284 lines
39 KiB
XML
284 lines
39 KiB
XML
<document id="4267DA6AD0B8410150484E4FF8216AFB" ID-DOI="10.24199/j.mmv.2004.61.13" ID-ISSN="1447-2554" ID-Zenodo-Dep="12209670" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1718990350741" checkinUser="felipe" docAuthor="Holmes, Francis C." docDate="2004" docId="5E4CE76FFFBF9E7C4386B8F1FE27FA8D" docLanguage="en" docName="MemMusVictoria.61.2.209-216.pdf" docOrigin="Memoirs of Museum Victoria 61 (2)" docSource="https://museumsvictoria.com.au/collections-research/journals/memoirs-of-museum-victoria/volume-61-issue-2-2004/pages-209-216/" docStyle="DocumentStyle:5B3B5C052A93BFEEE334AD6097A94E45.2:MemMusVictoria.2003-.journal_article.type1" docStyleId="5B3B5C052A93BFEEE334AD6097A94E45" docStyleName="MemMusVictoria.2003-.journal_article.type1" docStyleVersion="2" docTitle="Rhyncopygus janchrisorum Holmes, 2004, sp. nov." docType="treatment" docVersion="2" lastPageNumber="213" masterDocId="A2759F17FFBD9E78430DBD79FFBDFFA7" masterDocTitle="A new Late Eocene cassiduloid (Echinoidea) from Yorke Peninsula, South Australia" masterLastPageNumber="216" masterPageNumber="209" pageNumber="211" updateTime="1718999035966" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="E310188131849F3D45A970140DC347AC">
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<mods:title id="A3E8329F9A1205162EC53FA0AEFACA79">A new Late Eocene cassiduloid (Echinoidea) from Yorke Peninsula, South Australia</mods:title>
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<mods:name id="7EF9055F41A7B6C11C31535364409453" type="personal">
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<mods:roleTerm id="714F4FCD2736CD316385D0EA3666B282">Author</mods:roleTerm>
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<mods:namePart id="412EF9F9E3F6DFBDB145E7FAB4701028">Holmes, Francis C.</mods:namePart>
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<mods:typeOfResource id="A80CFA4ED982575623640C05242ED3D0">text</mods:typeOfResource>
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<mods:title id="B9C66DBAFD7690B5E989C824E2E07806">Memoirs of Museum Victoria</mods:title>
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</mods:titleInfo>
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<mods:part id="F6A47EEC54036A1FE503295CA16E7D66">
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<mods:date id="3ACB2238C9F723FC1C0AFDE6FCA80AEB">2004</mods:date>
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<mods:detail id="DB0416535EB6D430C048CDD403502972" type="pubDate">
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<mods:number id="C4AD29B21E71C89ACD2BBA7803F9B1F0">2004-12-31</mods:number>
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<mods:detail id="90E823419639EAD4954B54D626258CE8" type="volume">
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<mods:number id="C6AA152718F661CBD65D3A82B5E5B3E7">61</mods:number>
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<mods:number id="CBCE6D7ED0641CAC6C0653C9B5744F26">2</mods:number>
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<mods:start id="B3661CEC1440FD9B3DBC252A36E80C0B">209</mods:start>
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<mods:location id="31F9509C12AAC2EB63490E164F72E605">
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<mods:url id="4AECE18346A901C15B4D2940D8A5412F">https://museumsvictoria.com.au/collections-research/journals/memoirs-of-museum-victoria/volume-61-issue-2-2004/pages-209-216/</mods:url>
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</mods:location>
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<mods:classification id="C3CA4783B58819E7CF74B27CCA3B1839">journal article</mods:classification>
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<mods:identifier id="0963ECACC217F03B9E11E1012B3D40F5" type="DOI">10.24199/j.mmv.2004.61.13</mods:identifier>
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<mods:identifier id="79D4C36CC7BF63304188102BA3DF9ABE" type="ISSN">1447-2554</mods:identifier>
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<mods:identifier id="B86E20B628335153BCF25102AFABB9BE" type="Zenodo-Dep">12209670</mods:identifier>
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</mods:mods>
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<treatment id="5E4CE76FFFBF9E7C4386B8F1FE27FA8D" LSID="urn:lsid:plazi:treatment:5E4CE76FFFBF9E7C4386B8F1FE27FA8D" httpUri="http://treatment.plazi.org/id/5E4CE76FFFBF9E7C4386B8F1FE27FA8D" lastPageId="4" lastPageNumber="213" pageId="2" pageNumber="211">
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<subSubSection id="9EFF05F2FFBF9E7A4386B8F1FDADFA39" box="[139,528,1415,1438]" pageId="2" pageNumber="211" type="nomenclature">
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<paragraph id="D65A5679FFBF9E7A4386B8F1FDADFA39" blockId="2.[139,528,1415,1438]" box="[139,528,1415,1438]" pageId="2" pageNumber="211">
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<heading id="8D12E115FFBF9E7A4386B8F1FDADFA39" box="[139,528,1415,1438]" fontSize="9" level="2" pageId="2" pageNumber="211" reason="5">
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<taxonomicName id="11E52DFAFFBF9E7A4386B8F1FE07FA39" authorityName="Holmes" authorityYear="2004" box="[139,442,1415,1438]" class="Echinoidea" family="Cassidulidae" genus="Rhyncopygus" kingdom="Animalia" order="Echinolampadacea" pageId="2" pageNumber="211" phylum="Echinodermata" rank="species" species="janchrisorum" status="sp. nov.">
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<emphasis id="E4918A6BFFBF9E7A4386B8F1FE07FA39" bold="true" box="[139,442,1415,1438]" pageId="2" pageNumber="211">
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<emphasis id="E4918A6BFFBF9E7A4386B8F1FEA5FA39" bold="true" box="[139,280,1416,1438]" italics="true" pageId="2" pageNumber="211">Rhyncopygus</emphasis>
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?
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<emphasis id="E4918A6BFFBF9E7A4221B8F1FE07FA39" bold="true" box="[300,442,1416,1438]" italics="true" pageId="2" pageNumber="211">janchrisorum</emphasis>
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</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="FFA23710FFBF9E7A42CCB8F1FDADFA39" box="[449,528,1416,1438]" pageId="2" pageNumber="211" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="9EFF05F2FFBF9E7C4386B8CCFE27FA8D" lastPageId="4" lastPageNumber="213" pageId="2" pageNumber="211" type="description">
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<paragraph id="D65A5679FFBF9E7A4386B8CCFE4CFA6C" blockId="2.[139,497,1461,1483]" box="[139,497,1461,1483]" pageId="2" pageNumber="211">
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<figureCitation id="4EDE4AFCFFBF9E7A4386B8CCFEA5FA6C" box="[139,280,1461,1483]" captionStart="Figure 2" captionStartId="3.[131,190,1086,1106]" captionTargetBox="[130,1451,222,1067]" captionTargetId="figure-11@3.[130,1452,222,1067]" captionTargetPageId="3" captionText="Figure 2. Rhynchopygus? janchrisorum sp. nov. A–C, adapical, right lateral and posterior views of holotype NMV P145616; D, adoral view of paratype NMV P312115; E, adoral view of paratype NMV P312114; F, adapical view of paratype NMV P312113. All specimens form the Late Eocene Muloowurtie Formation, Yorke Peninsula, South Australia. Scale bar 10 mm." figureDoi="http://doi.org/10.5281/zenodo.12209677" httpUri="https://zenodo.org/record/12209677/files/figure.png" pageId="2" pageNumber="211">Figures 2A–F</figureCitation>
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,
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<figureCitation id="4EDE4AFCFFBF9E7A422EB8CCFEDEFA6C" box="[291,355,1461,1483]" captionStart="Figure 3" captionStartId="3.[131,190,1812,1832]" captionTargetBox="[130,754,1294,1793]" captionTargetId="figure-163@3.[130,528,1294,1793]" captionTargetPageId="3" captionText="Figure 3. Apical plate structure. A and B, Rhynchopygus? janchrisorum sp. nov., Late Eocene holotype NMV P145616; C, R. marmini, Late Cretaceous USNM 19559 from Port Brechay, La Manche, France (drawing adapted from Kier, 1962). Scale bars 1mm." figureDoi="http://doi.org/10.5281/zenodo.12209681" httpUri="https://zenodo.org/record/12209681/files/figure.png" pageId="2" pageNumber="211">3A, B</figureCitation>
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,
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<figureCitation id="4EDE4AFCFFBF9E7A4263B8CCFE10FA6C" box="[366,429,1461,1483]" captionStart="Figure 4" captionStartId="3.[811,870,1675,1695]" captionTargetBox="[813,1452,1294,1649]" captionTargetId="figure-124@3.[808,1452,1293,1657]" captionTargetPageId="3" captionText="Figure 4. Rhynchopygus? janchrisorum sp. nov. A–C, reconstruction of adapical, right lateral and posterior characteristics of holotype NMV P145616 (actual specimen outline shown with broken line). Scale bar 10 mm." figureDoi="http://doi.org/10.5281/zenodo.12209679" httpUri="https://zenodo.org/record/12209679/files/figure.png" pageId="2" pageNumber="211">4A–C</figureCitation>
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,
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<figureCitation id="4EDE4AFCFFBF9E7A42B5B8CCFE7BFA6C" box="[440,454,1461,1483]" captionStart="Figure 5" captionStartId="4.[139,198,632,652]" captionTargetBox="[250,661,224,610]" captionTargetId="figure-614@4.[178,780,220,614]" captionTargetPageId="4" captionText="Figure 5. Rhynchopygus? janchrisorum sp. nov. Drawing of adapical surface of NMV P312113 showing plate structure distal to petals. Scale bar 10 mm." figureDoi="http://doi.org/10.5281/zenodo.12209683" httpUri="https://zenodo.org/record/12209683/files/figure.png" pageId="2" pageNumber="211">5</figureCitation>
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,
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<figureCitation id="4EDE4AFCFFBF9E7A42DFB8CCFE4CFA6C" box="[466,497,1461,1483]" captionStart="Figure 6" captionStartId="4.[141,200,1789,1809]" captionTargetBox="[137,1460,1355,1763]" captionTargetId="figure-522@4.[136,1460,1345,1771]" captionTargetPageId="4" captionText="Figure 6. Comparative drawings of A, Late Eocene Rhynchopygus? janchrisorum sp. nov. from Yorke Peninsula, South Australia; B, Late Cretaceous (Late Maastrichtian) R. donetzensis from the Severny Donetz Basin, Ukraine (adapted from Smith and Jeffery, 2000); C1, Late Cretaceous (Late? Santonian) R. marmini from La Richardière, Dissay, France; C2, posterior profile of Late Cretaceous (Late Campanian) R. marmini from near Royan and St Palais-sur-Mer, France. Based on published illustrations, latter profile typical of Maastrichtian R. marmini specimens in general. Scale bars 5 mm." figureDoi="http://doi.org/10.5281/zenodo.12209687" httpUri="https://zenodo.org/record/12209687/files/figure.png" pageId="2" pageNumber="211">6A</figureCitation>
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</paragraph>
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<paragraph id="D65A5679FFBF9E7A4386B8A6FD18F9F9" blockId="2.[139,780,1503,1630]" pageId="2" pageNumber="211">
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<emphasis id="E4918A6BFFBF9E7A4386B8A6FEB5FA54" box="[139,264,1503,1523]" italics="true" pageId="2" pageNumber="211">Type material</emphasis>
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.
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<materialsCitation id="668D5C24FFBF9E7A421EB8A6FE37F98F" collectionCode="NMV" county="Late Eocene" location="Yorke Peninsula" municipality="Muloowurtie Formation" pageId="2" pageNumber="211" specimenCode="P145616" specimenCount="1" stateProvince="South Australia" typeStatus="holotype">
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<typeStatus id="095EE8DBFFBF9E7A421EB8A6FED7FA54" box="[275,362,1503,1523]" pageId="2" pageNumber="211" type="holotype">Holotype</typeStatus>
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,
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<collectionCode id="B0F4CEBCFFBF9E7A427EB8A6FE15FA54" box="[371,424,1503,1523]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="2" pageNumber="211" type="Museum">NMV</collectionCode>
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<specimenCode id="8643FE02FFBF9E7A42A1B8A6FE40FA54" box="[428,509,1503,1523]" collectionCode="NMV" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="2" pageNumber="211" type="Museum">P145616</specimenCode>
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from
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<collectingCounty id="3F3B2EF5FFBF9E7A4139B8A6FD18FA54" box="[564,677,1503,1523]" pageId="2" pageNumber="211">Late Eocene</collectingCounty>
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(Aldingan, Priabonian)
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<collectingMunicipality id="363ECC03FFBF9E7A4201B883FE42F9A9" box="[268,511,1530,1550]" pageId="2" pageNumber="211">Muloowurtie Formation</collectingMunicipality>
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, “Sliding Rocks”,
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<location id="D33A00A2FFBF9E7A41DEB883FF58F98F" LSID="urn:lsid:plazi:treatment:5E4CE76FFFBF9E7C4386B8F1FE27FA8D:D33A00A2FFBF9E7A41DEB883FF58F98F" county="Late Eocene" municipality="Muloowurtie Formation" name="Yorke Peninsula" pageId="2" pageNumber="211" stateProvince="South Australia">Yorke Peninsula</location>
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,
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<collectingRegion id="1421989BFFBF9E7A43F8BB6DFE37F98F" box="[245,394,1556,1576]" country="Australia" name="South Australia" pageId="2" pageNumber="211">South Australia</collectingRegion>
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</materialsCitation>
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.
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<materialsCitation id="668D5C24FFBF9E7A4294BB6DFD0DF98F" box="[409,688,1556,1576]" collectionCode="NMV" pageId="2" pageNumber="211" specimenCode="P312113" specimenCount="1" stateProvince="South Australia" typeStatus="paratype">
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<typeStatus id="095EE8DBFFBF9E7A4294BB6DFE49F98F" box="[409,500,1556,1576]" pageId="2" pageNumber="211" type="paratype">Paratypes</typeStatus>
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.
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<collectionCode id="B0F4CEBCFFBF9E7A410FBB6DFD8AF98F" box="[514,567,1556,1576]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="2" pageNumber="211" type="Museum">NMV</collectionCode>
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<specimenCode id="8643FE02FFBF9E7A414EBB6DFD2FF98F" box="[579,658,1556,1576]" collectionCode="NMV" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="2" pageNumber="211" type="Museum">P312113</specimenCode>
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to
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</materialsCitation>
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<materialsCitation id="668D5C24FFBF9E7A41B1BB6DFD1CF9F9" collectionCode="NMV" latitude="-34.5695" longLatPrecision="11" longitude="137.89" pageId="2" pageNumber="211" specimenCode="P312115" specimenCount="1" stateProvince="South Australia" typeStatus="paratype">
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<specimenCode id="8643FE02FFBF9E7A41B1BB6DFCB6F98F" box="[700,779,1556,1576]" collectionCode="NMV" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="2" pageNumber="211" type="Museum">P312115</specimenCode>
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from the uppermost bed of the same formation, NNE of “Sliding Rocks” at approximately
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<geoCoordinate id="B3D130BEFFBF9E7A4279BB33FE67F9F9" box="[372,474,1608,1630]" degrees="34" direction="south" minutes="34.17" orientation="latitude" pageId="2" pageNumber="211" precision="9" value="-34.5695">34°34.17'S</geoCoordinate>
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,
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<geoCoordinate id="B3D130BEFFBF9E7A42E8BB33FDE8F9F9" box="[485,597,1610,1630]" degrees="137" direction="east" minutes="53.40" orientation="longitude" pageId="2" pageNumber="211" precision="9" value="137.89">137°53.40'E</geoCoordinate>
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(
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<figureCitation id="4EDE4AFCFFBF9E7A416FBB33FD25F9F9" box="[610,664,1610,1630]" captionStart="Figure 1" captionStartId="1.[131,190,953,973]" captionTargetBox="[129,1451,221,935]" captionTargetId="figure-462@1.[129,1452,221,935]" captionTargetPageId="1" captionText="Figure 1. A and B, general location maps; C, map of St Vincent Basin in relation to Yorke Peninsula, Kangaroo Island and the Fleurieu Peninsula (Maslin Bay), South Australia; D, location of the northernmost exposure (1), and southernmost exposure (2), of the Muloowurtie Formation, west side of Gulf St Vincent." figureDoi="http://doi.org/10.5281/zenodo.12209672" httpUri="https://zenodo.org/record/12209672/files/figure.png" pageId="2" pageNumber="211">Fig. 1</figureCitation>
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)
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="D65A5679FFBF9E7A4386BB0DFD4DF89D" blockId="2.[139,780,1652,1908]" pageId="2" pageNumber="211">
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<emphasis id="E4918A6BFFBF9E7A4386BB0DFEBEF92D" box="[139,259,1652,1674]" italics="true" pageId="2" pageNumber="211">Description</emphasis>
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. Test moderately small, oval in outline at the ambitus, anterior and posterior evenly rounded, widest point central. Aboral surface moderately inflated, except for a depression posterior to the periproct, with the apex just anteri- or of the apical disk at the proximal end of slightly swollen ambulacrum III. Adoral surface slightly depressed around peristome and along the posterior paired ambulacra I and V.
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</paragraph>
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<paragraph id="D65A5679FFBF9E7A43A6BA38FACBFE97" blockId="2.[139,780,1652,1908]" lastBlockId="2.[817,1458,224,1331]" pageId="2" pageNumber="211">
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Aboral tubercules very small, about
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<quantity id="111DFB9CFFBF9E7A4128BA38FDC5F8F0" box="[549,632,1857,1879]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="211" unit="mm" value="0.15">0.15mm</quantity>
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diameter, and closely spaced with a density of about 12–15 tubercles per mm2. Adoral tubercles, where visible near the margin, are also closely spaced but larger, about
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<quantity id="111DFB9CFFBF9E7A4792BD84FB49FEB4" box="[1183,1268,253,275]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="211" unit="mm" value="0.6">0.6 mm</quantity>
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diameter. Naked granular zone in ambulacrum III and interambulacrum 5.
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</paragraph>
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<paragraph id="D65A5679FFBF9E7A405CBC41FC40FE47" blockId="2.[817,1458,224,1331]" pageId="2" pageNumber="211">
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Apical system monobasal, centre of disk 42–45%TL from anterior margin, with 4 gonopores in contact with the apical disk but extending into the first pair of interambulacral plates, anterior pair closer together than posterior pair. Ocular plates relatively small and about equal in size. Approximately 80 hydropores (
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<figureCitation id="4EDE4AFCFFBF9E7A40BFBCB3FC4DFE47" box="[946,1008,458,480]" captionStart="Figure 3" captionStartId="3.[131,190,1812,1832]" captionTargetBox="[130,754,1294,1793]" captionTargetId="figure-163@3.[130,528,1294,1793]" captionTargetPageId="3" captionText="Figure 3. Apical plate structure. A and B, Rhynchopygus? janchrisorum sp. nov., Late Eocene holotype NMV P145616; C, R. marmini, Late Cretaceous USNM 19559 from Port Brechay, La Manche, France (drawing adapted from Kier, 1962). Scale bars 1mm." figureDoi="http://doi.org/10.5281/zenodo.12209681" httpUri="https://zenodo.org/record/12209681/files/figure.png" pageId="2" pageNumber="211">Fig. 3</figureCitation>
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).
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</paragraph>
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<paragraph id="D65A5679FFBF9E7A405CBC91FC3BFC11" blockId="2.[817,1458,224,1331]" pageId="2" pageNumber="211">Petals moderately short, broad and unequal in length. Longest in ambulacrum III (approx. 62% radius with about 30 pores per tract) and shortest in anterior pair II and IV (approx. 50% radius with about 20 pores per tract). Inner pores of pore pairs oval, outer pores slot like, with alignment noticeably oblique in ambulacra II and IV. At widest point interporiferous zone in ambulacra II, III and IV about equal in width to zones of pore pairs, parallel sided and open ended in ambulcrum III and narrowing distally in II and IV. Posterior pair of petals have narrower poriferous zones curving outwards distally with poriferous zones Ib and Va noticeably wider than Ia and Vb. Poriferous zones in individual petals equal in length. Anterior pair of petals diverge between 134° and 140°, posterior pair between 306° and 310°. The aboral swelling in ambulacrum III forms a low ridge along line of perradial suture for full length of petal.</paragraph>
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<paragraph id="D65A5679FFBF9E7A405CBEC4FC44FBC1" blockId="2.[817,1458,224,1331]" pageId="2" pageNumber="211">Periproct supramarginal, transverse, situated at the anterior end of a pronounced posterior anal depression in interambulacrum 5, beneath a wide semi-circular projecting lip (about 19%TW) extending to nearly 20%TL from the posterior margin. The anal depression diminishes posteriorly and barely reaches the margin.</paragraph>
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<paragraph id="D65A5679FFBF9E7A405CB914FADBFA94" blockId="2.[817,1458,224,1331]" pageId="2" pageNumber="211">Peristome small, pentagonal, centre of opening situated 37-40%TL from the anterior margin, Floscelle well developed with very pronounced pointed and inflated bourrelets, anterior pair wide and wedge-shaped, posterior single and pair elongated and near parallel sided. Phyllodes deeply sunken proximally but with pronounced ridge between bourrelets at edge of peristomal opening. Detail of phyllode pores unknown.</paragraph>
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<paragraph id="D65A5679FFBF9E7A403CB83DFC4AFAD0" blockId="2.[817,1457,1348,1399]" pageId="2" pageNumber="211">
|
||
<emphasis id="E4918A6BFFBF9E7A403CB83DFC20FAFD" box="[817,925,1348,1370]" italics="true" pageId="2" pageNumber="211">Etymology</emphasis>
|
||
. For Janice Krause and Christopher Ah Yee of Hamilton,
|
||
<collectingRegion id="1421989BFFBF9E7A4093B818FC4EFAD0" box="[926,1011,1377,1399]" country="Australia" name="Victoria" pageId="2" pageNumber="211">Victoria</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="D65A5679FFBF9E7C403CB8F1FE14FC37" blockId="2.[817,1457,1416,1907]" lastBlockId="4.[139,780,743,1322]" lastPageId="4" lastPageNumber="213" pageId="2" pageNumber="211">
|
||
<emphasis id="E4918A6BFFBF9E7A403CB8F1FC37FA39" box="[817,906,1416,1438]" italics="true" pageId="2" pageNumber="211">Remarks</emphasis>
|
||
. The description is based on
|
||
<specimenCount id="C0E39DF0FFBF9E7A47C4B8F1FAD7FA39" box="[1225,1386,1416,1438]" count="4" pageId="2" pageNumber="211" type="generic">four specimens</specimenCount>
|
||
, all of which have been subject to diagenetic compression resulting in radial cracks along adradial and interradial sutures. In the case of the
|
||
<typeStatus id="095EE8DBFFBF9E7A407DB899FC74FA51" box="[880,969,1504,1526]" pageId="2" pageNumber="211" type="holotype">holotype</typeStatus>
|
||
these cracks extend between half and two-thirds distance between ambitus and distal end of petals. Compared with a similarly compressed specimen of
|
||
<taxonomicName id="11E52DFAFFBF9E7A47FDBB62FCC8F9E9" baseAuthorityName="Gregory" baseAuthorityYear="1890" class="Echinoidea" family="Faujasiidae" genus="Australanthus" kingdom="Animalia" order="Clypeasteroida" pageId="2" pageNumber="211" phylum="Echinodermata" rank="species" species="longianus">
|
||
<emphasis id="E4918A6BFFBF9E7A47FDBB62FCC8F9E9" italics="true" pageId="2" pageNumber="211">Australanthus longianus</emphasis>
|
||
</taxonomicName>
|
||
from the same locality, it is estimated that the
|
||
<typeStatus id="095EE8DBFFBF9E7A4655BB41FA0CF9E9" box="[1368,1457,1592,1614]" pageId="2" pageNumber="211" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="11E52DFAFFBF9E7A4041BB2FFBD0F9CB" authorityName="Holmes" authorityYear="2004" box="[844,1133,1621,1644]" class="Echinoidea" family="Cassidulidae" genus="Rhyncopygus" kingdom="Animalia" order="Echinolampadacea" pageId="2" pageNumber="211" phylum="Echinodermata" rank="species" species="janchrisorum" status="sp. nov.">
|
||
<emphasis id="E4918A6BFFBF9E7A4041BB2FFC6EF9CB" box="[844,979,1622,1644]" italics="true" pageId="2" pageNumber="211">Rhyncopygus</emphasis>
|
||
?
|
||
<emphasis id="E4918A6BFFBF9E7A40E8BB2FFBD0F9CB" box="[997,1133,1622,1644]" italics="true" pageId="2" pageNumber="211">janchrisorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="FFA23710FFBF9E7A477EBB2CFB7FF9CC" box="[1139,1218,1621,1643]" pageId="2" pageNumber="211" rank="species">sp. nov.</taxonomicNameLabel>
|
||
would have been about 24.0 mm long,
|
||
<quantity id="111DFB9CFFBF9E7A40D9BB0AFB8DF92E" box="[980,1072,1651,1673]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" pageId="2" pageNumber="211" unit="mm" value="20.5">20.5 mm</quantity>
|
||
(85.5%TL) wide and a minimum of
|
||
<quantity id="111DFB9CFFBF9E7A403CBBE9FC36F901" box="[817,907,1680,1702]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="2" pageNumber="211" unit="mm" value="10.5">10.5 mm</quantity>
|
||
(44%TL) high compared with the
|
||
<quantity id="111DFB9CFFBF9E7A47E6BBE9FAF8F901" box="[1259,1349,1680,1702]" metricMagnitude="-2" metricUnit="m" metricValue="2.68" pageId="2" pageNumber="211" unit="mm" value="26.8">26.8 mm</quantity>
|
||
long,
|
||
<quantity id="111DFB9CFFBF9E7A4688BBE9FCE5F964" metricMagnitude="-2" metricUnit="m" metricValue="2.29" pageId="2" pageNumber="211" unit="mm" value="22.9">22.9 mm</quantity>
|
||
wide and
|
||
<quantity id="111DFB9CFFBF9E7A40D9BBD4FB9AF964" box="[980,1063,1709,1731]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="2" pageNumber="211" unit="mm" value="7.8">7.8 mm</quantity>
|
||
high dimensions of the compressed fossil test (
|
||
<figureCitation id="4EDE4AFCFFBF9E7A40ABBBB2FC55F946" box="[934,1000,1739,1761]" captionStart="Figure 4" captionStartId="3.[811,870,1675,1695]" captionTargetBox="[813,1452,1294,1649]" captionTargetId="figure-124@3.[808,1452,1293,1657]" captionTargetPageId="3" captionText="Figure 4. Rhynchopygus? janchrisorum sp. nov. A–C, reconstruction of adapical, right lateral and posterior characteristics of holotype NMV P145616 (actual specimen outline shown with broken line). Scale bar 10 mm." figureDoi="http://doi.org/10.5281/zenodo.12209679" httpUri="https://zenodo.org/record/12209679/files/figure.png" pageId="2" pageNumber="211">Fig. 4</figureCitation>
|
||
).
|
||
<taxonomicName id="11E52DFAFFBF9E7A40F3BBB2FA9FF946" authorityName="Holmes" authorityYear="2004" box="[1022,1314,1739,1761]" class="Echinoidea" family="Cassidulidae" genus="Rhyncopygus" kingdom="Animalia" order="Echinolampadacea" pageId="2" pageNumber="211" phylum="Echinodermata" rank="species" species="janchrisorum">
|
||
<emphasis id="E4918A6BFFBF9E7A40F3BBB2FB38F946" box="[1022,1157,1739,1761]" italics="true" pageId="2" pageNumber="211">Rhyncopygus</emphasis>
|
||
?
|
||
<emphasis id="E4918A6BFFBF9E7A4797BBB2FA9FF946" box="[1178,1314,1739,1761]" italics="true" pageId="2" pageNumber="211">janchrisorum</emphasis>
|
||
</taxonomicName>
|
||
is easily distinguished from the type species of the genus,
|
||
<taxonomicName id="11E52DFAFFBF9E7A4617BB91FA33F959" authorityName="Agassiz, in Agassiz and Desor" authorityYear="1847" box="[1306,1422,1768,1790]" class="Echinoidea" family="Cassidulidae" genus="Cassidulus" kingdom="Animalia" order="Echinolampadacea" pageId="2" pageNumber="211" phylum="Echinodermata" rank="species" species="marmini">
|
||
<emphasis id="E4918A6BFFBF9E7A4617BB91FA33F959" box="[1306,1422,1768,1790]" italics="true" pageId="2" pageNumber="211">R. marmini</emphasis>
|
||
</taxonomicName>
|
||
, in having a larger, less elongated and less inflated test, a monobasal apical disk, far more pronounced and broader petals, and prominent bourrelets. It differs from
|
||
<taxonomicName id="11E52DFAFFBF9E7A4617BA39FA16F8F1" authorityName="Faas" authorityYear="1918" box="[1306,1451,1856,1878]" class="Echinoidea" family="Faujasiidae" genus="Rhynchopygus" kingdom="Animalia" order="Clypeasteroida" pageId="2" pageNumber="211" phylum="Echinodermata" rank="species" species="donetzensis">
|
||
<emphasis id="E4918A6BFFBF9E7A4617BA39FA16F8F1" box="[1306,1451,1856,1878]" italics="true" pageId="2" pageNumber="211">R. donetzensis</emphasis>
|
||
</taxonomicName>
|
||
, based on the illustrations in
|
||
<bibRefCitation id="B2742B88FFBF9E7A4785BA24FA10F8D4" author="Smith, A. B. & Jeffery, C. H." box="[1160,1453,1885,1907]" pageId="2" pageNumber="211" refId="ref5261" refString="Smith, A. B., and Jeffery, C. H., 2000. Maastrichtian and Palaeocene echinoids: a key to world faunas. Special papers in palaeontology 63. The Palaeontological Association: London. 406 pp." type="book" year="2000">Smith and Jeffery (2000)</bibRefCitation>
|
||
, primarily in being far less tumid and having a longer and broader petal in ambulacrum III, shorter posterior petals and considerably wider ambulacral plates at the ambitus (
|
||
<figureCitation id="4EDE4AFCFFB99E7C41B0BE5BFD43FC9F" box="[701,766,802,824]" captionStart="Figure 5" captionStartId="4.[139,198,632,652]" captionTargetBox="[250,661,224,610]" captionTargetId="figure-614@4.[178,780,220,614]" captionTargetPageId="4" captionText="Figure 5. Rhynchopygus? janchrisorum sp. nov. Drawing of adapical surface of NMV P312113 showing plate structure distal to petals. Scale bar 10 mm." figureDoi="http://doi.org/10.5281/zenodo.12209683" httpUri="https://zenodo.org/record/12209683/files/figure.png" pageId="4" pageNumber="213">Fig. 5</figureCitation>
|
||
). The test of
|
||
<taxonomicName id="11E52DFAFFB99E7C421CBE46FE7AFCF2" authorityName="Holmes" authorityYear="2004" box="[273,455,831,853]" class="Echinoidea" family="Cassidulidae" genus="Rhyncopygus" kingdom="Animalia" order="Echinolampadacea" pageId="4" pageNumber="213" phylum="Echinodermata" rank="species" species="janchrisorum">
|
||
<emphasis id="E4918A6BFFB99E7C421CBE46FE9AFCF2" box="[273,295,831,853]" italics="true" pageId="4" pageNumber="213">R.</emphasis>
|
||
?
|
||
<emphasis id="E4918A6BFFB99E7C4232BE46FE7AFCF2" box="[319,455,831,853]" italics="true" pageId="4" pageNumber="213">janchrisorum</emphasis>
|
||
</taxonomicName>
|
||
is longer and proportionately wider than the other two species and has a more anterior periproct and projecting lip.
|
||
</paragraph>
|
||
<caption id="829A06F1FFBE9E7B438EB947FC30FB20" ID-DOI="http://doi.org/10.5281/zenodo.12209677" ID-Zenodo-Dep="12209677" httpUri="https://zenodo.org/record/12209677/files/figure.png" pageId="3" pageNumber="212" startId="3.[131,190,1086,1106]" targetBox="[130,1451,222,1067]" targetPageId="3" targetType="figure">
|
||
<paragraph id="D65A5679FFBE9E7B438EB947FC30FB20" blockId="3.[131,1449,1086,1159]" pageId="3" pageNumber="212">
|
||
Figure 2.
|
||
<taxonomicName id="11E52DFAFFBE9E7B43D0B947FE51FBF5" authorityName="Holmes" authorityYear="2004" box="[221,492,1086,1106]" class="Echinoidea" family="Faujasiidae" genus="Rhynchopygus" kingdom="Animalia" order="Clypeasteroida" pageId="3" pageNumber="212" phylum="Echinodermata" rank="species" species="janchrisorum" status="sp. nov.">
|
||
<emphasis id="E4918A6BFFBE9E7B43D0B947FEDCFBF5" box="[221,353,1086,1106]" italics="true" pageId="3" pageNumber="212">Rhynchopygus</emphasis>
|
||
?
|
||
<emphasis id="E4918A6BFFBE9E7B427FB947FE51FBF5" box="[370,492,1086,1106]" italics="true" pageId="3" pageNumber="212">janchrisorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="FFA23710FFBE9E7B42FEB947FD86FBF5" box="[499,571,1086,1106]" pageId="3" pageNumber="212" rank="species">sp. nov.</taxonomicNameLabel>
|
||
A–C, adapical, right lateral and posterior views of holotype NMV P145616; D, adoral view of paratype NMV P312115; E, adoral view of paratype NMV P312114; F, adapical view of paratype NMV P312113. All specimens form the Late Eocene Muloowurtie Formation, Yorke Peninsula, South Australia. Scale bar 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="829A06F1FFBE9E7B4026BBF2FCCEF948" ID-DOI="http://doi.org/10.5281/zenodo.12209679" ID-Zenodo-Dep="12209679" httpUri="https://zenodo.org/record/12209679/files/figure.png" pageId="3" pageNumber="212" startId="3.[811,870,1675,1695]" targetBox="[813,1452,1294,1649]" targetPageId="3" targetType="figure">
|
||
<paragraph id="D65A5679FFBE9E7B4026BBF2FCCEF948" blockId="3.[811,1449,1675,1775]" pageId="3" pageNumber="212">
|
||
Figure 4.
|
||
<taxonomicName id="11E52DFAFFBE9E7B4085BBF5FB25F907" authorityName="Holmes" authorityYear="2004" box="[904,1176,1675,1696]" class="Echinoidea" family="Faujasiidae" genus="Rhynchopygus" kingdom="Animalia" order="Clypeasteroida" pageId="3" pageNumber="212" phylum="Echinodermata" rank="species" species="janchrisorum" status="sp. nov.">
|
||
<emphasis id="E4918A6BFFBE9E7B4085BBF5FBB1F907" box="[904,1036,1676,1696]" italics="true" pageId="3" pageNumber="212">Rhynchopygus</emphasis>
|
||
?
|
||
<emphasis id="E4918A6BFFBE9E7B4713BBF5FB25F907" box="[1054,1176,1676,1696]" italics="true" pageId="3" pageNumber="212">janchrisorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="FFA23710FFBE9E7B47ADBBF2FB54F938" box="[1184,1257,1675,1695]" pageId="3" pageNumber="212" rank="species">sp. nov.</taxonomicNameLabel>
|
||
A–C, reconstruction of adapical, right lateral and posterior characteristics of holotype NMV P145616 (actual specimen outline shown with broken line). Scale bar 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="829A06F1FFBE9E7B438EBA6DFDDAF8DF" ID-DOI="http://doi.org/10.5281/zenodo.12209681" ID-Zenodo-Dep="12209681" httpUri="https://zenodo.org/record/12209681/files/figure.png" pageId="3" pageNumber="212" startId="3.[131,190,1812,1832]" targetBox="[130,754,1294,1793]" targetPageId="3" targetType="figure">
|
||
<paragraph id="D65A5679FFBE9E7B438EBA6DFDDAF8DF" blockId="3.[131,769,1812,1912]" pageId="3" pageNumber="212">
|
||
Figure 3. Apical plate structure. A and B,
|
||
<taxonomicName id="11E52DFAFFBE9E7B4103BA6CFF15F8E4" authorityName="Holmes" authorityYear="2004" class="Echinoidea" family="Faujasiidae" genus="Rhynchopygus" kingdom="Animalia" order="Clypeasteroida" pageId="3" pageNumber="212" phylum="Echinodermata" rank="species" species="janchrisorum" status="sp. nov.">
|
||
<emphasis id="E4918A6BFFBE9E7B4103BA6CFD2FF88E" box="[526,658,1813,1833]" italics="true" pageId="3" pageNumber="212">Rhynchopygus</emphasis>
|
||
?
|
||
<emphasis id="E4918A6BFFBE9E7B41A9BA6CFF15F8E4" italics="true" pageId="3" pageNumber="212">janchrisorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="FFA23710FFBE9E7B43BDBA56FF46F8E4" box="[176,251,1839,1859]" pageId="3" pageNumber="212" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, Late Eocene holotype NMV P145616; C,
|
||
<taxonomicName id="11E52DFAFFBE9E7B419FBA56FEE1F8F9" authority=", Late Cretaceous USNM" authorityName="Late Cretaceous USNM" class="Echinoidea" family="Cassidulidae" genus="Cassidulus" kingdom="Animalia" order="Echinolampadacea" pageId="3" pageNumber="212" phylum="Echinodermata" rank="species" species="marmini">
|
||
<emphasis id="E4918A6BFFBE9E7B419FBA56FD46F8E4" box="[658,763,1839,1859]" italics="true" pageId="3" pageNumber="212">R. marmini</emphasis>
|
||
, Late Cretaceous USNM
|
||
</taxonomicName>
|
||
19559 from Port Brechay, La Manche, France (drawing adapted from
|
||
<bibRefCitation id="B2742B88FFBE9E7B4257BA1DFE03F8DF" author="Kier, P. M." box="[346,446,1892,1912]" pageId="3" pageNumber="212" pagination="1 - 262" refId="ref4571" refString="Kier, P. M. 1962. Revision of the cassiduloid echinoids. Smithsonian Miscellaneous Collections 144 (3): 1 - 262, pls 1 - 44." type="journal article" year="1962">Kier, 1962</bibRefCitation>
|
||
). Scale bars 1mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="829A06F1FFB99E7C4386BF01FE90FD65" ID-DOI="http://doi.org/10.5281/zenodo.12209683" ID-Zenodo-Dep="12209683" httpUri="https://zenodo.org/record/12209683/files/figure.png" pageId="4" pageNumber="213" startId="4.[139,198,632,652]" targetBox="[250,661,224,610]" targetPageId="4" targetType="figure">
|
||
<paragraph id="D65A5679FFB99E7C4386BF01FE90FD65" blockId="4.[139,778,632,706]" pageId="4" pageNumber="213">
|
||
Figure 5.
|
||
<taxonomicName id="11E52DFAFFB99E7C43EABF01FE4BFD2B" authorityName="Holmes" authorityYear="2004" box="[231,502,632,652]" class="Echinoidea" family="Faujasiidae" genus="Rhynchopygus" kingdom="Animalia" order="Clypeasteroida" pageId="4" pageNumber="213" phylum="Echinodermata" rank="species" species="janchrisorum" status="sp. nov.">
|
||
<emphasis id="E4918A6BFFB99E7C43EABF01FED6FD2B" box="[231,363,632,652]" italics="true" pageId="4" pageNumber="213">Rhynchopygus</emphasis>
|
||
?
|
||
<emphasis id="E4918A6BFFB99E7C4271BF01FE4BFD2B" box="[380,502,632,652]" italics="true" pageId="4" pageNumber="213">janchrisorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="FFA23710FFB99E7C42F0BF01FDFBFD2B" box="[509,582,632,652]" pageId="4" pageNumber="213" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Drawing of adapical surface of NMV P312113 showing plate structure distal to petals. Scale bar 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="D65A5679FFB99E7C43A6BEEEFE27FA8D" blockId="4.[139,780,743,1322]" pageId="4" pageNumber="213">
|
||
The occurrence of a similar but much less pronounced canopy above the periproct is also present in
|
||
<taxonomicName id="11E52DFAFFB99E7C419BBECCFE06FC4F" authority="Lambert" authorityName="Lambert" class="Echinoidea" family="Faujasiidae" genus="Hardouinia" kingdom="Animalia" order="Clypeasteroida" pageId="4" pageNumber="213" phylum="Echinodermata" rank="species" species="priscus" subGenus="Fauraster">
|
||
<emphasis id="E4918A6BFFB99E7C419BBECCFCB6FC6C" box="[662,779,949,971]" italics="true" pageId="4" pageNumber="213">Hardouinia</emphasis>
|
||
(
|
||
<emphasis id="E4918A6BFFB99E7C4399BEABFF4AFC4F" box="[148,247,978,1000]" italics="true" pageId="4" pageNumber="213">Fauraster</emphasis>
|
||
)
|
||
<emphasis id="E4918A6BFFB99E7C4201BEABFEE8FC4F" box="[268,341,978,1000]" italics="true" pageId="4" pageNumber="213">priscus</emphasis>
|
||
Lambert
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="E4918A6BFFB99E7C42C7BEABFE63FC4F" box="[458,478,978,1000]" italics="true" pageId="4" pageNumber="213">in</emphasis>
|
||
Lambert and Thiéry, 1924.
|
||
<bibRefCitation id="B2742B88FFB99E7C4386BE96FE8AFBA2" author="Kier, P. M." box="[139,311,1007,1029]" pageId="4" pageNumber="213" pagination="1 - 262" refId="ref4571" refString="Kier, P. M. 1962. Revision of the cassiduloid echinoids. Smithsonian Miscellaneous Collections 144 (3): 1 - 262, pls 1 - 44." type="journal article" year="1962">Kier (1962: 143)</bibRefCitation>
|
||
, in his description of the poorly preserved and compressed test, noted that the periproct is very wide and low; however,
|
||
<bibRefCitation id="B2742B88FFB99E7C43FCB953FD96FBE7" author="Smith, A. B. & Jeffery, C. H." box="[241,555,1066,1088]" pageId="4" pageNumber="213" refId="ref5261" refString="Smith, A. B., and Jeffery, C. H., 2000. Maastrichtian and Palaeocene echinoids: a key to world faunas. Special papers in palaeontology 63. The Palaeontological Association: London. 406 pp." type="book" year="2000">Smith and Jeffery (2000: 205)</bibRefCitation>
|
||
referred to this as an external feature that funnels into an opening only a little wider than tall and list
|
||
<taxonomicName id="11E52DFAFFB99E7C424EB91CFDA5FBDC" authorityName="Holland & Feldmann" authorityYear="1967" box="[323,536,1125,1147]" class="Echinoidea" family="Faujasiidae" genus="Hardouinia" kingdom="Animalia" order="Clypeasteroida" pageId="4" pageNumber="213" phylum="Echinodermata" rank="species" species="waageni">
|
||
<emphasis id="E4918A6BFFB99E7C424EB91CFDA5FBDC" box="[323,536,1125,1147]" italics="true" pageId="4" pageNumber="213">Hardouinia waageni</emphasis>
|
||
</taxonomicName>
|
||
<bibRefCitation id="B2742B88FFB99E7C412FB91DFF7DFB3F" author="Holland, F. D. & Feldman, R. M." pageId="4" pageNumber="213" pagination="252 - 255" refId="ref4452" refString="Holland, F. D., and Feldman, R. M. 1967. A new species of cassiduloid echinoid from the Fox Hills Formation (Upper Cretaceous) of North Dakota. Journal of Paleontology 41: 252 - 255." type="journal article" year="1967">
|
||
<collectingCountry id="AEF216E9FFB99E7C412FB91DFDC9FBDD" box="[546,628,1124,1146]" name="Netherlands" pageId="4" pageNumber="213">Holland</collectingCountry>
|
||
and Feldman, 1967
|
||
</bibRefCitation>
|
||
, and
|
||
<taxonomicName id="11E52DFAFFB99E7C43FBB9FBFDC3FB3F" authority="Moskvin, 1984" authorityName="Moskvin" authorityYear="1984" box="[246,638,1154,1176]" class="Echinoidea" family="Faujasiidae" genus="Hardouinia" kingdom="Animalia" order="Clypeasteroida" pageId="4" pageNumber="213" phylum="Echinodermata" rank="species" species="nuratensis">
|
||
<emphasis id="E4918A6BFFB99E7C43FBB9FBFE66FB3F" box="[246,475,1154,1176]" italics="true" pageId="4" pageNumber="213">Hardouinia nuratensis</emphasis>
|
||
<bibRefCitation id="B2742B88FFB99E7C42EFB9FBFDC3FB3F" author="Moskvin, M. M." box="[482,638,1154,1176]" pageId="4" pageNumber="213" pagination="63 - 68" refId="ref4931" refString="Moskvin, M. M. 1984. [Echinoids Domechinus and Hardouinia (Echinoidea) from the Upper Cretaceous of Middle Asia]. Paleontologicheskii Sbornik 21, 63 - 68, 1 pl. [In Russian]." type="journal article" year="1984">Moskvin, 1984</bibRefCitation>
|
||
</taxonomicName>
|
||
, as synonyms of this species. Apart from this feature,
|
||
<taxonomicName id="11E52DFAFFB99E7C4121B9E6FD7CFB12" box="[556,705,1183,1205]" class="Echinoidea" family="Faujasiidae" genus="Hardouinia" kingdom="Animalia" order="Clypeasteroida" pageId="4" pageNumber="213" phylum="Echinodermata" rank="species" species="priscus" subGenus="Fauraster">H. (F.) priscus</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="11E52DFAFFB99E7C43C9B9C4FEC9FB74" authorityName="Holmes" authorityYear="2004" box="[196,372,1212,1235]" class="Echinoidea" family="Faujasiidae" genus="Rhynchopygus" kingdom="Animalia" order="Clypeasteroida" pageId="4" pageNumber="213" phylum="Echinodermata" rank="species" species="janchrisorum">
|
||
<emphasis id="E4918A6BFFB99E7C43C9B9C4FF6EFB74" box="[196,211,1213,1235]" italics="true" pageId="4" pageNumber="213">R</emphasis>
|
||
.?
|
||
<emphasis id="E4918A6BFFB99E7C43E1B9C4FEC9FB74" box="[236,372,1213,1235]" italics="true" pageId="4" pageNumber="213">janchrisorum</emphasis>
|
||
</taxonomicName>
|
||
in its near circular outline with slightly truncated posterior, its tetrabasal apical system and smaller and equal sized petals with narrower poriferous tracts and much wider interporiferous zone
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |