153 lines
12 KiB
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153 lines
12 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/JHR.44.4727" ID-GBIF-Dataset="5335d345-00bc-40f6-a476-0f2efddeef76" ID-Pensoft-Pub="1314-2607-44-57" ID-Pensoft-UUID="5F4F7145FFD80F29FFE2FFA3FFADFFC0" ID-ZBK="8C4FCBF6E5494987B3D22C343258CD8B" ID-Zenodo-Dep="575048" ID-ZooBank="8C4FCBF6E5494987B3D22C343258CD8B" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1314-2607-44-57" ModsDocOrigin="Journal of Hymenoptera Research 44" ModsDocTitle="Colemanuskeeleyorum (Braconidae, Ichneutinae s. l.): a new genus and species of Eocene wasp from the Green River Formation of western North America" checkinTime="1451252199124" checkinUser="pensoft" docAuthor="Fisher, J. Ray, Tucker, Erika M. & Sharkey, Michael J." docDate="2015" docId="C91577AF43A3AD7CBC5B3A249DA7E31A" docLanguage="en" docName="JourHymenoptRes 44: 57-67" docOrigin="Journal of Hymenoptera Research 44" docPubDate="2015-06-11" docSource="http://dx.doi.org/10.3897/JHR.44.4727" docTitle="Colemanus keeleyorum Fisher 2015, sp. n." docType="treatment" docUuid="AC0F5EA3-4526-43FD-85F3-4544FFC0EF5E" docUuidSource="ZooBank" docVersion="4" id="5F4F7145FFD80F29FFE2FFA3FFADFFC0" lastPageNumber="64" masterDocId="5F4F7145FFD80F29FFE2FFA3FFADFFC0" masterDocTitle="Colemanus keeleyorum (Braconidae, Ichneutinae s. l.): a new genus and species of Eocene wasp from the Green River Formation of western North America" masterLastPageNumber="67" masterPageNumber="57" pageNumber="61" updateTime="1678748503564" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Colemanus keeleyorum (Braconidae, Ichneutinae s. l.): a new genus and species of Eocene wasp from the Green River Formation of western North America</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fisher, J. Ray</mods:namePart>
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<mods:nameIdentifier type="email">jrfisher@uark.edu</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Tucker, Erika M.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sharkey, Michael J.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Journal of Hymenoptera Research</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2015</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2015-06-11</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>44</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>57</mods:start>
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<mods:end>67</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/JHR.44.4727</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/JHR.44.4727</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2607-44-57</mods:identifier>
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<mods:identifier type="ZooBank">8C4FCBF6E5494987B3D22C343258CD8B</mods:identifier>
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<mods:identifier type="Pensoft-UUID">5F4F7145FFD80F29FFE2FFA3FFADFFC0</mods:identifier>
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<mods:identifier type="Zenodo-Dep">575048</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152018110" LSID="urn:lsid:zoobank.org:act:AC0F5EA3-4526-43FD-85F3-4544FFC0EF5E" httpUri="http://treatment.plazi.org/id/C91577AF43A3AD7CBC5B3A249DA7E31A" lastPageId="7" lastPageNumber="64" pageId="4" pageNumber="61">
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<subSubSection pageId="4" pageNumber="61" type="nomenclature">
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<paragraph pageId="4" pageNumber="61">
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<taxonomicName LSID="http://zoobank.org/AC0F5EA3-4526-43FD-85F3-4544FFC0EF5E" authority="Fisher" authorityName="Fisher" authorityYear="2015" class="Insecta" family="Braconidae" genus="Colemanus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Colemanus keeleyorum" order="Hymenoptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="keeleyorum" status="sp. n.">Colemanus keeleyorum Fisher</taxonomicName>
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<taxonomicNameLabel pageId="4" pageNumber="61">sp. n.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="62" type="description">
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<paragraph pageId="5" pageNumber="62">
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Fossil of Colemanus keeleyorum sp. n.: A whole fossil B close-up of mesosoma C close-up of left pterostigma, note preserved microtrichia." figureDoi="10.3897/JHR.44.4727.figure1" httpUri="https://binary.pensoft.net/fig/45059" pageId="5" pageNumber="62">
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<pageBreakToken pageId="5" pageNumber="62" start="start">Figs</pageBreakToken>
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1
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</figureCitation>
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Reconstructions of Colemanus keeleyorum sp. n.: A dorsal mesosoma, note heavy sculpturing B wings, note last abscissa of forewing radial sector (3 RSb) is recurved." figureDoi="10.3897/JHR.44.4727.figure2" httpUri="https://binary.pensoft.net/fig/45060" pageId="5" pageNumber="62">, 2</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="62" type="diagnosis">
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<paragraph pageId="5" pageNumber="62">
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<pageBreakToken pageId="5" pageNumber="62" start="start">Diagnosis</pageBreakToken>
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.
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</paragraph>
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<paragraph pageId="5" pageNumber="62">
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<taxonomicName authorityName="Fisher" authorityYear="2015" class="Insecta" family="Braconidae" genus="Colemanus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Colemanus keeleyorum" order="Hymenoptera" pageId="5" pageNumber="62" phylum="Arthropoda" rank="species" species="keeleyorum">
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<emphasis italics="true" pageId="5" pageNumber="62">Colemanus keeleyorum</emphasis>
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</taxonomicName>
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can be distinguished from
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<taxonomicName class="Reptilia" kingdom="Animalia" lsidName="C. contortus" order="Hymenoptera" pageId="5" pageNumber="62" phylum="Chordata" rank="species" species="contortus">
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<emphasis italics="true" pageId="5" pageNumber="62">C. contortus</emphasis>
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</taxonomicName>
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(Brues, 1933) (new combination; see below) by having curved (RS+M)a and 2RS veins in the fore wing (straight in
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<taxonomicName class="Reptilia" kingdom="Animalia" lsidName="C. contortus" order="Hymenoptera" pageId="5" pageNumber="62" phylum="Chordata" rank="species" species="contortus">
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<emphasis italics="true" pageId="5" pageNumber="62">C. contortus</emphasis>
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</taxonomicName>
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). Also,
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<taxonomicName class="Reptilia" kingdom="Animalia" lsidName="C. keeleyorum" order="Hymenoptera" pageId="5" pageNumber="62" phylum="Chordata" rank="species" species="keeleyorum">
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<emphasis italics="true" pageId="5" pageNumber="62">C. keeleyorum</emphasis>
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</taxonomicName>
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is only known from the western United States (Eocene) and
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<taxonomicName class="Reptilia" kingdom="Animalia" lsidName="C. contortus" order="Hymenoptera" pageId="5" pageNumber="62" phylum="Chordata" rank="species" species="contortus">
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<emphasis italics="true" pageId="5" pageNumber="62">C. contortus</emphasis>
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</taxonomicName>
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is only known from the Baltic region (lower Oligocene).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="62" type="description">
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<paragraph pageId="5" pageNumber="62">Description.</paragraph>
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<paragraph pageId="5" pageNumber="62">
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Holotype (n = 1): body length 9 mm (estimated due to incomplete metasoma); sex unknown.
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<emphasis bold="true" pageId="5" pageNumber="62">Head</emphasis>
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(Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Fossil of Colemanus keeleyorum sp. n.: A whole fossil B close-up of mesosoma C close-up of left pterostigma, note preserved microtrichia." figureDoi="10.3897/JHR.44.4727.figure1" httpUri="https://binary.pensoft.net/fig/45059" pageId="5" pageNumber="62">1A</figureCitation>
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) 1.4 mm long and 1.7 mm wide. Antenna 7.2 mm long with 33-34 flagellomeres.
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<emphasis bold="true" pageId="5" pageNumber="62">Mesosoma</emphasis>
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(Figs
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Fossil of Colemanus keeleyorum sp. n.: A whole fossil B close-up of mesosoma C close-up of left pterostigma, note preserved microtrichia." figureDoi="10.3897/JHR.44.4727.figure1" httpUri="https://binary.pensoft.net/fig/45059" pageId="5" pageNumber="62">1B</figureCitation>
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,
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Reconstructions of Colemanus keeleyorum sp. n.: A dorsal mesosoma, note heavy sculpturing B wings, note last abscissa of forewing radial sector (3 RSb) is recurved." figureDoi="10.3897/JHR.44.4727.figure2" httpUri="https://binary.pensoft.net/fig/45060" pageId="5" pageNumber="62">2A</figureCitation>
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) 3.8 mm long and 3.3 mm wide; robust and heavily sculptured; notauli deeply crenulate; scutellum with crenulate depression medio-posteriorly; side of scutellum deeply hollowed with crenulate carinae and bordered by prominent carinae; metanotum with median, raised, rectangular tubercle having radiating carinae; scutellar sulcus with carinae; propodeum with areolate sculpturing.
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<emphasis bold="true" pageId="5" pageNumber="62">Wings</emphasis>
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(Figs
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Fossil of Colemanus keeleyorum sp. n.: A whole fossil B close-up of mesosoma C close-up of left pterostigma, note preserved microtrichia." figureDoi="10.3897/JHR.44.4727.figure1" httpUri="https://binary.pensoft.net/fig/45059" pageId="5" pageNumber="62">1A,C</figureCitation>
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,
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Reconstructions of Colemanus keeleyorum sp. n.: A dorsal mesosoma, note heavy sculpturing B wings, note last abscissa of forewing radial sector (3 RSb) is recurved." figureDoi="10.3897/JHR.44.4727.figure2" httpUri="https://binary.pensoft.net/fig/45060" pageId="5" pageNumber="62">2B</figureCitation>
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) with last abscissa of fore wing radial sector (3RSb) recurved and tubular as it reaches the wing margin; (RS+M)b short, nearly vertical; 1cu-a originating far distal to M and curving downward, thus not angling toward wing apex; fore wing M curved; tubular portion of 1a long; C+SC and R closely fused; second abscissa of RS strongly curved; parastigma well-developed; hind wing M+Cu longer than first abscissa of M; hind wing r-crossvein absent; hind wing M+Cu in posterior half of wing; 1A strongly developed.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="62" type="biology">
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<paragraph pageId="5" pageNumber="62">Biology.</paragraph>
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<paragraph pageId="5" pageNumber="62">
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Unknown. However, placement within
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<taxonomicName authorityName="van Achterberg" authorityYear="1976" lsidName="" pageId="5" pageNumber="62" rank="tribe" tribe="Proteropini">Proteropini</taxonomicName>
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is suggestive of shared biology, koinobiont endoparasitoids of sawflies.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="62" type="remarks">
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<paragraph pageId="5" pageNumber="62">Remarks.</paragraph>
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<paragraph pageId="5" pageNumber="62">Wings, antennae, and dorsal mesosoma are overall well-preserved; metasoma and legs either did not completely fossilize or are obscured by the rock matrix; head is crushed.</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="63" type="etymology">
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<paragraph pageId="6" pageNumber="63">
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<pageBreakToken pageId="6" pageNumber="63" start="start">Etymology</pageBreakToken>
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.
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</paragraph>
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<paragraph pageId="6" pageNumber="63">
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Named for Dr. Jack and Flo Keeley, who, together with their daughter and her husband (first
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<normalizedToken originalValue="author’s">author's</normalizedToken>
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mother & father), were largely responsible for the first
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<normalizedToken originalValue="author’s">author's</normalizedToken>
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pursuit of the natural sciences.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="64" type="material examined">
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<paragraph pageId="7" pageNumber="64">
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<pageBreakToken pageId="7" pageNumber="64" start="start">Material</pageBreakToken>
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examined.
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</paragraph>
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<paragraph pageId="7" pageNumber="64">HOLOTYPE: USA, Colorado, Piceance Creek Basin, Parachute Member, 2005. Deposited with the David Kohls collection in the Smithsonian Institution, Museum of Natural History, Washington D.C.</paragraph>
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</subSubSection>
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</treatment>
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</document> |