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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1145.96724" ID-Pensoft-Pub="1313-2970-1145-1" ID-Pensoft-UUID="ACBBD138B7375B0D9F63CE792A82F653" ID-ZooBank="830628C276CD4641BFC6144CD775ED6B" ModsDocID="1313-2970-1145-1" checkinTime="1675506069289" checkinUser="pensoft" docAuthor="Hedin, Marshal &amp; Milne, Marc A." docDate="2023" docId="268D87D2F4DE58C996F2140C6FC87350" docLanguage="en" docName="ZooKeys 1145: 1-130" docOrigin="ZooKeys 1145" docPubDate="2023-02-03" docSource="http://dx.doi.org/10.3897/zookeys.1145.96724" docTitle="Nesticus tennesseensis" docType="treatment" docVersion="1" id="ACBBD138B7375B0D9F63CE792A82F653" lastPageNumber="1" masterDocId="ACBBD138B7375B0D9F63CE792A82F653" masterDocTitle="New species in old mountains: integrative taxonomy reveals ten new species and extensive short-range endemism in Nesticus spiders (Araneae, Nesticidae) from the southern Appalachian Mountains" masterLastPageNumber="130" masterPageNumber="1" pageNumber="1" updateTime="1675506069289" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>New species in old mountains: integrative taxonomy reveals ten new species and extensive short-range endemism in Nesticus spiders (Araneae, Nesticidae) from the southern Appalachian Mountains</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hedin, Marshal</mods:namePart>
<mods:affiliation>Department of Biology, San Diego State University, San Diego, California 92182 - 4614, USA</mods:affiliation>
<mods:nameIdentifier type="email">mhedin@sdsu.edu</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Milne, Marc A.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-1943-0161</mods:nameIdentifier>
<mods:affiliation>Department of Biology, University of Indianapolis, Indianapolis, Indiana 46227, USA</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-02-03</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1145</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>130</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1145.96724</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1145.96724</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1145-1</mods:identifier>
<mods:identifier type="ZooBank">830628C276CD4641BFC6144CD775ED6B</mods:identifier>
<mods:identifier type="Pensoft-UUID">ACBBD138B7375B0D9F63CE792A82F653</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:268D87D2F4DE58C996F2140C6FC87350" httpUri="http://treatment.plazi.org/id/268D87D2F4DE58C996F2140C6FC87350" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName ID-CoL="478ZF" ID-ENA="44403" LSID="268D87D2-F4DE-58C9-96F2-140C6FC87350" authority="(Petrunkevitch, 1925)" baseAuthorityName="Petrunkevitch" baseAuthorityYear="1925" class="Arachnida" family="Nesticidae" genus="Nesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nesticus tennesseensis" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tennesseensis">Nesticus tennesseensis (Petrunkevitch, 1925)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Nesticus tennesseensis ♂ palps, ventral view A West Virginia, Raleigh Co., Grandview SP, MCH specimen # 1360 B Virginia, Tazewell Co., Fallen Rock Cave, MCH specimen # 1808 C Fallen Rock Cave, MCH specimen # 1807. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1145.96724.figure21" httpUri="https://binary.pensoft.net/fig/804472" pageId="0" pageNumber="1">Figs 21A-C</figureCitation>
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Nesticus tennesseensis epigynal variation. Tennessee, Grainger Co., Indian Cave, MCH specimen # 1010, ventral (A), dorsal (B). North Carolina, Surry Co., near Fisher Peak, MCH specimen # N 1126, ventral (C), dorsal (D). Tennessee, Carter Co., near Watauga Lake, MCH 05 _ 078, ventral (E), dorsal (F). West Virginia, Raleigh Co., Grandview SP, MCH specimen # 1362, ventral (G)." figureDoi="10.3897/zookeys.1145.96724.figure22" httpUri="https://binary.pensoft.net/fig/804473" pageId="0" pageNumber="1">, 22A-G</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Magnoliopsida" family="Nesticidae" genus="Ivesia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ivesia tennesseensis" order="Araneae" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="tennesseensis">Ivesia tennesseensis</taxonomicName>
Petrunkevitch, 1925: 321, pl. 20, figs 4, 7, 10.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName family="Nesticidae" genus="Yvesella" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Yvesella tennesseensis" order="Araneae" pageId="0" pageNumber="1" rank="species" species="tennesseensis">Yvesella tennesseensis</taxonomicName>
: Arndt 1928: 84.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Magnoliopsida" family="Nesticidae" genus="Ivesia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ivesia tennesseensis" order="Araneae" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="tennesseensis">Ivesia tennesseensis</taxonomicName>
: Bishop 1950: 10, figs 5-8.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="Petrunkevitch" baseAuthorityYear="1925" class="Arachnida" family="Nesticidae" genus="Nesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nesticus tennesseensis" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tennesseensis">Nesticus tennesseensis</taxonomicName>
:
<bibRefCitation author="Gertsch, WJ" journalOrPublisher="Texas Memorial Museum Bulletin" pageId="0" pageNumber="1" pagination="1 - 91" publicationUrl="http://hdl.handle.net/2152/29952" refId="B12" refString="Gertsch, WJ, 1984. The spider family Nesticidae (Araneae) in North America, Central America, and the West Indies. Texas Memorial Museum Bulletin 31: 1 - 91, http://hdl.handle.net/2152/29952" title="The spider family Nesticidae (Araneae) in North America, Central America, and the West Indies." url="http://hdl.handle.net/2152/29952" volume="31" year="1984">Gertsch 1984</bibRefCitation>
: 23, figs 58-63, 82-84.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
New collections from
<materialsCitation collectingDate="1991-09-25" collectorName="M. Hedin, K. Crandall" county="Grainger Co." location="Indian Cave" municipality="Tennessee" specimenCount="4" specimenCount-female="2" specimenCount-male="2" typeStatus="type">
<typeStatus>type</typeStatus>
locality: -
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingMunicipality>Tennessee</collectingMunicipality>
,
<collectingCounty>Grainger Co.</collectingCounty>
</emphasis>
<specimenCount type="male">2♂</specimenCount>
,
<specimenCount type="female">2♀</specimenCount>
;
<location LSID="urn:lsid:plazi:treatment:268D87D2F4DE58C996F2140C6FC87350:9DD8F6D1EDC3452B1A73F8DD4BD0AF75" county="Grainger Co." municipality="Tennessee" name="Indian Cave">Indian Cave</location>
,
<locationDeviation location="Blaine">E of Blaine</locationDeviation>
;
<collectingDate value="1991-09-25">25 Sep. 1991</collectingDate>
;
<collectorName>M. Hedin</collectorName>
,
<collectorName>K. Crandall</collectorName>
leg.
</materialsCitation>
</emphasis>
; •
<materialsCitation collectingDate="1992-09-21" collectorName="M. Hedin, S. O'Kane" location="Indian Cave" specimenCount="15" specimenCount-female="12" specimenCount-male="3">
<specimenCount type="male">3♂</specimenCount>
,
<specimenCount type="female">12♀</specimenCount>
;
<location LSID="urn:lsid:plazi:treatment:268D87D2F4DE58C996F2140C6FC87350:9B3A93DE488609DFB3A1A1F6F2D2293D" name="Indian Cave">Indian Cave</location>
,
<locationDeviation location="Blaine">E of Blaine</locationDeviation>
;
<collectingDate value="1992-09-21">21 Sep. 1992</collectingDate>
;
<collectorName>M. Hedin</collectorName>
,
<collectorName>
S.
<normalizedToken originalValue="OKane">O'Kane</normalizedToken>
</collectorName>
leg.
</materialsCitation>
; •
<materialsCitation collectingDate="2014-02-22" collectorName="M. L. Niemiller, A. S. Engel, S. Engel, A. Paterson" location="Indian Cave" specimenCount="♂">
<specimenCount></specimenCount>
,
<specimenCount></specimenCount>
, 5 imm;
<location LSID="urn:lsid:plazi:treatment:268D87D2F4DE58C996F2140C6FC87350:A1871C68A8FAF7EF3969894AABB2966F" name="Indian Cave">Indian Cave</location>
, TGA4;
<collectingDate value="2014-02-22">22 Feb. 2014</collectingDate>
;
<collectorName>M.L. Niemiller</collectorName>
,
<collectorName>A.S. Engel</collectorName>
,
<collectorName>S. Engel</collectorName>
,
<collectorName>A. Paterson</collectorName>
leg.
</materialsCitation>
; MLN 14-010.7;
<emphasis bold="true" pageId="0" pageNumber="1">
Non
<materialsCitation collectingDate="2007-08-12" collectorName="M. Hedin, R. Keith" country="USA" county="Surry Co." latitude="36.559" location="Fisher Peak" longLatPrecision="55" longitude="-80.8276" specimenCount="♀" stateProvince="North Carolina" typeStatus="type">
<typeStatus>type</typeStatus>
material:
<collectingCountry name="United States of America">USA</collectingCountry>
-
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingRegion country="United States of America" name="North Carolina">North Carolina</collectingRegion>
,
<collectingCounty>Surry Co.</collectingCounty>
</emphasis>
<specimenCount></specimenCount>
, 1 imm; vic
<location LSID="urn:lsid:plazi:treatment:268D87D2F4DE58C996F2140C6FC87350:6DAB0D4B6F358B61FBF7ED86134536E1" country="USA" county="Surry Co." latitude="36.559" longLatPrecision="55" longitude="-80.8276" name="Fisher Peak" stateProvince="North Carolina">Fisher Peak</location>
lookout tower,
<locationDeviation location="Blue Ridge Parkway">SE of Blue Ridge Parkway</locationDeviation>
;
<geoCoordinate degrees="36.559" direction="north" orientation="latitude" precision="55" value="36.559">36.559°N</geoCoordinate>
, -
<geoCoordinate degrees="80.8276" direction="west" orientation="longitude" precision="5" value="-80.8276">80.8276°W</geoCoordinate>
;
<collectingDate value="2007-08-12">12 Aug. 2007</collectingDate>
;
<collectorName>M. Hedin</collectorName>
,
<collectorName>R. Keith</collectorName>
leg.
</materialsCitation>
</emphasis>
;
<materialsCitation collectingDate="2005-08-23" collectorName="M. Hedin, R. Keith, J. Starrett, S. Thomas" country="United States of America" county="Carter Co." latitude="36.309" location="Watauga Lake" longLatPrecision="55" longitude="-82.0732" municipality="Hwy" specimenCount="3" specimenCount-female="3" stateProvince="Tennessee">
MCH 07_091; -
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingRegion country="United States of America" name="Tennessee">Tennessee</collectingRegion>
,
<collectingCounty>Carter Co.</collectingCounty>
</emphasis>
<specimenCount type="female">3♀</specimenCount>
; off
<collectingMunicipality>Hwy</collectingMunicipality>
167/321, near
<location LSID="urn:lsid:plazi:treatment:268D87D2F4DE58C996F2140C6FC87350:735BA4690984060D58FDD1D75FAC1CA0" country="United States of America" county="Carter Co." latitude="36.309" longLatPrecision="55" longitude="-82.0732" municipality="Hwy" name="Watauga Lake" stateProvince="Tennessee">Watauga Lake</location>
, along
<location LSID="urn:lsid:plazi:treatment:268D87D2F4DE58C996F2140C6FC87350:895CFF0535068F10EB63CC6B3EC14960" country="United States of America" county="Carter Co." latitude="36.309" longLatPrecision="55" longitude="-82.0732" municipality="Hwy" name="Little Stony Creek" stateProvince="Tennessee">Little Stony Creek</location>
;
<geoCoordinate degrees="36.309" direction="north" orientation="latitude" precision="55" value="36.309">36.309°N</geoCoordinate>
, -
<geoCoordinate degrees="82.0732" direction="west" orientation="longitude" precision="5" value="-82.0732">82.0732°W</geoCoordinate>
;
<collectingDate value="2005-08-23">23 Aug. 2005</collectingDate>
;
<collectorName>M. Hedin</collectorName>
,
<collectorName>R. Keith</collectorName>
,
<collectorName>J. Starrett</collectorName>
,
<collectorName>S. Thomas</collectorName>
leg.
</materialsCitation>
;
<materialsCitation collectingDate="2014-06-28" country="United States of America" county="Roane Co." location="Berry Cave" specimenCount="5" specimenCount-male="5" stateProvince="Tennessee">
MCH 05_078; -
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingRegion country="United States of America" name="Tennessee">Tennessee</collectingRegion>
,
<collectingCounty>Roane Co.</collectingCounty>
</emphasis>
<specimenCount type="male">5♂</specimenCount>
,
<specimenCount></specimenCount>
, 11 imm;
<location LSID="urn:lsid:plazi:treatment:268D87D2F4DE58C996F2140C6FC87350:454B479977575FA6D9BC2D045F7C40C2" country="United States of America" county="Roane Co." name="Berry Cave" stateProvince="Tennessee">Berry Cave</location>
, TRN3;
<collectingDate value="2014-06-28">28 Jun. 2014</collectingDate>
</materialsCitation>
;
<materialsCitation collectingDate="2020-06-12" collectorName="M. L. Niemiller, C. D. R. Stephen, A. S. Engel, T. Malabad, P. Tegelman Malabad, K. Malabad, A. Malabad" country="United States of America" county="Alleghany Co." location="Low Moor" municipality="Island Ford Cave" specimenCount="♂" stateProvince="Virginia">
<collectorName>M.L. Niemiller</collectorName>
,
<collectorName>C.D.R. Stephen</collectorName>
,
<collectorName>A.S. Engel</collectorName>
, et al. leg.; MLN 14-034.1; -
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingRegion country="United States of America" name="Virginia">Virginia</collectingRegion>
,
<collectingCounty>Alleghany Co.</collectingCounty>
</emphasis>
<specimenCount></specimenCount>
;
<collectingMunicipality>Island Ford Cave</collectingMunicipality>
, west of
<location LSID="urn:lsid:plazi:treatment:268D87D2F4DE58C996F2140C6FC87350:E75F1ED22356C3FEB240CF84A07C135A" country="United States of America" county="Alleghany Co." municipality="Island Ford Cave" name="Low Moor" stateProvince="Virginia">Low Moor</location>
, VA;
<collectingDate value="2020-06-12">12 Jun. 2020</collectingDate>
;
<collectorName>T. Malabad</collectorName>
,
<collectorName>P. Tegelman Malabad</collectorName>
,
<collectorName>K. Malabad</collectorName>
,
<collectorName>A. Malabad</collectorName>
leg.
</materialsCitation>
; •
<materialsCitation collectingDate="1992-09-16" collectorName="M. Hedin, S. O'Kane" country="United States of America" location="Callaghan" municipality="Rumbold's Cave" specimenCount="13" specimenCount-female="7" specimenCount-male="6" stateProvince="Virginia">
<specimenCount type="male">6♂</specimenCount>
,
<specimenCount type="female">7♀</specimenCount>
;
<collectingMunicipality>
<normalizedToken originalValue="Rumbolds">Rumbold's</normalizedToken>
Cave
</collectingMunicipality>
, near
<location LSID="urn:lsid:plazi:treatment:268D87D2F4DE58C996F2140C6FC87350:14D1AA8000D60246538C2BF21E57CB8C" country="United States of America" municipality="Rumbold's Cave" name="Callaghan" stateProvince="Virginia">Callaghan</location>
;
<collectingDate value="1992-09-16">16 Sep. 1992</collectingDate>
;
<collectorName>M. Hedin</collectorName>
,
<collectorName>
S.
<normalizedToken originalValue="OKane">O'Kane</normalizedToken>
</collectorName>
leg.
</materialsCitation>
; -
<materialsCitation collectingDate="1993-10-10" collectorName="M. Hedin, C. Phillips" country="United States of America" county="Craig Co." location="Newcastle" municipality="Walkthrough Cave" specimenCount="6" specimenCount-female="4" specimenCount-male="2" stateProvince="Virginia">
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingRegion country="United States of America" name="Virginia">Virginia</collectingRegion>
,
<collectingCounty>Craig Co.</collectingCounty>
</emphasis>
<specimenCount type="male">2♂</specimenCount>
,
<specimenCount type="female">4♀</specimenCount>
;
<collectingMunicipality>Walkthrough Cave</collectingMunicipality>
, sw of
<location LSID="urn:lsid:plazi:treatment:268D87D2F4DE58C996F2140C6FC87350:3C496D9F692E7EF9B8E2B84721EB638C" country="United States of America" county="Craig Co." municipality="Walkthrough Cave" name="Newcastle" stateProvince="Virginia">Newcastle</location>
;
<collectingDate value="1993-10-10">10 Oct. 1993</collectingDate>
;
<collectorName>M. Hedin</collectorName>
,
<collectorName>C. Phillips</collectorName>
leg.
</materialsCitation>
; -
<materialsCitation collectingDate="1993-10-09" collectorName="M. Hedin, C. Phillips" country="United States of America" county="Giles Co." location="Pearisburg" municipality="Ballard's Cave" specimenCount="13" specimenCount-female="7" specimenCount-male="6" stateProvince="Virginia">
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingRegion country="United States of America" name="Virginia">Virginia</collectingRegion>
,
<collectingCounty>Giles Co.</collectingCounty>
</emphasis>
<specimenCount type="male">6♂</specimenCount>
,
<specimenCount type="female">7♀</specimenCount>
;
<collectingMunicipality>
<normalizedToken originalValue="Ballards">Ballard's</normalizedToken>
Cave
</collectingMunicipality>
, just S
<location LSID="urn:lsid:plazi:treatment:268D87D2F4DE58C996F2140C6FC87350:D573D8B22B470A8AEA5EDA8FADEBC356" country="United States of America" county="Giles Co." municipality="Ballard's Cave" name="Pearisburg" stateProvince="Virginia">Pearisburg</location>
;
<collectingDate value="1993-10-09">9 Oct. 1993</collectingDate>
;
<collectorName>M. Hedin</collectorName>
,
<collectorName>C. Phillips</collectorName>
leg.
</materialsCitation>
; •
<materialsCitation collectingDate="2019-07-01" collectorName="T. Malabad" country="United States of America" location="Dead Doe Cave" specimenCount="6" specimenCount-female="4" specimenCount-male="2" stateProvince="Virginia">
<specimenCount type="male">2♂</specimenCount>
,
<specimenCount type="female">4♀</specimenCount>
;
<location LSID="urn:lsid:plazi:treatment:268D87D2F4DE58C996F2140C6FC87350:42D46BA83D92833299A669896A360AFE" country="United States of America" name="Dead Doe Cave" stateProvince="Virginia">Dead Doe Cave</location>
;
<collectingDate value="2019-07-01">1 Jul. 2019</collectingDate>
;
<collectorName>T. Malabad</collectorName>
leg.
</materialsCitation>
; •
<materialsCitation collectingDate="2019-08-16" collectorName="T. Malabad" country="United States of America" location="Doe Mountain Cave" specimenCount="♀" stateProvince="Virginia">
<specimenCount></specimenCount>
;
<location LSID="urn:lsid:plazi:treatment:268D87D2F4DE58C996F2140C6FC87350:B2553C851082F0988445FD7E6295171F" country="United States of America" name="Doe Mountain Cave" stateProvince="Virginia">Doe Mountain Cave</location>
;
<collectingDate value="2019-08-16">16 Aug. 2019</collectingDate>
;
<collectorName>T. Malabad</collectorName>
leg.
</materialsCitation>
; •
<materialsCitation collectingDate="2019-11-26" collectorName="T. Malabad, K. Kosic Ficco" country="United States of America" location="Pearisburg" municipality="Hodges Cave" specimenCount="♂" stateProvince="Virginia">
<specimenCount></specimenCount>
;
<collectingMunicipality>Hodges Cave</collectingMunicipality>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="5.632704" unit="mi" value="3.5">3.5 miles</quantity>
southwest of
<location LSID="urn:lsid:plazi:treatment:268D87D2F4DE58C996F2140C6FC87350:6F97FC216F5C426E1662E248CDC0CF5A" country="United States of America" municipality="Hodges Cave" name="Pearisburg" stateProvince="Virginia">Pearisburg</location>
, VA;
<collectingDate value="2019-11-26">26 Nov. 2019</collectingDate>
;
<collectorName>T. Malabad</collectorName>
,
<collectorName>
K.
<normalizedToken originalValue="Kosič">Kosic</normalizedToken>
Ficco
</collectorName>
leg.
</materialsCitation>
; •
<materialsCitation collectingDate="1993-10-10" collectorName="M. Hedin, C. Phillips" country="United States of America" latitude="37.3565" location="Little Stony Creek" longLatPrecision="6" longitude="-80.5921" specimenCount="♀" stateProvince="Virginia">
<specimenCount></specimenCount>
;
<location LSID="urn:lsid:plazi:treatment:268D87D2F4DE58C996F2140C6FC87350:B87C520453E53B049C208566DA801A65" country="United States of America" latitude="37.3565" longLatPrecision="6" longitude="-80.5921" name="Little Stony Creek" stateProvince="Virginia">Little Stony Creek</location>
,
<locationDeviation location="Pembroke">NE of Pembroke</locationDeviation>
;
<geoCoordinate degrees="37.3565" direction="north" orientation="latitude" precision="5" value="37.3565">37.3565°N</geoCoordinate>
, -
<geoCoordinate degrees="80.5921" direction="west" orientation="longitude" precision="5" value="-80.5921">80.5921°W</geoCoordinate>
;
<collectingDate value="1993-10-10">10 Oct. 1993</collectingDate>
;
<collectorName>M. Hedin</collectorName>
,
<collectorName>C. Phillips</collectorName>
leg.
</materialsCitation>
; •
<materialsCitation collectingDate="2013-07-11" collectorName="C Richart" country="United States of America" latitude="37.3755" location="Jefferson Nat Forest" longLatPrecision="6" longitude="-80.5158" municipality="Mountain Lake Biological Station" specimenCount="♂" stateProvince="Virginia">
<specimenCount></specimenCount>
;
<collectingMunicipality>Mountain Lake Biological Station</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:268D87D2F4DE58C996F2140C6FC87350:30EE57686039CC6B1A9868CCF4668451" country="United States of America" latitude="37.3755" longLatPrecision="6" longitude="-80.5158" municipality="Mountain Lake Biological Station" name="Jefferson Nat Forest" stateProvince="Virginia">Jefferson Nat Forest</location>
;
<geoCoordinate degrees="37.3755" direction="north" orientation="latitude" precision="5" value="37.3755">37.3755°N</geoCoordinate>
, -
<geoCoordinate degrees="80.5158" direction="west" orientation="longitude" precision="5" value="-80.5158">80.5158°W</geoCoordinate>
;
<collectingDate value="2013-07-11">11 Jul. 2013</collectingDate>
;
<collectorName>C Richart</collectorName>
leg.
</materialsCitation>
; •
<materialsCitation collectingDate="1993-10-10" collectorName="M. Hedin, C. Phillips" country="United States of America" county="Starne's Cave" location="Wilburn Valley" municipality="Pearisburg" specimenCount="14" specimenCount-female="10" specimenCount-male="4" stateProvince="Virginia">
<specimenCount type="male">4♂</specimenCount>
,
<specimenCount type="female">10♀</specimenCount>
;
<collectingCounty>
<normalizedToken originalValue="Starnes">Starne's</normalizedToken>
Cave
</collectingCounty>
, SW
<collectingMunicipality>Pearisburg</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:268D87D2F4DE58C996F2140C6FC87350:953658D474CE90B5D954CB0F9C52FA51" country="United States of America" county="Starne's Cave" municipality="Pearisburg" name="Wilburn Valley" stateProvince="Virginia">Wilburn Valley</location>
;
<collectingDate value="1993-10-10">10 Oct. 1993</collectingDate>
;
<collectorName>M. Hedin</collectorName>
,
<collectorName>C. Phillips</collectorName>
leg.
</materialsCitation>
; •
<materialsCitation collectingDate="2019-06-03" collectorName="T. Malabad" country="United States of America" location="Starnes Cave" specimenCount="♂" stateProvince="Virginia">
<specimenCount></specimenCount>
5 imm;
<location LSID="urn:lsid:plazi:treatment:268D87D2F4DE58C996F2140C6FC87350:534960B01AD04C3AE4DA14BD5C6BC825" country="United States of America" name="Starnes Cave" stateProvince="Virginia">Starnes Cave</location>
;
<collectingDate value="2019-06-03">3 Jun. 2019</collectingDate>
;
<collectorName>T. Malabad</collectorName>
leg.
</materialsCitation>
; •
<materialsCitation collectingDate="2019-08-09" collectorName="T. Malabad" country="United States of America" location="Yer Cave" specimenCount="4" specimenCount-female="4" stateProvince="Virginia">
<specimenCount></specimenCount>
,
<specimenCount type="female">4♀</specimenCount>
, 15 imm;
<location LSID="urn:lsid:plazi:treatment:268D87D2F4DE58C996F2140C6FC87350:902D89DE85C9539F0A973909504DB8F3" country="United States of America" name="Yer Cave" stateProvince="Virginia">Yer Cave</location>
;
<collectingDate value="2019-08-09">9 Aug. 2019</collectingDate>
;
<collectorName>T. Malabad</collectorName>
leg.
</materialsCitation>
; -
<materialsCitation collectingDate="2014-12-14" collectorName="E. Koertge" country="United States of America" county="Montgomery Co." location="Hancock Blowhole Cave" specimenCount="4" specimenCount-female="4" stateProvince="Virginia">
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingRegion country="United States of America" name="Virginia">Virginia</collectingRegion>
,
<collectingCounty>Montgomery Co.</collectingCounty>
</emphasis>
<specimenCount type="female">4♀</specimenCount>
, 1 imm;
<location LSID="urn:lsid:plazi:treatment:268D87D2F4DE58C996F2140C6FC87350:1A852DF2C72AB385CBAAA1DE1A997B89" country="United States of America" county="Montgomery Co." name="Hancock Blowhole Cave" stateProvince="Virginia">Hancock Blowhole Cave</location>
;
<collectingDate value="2014-12-14">14 Dec. 2014</collectingDate>
;
<collectorName>E. Koertge</collectorName>
leg.
</materialsCitation>
; -
<materialsCitation collectingDate="2015-09-15" collectorName="W. Orndorff" country="United States of America" county="Scott Co." location="Coley Cave" specimenCount="♀" stateProvince="Virginia">
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingRegion country="United States of America" name="Virginia">Virginia</collectingRegion>
,
<collectingCounty>Scott Co.</collectingCounty>
</emphasis>
<specimenCount></specimenCount>
, 6 imm;
<location LSID="urn:lsid:plazi:treatment:268D87D2F4DE58C996F2140C6FC87350:FBCAE98A4B81BEAAD7BDD5936DB565DE" country="United States of America" county="Scott Co." name="Coley Cave" stateProvince="Virginia">Coley Cave</location>
#2;
<collectingDate value="2015-09-15">15 Sep. 2015</collectingDate>
;
<collectorName>W. Orndorff</collectorName>
leg.
</materialsCitation>
; •
<materialsCitation collectingDate="2016-08-04" collectorName="T. Malabad" country="United States of America" location="Herrons Echo Hall" specimenCount="♀" stateProvince="Virginia">
<specimenCount></specimenCount>
;
<location LSID="urn:lsid:plazi:treatment:268D87D2F4DE58C996F2140C6FC87350:1C5304476F1767D43578E35747ADDC32" country="United States of America" name="Herrons Echo Hall" stateProvince="Virginia">Herrons Echo Hall</location>
;
<collectingDate value="2016-08-04">4 Aug. 2016</collectingDate>
;
<collectorName>T. Malabad</collectorName>
leg.
</materialsCitation>
; -
<materialsCitation collectingDate="1993-10-09" collectorName="M. Hedin, C. Phillips" country="United States of America" county="Tazewell Co." location="Burkes Garden" municipality="Cassell's Farm Cave" specimenCount="17" specimenCount-female="11" specimenCount-male="6" stateProvince="Virginia">
<collectingRegion country="United States of America" name="Virginia">Virginia</collectingRegion>
,
<collectingCounty>Tazewell Co.</collectingCounty>
<specimenCount type="male">6♂</specimenCount>
,
<specimenCount type="female">11♀</specimenCount>
;
<collectingMunicipality>
<normalizedToken originalValue="Cassells">Cassell's</normalizedToken>
Farm Cave
</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:268D87D2F4DE58C996F2140C6FC87350:2D86E826C1B77A5B7174D462FDDD04EB" country="United States of America" county="Tazewell Co." municipality="Cassell's Farm Cave" name="Burkes Garden" stateProvince="Virginia">Burkes Garden</location>
;
<collectingDate value="1993-10-09">9 Oct. 1993</collectingDate>
;
<collectorName>M. Hedin</collectorName>
,
<collectorName>C. Phillips</collectorName>
leg.
</materialsCitation>
; •
<materialsCitation collectingDate="2018-10-24" collectorName="T. Malabad" country="United States of America" location="Cauliflower Cave" specimenCount="2" specimenCount-female="2" stateProvince="Virginia">
<specimenCount type="female">2♀</specimenCount>
, 2 imm;
<location LSID="urn:lsid:plazi:treatment:268D87D2F4DE58C996F2140C6FC87350:B14A6B94C414F1E571F9A701A962CFA5" country="United States of America" name="Cauliflower Cave" stateProvince="Virginia">Cauliflower Cave</location>
;
<collectingDate value="2018-10-24">24 Oct. 2018</collectingDate>
;
<collectorName>T. Malabad</collectorName>
leg.
</materialsCitation>
; •
<materialsCitation collectingDate="2017-03-04" collectorName="K. Kosic Ficco" country="United States of America" location="Corkscrew Cave" specimenCount="1" stateProvince="Virginia">
2 imm;
<location LSID="urn:lsid:plazi:treatment:268D87D2F4DE58C996F2140C6FC87350:509E465487C4F8265AC594322BF86CA7" country="United States of America" name="Corkscrew Cave" stateProvince="Virginia">Corkscrew Cave</location>
;
<collectingDate value="2017-03-04">4 Mar. 2017</collectingDate>
;
<collectorName>
K.
<normalizedToken originalValue="Kosič">Kosic</normalizedToken>
Ficco
</collectorName>
leg.
</materialsCitation>
; •
<materialsCitation collectingDate="2018-10-27" collectorName="T. Malabad" country="United States of America" location="Corkscrew Cave" specimenCount="♀" stateProvince="Virginia">
<specimenCount></specimenCount>
, 4 imm;
<location LSID="urn:lsid:plazi:treatment:268D87D2F4DE58C996F2140C6FC87350:D5236C86A273C1E8588994EEC3ED5510" country="United States of America" name="Corkscrew Cave" stateProvince="Virginia">Corkscrew Cave</location>
;
<collectingDate value="2018-10-27">27 Oct. 2018</collectingDate>
;
<collectorName>T. Malabad</collectorName>
leg.
</materialsCitation>
; •
<materialsCitation collectingDate="1993-10-09" collectorName="M. Hedin, C. Phillips" country="United States of America" county="Fallen Rock Cave" location="Maiden Spring" municipality="Ward Cove" specimenCount="12" specimenCount-female="8" specimenCount-male="4" stateProvince="Virginia">
<specimenCount type="male">4♂</specimenCount>
,
<specimenCount type="female">8♀</specimenCount>
;
<collectingCounty>Fallen Rock Cave</collectingCounty>
,
<collectingMunicipality>Ward Cove</collectingMunicipality>
, S
<location LSID="urn:lsid:plazi:treatment:268D87D2F4DE58C996F2140C6FC87350:447ECD1C3D57791B57101BF7942890D2" country="United States of America" county="Fallen Rock Cave" municipality="Ward Cove" name="Maiden Spring" stateProvince="Virginia">Maiden Spring</location>
;
<collectingDate value="1993-10-09">9 Oct. 1993</collectingDate>
;
<collectorName>M. Hedin</collectorName>
,
<collectorName>C. Phillips</collectorName>
leg.
</materialsCitation>
; •
<materialsCitation collectingDate="2019-09-09" collectorName="T. Malabad, K. Kosic Ficco, A. Futrell" country="United States of America" location="Liberty" municipality="Gillespie Water Cave" specimenCount="♂" stateProvince="Virginia">
<specimenCount></specimenCount>
;
<collectingMunicipality>Gillespie Water Cave</collectingMunicipality>
, southwest of
<location LSID="urn:lsid:plazi:treatment:268D87D2F4DE58C996F2140C6FC87350:7261574DFB6029DAAD71F681CA93BD3B" country="United States of America" municipality="Gillespie Water Cave" name="Liberty" stateProvince="Virginia">Liberty</location>
, VA;
<collectingDate value="2019-09-09">9 Sep. 2019</collectingDate>
;
<collectorName>T. Malabad</collectorName>
,
<collectorName>
K.
<normalizedToken originalValue="Kosič">Kosic</normalizedToken>
Ficco
</collectorName>
,
<collectorName>A. Futrell</collectorName>
leg.
</materialsCitation>
; •
<materialsCitation collectingDate="2018-10-24" collectorName="T. Malabad" country="United States of America" location="Glenwood Church Cave" specimenCount="4" specimenCount-female="4" stateProvince="Virginia">
<specimenCount></specimenCount>
,
<specimenCount type="female">4♀</specimenCount>
, 9 imm;
<location LSID="urn:lsid:plazi:treatment:268D87D2F4DE58C996F2140C6FC87350:C9287A0A4E5CEF03DFB5E17D261E5329" country="United States of America" name="Glenwood Church Cave" stateProvince="Virginia">Glenwood Church Cave</location>
;
<collectingDate value="2018-10-24">24 Oct. 2018</collectingDate>
;
<collectorName>T. Malabad</collectorName>
leg.
</materialsCitation>
; •
<materialsCitation collectingDate="2019-07-22" collectorName="T. Malabad" country="United States of America" location="Gulley Cave" specimenCount="♀" stateProvince="Virginia">
<specimenCount></specimenCount>
, 5 imm;
<location LSID="urn:lsid:plazi:treatment:268D87D2F4DE58C996F2140C6FC87350:BE08159527072D3EF82F015BE6DF7268" country="United States of America" name="Gulley Cave" stateProvince="Virginia">Gulley Cave</location>
;
<collectingDate value="2019-07-22">22 Jul. 2019</collectingDate>
;
<collectorName>T. Malabad</collectorName>
leg.
</materialsCitation>
; •
<materialsCitation collectingDate="2018-11-29" collectorName="T. Malabad" country="United States of America" location="Little River Cave" specimenCount="2" specimenCount-female="2" stateProvince="Virginia">
<specimenCount type="female">2♀</specimenCount>
, 3 imm;
<location LSID="urn:lsid:plazi:treatment:268D87D2F4DE58C996F2140C6FC87350:1CE156D4C4FFCD7E339CAF28F5C9F00F" country="United States of America" name="Little River Cave" stateProvince="Virginia">Little River Cave</location>
;
<collectingDate value="2018-11-29">29 Nov. 2018</collectingDate>
;
<collectorName>T. Malabad</collectorName>
leg.
</materialsCitation>
; •
<materialsCitation collectingDate="2019-07-22" collectorName="T. Malabad" country="United States of America" location="Lost Mill Cave" specimenCount="3" specimenCount-female="3" stateProvince="Virginia">
<specimenCount type="female">3♀</specimenCount>
, 4 imm;
<location LSID="urn:lsid:plazi:treatment:268D87D2F4DE58C996F2140C6FC87350:5DE7B6204F5E498EC4C2FFA89D9945C2" country="United States of America" name="Lost Mill Cave" stateProvince="Virginia">Lost Mill Cave</location>
1;
<collectingDate value="2019-07-22">22 Jul. 2019</collectingDate>
;
<collectorName>T. Malabad</collectorName>
leg.
</materialsCitation>
; •
<materialsCitation collectingDate="2019-07-22" collectorName="T. Malabad" country="United States of America" location="Lost Mill Cave" specimenCount="2" specimenCount-female="2" stateProvince="Virginia">
<specimenCount type="female">2♀</specimenCount>
, 2 imm;
<location LSID="urn:lsid:plazi:treatment:268D87D2F4DE58C996F2140C6FC87350:C534C8105E37530E0B11F3739A0AD3DF" country="United States of America" name="Lost Mill Cave" stateProvince="Virginia">Lost Mill Cave</location>
2;
<collectingDate value="2019-07-22">22 Jul. 2019</collectingDate>
;
<collectorName>T. Malabad</collectorName>
leg.
</materialsCitation>
; •
<materialsCitation collectingDate="2021-05-05" collectorName="T. Malabad, K. Kosic Ficco, M. Ficco" country="United States of America" location="Claypool Hill" municipality="Stompbottom Cave" specimenCount="2" specimenCount-female="2" stateProvince="Virginia">
<specimenCount type="female">2♀</specimenCount>
;
<collectingMunicipality>Stompbottom Cave</collectingMunicipality>
, southeast of
<location LSID="urn:lsid:plazi:treatment:268D87D2F4DE58C996F2140C6FC87350:75BF669464DD086361C9617EEFCC24DE" country="United States of America" municipality="Stompbottom Cave" name="Claypool Hill" stateProvince="Virginia">Claypool Hill</location>
, VA;
<collectingDate value="2021-05-05">5 May. 2021</collectingDate>
;
<collectorName>T. Malabad</collectorName>
,
<collectorName>
K.
<normalizedToken originalValue="Kosič">Kosic</normalizedToken>
Ficco
</collectorName>
,
<collectorName>M. Ficco</collectorName>
leg.
</materialsCitation>
; •
<materialsCitation collectingDate="2019-01-10" collectorName="T. Malabad" country="United States of America" location="Stonley Cave" specimenCount="3" specimenCount-female="3" stateProvince="Virginia">
<specimenCount></specimenCount>
,
<specimenCount type="female">3♀</specimenCount>
;
<location LSID="urn:lsid:plazi:treatment:268D87D2F4DE58C996F2140C6FC87350:A2EA56669BDBF783B3F87890CFB56B83" country="United States of America" name="Stonley Cave" stateProvince="Virginia">Stonley Cave</location>
;
<collectingDate value="2019-01-10">10 Jan. 2019</collectingDate>
;
<collectorName>T. Malabad</collectorName>
leg.
</materialsCitation>
; -
<materialsCitation collectingDate="1992-09-15" collectorName="M. Hedin, S. O'Kane" country="United States of America" county="Raleigh Co." latitude="37.8321" location="River Gorge" longLatPrecision="6" longitude="-81.0614" municipality="Grandview State Park" specimenCount="5" specimenCount-female="5" stateProvince="West Virginia">
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingRegion country="United States of America" name="West Virginia">West Virginia</collectingRegion>
,
<collectingCounty>Raleigh Co.</collectingCounty>
</emphasis>
<specimenCount></specimenCount>
,
<specimenCount type="female">5♀</specimenCount>
;
<collectingMunicipality>Grandview State Park</collectingMunicipality>
, New
<location LSID="urn:lsid:plazi:treatment:268D87D2F4DE58C996F2140C6FC87350:ACB6DBD3FC82162CF82BDEE8611D2C1B" country="United States of America" county="Raleigh Co." latitude="37.8321" longLatPrecision="6" longitude="-81.0614" municipality="Grandview State Park" name="River Gorge" stateProvince="West Virginia">River Gorge</location>
;
<geoCoordinate degrees="37.8321" direction="north" orientation="latitude" precision="5" value="37.8321">37.8321°N</geoCoordinate>
, -
<geoCoordinate degrees="81.0614" direction="west" orientation="longitude" precision="5" value="-81.0614">81.0614°W</geoCoordinate>
;
<collectingDate value="1992-09-15">15 Sep. 1992</collectingDate>
;
<collectorName>M. Hedin</collectorName>
,
<collectorName>
S.
<normalizedToken originalValue="OKane">O'Kane</normalizedToken>
</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="Petrunkevitch" baseAuthorityYear="1925" class="Arachnida" family="Nesticidae" genus="Nesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nesticus tennesseensis" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tennesseensis">
<emphasis italics="true" pageId="0" pageNumber="1">Nesticus tennesseensis</emphasis>
</taxonomicName>
and
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="N. dilutus" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="dilutus">
<emphasis italics="true" pageId="0" pageNumber="1">N. dilutus</emphasis>
</taxonomicName>
are morphologically similar sister species. Male differences are noted in the Diagnosis of
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="N. dilutus" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="dilutus">
<emphasis italics="true" pageId="0" pageNumber="1">N. dilutus</emphasis>
</taxonomicName>
below. Male
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="N. tennesseensis" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tennesseensis">
<emphasis italics="true" pageId="0" pageNumber="1">N. tennesseensis</emphasis>
</taxonomicName>
may be differentiated from other members of this species group by the combination of palps with a paracymbium with a wide, broad ventral process, rectangular paradistal paracymbial process, a rectangular median apophysis with an anteriorly-pointed sclerotized edge, and a narrow, singularly-pointed tegular apophysis that extends to ~ half the length of the median apophysis (Fig.
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Nesticus tennesseensis ♂ palps, ventral view A West Virginia, Raleigh Co., Grandview SP, MCH specimen # 1360 B Virginia, Tazewell Co., Fallen Rock Cave, MCH specimen # 1808 C Fallen Rock Cave, MCH specimen # 1807. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1145.96724.figure21" httpUri="https://binary.pensoft.net/fig/804472" pageId="0" pageNumber="1">21A-C</figureCitation>
). Females may be differentiated from other members of this species group by an overall rounded epigynum with short, somewhat globular spermathecae that extend perpendicular to the median septum (Fig.
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Nesticus tennesseensis epigynal variation. Tennessee, Grainger Co., Indian Cave, MCH specimen # 1010, ventral (A), dorsal (B). North Carolina, Surry Co., near Fisher Peak, MCH specimen # N 1126, ventral (C), dorsal (D). Tennessee, Carter Co., near Watauga Lake, MCH 05 _ 078, ventral (E), dorsal (F). West Virginia, Raleigh Co., Grandview SP, MCH specimen # 1362, ventral (G)." figureDoi="10.3897/zookeys.1145.96724.figure22" httpUri="https://binary.pensoft.net/fig/804473" pageId="0" pageNumber="1">22A-G</figureCitation>
). Viewed dorsally, circular pockets lie above extended parallel separated medial margins that diverge posteriorly.
</paragraph>
<caption doi="10.3897/zookeys.1145.96724.figure21" httpUri="https://binary.pensoft.net/fig/804472" pageId="0" pageNumber="1" start="Figure 21" startId="F21">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 21.</emphasis>
<taxonomicName baseAuthorityName="Petrunkevitch" baseAuthorityYear="1925" class="Arachnida" family="Nesticidae" genus="Nesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nesticus tennesseensis" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tennesseensis">
<emphasis italics="true" pageId="0" pageNumber="1">Nesticus tennesseensis</emphasis>
</taxonomicName>
♂ palps, ventral view
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
West Virginia, Raleigh Co., Grandview SP, MCH specimen #1360
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
Virginia, Tazewell Co., Fallen Rock Cave, MCH specimen #1808
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
Fallen Rock Cave, MCH specimen #1807. Scale bar: 0.5 mm.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1145.96724.figure22" httpUri="https://binary.pensoft.net/fig/804473" pageId="0" pageNumber="1" start="Figure 22" startId="F22">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 22.</emphasis>
<taxonomicName baseAuthorityName="Petrunkevitch" baseAuthorityYear="1925" class="Arachnida" family="Nesticidae" genus="Nesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nesticus tennesseensis" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tennesseensis">
<emphasis italics="true" pageId="0" pageNumber="1">Nesticus tennesseensis</emphasis>
</taxonomicName>
epigynal variation. Tennessee, Grainger Co., Indian Cave, MCH specimen #1010, ventral (
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
), dorsal (
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
). North Carolina, Surry Co., near Fisher Peak, MCH specimen #N1126, ventral (
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
), dorsal (
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
). Tennessee, Carter Co., near Watauga Lake, MCH 05_078, ventral (
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
), dorsal (
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
). West Virginia, Raleigh Co., Grandview SP, MCH specimen #1362, ventral (
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="variation">
<paragraph pageId="0" pageNumber="1">Variation.</paragraph>
<paragraph pageId="0" pageNumber="1">
The shape of the tegular apophysis varies slightly across sample locations. Specimens from most locations (similar to type material from Indian Cave) possess a tegular apophysis with a broad, L-shaped base and an acute tip (see Gertsch, 1984: fig. 58), whereas other specimens have a narrower base with a gradually tapering tip (Fig.
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Nesticus tennesseensis ♂ palps, ventral view A West Virginia, Raleigh Co., Grandview SP, MCH specimen # 1360 B Virginia, Tazewell Co., Fallen Rock Cave, MCH specimen # 1808 C Fallen Rock Cave, MCH specimen # 1807. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1145.96724.figure21" httpUri="https://binary.pensoft.net/fig/804472" pageId="0" pageNumber="1">21A, C</figureCitation>
). Different specimens from the Fallen Rock Cave population exhibit both conditions (Fig.
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Nesticus tennesseensis ♂ palps, ventral view A West Virginia, Raleigh Co., Grandview SP, MCH specimen # 1360 B Virginia, Tazewell Co., Fallen Rock Cave, MCH specimen # 1808 C Fallen Rock Cave, MCH specimen # 1807. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1145.96724.figure21" httpUri="https://binary.pensoft.net/fig/804472" pageId="0" pageNumber="1">21B, C</figureCitation>
). We observed only minor variation in epigyna (Fig.
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Nesticus tennesseensis epigynal variation. Tennessee, Grainger Co., Indian Cave, MCH specimen # 1010, ventral (A), dorsal (B). North Carolina, Surry Co., near Fisher Peak, MCH specimen # N 1126, ventral (C), dorsal (D). Tennessee, Carter Co., near Watauga Lake, MCH 05 _ 078, ventral (E), dorsal (F). West Virginia, Raleigh Co., Grandview SP, MCH specimen # 1362, ventral (G)." figureDoi="10.3897/zookeys.1145.96724.figure22" httpUri="https://binary.pensoft.net/fig/804473" pageId="0" pageNumber="1">22A-G</figureCitation>
), even for geographically disjunct southeastern Surry and Carter County populations (Figs
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Distribution of Nesticus tennesseensis group species. Type localities designated with yellow circles. State boundaries and major cities shown for geographic context. Dashed lines circumscribe known species distributions." figureDoi="10.3897/zookeys.1145.96724.figure13" httpUri="https://binary.pensoft.net/fig/804464" pageId="0" pageNumber="1">13</figureCitation>
,
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Nesticus tennesseensis epigynal variation. Tennessee, Grainger Co., Indian Cave, MCH specimen # 1010, ventral (A), dorsal (B). North Carolina, Surry Co., near Fisher Peak, MCH specimen # N 1126, ventral (C), dorsal (D). Tennessee, Carter Co., near Watauga Lake, MCH 05 _ 078, ventral (E), dorsal (F). West Virginia, Raleigh Co., Grandview SP, MCH specimen # 1362, ventral (G)." figureDoi="10.3897/zookeys.1145.96724.figure22" httpUri="https://binary.pensoft.net/fig/804473" pageId="0" pageNumber="1">22C-F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution and natural history.</paragraph>
<paragraph pageId="0" pageNumber="1">
Known from both limestone caves (both shallow and deeper situations) and dark, cool, relatively moist near-surface habitats (e.g., rock piles, shallow cliff caves). Most known populations are from caves in the upper-central Appalachian Valley and Ridge, with a few peripheral montane surface populations (e.g., Surry County, NC; Carter County, TN; Raleigh County, WV). The southeastern Surry and Carter County populations appear disjunct, separated from the remainder of the
<normalizedToken originalValue="species">species'</normalizedToken>
range by regions occupied by other taxa in the species group (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Distribution of Nesticus tennesseensis group species. Type localities designated with yellow circles. State boundaries and major cities shown for geographic context. Dashed lines circumscribe known species distributions." figureDoi="10.3897/zookeys.1145.96724.figure13" httpUri="https://binary.pensoft.net/fig/804464" pageId="0" pageNumber="1">13</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation author="Gertsch, WJ" journalOrPublisher="Texas Memorial Museum Bulletin" pageId="0" pageNumber="1" pagination="1 - 91" publicationUrl="http://hdl.handle.net/2152/29952" refId="B12" refString="Gertsch, WJ, 1984. The spider family Nesticidae (Araneae) in North America, Central America, and the West Indies. Texas Memorial Museum Bulletin 31: 1 - 91, http://hdl.handle.net/2152/29952" title="The spider family Nesticidae (Araneae) in North America, Central America, and the West Indies." url="http://hdl.handle.net/2152/29952" volume="31" year="1984">Gertsch (1984</bibRefCitation>
: 24-25) incorrectly identified specimens from the following locations as
<taxonomicName baseAuthorityName="Petrunkevitch" baseAuthorityYear="1925" class="Arachnida" family="Nesticidae" genus="Nesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nesticus tennesseensis" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tennesseensis">
<emphasis italics="true" pageId="0" pageNumber="1">Nesticus tennesseensis</emphasis>
</taxonomicName>
: Sensabaugh Saltpeter Cave (3 imm) and &quot;Cave by Clinch River&quot; (one ♀) specimens belong to
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="N. paynei" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="paynei">
<emphasis italics="true" pageId="0" pageNumber="1">N. paynei</emphasis>
</taxonomicName>
.
<normalizedToken originalValue="Straleys">Straley's</normalizedToken>
Cave No. 1 (2 imm), Harris Cave (1 ♀), and
<normalizedToken originalValue="Potters">Potter's</normalizedToken>
Cave (1 ♂, 1 ♀) specimens belong to
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="N. mimus" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mimus">
<emphasis italics="true" pageId="0" pageNumber="1">N. mimus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>