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<document id="6DB35687B1208597A11DF0E30FE13451" ID-DOI="10.11646/zootaxa.5398.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="10494167" ID-ZooBank="E233F731-D5FA-4032-B3A4-CEFE5A809C49" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch,operationResults" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1705011554062" checkinUser="plazi" docAuthor="Van Soest, Rob W. M." docDate="2024" docId="BF4E397FFFBB310F9786FE27B9A803E8" docLanguage="en" docName="zootaxa.5398.1.1.pdf" docOrigin="Zootaxa 5398 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5398.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.18:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docTitle="Leucetta primigenia var. megalirrhaphis Ridley 1884" docType="treatment" docVersion="1" lastPageNumber="93" masterDocId="43774107FFE731539711FFD8BD62070B" masterDocTitle="Correcting sponge names: nomenclatural update of lower taxa level Porifera" masterLastPageNumber="122" masterPageNumber="1" pageNumber="93" updateTime="1706187678457" updateUser="felipe">
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<mods:title id="7231709ECF657676C5AACDA035671B89">Correcting sponge names: nomenclatural update of lower taxa level Porifera</mods:title>
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<mods:namePart id="3E96C4D2E1122FA85047E806182D0D96">Van Soest, Rob W. M.</mods:namePart>
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<mods:date id="D1240C302ED6ECDDE9B38AD4DEC21D95">2024</mods:date>
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<subSubSection id="7FFDDBE2FFBB310F9786FE27BE410513" box="[151,803,510,537]" pageId="92" pageNumber="93" type="nomenclature">
<paragraph id="37588869FFBB310F9786FE27BE410513" blockId="92.[151,803,510,537]" box="[151,803,510,537]" pageId="92" pageNumber="93">
<heading id="6C103F05FFBB310F9786FE27BE410513" bold="true" box="[151,803,510,537]" fontSize="11" level="2" pageId="92" pageNumber="93" reason="4">
<taxonomicName id="F0E7F3EAFFBB310F9786FE27BE410513" ID-CoL="7LVZS" authority="Ridley, 1884" authorityName="Ridley" authorityYear="1884" box="[151,803,510,537]" class="Calcarea" family="Leucettidae" genus="Leucetta" kingdom="Animalia" order="Clathrinida" pageId="92" pageNumber="93" phylum="Porifera" rank="variety" species="primigenia" variety="megalirrhaphis">
<emphasis id="0593547BFFBB310F9786FE27BE410513" bold="true" box="[151,803,510,537]" pageId="92" pageNumber="93">
<emphasis id="0593547BFFBB310F9786FE27BCE90512" bold="true" box="[151,395,511,537]" italics="true" pageId="92" pageNumber="93">Leucetta primigenia</emphasis>
var.
<emphasis id="0593547BFFBB310F96D9FE26BFE00512" bold="true" box="[456,642,510,537]" italics="true" pageId="92" pageNumber="93">megalirrhaphis</emphasis>
<bibRefCitation id="5376F598FFBB310F959BFE26BE410513" author="Ridley, S. O." box="[650,803,510,536]" pageId="92" pageNumber="93" pagination="582 - 630" refId="ref104302" refString="Ridley, S. O. (1884) Spongiida. In: Report on the Zoological Collections made in the Indo-Pacific Ocean during the Voyage of H. M. S. ' Alert', 1881 - 2. British Museum (Natural History), London, pp. 366 - 482, pls. 39 - 43, pp. 582 - 630, pls. 53 - 54. [https: // biodiversitylibrary. org / page / 12067365]" type="journal article" year="1884">Ridley, 1884</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="7FFDDBE2FFBB310F9786FD9CBEFB0550" box="[151,921,579,603]" pageId="92" pageNumber="93" type="reference_group">
<paragraph id="37588869FFBB310F9786FD9CBEFB0550" blockId="92.[151,921,579,603]" box="[151,921,579,603]" pageId="92" pageNumber="93">
<treatmentCitationGroup id="17F7AF47FFBB310F9786FD9CBEFB0550" box="[151,921,579,603]" pageId="92" pageNumber="93">
<treatmentCitation id="B646AE78FFBB310F9786FD9CBF8F0550" author="Ridley, S. O." box="[151,749,579,603]" page="628" pageId="92" pageNumber="93" year="1884">
<taxonomicName id="F0E7F3EAFFBB310F9786FD9CBF8F0550" ID-CoL="7LVZS" authority="Ridley, 1884: 628" authorityName="Ridley" authorityPageNumber="628" authorityYear="1884" box="[151,749,579,603]" class="Calcarea" family="Leucettidae" genus="Leucetta" kingdom="Animalia" order="Clathrinida" pageId="92" pageNumber="93" phylum="Porifera" rank="variety" species="primigenia" variety="megalirrhaphis">
<emphasis id="0593547BFFBB310F9786FD9CBC060550" box="[151,356,580,603]" italics="true" pageId="92" pageNumber="93">Leucetta primigenia</emphasis>
var.
<emphasis id="0593547BFFBB310F9687FD9BBF500550" box="[406,562,579,603]" italics="true" pageId="92" pageNumber="93">megalirrhaphis</emphasis>
<bibRefCitation id="5376F598FFBB310F9529FD9BBF8F0550" author="Ridley, S. O." box="[568,749,579,603]" pageId="92" pageNumber="93" pagination="582 - 630" refId="ref104302" refString="Ridley, S. O. (1884) Spongiida. In: Report on the Zoological Collections made in the Indo-Pacific Ocean during the Voyage of H. M. S. ' Alert', 1881 - 2. British Museum (Natural History), London, pp. 366 - 482, pls. 39 - 43, pp. 582 - 630, pls. 53 - 54. [https: // biodiversitylibrary. org / page / 12067365]" type="journal article" year="1884">Ridley, 1884: 628</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(no illustration).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="7FFDDBE2FFBB310F9786FD51B9A803E8" pageId="92" pageNumber="93" type="materials_examined">
<paragraph id="37588869FFBB310F9786FD51B9A803E8" blockId="92.[151,1437,649,1252]" pageId="92" pageNumber="93">
The variety was described by Ridley from the
<collectingCountry id="4FF0C8F9FFBB310F95B9FD51BE4105A8" box="[680,803,649,675]" name="Seychelles" pageId="92" pageNumber="93">Seychelles</collectingCountry>
, no further locality data, approximate coordinates
<geoCoordinate id="52D3EEAEFFBB310F924FFD51B8FE05A8" box="[1374,1436,649,675]" degrees="4.9" direction="south" orientation="latitude" pageId="92" pageNumber="93" precision="5555" value="-4.9">4.9°S</geoCoordinate>
<geoCoordinate id="52D3EEAEFFBB310F9786FD75BD8505C3" box="[151,231,685,712]" degrees="55.5" direction="east" orientation="longitude" pageId="92" pageNumber="93" precision="5555" value="55.5">55.5°E</geoCoordinate>
, depth
<specimenCount id="21E143E0FFBB310F9651FD75BCE705C3" box="[320,389,685,712]" count="22" pageId="92" pageNumber="93" type="generic">
<quantity id="F01F258CFFBB310F9651FD75BCE305C3" box="[320,385,685,712]" metricMagnitude="1" metricUnit="m" metricValue="2.2" pageId="92" pageNumber="93" unit="m" value="22.0">22 m</quantity>
.
</specimenCount>
This is a puzzling taxon, presumably represented by BMNH specimens labeled
<taxonomicName id="F0E7F3EAFFBB310F922DFD76BCBE05E0" ID-CoL="6Q3L8" authorityName="Haeckel" authorityYear="1872" class="Calcarea" family="Leucettidae" genus="Leucetta" kingdom="Animalia" order="Clathrinida" pageId="92" pageNumber="93" phylum="Porifera" rank="variety" species="primigenia" variety="megaraphis">
<emphasis id="0593547BFFBB310F922DFD76BC7005E0" italics="true" pageId="92" pageNumber="93">Leucetta primigenia</emphasis>
var.
<emphasis id="0593547BFFBB310F9649FD09BCBE05E0" box="[344,476,721,747]" italics="true" pageId="92" pageNumber="93">megaraphis</emphasis>
</taxonomicName>
, reg nr. 1882.10.17.59 (cf.
<bibRefCitation id="5376F598FFBB310F9423FD0ABEAD05E0" author="Burton, M." box="[818,975,721,748]" pageId="92" pageNumber="93" pagination="1 - 693" refId="ref90268" refString="Burton, M. (1963) A revision of the Classification of the Calcareous Sponges. With a Catalogue of the specimens in the British Museum (Natural History). W. Clowes and Sons Ltd., London, pp. 1 - 693." type="book chapter" year="1963">Burton, 1963</bibRefCitation>
) and a wet specimen kept in BMNH 1955.11.2.122 (according to the BMNH specimens Data Portal https://data.nhm.ac.uk/dataset/56e711e6-c847-4f99- 915a-6894bb5c5dea/resource/05ff2255-c38a-40c9-b657-4ccb55ab2feb/record/2760023, status unclear). Ridley erected the name
<emphasis id="0593547BFFBB310F9677FCE6BCA9045C" box="[358,459,830,855]" italics="true" pageId="92" pageNumber="93">
<taxonomicName id="F0E7F3EAFFBB310F9677FCE6BCA5045C" ID-CoL="84PNJ" authorityName="Haeckel" authorityYear="1872" box="[358,455,830,855]" class="Calcarea" family="Leucettidae" genus="Leucetta" kingdom="Animalia" order="Clathrinida" pageId="92" pageNumber="93" phylum="Porifera" rank="genus">Leucetta</taxonomicName>
.
</emphasis>
<taxonomicName id="F0E7F3EAFFBB310F96DBFCE6BE51045C" ID-CoL="7LVZS" authorityName="Ridley" authorityYear="1884" box="[458,819,829,856]" class="Calcarea" family="Leucettidae" genus="Leucetta" kingdom="Animalia" order="Clathrinida" pageId="92" pageNumber="93" phylum="Porifera" rank="variety" species="primigenia" variety="megalirrhaphis">
<emphasis id="0593547BFFBB310F96DBFCE6BF2A045C" box="[458,584,830,855]" italics="true" pageId="92" pageNumber="93">primigenia</emphasis>
var.
<emphasis id="0593547BFFBB310F9596FCE5BE51045C" box="[647,819,829,855]" italics="true" pageId="92" pageNumber="93">megalirrhaphis</emphasis>
</taxonomicName>
for
<specimenCount id="21E143E0FFBB310F9479FCE6B970045C" box="[872,1042,829,856]" count="2" pageId="92" pageNumber="93" type="generic">two specimens</specimenCount>
of brown colour (in alcohol), but cited
<taxonomicName id="F0E7F3EAFFBB310F97C4FCBABFC20470" ID-CoL="6Q3L8" authority="Haeckel, 1872: 119" authorityName="Haeckel" authorityPageNumber="119" authorityYear="1872" box="[213,672,865,892]" class="Calcarea" family="Leucettidae" genus="Leucetta" kingdom="Animalia" order="Clathrinida" pageId="92" pageNumber="93" phylum="Porifera" rank="variety" species="primigenia" variety="megaraphis">
<emphasis id="0593547BFFBB310F97C4FCBABC630470" box="[213,257,866,891]" italics="true" pageId="92" pageNumber="93">L.p.</emphasis>
var.
<emphasis id="0593547BFFBB310F9628FCB9BCDF0470" box="[313,445,865,891]" italics="true" pageId="92" pageNumber="93">megaraphis</emphasis>
<bibRefCitation id="5376F598FFBB310F96D4FCB9BFC20470" author="Haeckel, E." box="[453,672,865,891]" pageId="92" pageNumber="93" refId="ref96954" refString="Haeckel, E. (1872) Die Kalkschwamme. Eine Monographie in zwei Banden Text und einem Atlas mit 60 Tafeln Abbildungen. Vols. 1 - 3. G. Reimer, Berlin, 484 pp., 418 pp. &amp; 60 pls. [https: // www. biodiversitylibrary. org / page / 12042750]" type="book" year="1872">Haeckel, 1872: 119</bibRefCitation>
</taxonomicName>
as a presumed synonym. Ridley did not explain why he erected the new name (it could be a misspelling), but possibly he acknowledged by this that the
<collectingCountry id="4FF0C8F9FFBB310F9353FC5DB9D80494" box="[1090,1210,901,927]" name="Seychelles" pageId="92" pageNumber="93">Seychelles</collectingCountry>
specimens were not likely to be the same variety as Haeckels
<typeStatus id="E85C36CBFFBB310F9565FC72BFC604CF" box="[628,676,938,964]" pageId="92" pageNumber="93">type</typeStatus>
from the Adriatic (no further locality data).
<bibRefCitation id="5376F598FFBB310F9389FC72BDCD04EC" author="Von Lendenfeld, R." pageId="92" pageNumber="93" refId="ref111061" refString="Von Lendenfeld, R. (1888) Descriptive Catalogue of the Sponges in the Australian Museum, Sidney. Taylor &amp; Francis, London, xiv + 260 pp., 12 pls. [https: // biodiversitylibrary. org / page / 12762613]" type="book" year="1888">Von Lendenfeld (1888: 7)</bibRefCitation>
reported
<taxonomicName id="F0E7F3EAFFBB310F9633FC16BF4804EC" authorityName="Haeckel" authorityYear="1872" box="[290,554,973,999]" class="Calcarea" family="Leucettidae" genus="Leucetta" kingdom="Animalia" order="Clathrinida" pageId="92" pageNumber="93" phylum="Porifera" rank="species" species="megarrhaphis">
<emphasis id="0593547BFFBB310F9633FC16BF4804EC" box="[290,554,973,999]" italics="true" pageId="92" pageNumber="93">Leucetta megarrhaphis</emphasis>
</taxonomicName>
(sic) from Broughton Island, north of Port Jackson,
<collectingCountry id="4FF0C8F9FFBB310F9389FC15B86004EC" box="[1176,1282,973,999]" name="Australia" pageId="92" pageNumber="93">Australia</collectingCountry>
, approximate coordinates
<geoCoordinate id="52D3EEAEFFBB310F9630FC29BC0F0300" box="[289,365,1009,1035]" degrees="32.6" direction="south" orientation="latitude" pageId="92" pageNumber="93" precision="5555" value="-32.6">32.6°S</geoCoordinate>
<geoCoordinate id="52D3EEAEFFBB310F9667FC29BCB60307" box="[374,468,1009,1036]" degrees="152,3" direction="east" orientation="longitude" pageId="92" pageNumber="93" precision="5555" value="152.3">152,3°E</geoCoordinate>
, no further data, specimen BMNH 1886.6.7.41, cf.
<bibRefCitation id="5376F598FFBB310F9334FC2AB98D0307" author="Burton, M." box="[1061,1263,1009,1036]" pageId="92" pageNumber="93" pagination="1 - 693" refId="ref90268" refString="Burton, M. (1963) A revision of the Classification of the Calcareous Sponges. With a Catalogue of the specimens in the British Museum (Natural History). W. Clowes and Sons Ltd., London, pp. 1 - 693." type="book chapter" year="1963">Burton 1963: 596</bibRefCitation>
and
<bibRefCitation id="5376F598FFBB310F9238FC2ABCD20324" author="Hooper, J. N. A. &amp; Wiedenmayer, F." pageId="92" pageNumber="93" pagination="1 - 624" refId="ref98274" refString="Hooper, J. N. A. &amp; Wiedenmayer, F. (1994) Porifera. In: Wells, A. (Ed.), Zoological Catalogue of Australia. Vol. 12. CSIRO, Melbourne, pp. i - xiii + 1 - 624." type="book chapter" year="1994">Hooper &amp; Wiedenmayer 1994: 488</bibRefCitation>
), an obvious misapplication of an Adriatic species. Subsequent authors, including
<bibRefCitation id="5376F598FFBB310F925FFBCEBD800358" author="Burton, M." pageId="92" pageNumber="93" pagination="1 - 693" refId="ref90268" refString="Burton, M. (1963) A revision of the Classification of the Calcareous Sponges. With a Catalogue of the specimens in the British Museum (Natural History). W. Clowes and Sons Ltd., London, pp. 1 - 693." type="book chapter" year="1963">Burton (1963)</bibRefCitation>
and
<bibRefCitation id="5376F598FFBB310F9606FBE2BF0B0358" author="Van Soest, R. W. M. &amp; De Voogd, N. J." box="[279,617,1081,1108]" pageId="92" pageNumber="93" pagination="1 - 160" refId="ref110534" refString="Van Soest, R. W. M. &amp; De Voogd, N. J. (2018) Calcareous sponges of the Western Indian Ocean and Red Sea. Zootaxa, 4426 (1), 1 - 160. https: // doi. org / 10.11646 / zootaxa. 4426.1.1" type="journal article" year="2018">Van Soest &amp; De Voogd (2018)</bibRefCitation>
ignored the name
<emphasis id="0593547BFFBB310F9429FBE1BE860358" box="[824,996,1081,1107]" italics="true" pageId="92" pageNumber="93">megalirrhaphis</emphasis>
. Ridleys description is inadequate, but assuming we are dealing with a
<taxonomicName id="F0E7F3EAFFBB310F9511FB86BFDE037C" box="[512,700,1117,1143]" pageId="92" pageNumber="93">
<emphasis id="0593547BFFBB310F9511FB86BF02037C" box="[512,608,1118,1143]" italics="true" pageId="92" pageNumber="93">Leucetta</emphasis>
species
</taxonomicName>
, the brown colour in alcohol indicates that it could very well be a member of what is so far named
<taxonomicName id="F0E7F3EAFFBB310F9517FB5ABEFF0390" ID-CoL="723L8" authority="Haeckel, 1872" authorityName="Haeckel" authorityYear="1872" box="[518,925,1153,1179]" class="Calcarea" family="Leucettidae" genus="Leucetta" kingdom="Animalia" order="Clathrinida" pageId="92" pageNumber="93" phylum="Porifera" rank="species" species="microraphis">
<emphasis id="0593547BFFBB310F9517FB5ABF970390" box="[518,757,1153,1179]" italics="true" pageId="92" pageNumber="93">Leucetta microraphis</emphasis>
<bibRefCitation id="5376F598FFBB310F95EAFB59BEFF0390" author="Haeckel, E." box="[763,925,1153,1179]" pageId="92" pageNumber="93" refId="ref96954" refString="Haeckel, E. (1872) Die Kalkschwamme. Eine Monographie in zwei Banden Text und einem Atlas mit 60 Tafeln Abbildungen. Vols. 1 - 3. G. Reimer, Berlin, 484 pp., 418 pp. &amp; 60 pls. [https: // www. biodiversitylibrary. org / page / 12042750]" type="book" year="1872">
Haeckel,
<quantity id="F01F258CFFBB310F9474FB59BEFF0390" box="[869,925,1153,1179]" metricMagnitude="1" metricUnit="m" metricValue="4.75488" pageId="92" pageNumber="93" unit="in" value="1872.0">1872</quantity>
</bibRefCitation>
</taxonomicName>
in the sense of
<bibRefCitation id="5376F598FFBB310F935BFB5ABDD703CB" author="Van Soest, R. W. M. &amp; De Voogd, N. J." pageId="92" pageNumber="93" pagination="1 - 105" refId="ref110486" refString="Van Soest, R. W. M. &amp; De Voogd, N. J. (2015) Calcareous sponges of Indonesia. Zootaxa, 3951 (1), 1 - 105. https: // doi. org / 10.11646 / zootaxa. 3951.1.1" type="journal article" year="2015">Van Soest &amp; De Voogd (2015: 54</bibRefCitation>
, figs. 39ad, 40ad; 2018: 80, figs. 45af, 46ae). Accordingly, I reassign this variety to the synonymy of that species, until a proper and very necessary revision of
<taxonomicName id="F0E7F3EAFFBB310F95FEFB12BEBD03E8" ID-CoL="723L8" authorityName="Haeckel" authorityYear="1872" box="[751,991,1225,1251]" class="Calcarea" family="Leucettidae" genus="Leucetta" kingdom="Animalia" order="Clathrinida" pageId="92" pageNumber="93" phylum="Porifera" rank="species" species="microraphis">
<emphasis id="0593547BFFBB310F95FEFB12BEBD03E8" box="[751,991,1225,1251]" italics="true" pageId="92" pageNumber="93">Leucetta microraphis</emphasis>
</taxonomicName>
has been completed.
</paragraph>
</subSubSection>
</treatment>
</document>