325 lines
38 KiB
XML
325 lines
38 KiB
XML
<document id="F16275171CDCA9DAF059E873B4B22018" ID-CLB-Dataset="26002" ID-DOI="10.11646/zootaxa.4657.1.2" ID-GBIF-Dataset="a7c3cd9d-d4be-4686-a198-3255c948beb8" ID-ISSN="1175-5326" ID-Zenodo-Dep="3369801" ID-ZooBank="F27CB967-C001-436A-9F25-184D5484363C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1565968506341" checkinUser="plazi" docAuthor="Perkins, Philip D. & Bergsten, Johannes" docDate="2019" docId="AD076747FFD5FFC9FF6DF88FB63B905F" docLanguage="en" docName="zootaxa.4657.1.2.pdf" docOrigin="Zootaxa 4657 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Incoltorrida madagassica Steffan 1973" docType="treatment" docVersion="12" lastPageNumber="67" masterDocId="513E1F3FFFD2FFC3FFFAFFB9B27A9138" masterDocTitle="New Myxophagan water beetles from Madagascar (Coleoptera: Torridincolidae, Hydroscaphidae)" masterLastPageNumber="96" masterPageNumber="57" pageNumber="64" updateTime="1698751487288" updateUser="ExternalLinkService">
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<mods:titleInfo id="A8291F10F9719ECC7A04D558AB2D7EEC">
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<mods:title id="A4E2A0750DF4706B6110B61340E9EDFC">New Myxophagan water beetles from Madagascar (Coleoptera: Torridincolidae, Hydroscaphidae)</mods:title>
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<mods:namePart id="2D112BA11F031E7B069B0A1E51C59A2D">Perkins, Philip D.</mods:namePart>
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<mods:namePart id="729E2492C7701900B2494F72C8FF9925">Bergsten, Johannes</mods:namePart>
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<mods:title id="010B023BF18B0E0B897459C673FDF660">Zootaxa</mods:title>
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<mods:part id="BB0926CA53417B758CB238F9658511AD">
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<mods:date id="E9604D716E4EBDCE4DE8DE2DA457EBFF">2019</mods:date>
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<mods:number id="90677D30C5E03208122B1D3689704CB2">2019-08-16</mods:number>
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<mods:number id="F3FE575CDD8C04D8C6B6838E83DFFE69">4657</mods:number>
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<mods:number id="C0E6CC31A160CB9ECE76E1ED981FEE49">1</mods:number>
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<treatment id="AD076747FFD5FFC9FF6DF88FB63B905F" ID-DOI="http://doi.org/10.5281/zenodo.5934249" ID-GBIF-Taxon="159131434" ID-Zenodo-Dep="5934249" LSID="urn:lsid:plazi:treatment:AD076747FFD5FFC9FF6DF88FB63B905F" httpUri="http://treatment.plazi.org/id/AD076747FFD5FFC9FF6DF88FB63B905F" lastPageId="10" lastPageNumber="67" pageId="7" pageNumber="64">
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<subSubSection id="6DB485DAFFD5FFC4FF6DF88FB0579668" box="[151,557,1846,1873]" pageId="7" pageNumber="64" type="nomenclature">
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<paragraph id="2511D651FFD5FFC4FF6DF88FB0579668" blockId="7.[151,915,1846,1909]" box="[151,557,1846,1873]" pageId="7" pageNumber="64">
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<heading id="7E59613DFFD5FFC4FF6DF88FB0579668" bold="true" box="[151,557,1846,1873]" fontSize="11" level="1" pageId="7" pageNumber="64" reason="1">
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<taxonomicName id="E2AEADD2FFD5FFC4FF6DF88FB0579668" authority="Steffan" authorityName="Steffan" authorityYear="1973" box="[151,557,1846,1873]" class="Insecta" family="Torridincolidae" genus="Incoltorrida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="64" phylum="Arthropoda" rank="species" species="madagassica">
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<emphasis id="17DA0A43FFD5FFC4FF6DF88FB0579668" bold="true" box="[151,557,1846,1873]" pageId="7" pageNumber="64">
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<emphasis id="17DA0A43FFD5FFC4FF6DF88FB3B69669" bold="true" box="[151,460,1846,1873]" italics="true" pageId="7" pageNumber="64">Incoltorrida madagassica</emphasis>
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Steffan
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="6DB485DAFFD5FFC4FF6DF8E3B1E9964D" box="[151,915,1881,1909]" pageId="7" pageNumber="64" type="description">
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<paragraph id="2511D651FFD5FFC4FF6DF8E3B1E9964D" blockId="7.[151,915,1846,1909]" box="[151,915,1881,1909]" pageId="7" pageNumber="64">
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<figureCitation id="BD95CAD4FFD5FFC4FF6DF8E3B29F964C" box="[151,229,1881,1908]" captionStart="FIGURE 5" captionStartId="5.[151,250,1790,1815]" captionTargetBox="[173,1411,243,1756]" captionTargetId="figure@5.[151,1435,179,1769]" captionTargetPageId="5" captionText="FIGURE 5. Incoltorrida madagassica Steffan, habitus views of non-type with attached Platycola epibionts." httpUri="https://zenodo.org/record/3369811/files/figure.png" pageId="7" pageNumber="64">Figs. 5</figureCitation>
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,
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<figureCitation id="BD95CAD4FFD5FFC4FF0BF8E3B285964C" box="[241,255,1882,1908]" captionStart="FIGURE 6" captionStartId="8.[151,250,2006,2031]" captionTargetBox="[151,1435,1278,1984]" captionTargetId="figure@8.[151,1435,1278,1984]" captionTargetPageId="8" captionText="FIGURE 6. Incoltorrida madagassica Steffan, fragmentary holotype and labels." httpUri="https://zenodo.org/record/3369813/files/figure.png" pageId="7" pageNumber="64">6</figureCitation>
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(habitus); 14, 15, 19 (genitalia); 38 (map); 48, 49 (habitat)
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</paragraph>
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</subSubSection>
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<subSubSection id="6DB485DAFFD5FFC4FF6DF827B01A968F" box="[151,608,1950,1975]" pageId="7" pageNumber="64" type="reference_group">
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<paragraph id="2511D651FFD5FFC4FF6DF827B01A968F" blockId="7.[151,608,1950,1975]" box="[151,608,1950,1975]" pageId="7" pageNumber="64">
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<treatmentCitationGroup id="05BEF17FFFD5FFC4FF6DF827B01A968F" box="[151,608,1950,1975]" pageId="7" pageNumber="64">
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<taxonomicName id="E2AEADD2FFD5FFC4FF6DF827B026968F" authority="Steffan 1973: 635" authorityName="Steffan" authorityPageNumber="635" authorityYear="1973" box="[151,604,1950,1975]" class="Insecta" family="Torridincolidae" genus="Incoltorrida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="64" phylum="Arthropoda" rank="species" species="madagassica">
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||
<emphasis id="17DA0A43FFD5FFC4FF6DF827B3E7968E" box="[151,413,1950,1974]" italics="true" pageId="7" pageNumber="64">Incoltorrida madagassica</emphasis>
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<treatmentCitation id="A40FF040FFD5FFC4FE5EF827B026968F" author="Steffan, A. W." box="[420,604,1950,1975]" page="635" pageId="7" pageNumber="64" year="1973">
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<bibRefCitation id="413FABA0FFD5FFC4FE5EF827B026968F" author="Steffan, A. W." box="[420,604,1950,1975]" pageId="7" pageNumber="64" pagination="633 - 646" refId="ref13951" refString="Steffan, A. W. (1973) Torridincoidarum tres novae species e regione aethiopica (Coleoptera: Sphaeroidea). Revue de Zoologie et de Botanique Africaines, 87, 633 - 646." type="journal article" year="1973">Steffan 1973: 635</bibRefCitation>
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</treatmentCitation>
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</taxonomicName>
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.
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||
</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="6DB485DAFFD5FFC9FF6DF859B63B905F" lastPageId="10" lastPageNumber="67" pageId="7" pageNumber="64" type="materials_examined">
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<paragraph id="2511D651FFD5FFCBFF6DF859B39991C3" blockId="7.[151,1436,2015,2042]" lastBlockId="8.[151,1437,153,1260]" lastPageId="8" lastPageNumber="65" pageId="7" pageNumber="64">
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<emphasis id="17DA0A43FFD5FFC4FF6DF859B33296C2" bold="true" box="[151,328,2015,2042]" pageId="7" pageNumber="64">Type material:</emphasis>
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<typeStatus id="FA1568F3FFD5FFC4FEB5F866B3CD96C1" box="[335,439,2015,2041]" pageId="7" pageNumber="64" type="holotype">Holotype</typeStatus>
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(male) with labels: “
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<materialsCitation id="95C6DC0CFFD5FFCBFD58F859B280918B" ID-GBIF-Occurrence="2332527079" collectionCode="BER-" collectorName="Cap Ambalasandra H." country="Madagascar" lastPageId="8" lastPageNumber="65" location="Madagascar" pageId="7" pageNumber="64" specimenCount="1" typeStatus="holotype">
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<typeStatus id="FA1568F3FFD5FFC4FD58F859B13696C2" box="[674,844,2016,2042]" pageId="7" pageNumber="64" type="holotype">HOLOTYPUS</typeStatus>
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[red]//
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<collectingCountry id="5DB996C1FFD5FFC4FC6CF866B65A96C2" box="[918,1056,2015,2042]" name="Madagascar" pageId="7" pageNumber="64">Madagascar</collectingCountry>
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(65)
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<collectorName id="885BB387FFD5FFC4FBA4F859B72F96C2" box="[1118,1365,2015,2042]" pageId="7" pageNumber="64">Cap Ambalasandra H.</collectorName>
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<materialsCitation id="95C6DC0CFFD5FFCBFAA6F859B280918B" ID-GBIF-Occurrence="2332527076" collectingDate="1960-03-29" collectionCode="TRAND" lastPageId="8" lastPageNumber="65" pageId="7" pageNumber="64" specimenCount="1">
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<collectionCode id="43BF4E94FFD5FFCBFAA6F859B280918B" lastPageId="8" lastPageNumber="65" pageId="7" pageNumber="64">BERTRAND</collectionCode>
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</materialsCitation>
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</materialsCitation>
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,
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<date id="5110F091FFDAFFCBFEF2FF20B31D918B" box="[264,359,153,179]" pageId="8" pageNumber="65" value="1960-03-29">
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||
<collectingDate id="41540979FFDAFFCBFEF2FF20B31D918B" box="[264,359,153,179]" pageId="8" pageNumber="65" value="1960-03-29">29.III.60</collectingDate>
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</date>
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//MUS. ROY. AFR.
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<materialsCitation id="95C6DC0CFFDAFFCBFDAEFF20B29191C3" ID-GBIF-Occurrence="2332527083" collectionCode="CENTR" pageId="8" pageNumber="65" specimenCount="1" typeStatus="holotype">
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<collectionCode id="43BF4E94FFDAFFCBFDAEFF20B0CE918B" box="[596,692,153,179]" pageId="8" pageNumber="65">CENTR</collectionCode>
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||
. caput fehlt! TO-ma-03. mikr prap: 03.01. = genital//
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||
<taxonomicName id="E2AEADD2FFDAFFCBFAE2FF20B39B91EF" authority="STEFFAN 1973" authorityName="Steffan" authorityYear="1973" class="Insecta" family="Torridincolidae" genus="Incoltorrida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="64" phylum="Arthropoda" rank="species" species="madagassica">
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Incoltorrida madagassica
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||
<bibRefCitation id="413FABA0FFDAFFCBFED1FF04B39B91EF" author="Steffan, A. W." box="[299,481,189,215]" pageId="8" pageNumber="65" pagination="633 - 646" refId="ref13951" refString="Steffan, A. W. (1973) Torridincoidarum tres novae species e regione aethiopica (Coleoptera: Sphaeroidea). Revue de Zoologie et de Botanique Africaines, 87, 633 - 646." type="journal article" year="1973">STEFFAN 1973</bibRefCitation>
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</taxonomicName>
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<typeStatus id="FA1568F3FFDAFFCBFE1DFF04B0F791EF" box="[487,653,189,215]" pageId="8" pageNumber="65" type="holotype">HOLOTYPUS</typeStatus>
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||
male [red].” Now deposited, with permanent microslide of male genitalia, in
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</materialsCitation>
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<collectionCode id="43BF4E94FFDAFFCBFF08FF58B33191C3" box="[242,331,225,251]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="8" pageNumber="65">MNHN</collectionCode>
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; see remarks.
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</paragraph>
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<paragraph id="2511D651FFDAFFCBFF3DFEBDB1819097" blockId="8.[151,1437,153,1260]" pageId="8" pageNumber="65">
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<emphasis id="17DA0A43FFDAFFCBFF3DFEBDB39D9027" bold="true" box="[199,487,260,287]" pageId="8" pageNumber="65">Material examined (52):</emphasis>
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<materialsCitation id="95C6DC0CFFDAFFCBFE17FEBDB1D6905F" ID-GBIF-Occurrence="2332527091" collectingDate="2014-11-01" collectionCode="NHRS" collectorName="J. Bergsten & T. Ranarilalatiana & S. Holmgren" county="Amoron'i Mania" elevation="1700" latitude="-20.7722" location="Ankazomivady forest" longLatPrecision="7" longitude="47.1809" municipality="Ambalamanakana" pageId="8" pageNumber="65" specimenCode="MAD14-02" specimenCount="1" stateProvince="Fianarantsoa">
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<emphasis id="17DA0A43FFDAFFCBFE17FEBDB0F69027" bold="true" box="[493,652,260,287]" pageId="8" pageNumber="65">
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<collectingRegion id="E76A18B3FFDAFFCBFE17FEBDB0F69027" box="[493,652,260,287]" country="Madagascar" name="Fianarantsoa" pageId="8" pageNumber="65">Fianarantsoa</collectingRegion>
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</emphasis>
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,
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<geoCoordinate id="409AB096FFDAFFCBFD63FEBCB17E9027" box="[665,772,261,287]" degrees="20.7722" direction="south" orientation="latitude" pageId="8" pageNumber="65" precision="5" value="-20.7722">20.7722S</geoCoordinate>
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<geoCoordinate id="409AB096FFDAFFCBFCF0FEBCB1009027" box="[778,890,261,287]" degrees="47.1809" direction="east" orientation="longitude" pageId="8" pageNumber="65" precision="5" value="47.1809">47.1809E</geoCoordinate>
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;
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<collectingCounty id="CC70AEDDFFDAFFCBFC79FEBCB63B9027" box="[899,1089,261,287]" pageId="8" pageNumber="65">Amoron’i Mania</collectingCounty>
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,
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<quantity id="E2567BB4FFDAFFCBFBB6FEBCB6049027" box="[1100,1150,261,287]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="65" unit="km" value="3.0">3km</quantity>
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south of
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||
<collectingMunicipality id="C5754C2BFFDAFFCBFB19FEBCB2CB907B" pageId="8" pageNumber="65">Ambalamanakana</collectingMunicipality>
|
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next to RN7,
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<location id="2071808AFFDAFFCBFEACFE90B037907B" LSID="urn:lsid:plazi:treatment:AD076747FFD5FFC9FF6DF88FB63B905F:2071808AFFDAFFCBFEACFE90B037907B" box="[342,589,297,323]" county="Amoron'i Mania" latitude="-20.7722" longLatPrecision="7" longitude="47.1809" municipality="Ambalamanakana" name="Ankazomivady forest" pageId="8" pageNumber="65" stateProvince="Fianarantsoa">Ankazomivady forest</location>
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, hygropetric rocks and marsh with vegetation, elev.
|
||
<quantity id="E2567BB4FFDAFFCBFB5FFE90B685907B" box="[1189,1279,297,323]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="8" pageNumber="65" unit="m" value="1700.0">
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<elevation id="AE833162FFDAFFCBFB5FFE90B685907B" box="[1189,1279,297,323]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="8" pageNumber="65" unit="m" value="1700.0">1700 m</elevation>
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</quantity>
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,
|
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<date id="5110F091FFDAFFCBFAF6FE90B707907B" box="[1292,1405,297,323]" pageId="8" pageNumber="65" value="2014-11-01">
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||
<collectingDate id="41540979FFDAFFCBFAF6FE90B707907B" box="[1292,1405,297,323]" pageId="8" pageNumber="65" value="2014-11-01">1 xi 2014</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="885BB387FFDAFFCBFA70FE90B287905F" pageId="8" pageNumber="65">J. Bergsten</collectorName>
|
||
,
|
||
<collectorName id="885BB387FFDAFFCBFEF0FEF4B3A9905F" box="[266,467,333,359]" pageId="8" pageNumber="65">T. Ranarilalatiana</collectorName>
|
||
&
|
||
<collectorName id="885BB387FFDAFFCBFE01FEF4B0F7905F" box="[507,653,333,359]" pageId="8" pageNumber="65">S. Holmgren</collectorName>
|
||
(
|
||
<specimenCode id="75087E2AFFDAFFCBFD64FEF4B159905F" box="[670,803,333,359]" collectionCode="MAD" country="India" httpUri="http://grbio.org/cool/dqz8-sfkc" name="Madras Museum" pageId="8" pageNumber="65">
|
||
<collectionCode id="43BF4E94FFDAFFCBFD64FEF4B09E905F" box="[670,740,333,359]" country="India" httpUri="http://grbio.org/cool/dqz8-sfkc" name="Madras Museum" pageId="8" pageNumber="65">MAD</collectionCode>
|
||
14-02
|
||
</specimenCode>
|
||
) (3
|
||
<collectionCode id="43BF4E94FFDAFFCBFCAFFEF4B1DF905F" box="[853,933,333,359]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="8" pageNumber="65">NHRS</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="95C6DC0CFFDAFFCBFC43FEF5B2CF9097" ID-GBIF-Occurrence="2332527084" collectingDate="2018-02-20" collectionCode="MAD" county="Analanjirofo" latitude="-15.5056" location="Masoala National Park" longLatPrecision="7" longitude="49.7571" municipality="Nosy Mangabe" pageId="8" pageNumber="65" specimenCode="MAD18- 60" specimenCount="1" stateProvince="Toamasina">
|
||
<emphasis id="17DA0A43FFDAFFCBFC43FEF5B641905F" bold="true" box="[953,1083,332,359]" pageId="8" pageNumber="65">
|
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<collectingRegion id="E76A18B3FFDAFFCBFC43FEF5B641905F" box="[953,1083,332,359]" country="Madagascar" name="Toamasina" pageId="8" pageNumber="65">Toamasina</collectingRegion>
|
||
</emphasis>
|
||
:
|
||
<collectingCounty id="CC70AEDDFFDAFFCBFBB0FEF4B6A4905F" box="[1098,1246,333,359]" pageId="8" pageNumber="65">Analanjirofo</collectingCounty>
|
||
: S. side of
|
||
<collectingMunicipality id="C5754C2BFFDAFFCBFA9BFEF4B37890B3" pageId="8" pageNumber="65">Nosy Mangabe</collectingMunicipality>
|
||
,
|
||
<location id="2071808AFFDAFFCBFEF6FEC8B07690B3" LSID="urn:lsid:plazi:treatment:AD076747FFD5FFC9FF6DF88FB63B905F:2071808AFFDAFFCBFEF6FEC8B07690B3" box="[268,524,369,395]" county="Analanjirofo" latitude="-15.5056" longLatPrecision="7" longitude="49.7571" municipality="Nosy Mangabe" name="Masoala National Park" pageId="8" pageNumber="65" stateProvince="Toamasina">Masoala National Park</location>
|
||
, hygropetric rocks by the sea,
|
||
<geoCoordinate id="409AB096FFDAFFCBFCA2FEC9B1BC90B3" box="[856,966,368,395]" degrees="15.5056" direction="south" orientation="latitude" pageId="8" pageNumber="65" precision="5" value="-15.5056">15.5056S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="409AB096FFDAFFCBFC2AFEC9B64490B3" box="[976,1086,368,395]" degrees="49.7571" direction="east" orientation="longitude" pageId="8" pageNumber="65" precision="5" value="49.7571">49.7571E</geoCoordinate>
|
||
, sea level,
|
||
<date id="5110F091FFDAFFCBFB4EFEC8B75690B3" box="[1204,1324,369,395]" pageId="8" pageNumber="65" value="2018-02-20">
|
||
<collectingDate id="41540979FFDAFFCBFB4EFEC8B75690B3" box="[1204,1324,369,395]" pageId="8" pageNumber="65" value="2018-02-20">20.II.2018</collectingDate>
|
||
</date>
|
||
,
|
||
<specimenCode id="75087E2AFFDAFFCBFACCFEC8B2CF9097" collectionCode="MAD" country="India" httpUri="http://grbio.org/cool/dqz8-sfkc" name="Madras Museum" pageId="8" pageNumber="65">
|
||
<collectionCode id="43BF4E94FFDAFFCBFACCFEC8B70690B3" box="[1334,1404,369,395]" country="India" httpUri="http://grbio.org/cool/dqz8-sfkc" name="Madras Museum" pageId="8" pageNumber="65">MAD</collectionCode>
|
||
18- 60
|
||
</specimenCode>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="95C6DC0CFFDAFFCBFF3BFE2CB18D9097" ID-GBIF-Occurrence="2332527070" box="[193,1015,405,431]" collectionCode="NHRS, BMNH, MCZ, PBZT, MBC" collectorName="J. Bergsten & T. Ranarilalatiana" pageId="8" pageNumber="65" specimenCount="1" stateProvince="Toamasina">
|
||
<collectorName id="885BB387FFDAFFCBFF3BFE2CB3479097" box="[193,317,405,431]" pageId="8" pageNumber="65">J. Bergsten</collectorName>
|
||
&
|
||
<collectorName id="885BB387FFDAFFCBFE9AFE2CB05D9097" box="[352,551,405,431]" pageId="8" pageNumber="65">T. Ranarilalatiana</collectorName>
|
||
(49
|
||
<collectionCode id="43BF4E94FFDAFFCBFDA0FE2CB0DD9097" box="[602,679,405,431]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="8" pageNumber="65">NHRS</collectionCode>
|
||
,
|
||
<collectionCode id="43BF4E94FFDAFFCBFD49FE2CB1709097" box="[691,778,405,431]" pageId="8" pageNumber="65">BMNH</collectionCode>
|
||
,
|
||
<collectionCode id="43BF4E94FFDAFFCBFCEFFE2CB12F9097" box="[789,853,405,431]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="8" pageNumber="65" type="Museum">MCZ</collectionCode>
|
||
,
|
||
<collectionCode id="43BF4E94FFDAFFCBFC9AFE2CB1DD9097" box="[864,935,405,431]" country="Madagascar" httpUri="http://grbio.org/cool/a6tz-5rcd" name="Parc Botanique et Zoologique de Tsimbazaza" pageId="8" pageNumber="65">PBZT</collectionCode>
|
||
/
|
||
<collectionCode id="43BF4E94FFDAFFCBFC4AFE2CB18A9097" box="[944,1008,405,431]" country="USA" httpUri="http://grbio.org/cool/tqbp-wnpd" name="Montgomery Botanical Center" pageId="8" pageNumber="65">MBC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="2511D651FFDAFFCBFF3DFE01B64793F7" blockId="8.[151,1437,153,1260]" pageId="8" pageNumber="65">
|
||
<emphasis id="17DA0A43FFDAFFCBFF3DFE01B3AE90EB" bold="true" box="[199,468,440,467]" pageId="8" pageNumber="65">Differential Diagnosis.</emphasis>
|
||
Similar in some characters to
|
||
<taxonomicName id="E2AEADD2FFDAFFCBFCCBFE03B1EF90EB" authorityName="Perkins & Bergsten" authorityYear="2019" box="[817,917,442,467]" class="Insecta" family="Torridincolidae" genus="Incoltorrida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="65" phylum="Arthropoda" rank="species" species="magna">
|
||
<emphasis id="17DA0A43FFDAFFCBFCCBFE03B1EF90EB" box="[817,917,442,467]" italics="true" pageId="8" pageNumber="65">I. magna</emphasis>
|
||
</taxonomicName>
|
||
; differing therefrom by smaller size (ca. 2.31 vs.
|
||
<quantity id="E2567BB4FFDAFFCBFF3AFE64B35390CF" box="[192,297,477,503]" metricMagnitude="-3" metricUnit="m" metricValue="2.62" pageId="8" pageNumber="65" unit="mm" value="2.62">2.62 mm</quantity>
|
||
), less convex elytra with a more gradual posterior declivity, slightly narrower elytral costae, smaller elytral serial punctures, more sinuate fifth elytral costa, and especially differing by having two carinae and a distinct impression between the carinae on the basal 1/2 of the pronotum. The elytron of
|
||
<taxonomicName id="E2AEADD2FFDAFFCBFBE7FD9FB6BD9307" authorityName="Steffan" authorityYear="1973" box="[1053,1223,549,575]" class="Insecta" family="Torridincolidae" genus="Incoltorrida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="65" phylum="Arthropoda" rank="species" species="madagassica">
|
||
<emphasis id="17DA0A43FFDAFFCBFBE7FD9FB6BD9307" box="[1053,1223,549,575]" italics="true" pageId="8" pageNumber="65">I. madagassica</emphasis>
|
||
</taxonomicName>
|
||
has two transverse ridges, offset from one another, one linking costae #3 and #5 (located just behind the humeral umbo), and one linking costae #5 and #8 (located at the sinuation dip of costa #5). In
|
||
<taxonomicName id="E2AEADD2FFDAFFCBFC8EFDD7B1AD93BF" authorityName="Perkins & Bergsten" authorityYear="2019" box="[884,983,622,647]" class="Insecta" family="Torridincolidae" genus="Incoltorrida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="65" phylum="Arthropoda" rank="species" species="magna">
|
||
<emphasis id="17DA0A43FFDAFFCBFC8EFDD7B1AD93BF" box="[884,983,622,647]" italics="true" pageId="8" pageNumber="65">I. magna</emphasis>
|
||
</taxonomicName>
|
||
these two ridges are located in the same plane, not offset from one another. The male genitalia of
|
||
<taxonomicName id="E2AEADD2FFDAFFCBFCE7FD2BB1BD9393" authorityName="Steffan" authorityYear="1973" box="[797,967,657,683]" class="Insecta" family="Torridincolidae" genus="Incoltorrida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="65" phylum="Arthropoda" rank="species" species="madagassica">
|
||
<emphasis id="17DA0A43FFDAFFCBFCE7FD2BB1BD9393" box="[797,967,657,683]" italics="true" pageId="8" pageNumber="65">I. madagassica</emphasis>
|
||
</taxonomicName>
|
||
differ from that of
|
||
<taxonomicName id="E2AEADD2FFDAFFCBFB59FD2BB77D9393" authorityName="Perkins & Bergsten" authorityYear="2019" box="[1187,1287,658,683]" class="Insecta" family="Torridincolidae" genus="Incoltorrida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="65" phylum="Arthropoda" rank="species" species="magna">
|
||
<emphasis id="17DA0A43FFDAFFCBFB59FD2BB77D9393" box="[1187,1287,658,683]" italics="true" pageId="8" pageNumber="65">I. magna</emphasis>
|
||
</taxonomicName>
|
||
in being less arcuate in lateral view, and shaped differently in ventral view (
|
||
<figureCitation id="BD95CAD4FFDAFFCBFCA9FD0CB1A193F7" box="[851,987,692,719]" captionStart="FIGURES 14–18" captionStartId="16.[151,264,1745,1770]" captionTargetBox="[177,1384,429,1687]" captionTargetId="figure@16.[177,1411,345,1726]" captionTargetPageId="16" captionText="FIGURES 14–18. Incoltorrida spp.; (14) I. madagassica, male genitalia of holotype (after Steffan 1973, permanent slide mount). (15–17) male genitalia apex showing pores of I. madagassica and spines of I. benesculpta and I. quintacostata; (18) I. quintacostata, aedeagus of holotype." httpUri="https://zenodo.org/record/3369827/files/figure.png" pageId="8" pageNumber="65">Figs. 14, 15</figureCitation>
|
||
,
|
||
<figureCitation id="BD95CAD4FFDAFFCBFC1DFD0CB67F93F7" box="[999,1029,693,719]" captionStart="FIGURES 19–20" captionStartId="17.[151,264,1761,1786]" captionTargetBox="[235,1353,486,1744]" captionTargetId="figure@17.[235,1353,486,1744]" captionTargetPageId="17" captionText="FIGURES 19–20. (19) Incoltorrida madagassica, male genitalia of non-type; (20) Incoltorrida benesculpta, male genitalia of holotype." httpUri="https://zenodo.org/record/3369829/files/figure.png" pageId="8" pageNumber="65">19</figureCitation>
|
||
,
|
||
<figureCitation id="BD95CAD4FFDAFFCBFBEBFD0CB64A93F7" box="[1041,1072,693,719]" captionStart="FIGURES 21–22" captionStartId="18.[151,264,1508,1533]" captionTargetBox="[151,1410,214,1442]" captionTargetId="figure@18.[151,1436,204,1459]" captionTargetPageId="18" captionText="FIGURES 21–22. Incoltorrida, male genitalia of holotypes. (21) I. magna; (22) I. marojejy." httpUri="https://zenodo.org/record/3369831/files/figure.png" pageId="8" pageNumber="65">21</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="2511D651FFDAFFCBFF3DFD61B1699203" blockId="8.[151,1437,153,1260]" pageId="8" pageNumber="65">
|
||
<emphasis id="17DA0A43FFDAFFCBFF3DFD61B32B93CB" bold="true" box="[199,337,728,755]" pageId="8" pageNumber="65">Description</emphasis>
|
||
. Size:
|
||
<typeStatus id="FA1568F3FFDAFFCBFE5EFD60B07C93CB" box="[420,518,729,755]" pageId="8" pageNumber="65" type="holotype">holotype</typeStatus>
|
||
(length/width, mm): body (length to elytral apices) 2.31/1.29; head width 0.46; pronotum 0.50/0.96; elytra 1.51/1.29. Dorsum dark brown, venter brown to reddish brown, legs brown to reddish brown except tarsi and femoral-tibial articulations black.
|
||
</paragraph>
|
||
<paragraph id="2511D651FFDAFFCBFF3DFCFCB0F992BB" blockId="8.[151,1437,153,1260]" pageId="8" pageNumber="65">Sides of frontoclypeal shield slightly arcuate toward midline at frontoclypeal suture, narrowest part slightly past midlength of shield, at frontoclypeal suture.</paragraph>
|
||
<paragraph id="2511D651FFDAFFCBFF3DFC34B7E092F3" blockId="8.[151,1437,153,1260]" pageId="8" pageNumber="65">Short oblique carinae on the anterior 1/3 of the pronotum are well developed, but there is no indication of a midlongitudinal carina. Two short carinae and a distinct impression between carinae located on basal 1/2 of pronotum.</paragraph>
|
||
<paragraph id="2511D651FFDAFFCBFF3DFC6CB655950F" blockId="8.[151,1437,153,1260]" pageId="8" pageNumber="65">Elytron with eight costae; fourth costa interrupted by strong punctures; fifth costa strongly bisinuate; sixth and seventh costae very indistinct. Two transverse ridges, one linking costae #3 and #5 (located just behind the humeral umbo), and one linking costae #5 and #8 (located at the sinuation dip of costa #5).</paragraph>
|
||
<paragraph id="2511D651FFDAFFCBFF3DFBF8B14C9547" blockId="8.[151,1437,153,1260]" pageId="8" pageNumber="65">Posterior 1/2 of metaventral tabella with closely spaced transverse grooves. Midlongitudinal groove in posterior 1/2 of metaventral tabella shallow, with ill-defined margins.</paragraph>
|
||
<paragraph id="2511D651FFDAFFCBFF3DFB30B6CD959B" blockId="8.[151,1437,153,1260]" box="[199,1207,1161,1187]" pageId="8" pageNumber="65">Midlongitudinal carina of first abdominal ventrite is strong and extends length of ventrite.</paragraph>
|
||
<paragraph id="2511D651FFDAFFCBFF3DFB14B70495D3" blockId="8.[151,1437,153,1260]" pageId="8" pageNumber="65">
|
||
Male genitalia (
|
||
<figureCitation id="BD95CAD4FFDAFFCBFE82FB14B07A95FF" box="[376,512,1196,1223]" captionStart="FIGURES 14–18" captionStartId="16.[151,264,1745,1770]" captionTargetBox="[177,1384,429,1687]" captionTargetId="figure@16.[177,1411,345,1726]" captionTargetPageId="16" captionText="FIGURES 14–18. Incoltorrida spp.; (14) I. madagassica, male genitalia of holotype (after Steffan 1973, permanent slide mount). (15–17) male genitalia apex showing pores of I. madagassica and spines of I. benesculpta and I. quintacostata; (18) I. quintacostata, aedeagus of holotype." httpUri="https://zenodo.org/record/3369827/files/figure.png" pageId="8" pageNumber="65">Figs. 14, 15</figureCitation>
|
||
,
|
||
<figureCitation id="BD95CAD4FFDAFFCBFDF6FB14B05395FF" box="[524,553,1197,1223]" captionStart="FIGURES 19–20" captionStartId="17.[151,264,1761,1786]" captionTargetBox="[235,1353,486,1744]" captionTargetId="figure@17.[235,1353,486,1744]" captionTargetPageId="17" captionText="FIGURES 19–20. (19) Incoltorrida madagassica, male genitalia of non-type; (20) Incoltorrida benesculpta, male genitalia of holotype." httpUri="https://zenodo.org/record/3369829/files/figure.png" pageId="8" pageNumber="65">19</figureCitation>
|
||
) in lateral view slightly arcuate, aedeagal tips wide, apically rounded and with many pores (apparently without microsetae); in ventral view aedeagus nearly parallel-sided, with narrowed tips.
|
||
</paragraph>
|
||
<caption id="71D186D9FFDAFFCBFF6DF86FB1B096D7" box="[151,970,2006,2031]" httpUri="https://zenodo.org/record/3369813/files/figure.png" pageId="8" pageNumber="65" startId="8.[151,250,2006,2031]" targetBox="[151,1435,1278,1984]" targetPageId="8">
|
||
<paragraph id="2511D651FFDAFFCBFF6DF86FB1B096D7" blockId="8.[151,970,2006,2031]" box="[151,970,2006,2031]" pageId="8" pageNumber="65">
|
||
<emphasis id="17DA0A43FFDAFFCBFF6DF86FB36996D7" bold="true" box="[151,275,2006,2031]" pageId="8" pageNumber="65">FIGURE 6.</emphasis>
|
||
<taxonomicName id="E2AEADD2FFDAFFCBFEE3F86EB00B96D7" authority="Steffan" authorityName="Steffan" authorityYear="1973" box="[281,625,2006,2031]" class="Insecta" family="Torridincolidae" genus="Incoltorrida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="65" phylum="Arthropoda" rank="species" species="madagassica">
|
||
<emphasis id="17DA0A43FFDAFFCBFEE3F86EB05A96D7" box="[281,544,2007,2031]" italics="true" pageId="8" pageNumber="65">Incoltorrida madagassica</emphasis>
|
||
Steffan
|
||
</taxonomicName>
|
||
, fragmentary holotype and labels.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="71D186D9FFDBFFCAFF6DF918B1459782" box="[151,831,1697,1722]" httpUri="https://zenodo.org/record/3369815/files/figure.png" pageId="9" pageNumber="66" startId="9.[151,250,1697,1722]" targetBox="[182,1398,192,1651]" targetPageId="9">
|
||
<paragraph id="2511D651FFDBFFCAFF6DF918B1459782" blockId="9.[151,831,1697,1722]" box="[151,831,1697,1722]" pageId="9" pageNumber="66">
|
||
<emphasis id="17DA0A43FFDBFFCAFF6DF918B3699781" bold="true" box="[151,275,1697,1722]" pageId="9" pageNumber="66">FIGURE 7.</emphasis>
|
||
<taxonomicName id="E2AEADD2FFDBFFCAFEE3F918B05C9781" authorityName="Perkins & Bergsten" authorityYear="2019" box="[281,550,1697,1721]" class="Insecta" family="Torridincolidae" genus="Incoltorrida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="66" phylum="Arthropoda" rank="species" species="quintacostata">
|
||
<emphasis id="17DA0A43FFDBFFCAFEE3F918B05C9781" box="[281,550,1697,1721]" italics="true" pageId="9" pageNumber="66">Incoltorrida quintacostata</emphasis>
|
||
</taxonomicName>
|
||
, habitus views of holotype.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="2511D651FFDBFFCAFF3DF951B64C9656" blockId="9.[151,1437,1768,2011]" pageId="9" pageNumber="66">
|
||
<emphasis id="17DA0A43FFDBFFCAFF3DF951B341963A" bold="true" box="[199,315,1768,1794]" pageId="9" pageNumber="66">Remarks.</emphasis>
|
||
In the original description,
|
||
<bibRefCitation id="413FABA0FFDBFFCAFD7BF951B15C963A" author="Steffan, A. W." box="[641,806,1768,1794]" pageId="9" pageNumber="66" pagination="633 - 646" refId="ref13951" refString="Steffan, A. W. (1973) Torridincoidarum tres novae species e regione aethiopica (Coleoptera: Sphaeroidea). Revue de Zoologie et de Botanique Africaines, 87, 633 - 646." type="journal article" year="1973">Steffan (1973)</bibRefCitation>
|
||
stated that the
|
||
<typeStatus id="FA1568F3FFDBFFCAFC26F951B644963A" box="[988,1086,1768,1794]" pageId="9" pageNumber="66" type="holotype">holotype</typeStatus>
|
||
was deposited in the
|
||
<collectionCode id="43BF4E94FFDBFFCAFABAF951B7E3963A" box="[1344,1433,1768,1794]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="66">MNHN</collectionCode>
|
||
. However,
|
||
<collectionCode id="43BF4E94FFDBFFCAFEF1F8B5B31B961E" box="[267,353,1804,1830]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="66">MNHN</collectionCode>
|
||
collection staff were unable to find the type. Search at other museums determined that the
|
||
<typeStatus id="FA1568F3FFDBFFCAFA9BF8B5B2BD9672" pageId="9" pageNumber="66" type="holotype">holotype</typeStatus>
|
||
was still with the
|
||
<typeStatus id="FA1568F3FFDBFFCAFE6AF889B3849672" box="[400,510,1840,1866]" pageId="9" pageNumber="66" type="paratype">paratypes</typeStatus>
|
||
, at the
|
||
<collectionCode id="43BF4E94FFDBFFCAFDB6F889B0D89672" box="[588,674,1840,1866]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="9" pageNumber="66" type="Museum">MRAC</collectionCode>
|
||
. Having permission of that museum, the
|
||
<typeStatus id="FA1568F3FFDBFFCAFB92F889B6B09672" box="[1128,1226,1840,1866]" pageId="9" pageNumber="66" type="holotype">holotype</typeStatus>
|
||
(fragmentary;
|
||
<figureCitation id="BD95CAD4FFDBFFCAFA8AF889B2DC9657" captionStart="FIGURE 6" captionStartId="8.[151,250,2006,2031]" captionTargetBox="[151,1435,1278,1984]" captionTargetId="figure@8.[151,1435,1278,1984]" captionTargetPageId="8" captionText="FIGURE 6. Incoltorrida madagassica Steffan, fragmentary holotype and labels." httpUri="https://zenodo.org/record/3369813/files/figure.png" pageId="9" pageNumber="66">Fig. 6</figureCitation>
|
||
) and the permanent slide mounted male genitalia were transferred to the
|
||
<collectionCode id="43BF4E94FFDBFFCAFC23F8EDB6489656" box="[985,1074,1876,1902]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="66">MNHN</collectionCode>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="2511D651FFDBFFC9FF3DF8C1B28A91EF" blockId="9.[151,1437,1768,2011]" lastBlockId="10.[151,1436,153,359]" lastPageId="10" lastPageNumber="67" pageId="9" pageNumber="66">
|
||
In the original description (
|
||
<bibRefCitation id="413FABA0FFDBFFCAFDF9F8C1B0E096AA" author="Steffan, A. W." box="[515,666,1912,1938]" pageId="9" pageNumber="66" pagination="633 - 646" refId="ref13951" refString="Steffan, A. W. (1973) Torridincoidarum tres novae species e regione aethiopica (Coleoptera: Sphaeroidea). Revue de Zoologie et de Botanique Africaines, 87, 633 - 646." type="journal article" year="1973">Steffan 1973</bibRefCitation>
|
||
), it is noted by the author that the head of the
|
||
<typeStatus id="FA1568F3FFDBFFCAFB47F8C1B76596AA" box="[1213,1311,1912,1938]" pageId="9" pageNumber="66" type="holotype">holotype</typeStatus>
|
||
is missing (“caput fehlt”). However, both the head and the prothorax are missing (
|
||
<figureCitation id="BD95CAD4FFDBFFCAFC3CF825B676968F" box="[966,1036,1948,1975]" captionStart="FIGURE 6" captionStartId="8.[151,250,2006,2031]" captionTargetBox="[151,1435,1278,1984]" captionTargetId="figure@8.[151,1435,1278,1984]" captionTargetPageId="8" captionText="FIGURE 6. Incoltorrida madagassica Steffan, fragmentary holotype and labels." httpUri="https://zenodo.org/record/3369813/files/figure.png" pageId="9" pageNumber="66">Fig. 6</figureCitation>
|
||
). The two
|
||
<typeStatus id="FA1568F3FFDBFFCAFB73F825B68D968E" box="[1161,1271,1948,1974]" pageId="9" pageNumber="66" type="paratype">paratypes</typeStatus>
|
||
, both females, are from a different locality than the
|
||
<typeStatus id="FA1568F3FFDBFFCAFDCAF879B0E896E2" box="[560,658,1984,2010]" pageId="9" pageNumber="66" type="holotype">holotype</typeStatus>
|
||
(“Zentrales
|
||
<collectingRegion id="E76A18B3FFDBFFCAFCE6F879B1F396E2" box="[796,905,1984,2010]" country="United Kingdom" name="Highland" pageId="9" pageNumber="66">Hochland</collectingRegion>
|
||
auf der Strecke Tananarie - Ambositra”). These
|
||
<typeStatus id="FA1568F3FFD8FFC9FF6DFF20B378918B" box="[151,258,153,179]" pageId="10" pageNumber="67" type="paratype">paratypes</typeStatus>
|
||
are larger than the
|
||
<typeStatus id="FA1568F3FFD8FFC9FE12FF20B030918B" box="[488,586,153,179]" pageId="10" pageNumber="67" type="holotype">holotype</typeStatus>
|
||
and have some sculptural differences; they probably represent a different species.
|
||
</paragraph>
|
||
<paragraph id="2511D651FFD8FFC9FF3DFF58B0B7907B" blockId="10.[151,1436,153,359]" pageId="10" pageNumber="67">
|
||
The original description gives “Cap Ambalasandra” as the
|
||
<typeStatus id="FA1568F3FFD8FFC9FCA9FF58B1F991C3" box="[851,899,225,251]" pageId="10" pageNumber="67">type</typeStatus>
|
||
locality. Multiple searches online gave only one record with that geographical name, in a table in a paper on geological deposits. That locality is very near where the Mananara River enters the Indian Ocean (
|
||
<figureCitation id="BD95CAD4FFD8FFC9FD96FE90B0BA907B" box="[620,704,297,323]" captionStart="FIGURES 38–39" captionStartId="27.[151,264,1518,1543]" captionTargetBox="[151,1435,414,1495]" captionTargetId="figure@27.[151,1435,414,1495]" captionTargetPageId="27" captionText="FIGURES 38–39. Distribution maps. (38) Incoltorrida madagassica; (39) Hydroscapha species." httpUri="https://zenodo.org/record/3369845/files/figure.png" pageId="10" pageNumber="67">Fig. 38</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="2511D651FFD8FFC9FF3DFEF4B63B905F" blockId="10.[151,1436,153,359]" box="[199,1089,333,359]" pageId="10" pageNumber="67">
|
||
Thirty-three larvae were collected in association with adults at site
|
||
<collectionCode id="43BF4E94FFD8FFC9FC4CFEF4B187905F" box="[950,1021,333,359]" country="India" httpUri="http://grbio.org/cool/dqz8-sfkc" name="Madras Museum" pageId="10" pageNumber="67">MAD</collectionCode>
|
||
18-60.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |