId="11" pageNumber="12">
Second pair of raptorial appendages consisting of a proximal part with two podomeres (
<figureCitation id="D1BD2A41997DFFD92B8AFCC6FC65F53E" box="[1250,1384,778,804]" captionStart="Fig" captionStartId="12.[189,231,1716,1742]" captionTargetBox="[269,1317,265,1679]" captionTargetId="figure-12@12.[269,1318,265,1679]" captionTargetPageId="12" captionText="Fig. 3. Austriocaris secretanae sp. nov. AF. Holotype MNHN.F.R50963. A. General view (part). B. General view (counter-part). C. Drawing (part). D. Drawing (counter-part). E. Close-up of pores. F. Close-up of muscles of raptorial appendages. G. Paratype MNHN.F.A29257. HI. Paratype MNHN.F.R06203. H. General view. I. Drawing. Abbreviations: am = anterior margin; ce = compound eye; dlc = dorso-lateral carina; dm = dorsal midline; g = gills; mlc = medio-lateral carina; os = oval structure; p15 = podomeres of raptorial appendages; pm = posterior margin; po = pores; ra 13 = raptorial appendages; sp = spines; st = stalk; t = posterior trunk; ta 8 = posterior trunk appendage 8; tp 111 = pleural parts of posterior trunk; vlg = ventro-lateral groove; vm = ventral margin. Scale bars: AD, GI = 10 mm; E = 1 mm; F = 5 mm. Photos: AB, G: L. Cazes; EF: T. Laville. H: C. Haug &amp; Joachim T. Haug." figureDoi="http://doi.org/10.5281/zenodo.8413508" httpUri="https://zenodo.org/record/8413508/files/figure.png" pageId="11" pageNumber="12">Fig. 3A, C</figureCitation>
): podomere 1 being 4.7 times as long as wide; podomere 2, the longest, being 5.3 times as long as wide. Distal part of the appendage consisting of three podomeres: podomere 3 being 3.2 times as long as wide; podomere 4 being 3.1 times as long as wide; podomere 5 being 1.5 times as long as wide and ending distally in a cluster of five spines.
</paragraph>
<paragraph id="493936C4997DFFD92FD5FC14FC44F2DF" blockId="11.[187,1400,984,1222]" pageId="11" pageNumber="12">
Third pair of raptorial appendages, the longest, consisting of a proximal part formed by two rectangular podomeres (
<figureCitation id="D1BD2A41997DFFD92E27FC37F8D9F20C" box="[335,468,1019,1046]" captionStart="Fig" captionStartId="12.[189,231,1716,1742]" captionTargetBox="[269,1317,265,1679]" captionTargetId="figure-12@12.[269,1318,265,1679]" captionTargetPageId="12" captionText="Fig. 3. Austriocaris secretanae sp. nov. AF. Holotype MNHN.F.R50963. A. General view (part). B. General view (counter-part). C. Drawing (part). D. Drawing (counter-part). E. Close-up of pores. F. Close-up of muscles of raptorial appendages. G. Paratype MNHN.F.A29257. HI. Paratype MNHN.F.R06203. H. General view. I. Drawing. Abbreviations: am = anterior margin; ce = compound eye; dlc = dorso-lateral carina; dm = dorsal midline; g = gills; mlc = medio-lateral carina; os = oval structure; p15 = podomeres of raptorial appendages; pm = posterior margin; po = pores; ra 13 = raptorial appendages; sp = spines; st = stalk; t = posterior trunk; ta 8 = posterior trunk appendage 8; tp 111 = pleural parts of posterior trunk; vlg = ventro-lateral groove; vm = ventral margin. Scale bars: AD, GI = 10 mm; E = 1 mm; F = 5 mm. Photos: AB, G: L. Cazes; EF: T. Laville. H: C. Haug &amp; Joachim T. Haug." figureDoi="http://doi.org/10.5281/zenodo.8413508" httpUri="https://zenodo.org/record/8413508/files/figure.png" pageId="11" pageNumber="12">Figs 3AC</figureCitation>
,
<figureCitation id="D1BD2A41997DFFD92E8AFC30FB17F20C" box="[482,538,1020,1046]" captionStart="Fig" captionStartId="13.[187,230,1704,1730]" captionTargetBox="[189,1398,268,1649]" captionTargetId="figure-11@13.[189,1398,268,1649]" captionTargetPageId="13" captionText="Fig. 4. Austriocaris secretanae sp. nov. A, C, F. Paratype MNHN.F.A29286. A. General view. C. Closeup of ornamentation. F. Close-up of compound eye. B. Paratype MNHN.F.R50971. D, G. Paratype MNHN.F.A29249. D. General view. G. Drawing. EH. Paratype MNHN.F.A84029. E. General view. H. Close-up of posterior trunk appendages. IJ. Paratype MSNM i7591. I. General view. J. Close-up of raptorial appendages. Abbreviations: cs = circular structure; dm = dorsal midline; os = oval structure; pm = posterior margin; t = posterior trunk; tp111 = pleural parts of posterior trunk appendages. Scale bars: AB, DE, G, I = 10 mm; C = 1 mm; F = 400 µm; H, J = 5 mm. Photos: A, CF, HJ: T. Laville; B: L. Cazes." figureDoi="http://doi.org/10.5281/zenodo.8413512" httpUri="https://zenodo.org/record/8413512/files/figure.png" pageId="11" pageNumber="12">4IJ</figureCitation>
): podomere 1 being 3.2 times as long as wide; podomere 2, the longest, being 7.9 times as long as wide. Distal part consisting of three podomeres: podomere 3 being 5.2 times as long as wide; podomere 4 being 4.5 times as long as wide and bearing eight spines on its medial margin, the first two being close to the proximal margin (
<figureCitation id="D1BD2A41997DFFD92D9DFBA9FA6EF265" box="[757,867,1125,1151]" captionStart="Fig" captionStartId="13.[187,230,1704,1730]" captionTargetBox="[189,1398,268,1649]" captionTargetId="figure-11@13.[189,1398,268,1649]" captionTargetPageId="13" captionText="Fig. 4. Austriocaris secretanae sp. nov. A, C, F. Paratype MNHN.F.A29286. A. General view. C. Closeup of ornamentation. F. Close-up of compound eye. B. Paratype MNHN.F.R50971. D, G. Paratype MNHN.F.A29249. D. General view. G. Drawing. EH. Paratype MNHN.F.A84029. E. General view. H. Close-up of posterior trunk appendages. IJ. Paratype MSNM i7591. I. General view. J. Close-up of raptorial appendages. Abbreviations: cs = circular structure; dm = dorsal midline; os = oval structure; pm = posterior margin; t = posterior trunk; tp111 = pleural parts of posterior trunk appendages. Scale bars: AB, DE, G, I = 10 mm; C = 1 mm; F = 400 µm; H, J = 5 mm. Photos: A, CF, HJ: T. Laville; B: L. Cazes." figureDoi="http://doi.org/10.5281/zenodo.8413512" httpUri="https://zenodo.org/record/8413512/files/figure.png" pageId="11" pageNumber="12">Fig. 4IJ</figureCitation>
), followed by two stout spines, surrounding a more gracile one, and then by three short spines and a long, stout spine. Distal part of the third raptorial appendages ending with a square-shaped podomere (podomere 5), bearing a cluster of three spines.
</paragraph>
<paragraph id="493936C4997DFFD92F82FB21F8ACF31D" blockId="11.[188,1399,1261,1393]" box="[234,417,1261,1287]" pageId="11" pageNumber="12">
<emphasis id="7BF2EAD6997DFFD92F82FB21F8ACF31D" bold="true" box="[234,417,1261,1287]" pageId="11" pageNumber="12">Oval structure</emphasis>
</paragraph>
<paragraph id="493936C4997DFFD92FD5FADDFB9EF36B" blockId="11.[188,1399,1261,1393]" pageId="11" pageNumber="12">
Long, oval structure protruding from the anterior part of the ventral margin, between raptorial and trunk appendages (
<figureCitation id="D1BD2A41997DFFD92E3EFAFFFB1AF354" box="[342,535,1331,1358]" captionStart="Fig" captionStartId="12.[189,231,1716,1742]" captionTargetBox="[269,1317,265,1679]" captionTargetId="figure-12@12.[269,1318,265,1679]" captionTargetPageId="12" captionText="Fig. 3. Austriocaris secretanae sp. nov. AF. Holotype MNHN.F.R50963. A. General view (part). B. General view (counter-part). C. Drawing (part). D. Drawing (counter-part). E. Close-up of pores. F. Close-up of muscles of raptorial appendages. G. Paratype MNHN.F.A29257. HI. Paratype MNHN.F.R06203. H. General view. I. Drawing. Abbreviations: am = anterior margin; ce = compound eye; dlc = dorso-lateral carina; dm = dorsal midline; g = gills; mlc = medio-lateral carina; os = oval structure; p15 = podomeres of raptorial appendages; pm = posterior margin; po = pores; ra 13 = raptorial appendages; sp = spines; st = stalk; t = posterior trunk; ta 8 = posterior trunk appendage 8; tp 111 = pleural parts of posterior trunk; vlg = ventro-lateral groove; vm = ventral margin. Scale bars: AD, GI = 10 mm; E = 1 mm; F = 5 mm. Photos: AB, G: L. Cazes; EF: T. Laville. H: C. Haug &amp; Joachim T. Haug." figureDoi="http://doi.org/10.5281/zenodo.8413508" httpUri="https://zenodo.org/record/8413508/files/figure.png" pageId="11" pageNumber="12">Fig. 3AD, HI</figureCitation>
). Two elongate and sinuous rectangular structures visible on this feature. The nature of this structure is unknown.
</paragraph>
<paragraph id="493936C4997DFFD92F82FA54F8A0F3A8" blockId="11.[187,1400,1432,1635]" box="[234,429,1432,1458]" pageId="11" pageNumber="12">
<emphasis id="7BF2EAD6997DFFD92F82FA54F8A0F3A8" bold="true" box="[234,429,1432,1458]" pageId="11" pageNumber="12">Posterior trunk</emphasis>
</paragraph>
<paragraph id="493936C4997DFFD92FD5FA77FC3AF079" blockId="11.[187,1400,1432,1635]" pageId="11" pageNumber="12">
Posterior trunk consisting of eleven elongate, rectangular structures (i.e., pleural parts of the segments;
<figureCitation id="D1BD2A41997DFFD92FD5FA12F832F3E3" box="[189,319,1502,1529]" captionStart="Fig" captionStartId="12.[189,231,1716,1742]" captionTargetBox="[269,1317,265,1679]" captionTargetId="figure-12@12.[269,1318,265,1679]" captionTargetPageId="12" captionText="Fig. 3. Austriocaris secretanae sp. nov. AF. Holotype MNHN.F.R50963. A. General view (part). B. General view (counter-part). C. Drawing (part). D. Drawing (counter-part). E. Close-up of pores. F. Close-up of muscles of raptorial appendages. G. Paratype MNHN.F.A29257. HI. Paratype MNHN.F.R06203. H. General view. I. Drawing. Abbreviations: am = anterior margin; ce = compound eye; dlc = dorso-lateral carina; dm = dorsal midline; g = gills; mlc = medio-lateral carina; os = oval structure; p15 = podomeres of raptorial appendages; pm = posterior margin; po = pores; ra 13 = raptorial appendages; sp = spines; st = stalk; t = posterior trunk; ta 8 = posterior trunk appendage 8; tp 111 = pleural parts of posterior trunk; vlg = ventro-lateral groove; vm = ventral margin. Scale bars: AD, GI = 10 mm; E = 1 mm; F = 5 mm. Photos: AB, G: L. Cazes; EF: T. Laville. H: C. Haug &amp; Joachim T. Haug." figureDoi="http://doi.org/10.5281/zenodo.8413508" httpUri="https://zenodo.org/record/8413508/files/figure.png" pageId="11" pageNumber="12">Figs 3AD</figureCitation>
,
<figureCitation id="D1BD2A41997DFFD92E21FA13F89CF3E3" box="[329,401,1503,1529]" captionStart="Fig" captionStartId="13.[187,230,1704,1730]" captionTargetBox="[189,1398,268,1649]" captionTargetId="figure-11@13.[189,1398,268,1649]" captionTargetPageId="13" captionText="Fig. 4. Austriocaris secretanae sp. nov. A, C, F. Paratype MNHN.F.A29286. A. General view. C. Closeup of ornamentation. F. Close-up of compound eye. B. Paratype MNHN.F.R50971. D, G. Paratype MNHN.F.A29249. D. General view. G. Drawing. EH. Paratype MNHN.F.A84029. E. General view. H. Close-up of posterior trunk appendages. IJ. Paratype MSNM i7591. I. General view. J. Close-up of raptorial appendages. Abbreviations: cs = circular structure; dm = dorsal midline; os = oval structure; pm = posterior margin; t = posterior trunk; tp111 = pleural parts of posterior trunk appendages. Scale bars: AB, DE, G, I = 10 mm; C = 1 mm; F = 400 µm; H, J = 5 mm. Photos: A, CF, HJ: T. Laville; B: L. Cazes." figureDoi="http://doi.org/10.5281/zenodo.8413512" httpUri="https://zenodo.org/record/8413512/files/figure.png" pageId="11" pageNumber="12">4D, G</figureCitation>
), slightly curved upward at their distal end and surrounding a long rectangular structure (i.e., tergal part of posterior trunk segments). Last pleural part thinner than the other ones. Presence of a square-shaped element (i.e., telson) at the posterior extremity of the trunk, bearing a circular structure formed by two circular parts: an outer one, delimiting the entire feature, and an inner smaller one.
</paragraph>
<paragraph id="493936C4997DFFD92F82F945FB4AF0BE" blockId="11.[189,1398,1673,1770]" box="[234,583,1673,1700]" pageId="11" pageNumber="12">
<emphasis id="7BF2EAD6997DFFD92F82F945FB4AF0BE" bold="true" box="[234,583,1673,1700]" pageId="11" pageNumber="12">Posterior trunk appendages</emphasis>
</paragraph>
<paragraph id="493936C4997DFFD92FD5F961FB87F0F0" blockId="11.[189,1398,1673,1770]" pageId="11" pageNumber="12">
Eleven pairs of rod-like trunk appendages posteriorly oriented and protruding from the posterior part of the ventral margin (
<figureCitation id="D1BD2A41997DFFD92ECBF91CFB25F0F0" box="[419,552,1744,1770]" captionStart="Fig" captionStartId="12.[189,231,1716,1742]" captionTargetBox="[269,1317,265,1679]" captionTargetId="figure-12@12.[269,1318,265,1679]" captionTargetPageId="12" captionText="Fig. 3. Austriocaris secretanae sp. nov. AF. Holotype MNHN.F.R50963. A. General view (part). B. General view (counter-part). C. Drawing (part). D. Drawing (counter-part). E. Close-up of pores. F. Close-up of muscles of raptorial appendages. G. Paratype MNHN.F.A29257. HI. Paratype MNHN.F.R06203. H. General view. I. Drawing. Abbreviations: am = anterior margin; ce = compound eye; dlc = dorso-lateral carina; dm = dorsal midline; g = gills; mlc = medio-lateral carina; os = oval structure; p15 = podomeres of raptorial appendages; pm = posterior margin; po = pores; ra 13 = raptorial appendages; sp = spines; st = stalk; t = posterior trunk; ta 8 = posterior trunk appendage 8; tp 111 = pleural parts of posterior trunk; vlg = ventro-lateral groove; vm = ventral margin. Scale bars: AD, GI = 10 mm; E = 1 mm; F = 5 mm. Photos: AB, G: L. Cazes; EF: T. Laville. H: C. Haug &amp; Joachim T. Haug." figureDoi="http://doi.org/10.5281/zenodo.8413508" httpUri="https://zenodo.org/record/8413508/files/figure.png" pageId="11" pageNumber="12">Figs 3AD</figureCitation>
,
<figureCitation id="D1BD2A41997DFFD92D5DF91CFB70F0F0" box="[565,637,1744,1770]" captionStart="Fig" captionStartId="13.[187,230,1704,1730]" captionTargetBox="[189,1398,268,1649]" captionTargetId="figure-11@13.[189,1398,268,1649]" captionTargetPageId="13" captionText="Fig. 4. Austriocaris secretanae sp. nov. A, C, F. Paratype MNHN.F.A29286. A. General view. C. Closeup of ornamentation. F. Close-up of compound eye. B. Paratype MNHN.F.R50971. D, G. Paratype MNHN.F.A29249. D. General view. G. Drawing. EH. Paratype MNHN.F.A84029. E. General view. H. Close-up of posterior trunk appendages. IJ. Paratype MSNM i7591. I. General view. J. Close-up of raptorial appendages. Abbreviations: cs = circular structure; dm = dorsal midline; os = oval structure; pm = posterior margin; t = posterior trunk; tp111 = pleural parts of posterior trunk appendages. Scale bars: AB, DE, G, I = 10 mm; C = 1 mm; F = 400 µm; H, J = 5 mm. Photos: A, CF, HJ: T. Laville; B: L. Cazes." figureDoi="http://doi.org/10.5281/zenodo.8413512" httpUri="https://zenodo.org/record/8413512/files/figure.png" pageId="11" pageNumber="12">4E, H</figureCitation>
).
</paragraph>
<paragraph id="493936C4997DFFD92F82F8DDF828F131" blockId="11.[189,1399,1809,1906]" box="[234,293,1809,1835]" pageId="11" pageNumber="12">
<emphasis id="7BF2EAD6997DFFD92F82F8DDF828F131" bold="true" box="[234,293,1809,1835]" pageId="11" pageNumber="12">Gills</emphasis>
</paragraph>
<paragraph id="493936C4997DFFD92FD5F8F9F846F168" blockId="11.[189,1399,1809,1906]" pageId="11" pageNumber="12">
Four long lamellar gills preserved under the posterior area of the shield below the medio-lateral carina (
<figureCitation id="D1BD2A41997DFFD92FAEF894F833F168" box="[198,318,1880,1906]" captionStart="Fig" captionStartId="12.[189,231,1716,1742]" captionTargetBox="[269,1317,265,1679]" captionTargetId="figure-12@12.[269,1318,265,1679]" captionTargetPageId="12" captionText="Fig. 3. Austriocaris secretanae sp. nov. AF. Holotype MNHN.F.R50963. A. General view (part). B. General view (counter-part). C. Drawing (part). D. Drawing (counter-part). E. Close-up of pores. F. Close-up of muscles of raptorial appendages. G. Paratype MNHN.F.A29257. HI. Paratype MNHN.F.R06203. H. General view. I. Drawing. Abbreviations: am = anterior margin; ce = compound eye; dlc = dorso-lateral carina; dm = dorsal midline; g = gills; mlc = medio-lateral carina; os = oval structure; p15 = podomeres of raptorial appendages; pm = posterior margin; po = pores; ra 13 = raptorial appendages; sp = spines; st = stalk; t = posterior trunk; ta 8 = posterior trunk appendage 8; tp 111 = pleural parts of posterior trunk; vlg = ventro-lateral groove; vm = ventral margin. Scale bars: AD, GI = 10 mm; E = 1 mm; F = 5 mm. Photos: AB, G: L. Cazes; EF: T. Laville. H: C. Haug &amp; Joachim T. Haug." figureDoi="http://doi.org/10.5281/zenodo.8413508" httpUri="https://zenodo.org/record/8413508/files/figure.png" pageId="11" pageNumber="12">Fig. 3HI</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="1DF9664C997AFFDE2FD5F978FC20F1F3" ID-DOI="http://doi.org/10.5281/zenodo.8413508" ID-Zenodo-Dep="8413508" httpUri="https://zenodo.org/record/8413508/files/figure.png" pageId="12" pageNumber="13" startId="12.[189,231,1716,1742]" targetBox="[269,1317,265,1679]" targetPageId="12">
<paragraph id="493936C4997AFFDE2FD5F978FC20F1F3" blockId="12.[188,1400,1716,2025]" pageId="12" pageNumber="13">
<emphasis id="7BF2EAD6997AFFDE2FD5F978F81DF0D5" bold="true" box="[189,272,1716,1743]" pageId="12" pageNumber="13">Fig. 3.</emphasis>
<taxonomicName id="8E864D47997AFFDE2E74F979FB3BF0D5" authority="Laville &amp; Forel &amp; Charbonnier, 2023" authorityName="Laville &amp; Forel &amp; Charbonnier" authorityYear="2023" box="[284,566,1717,1743]" family="Austriocarididae" genus="Austriocaris" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="secretanae" status="sp. nov.">
<emphasis id="7BF2EAD6997AFFDE2E74F979FB3BF0D5" box="[284,566,1717,1743]" italics="true" pageId="12" pageNumber="13">Austriocaris secretanae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="60C157AD997AFFDE2D2BF979FBAFF0D5" box="[579,674,1717,1743]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="7BF2EAD6997AFFDE2DC7F978FBE9F0D4" bold="true" box="[687,740,1716,1742]" pageId="12" pageNumber="13">AF</emphasis>
. Holotype MNHN.F.R50963.
<emphasis id="7BF2EAD6997AFFDE2B3FF978FD61F0D4" bold="true" box="[1111,1132,1716,1742]" pageId="12" pageNumber="13">A</emphasis>
. General view (part).
<emphasis id="7BF2EAD6997AFFDE2FD5F914F9DDF0E8" bold="true" box="[189,208,1752,1778]" pageId="12" pageNumber="13">B</emphasis>
. General view (counter-part).
<emphasis id="7BF2EAD6997AFFDE2D57F914FB59F0E8" bold="true" box="[575,596,1752,1778]" pageId="12" pageNumber="13">C</emphasis>
. Drawing (part).
<emphasis id="7BF2EAD6997AFFDE2C42F914FA32F0E8" bold="true" box="[810,831,1752,1778]" pageId="12" pageNumber="13">D</emphasis>
. Drawing (counter-part).
<emphasis id="7BF2EAD6997AFFDE2B1CF914FD8AF0E8" bold="true" box="[1140,1159,1752,1778]" pageId="12" pageNumber="13">E</emphasis>
. Close-up of pores.
<emphasis id="7BF2EAD6997AFFDE2FD5F937F9C2F10F" bold="true" box="[189,207,1787,1813]" pageId="12" pageNumber="13">F</emphasis>
. Close-up of muscles of raptorial appendages.
<emphasis id="7BF2EAD6997AFFDE2C5EF937FA40F10F" bold="true" box="[822,845,1787,1813]" pageId="12" pageNumber="13">G</emphasis>
. Paratype MNHN.F.A29257.
<emphasis id="7BF2EAD6997AFFDE2BA0F937FDF4F10F" bold="true" box="[1224,1273,1787,1813]" pageId="12" pageNumber="13">HI</emphasis>
. Paratype MNHN.F.R06203.
<emphasis id="7BF2EAD6997AFFDE2EF6F8D2F8B8F122" bold="true" box="[414,437,1822,1848]" pageId="12" pageNumber="13">H</emphasis>
. General view.
<emphasis id="7BF2EAD6997AFFDE2D1AF8D2FB71F122" bold="true" box="[626,636,1822,1848]" pageId="12" pageNumber="13">I</emphasis>
. Drawing. Abbreviations: am = anterior margin; ce = compound eye; dlc = dorso-lateral carina; dm = dorsal midline; g = gills; mlc = medio-lateral carina; os = oval structure; p
<subScript id="D5023481997AFFDE2E29F8BBF856F19F" attach="left" box="[321,347,1911,1925]" fontSize="6" pageId="12" pageNumber="13">15</subScript>
= podomeres of raptorial appendages; pm = posterior margin; po = pores; ra
<subScript id="D5023481997AFFDE2BB5F8B4FDFBF19C" attach="left" box="[1245,1270,1912,1926]" fontSize="6" pageId="12" pageNumber="13">13</subScript>
= raptorial appendages; sp = spines; st = stalk; t = posterior trunk; ta 8 = posterior trunk appendage 8; tp 111 = pleural parts of posterior trunk; vlg = ventro-lateral groove; vm = ventral margin. Scale bars: AD, GI = 10 mm; E = 1 mm; F = 5 mm. Photos: AB, G: L. Cazes; EF: T. Laville. H: C. Haug &amp; Joachim T. Haug.
</paragraph>
</caption>
<caption id="1DF9664C997BFFDF2FD3F964F840F1A3" ID-DOI="http://doi.org/10.5281/zenodo.8413512" ID-Zenodo-Dep="8413512" httpUri="https://zenodo.org/record/8413512/files/figure.png" pageId="13" pageNumber="14" startId="13.[187,230,1704,1730]" targetBox="[189,1398,268,1649]" targetPageId="13">
<paragraph id="493936C4997BFFDF2FD3F964F840F1A3" blockId="13.[187,1400,1704,1977]" pageId="13" pageNumber="14">
<emphasis id="7BF2EAD6997BFFDF2FD3F964F805F0D8" bold="true" box="[187,264,1704,1730]" pageId="13" pageNumber="14">Fig. 4.</emphasis>
<taxonomicName id="8E864D47997BFFDF2E67F965FB29F0D8" authority="Laville &amp; Forel &amp; Charbonnier, 2023" authorityName="Laville &amp; Forel &amp; Charbonnier" authorityYear="2023" box="[271,548,1705,1731]" family="Austriocarididae" genus="Austriocaris" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="secretanae" status="sp. nov.">
<emphasis id="7BF2EAD6997BFFDF2E67F965FB29F0D8" box="[271,548,1705,1731]" italics="true" pageId="13" pageNumber="14">Austriocaris secretanae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="60C157AD997BFFDF2D41F965FB8EF0D9" box="[553,643,1705,1731]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="7BF2EAD6997BFFDF2DE0F964FBEDF0D8" bold="true" box="[648,736,1704,1730]" pageId="13" pageNumber="14">A, C, F</emphasis>
. Paratype MNHN.F.A29286.
<emphasis id="7BF2EAD6997BFFDF2B53F964FD5DF0D8" bold="true" box="[1083,1104,1704,1730]" pageId="13" pageNumber="14">A</emphasis>
. General view.
<emphasis id="7BF2EAD6997BFFDF2A6EF964FC16F0D8" bold="true" box="[1286,1307,1704,1730]" pageId="13" pageNumber="14">C</emphasis>
. Closeup of ornamentation.
<emphasis id="7BF2EAD6997BFFDF2EABF900F8D8F0FC" bold="true" box="[451,469,1740,1766]" pageId="13" pageNumber="14">F</emphasis>
. Close-up of compound eye.
<emphasis id="7BF2EAD6997BFFDF2C2AF900FA5BF0FC" bold="true" box="[834,854,1740,1766]" pageId="13" pageNumber="14">B</emphasis>
. Paratype MNHN.F.R50971.
<emphasis id="7BF2EAD6997BFFDF2BD7F900FDF3F0FF" bold="true" box="[1215,1278,1739,1766]" pageId="13" pageNumber="14">D, G</emphasis>
. Paratype MNHN.F.A29249.
<emphasis id="7BF2EAD6997BFFDF2EC8F923F8B8F113" bold="true" box="[416,437,1775,1801]" pageId="13" pageNumber="14">D</emphasis>
. General view.
<emphasis id="7BF2EAD6997BFFDF2D1CF922FB84F112" bold="true" box="[628,649,1774,1800]" pageId="13" pageNumber="14">G</emphasis>
. Drawing.
<emphasis id="7BF2EAD6997BFFDF2C7AF923FA47F113" bold="true" box="[786,842,1775,1801]" pageId="13" pageNumber="14">EH</emphasis>
. Paratype MNHN.F.A84029.
<emphasis id="7BF2EAD6997BFFDF2BD8F923FDCFF113" bold="true" box="[1200,1218,1775,1801]" pageId="13" pageNumber="14">E</emphasis>
. General view.
<emphasis id="7BF2EAD6997BFFDF2FD5F8DEF9DFF136" bold="true" box="[189,210,1810,1836]" pageId="13" pageNumber="14">H</emphasis>
. Close-up of posterior trunk appendages.
<emphasis id="7BF2EAD6997BFFDF2DA1F8DEFBFDF136" bold="true" box="[713,752,1810,1836]" pageId="13" pageNumber="14">IJ</emphasis>
. Paratype MSNM i7591.
<emphasis id="7BF2EAD6997BFFDF2B4FF8DEFD3CF136" bold="true" box="[1063,1073,1810,1836]" pageId="13" pageNumber="14">I</emphasis>
. General view.
<emphasis id="7BF2EAD6997BFFDF2B87F8DEFDF1F136" bold="true" box="[1263,1276,1810,1836]" pageId="13" pageNumber="14">J</emphasis>
. Close-up of raptorial appendages. Abbreviations: cs = circular structure; dm = dorsal midline; os = oval structure; pm = posterior margin; t = posterior trunk; tp
<subScript id="D5023481997BFFDF2D89F8A7FA0EF163" attach="left" box="[737,771,1899,1913]" fontSize="6" pageId="13" pageNumber="14">111</subScript>
= pleural parts of posterior trunk appendages. Scale bars: AB, DE, G, I = 10 mm; C = 1 mm; F = 400 µm; H, J = 5 mm. Photos: A, CF, HJ: T. Laville; B: L. Cazes.
</paragraph>
</caption>
<subSubSection id="019C654F9978FFDC2FD5FEC5FA8CF418" pageId="14" pageNumber="15" type="discussion">
<paragraph id="493936C49978FFDC2FD5FEC5F822F739" blockId="14.[189,303,265,291]" box="[189,303,265,291]" pageId="14" pageNumber="15">
<heading id="127181A89978FFDC2FD5FEC5F822F739" bold="true" box="[189,303,265,291]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
<emphasis id="7BF2EAD69978FFDC2FD5FEC5F822F739" bold="true" box="[189,303,265,291]" pageId="14" pageNumber="15">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="493936C49978FFDC2FD5FEFBFA8CF418" blockId="14.[189,1400,311,514]" pageId="14" pageNumber="15">
The specimens described above display typical characters of
<taxonomicName id="8E864D479978FFDC2CF3FEFBFD21F74B" authorityName="Glaessner" authorityYear="1931" box="[923,1068,311,337]" family="Austriocarididae" genus="Austriocaris" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="7BF2EAD69978FFDC2CF3FEFBFD21F74B" box="[923,1068,311,337]" italics="true" pageId="14" pageNumber="15">Austriocaris</emphasis>
</taxonomicName>
such as the well-developed optic notch restricted to the ventral part of the anterior margin, the large triangular rostrum, a straight dorsal mid-line ending in a spinous postero-dorsal corner. They differ from the
<typeStatus id="963D88669978FFDC2B25FEB2FD8DF782" box="[1101,1152,382,408]" pageId="14" pageNumber="15">type</typeStatus>
species,
<taxonomicName id="8E864D479978FFDC2B8CFEB2F82DF7A0" authorityName="Glaessner" authorityYear="1931" family="Austriocarididae" genus="Austriocaris" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="carinata">
<emphasis id="7BF2EAD69978FFDC2B8CFEB2F82DF7A0" italics="true" pageId="14" pageNumber="15">Austriocaris carinata</emphasis>
</taxonomicName>
, by the hook-shaped and tuberculate dorso-lateral carina, by the presence of horseshoe-like ridges and by the absence of a Y-shaped ventro-lateral carina. Thus, we consider these specimens as belonging to a new species:
<taxonomicName id="8E864D479978FFDC2D61FE2BFA12F41B" authority="Laville &amp; Forel &amp; Charbonnier, 2023" authorityName="Laville &amp; Forel &amp; Charbonnier" authorityYear="2023" box="[521,799,487,513]" family="Austriocarididae" genus="Austriocaris" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="secretanae" status="sp. nov.">
<emphasis id="7BF2EAD69978FFDC2D61FE2BFA12F41B" box="[521,799,487,513]" italics="true" pageId="14" pageNumber="15">Austriocaris secretanae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="60C157AD9978FFDC2C4EFE24FA8CF418" box="[806,897,488,514]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>