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<document id="80B087253B773A2DF416B31268ABA1A1" ID-DOI="10.5281/zenodo.12587208" ID-ISSN="2064-2474" ID-Zenodo-Dep="12587208" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="admin" IM.treatments_approvedBy="felipe" checkinTime="1719597226178" checkinUser="felipe" docAuthor="Papp, L." docDate="2003" docId="A20987F8BF3AFFFEFD2AFE628D57B0CC" docLanguage="en" docName="ActaZoolAcadSciH.49.1.71-85.pdf" docOrigin="Acta Zoologica Academiae Scientiarum Hungaricae 49 (1)" docStyle="DocumentStyle:4FD749C6DF7B87D8940F4D8C7B0A0514.3:ActaZoolAcadSciH.2002-2014.journal_article" docStyleId="4FD749C6DF7B87D8940F4D8C7B0A0514" docStyleName="ActaZoolAcadSciH.2002-2014.journal_article" docStyleVersion="3" docTitle="Crumomyia topali Papp, 2003, sp. n." docType="treatment" docVersion="3" lastPageNumber="79" masterDocId="5E30FF80BF3CFFF6FFA9FFD5892AB468" masterDocTitle="New Oriental Species And Records Of Sphaerocerinae And Copromyzinae (Diptera: Sphaeroceridae)" masterLastPageNumber="85" masterPageNumber="71" pageNumber="77" updateTime="1719705059932" updateUser="admin" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="A5926DAE1B1A34B4DA54E9CE9B5A6C4F">New Oriental Species And Records Of Sphaerocerinae And Copromyzinae (Diptera: Sphaeroceridae)</mods:title>
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<mods:namePart id="C709952AD0B95A389DCCDDDD5BCA75D6">Papp, L.</mods:namePart>
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<mods:date id="005C7625A48C67007EF927C42B6FBE7D">2003</mods:date>
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<mods:number id="F5201F8FC756498D7367CB3608227B9B">49</mods:number>
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<mods:identifier id="3A42BABC841E1633FCF0692BCBD48EA6" type="DOI">10.5281/zenodo.12587208</mods:identifier>
<mods:identifier id="D80BA041B4C119A60D3C3258070D955C" type="ISSN">2064-2474</mods:identifier>
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<treatment id="A20987F8BF3AFFFEFD2AFE628D57B0CC" ID-DOI="http://doi.org/10.5281/zenodo.12587241" ID-Zenodo-Dep="12587241" LSID="urn:lsid:plazi:treatment:A20987F8BF3AFFFEFD2AFE628D57B0CC" httpUri="http://treatment.plazi.org/id/A20987F8BF3AFFFEFD2AFE628D57B0CC" lastPageId="8" lastPageNumber="79" pageId="6" pageNumber="77">
<subSubSection id="62BA6565BF3AFFF0FD2AFE628A85B5B9" box="[643,943,439,465]" pageId="6" pageNumber="77" type="nomenclature">
<paragraph id="2A1F36EEBF3AFFF0FD2AFE628A85B5B9" blockId="6.[643,943,439,465]" box="[643,943,439,465]" pageId="6" pageNumber="77">
<heading id="71578182BF3AFFF0FD2AFE628A85B5B9" box="[643,943,439,465]" centered="true" fontSize="11" level="2" pageId="6" pageNumber="77" reason="2">
<taxonomicName id="EDA04D6DBF3AFFF0FD2AFE628A40B5B9" ID-CoL="ZS2C" authorityName="Papp" authorityYear="2003" box="[643,874,439,465]" class="Insecta" family="Sphaeroceridae" genus="Crumomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="77" phylum="Arthropoda" rank="species" species="topali" status="sp. nov.">
<emphasis id="18D4EAFCBF3AFFF0FD2AFE628A40B5B9" bold="true" box="[643,874,439,465]" pageId="6" pageNumber="77">Crumomyia topali</emphasis>
</taxonomicName>
<taxonomicNameLabel id="03E75787BF3AFFF0FCD8FE628A85B5B9" box="[881,943,439,465]" pageId="6" pageNumber="77" rank="species">sp. n.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="62BA6565BF3AFFFEFD67FE0F8D57B0CC" lastPageId="8" lastPageNumber="79" pageId="6" pageNumber="77" type="description">
<paragraph id="2A1F36EEBF3AFFF0FD67FE0F8A4FB59C" blockId="6.[718,869,474,500]" box="[718,869,474,500]" pageId="6" pageNumber="77">
<heading id="71578182BF3AFFF0FD67FE0F8A4FB59C" box="[718,869,474,500]" centered="true" fontSize="11" level="3" pageId="6" pageNumber="77" reason="3">
(
<figureCitation id="B29B2A6BBF3AFFF0FD7FFE0F8A76B59C" box="[726,860,474,500]" captionStart="Figs 10–13" captionStartId="7.[317,360,1745,1767]" captionTargetBox="[326,1260,962,1711]" captionTargetId="figure-265@7.[323,1263,950,1716]" captionTargetPageId="7" captionText="Figs 10–13. Crumomyia topali sp. n., paratype male, genitalia: 10 = right halfofterminalia caudally, 11 = surstylus, broadest extension (setae on inner (medial) surface not depicted), 12 = phallus and" figureDoi="http://doi.org/10.5281/zenodo.12587214" httpUri="https://zenodo.org/record/12587214/files/figure.png" pageId="6" pageNumber="77">Figs 10–13</figureCitation>
)
</heading>
</paragraph>
<paragraph id="2A1F36EEBF3AFFF0FED7FDCB8856B638" blockId="6.[317,1271,542,1802]" pageId="6" pageNumber="77">
<materialsCitation id="9AC83CB3BF3AFFF0FED7FDCB8856B638" collectingDate="1967-01-01" collectingDateMax="1967-01-05" collectingDateMin="1967-01-01" collectionCode="HNHM" collectorName="Topal." country="India" location="Daitari" municipality="Jajpur-Keonjahr" pageId="6" pageNumber="77" specimenCount="1" typeStatus="holotype">
Material studied (
<collectionCode id="4CB1AE2BBF3AFFF0FD8EFDCB8B5DB65C" box="[551,631,542,564]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="6" pageNumber="77">HNHM</collectionCode>
):
<typeStatus id="F51B884CBF3AFFF0FD2FFDCB8BF0B65C" box="[646,730,542,564]" pageId="6" pageNumber="77" type="holotype">holotype</typeStatus>
male:
<collectingCountry id="52B7767EBF3AFFF0FCB3FDCB8A64B65C" box="[794,846,542,564]" name="India" pageId="6" pageNumber="77">India</collectingCountry>
,
<location id="2F7F6035BF3AFFF0FCFEFDCB8AB6B65C" LSID="urn:lsid:plazi:treatment:A20987F8BF3AFFFEFD2AFE628D57B0CC:2F7F6035BF3AFFF0FCFEFDCB8AB6B65C" box="[855,924,542,564]" country="India" municipality="Jajpur-Keonjahr" name="Daitari" pageId="6" pageNumber="77">Daitari</location>
,
<collectingMunicipality id="CA7BAC94BF3AFFF0FC0CFDCB8D6BB65C" box="[933,1089,542,564]" pageId="6" pageNumber="77">Jajpur-Keonjahr</collectingMunicipality>
–
<date id="5E1E102EBF3AFFF0FBF0FDCB8DEFB65C" box="[1113,1221,542,564]" pageId="6" pageNumber="77" value="1967-01-01" valueMax="1967-01-05" valueMin="1967-01-01">
<collectingDate id="4E5AE9C6BF3AFFF0FBF0FDCB8DEFB65C" box="[1113,1221,542,564]" pageId="6" pageNumber="77" value="1967-01-01" valueMax="1967-01-05" valueMin="1967-01-01">1–5.I.1967</collectingDate>
</date>
., leg.
<collectorName id="87555338BF3AFFF0FE94FDEF8856B638" box="[317,380,570,592]" pageId="6" pageNumber="77">Topál.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="2A1F36EEBF3AFFF0FED7FD828A75B6CF" blockId="6.[317,1271,542,1802]" pageId="6" pageNumber="77">
<materialsCitation id="9AC83CB3BF3AFFF0FED7FD828A70B6CF" collectingDate="1966-12-31" collectionCode="HNHM" pageId="6" pageNumber="77" specimenCount="1" typeStatus="paratype">
<typeStatus id="F51B884CBF3AFFF0FED7FD8288F8B605" box="[382,466,599,621]" pageId="6" pageNumber="77" type="paratype">Paratype</typeStatus>
male (
<collectionCode id="4CB1AE2BBF3AFFF0FDBBFD828B48B605" box="[530,610,599,621]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="6" pageNumber="77">HNHM</collectionCode>
): ibid.,
<date id="5E1E102EBF3AFFF0FD0EFD828A00B605" box="[679,810,599,621]" pageId="6" pageNumber="77" value="1966-12-31">
<collectingDate id="4E5AE9C6BF3AFFF0FD0EFD828A00B605" box="[679,810,599,621]" pageId="6" pageNumber="77" value="1966-12-31">31. XII. 1966</collectingDate>
</date>
. [left first flagellomere lost, abdomen with genitalia in a plastic microvial with glycerol, left wing of the specimen prepared between two pieces of cover glass glued to a paper card, below the specimen]
</materialsCitation>
.
</paragraph>
<paragraph id="2A1F36EEBF3AFFF0FED7FD7B8BB2B688" blockId="6.[317,1271,542,1802]" pageId="6" pageNumber="77">
Measurements in mm: body length 2.70 (
<typeStatus id="F51B884CBF3AFFF0FCA5FD7B8A4FB6AC" box="[780,869,686,708]" pageId="6" pageNumber="77" type="holotype">holotype</typeStatus>
), 3.19, wing length 2.60 (
<typeStatus id="F51B884CBF3AFFF0FBF5FD7B8D9FB6AC" box="[1116,1205,686,708]" pageId="6" pageNumber="77" type="holotype">holotype</typeStatus>
), 3.32, wing breadth 1.03 (
<typeStatus id="F51B884CBF3AFFF0FE55FD1F8B7FB688" box="[508,597,714,736]" pageId="6" pageNumber="77" type="holotype">holotype</typeStatus>
), 1.33.
</paragraph>
<paragraph id="2A1F36EEBF3AFFF0FED7FD328DECB695" blockId="6.[317,1271,542,1802]" box="[382,1222,743,765]" pageId="6" pageNumber="77">General colour dark to black; some parts ofbody with brownish grey microtomentum.</paragraph>
<paragraph id="2A1F36EEBF3AFFF0FED7FCD18BE5B0C5" blockId="6.[317,1271,542,1802]" pageId="6" pageNumber="77">Head slightly longer than high. Head with frons broad, frons mostly dark microtomentose, only orbital plates and narrow frontal triangle shiny.. Lunule microtomentose. Face (prefrons) shining black, with relatively deep and strongly microtomentose antennal concavities. Interantennal tubercle red. Parafacialia black, with glossy black margins. Vibrissal angle reddish microtomentose. Gena at genal seta 0.3 as high as longest eye diameter. Gena with a subquadrate microtomentose area on and above peristoma, which widens apically up to almost eye margin. Occiput and postgena black, wholly microtomentose. All cephalic macrosetae relatively thin; a pair ofsmall postocellar setulae only; outer and inner verticals long and subequal to ocellars in length; 2 subequal (or posterior slightly longer), closely set, lateroclinate orbitals; 6 fine interfrontals subequal in length; numerous additional setulae between orbits and interfrontalia, and a series of small, mostly lateroclinate setulae on orbits. Vibrissa long, subvibrissa about one-third ofits length and much thinner. Genal seta well-developed, long; peristomal setulae numerous and fine, postoculars numerous and unarranged. Eye not reduced, slightly oval, its longest (oblique) diameter about 1 1/3 times as long as shortest. Scape and pedicel black, first flagellomere rounded, dirty red to reddish brown. Arista long, about 4 times as long as antenna, with long cilia.</paragraph>
<paragraph id="2A1F36EEBF3AFFF0FED7FB618BCDB1FC" blockId="6.[317,1271,542,1802]" pageId="6" pageNumber="77">
Thorax black, dorsally mostly (except notopleuron and scutellum) dark microtomentose. Pleura bare, microtomentose pattern most resembling that of
<taxonomicName id="EDA04D6DBF3AFFF0FC2AFB048DD8B08F" authority="(MEIGEN, 1830)" baseAuthorityName="MEIGEN" baseAuthorityYear="1830" box="[899,1266,1233,1255]" class="Insecta" family="Sphaeroceridae" genus="Crumomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="77" phylum="Arthropoda" rank="species" species="fimetaria">
<emphasis id="18D4EAFCBF3AFFF0FC2AFB048D78B08F" box="[899,1106,1233,1255]" italics="true" pageId="6" pageNumber="77">Crumomyia fimetaria</emphasis>
(
<smallCapsWord id="2CF9A032BF3AFFF0FBF7FB048D86B08D" baselines="1249,1249" box="[1118,1196,1233,1255]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Meigen" pageId="6" pageNumber="77">MEIGEN</smallCapsWord>
, 1830)
</taxonomicName>
, i.e. microtomentose only behind halteres and on metanotum. All characteristic setae ofmesonotum long and thin: 1 proepisternal, 2 notopleural (anterior definitely longer than posterior), 1 short supra-alar and 1 very long postalar, 1 long presutural, 1+2 relatively weak dorsocentrals, 4 irregular rows ofsparse and fine acrostichal setulae, 2 very long scutellars (basal only slightly shorter than api - cal, no additional scutellars). Propleuron finely setulose, anepisternum without setae. Katepisternum with 4–5 fine pale setulae at dorsal margin.
</paragraph>
<paragraph id="2A1F36EEBF3AFFF0FED7FA4F8A90B2DC" blockId="6.[317,1271,542,1802]" pageId="6" pageNumber="77">
Legs as in
<taxonomicName id="EDA04D6DBF3AFFF0FE4FFA4F8B14B1D8" baseAuthorityName="MEIGEN" baseAuthorityYear="1830" box="[486,574,1434,1456]" class="Insecta" family="Sphaeroceridae" genus="Crumomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="77" phylum="Arthropoda" rank="species" species="fimetaria">
<emphasis id="18D4EAFCBF3AFFF0FE4FFA4F8B14B1D8" box="[486,574,1434,1456]" italics="true" pageId="6" pageNumber="77">fimetaria</emphasis>
</taxonomicName>
, black, but trochanters, knees, apices oftibiae, and tarsi yellowish. Fore femur somewhat thickened, with longer but hair-like setae in posteroventral and posterodorsal rows. Mid femur with 4 thick preapical setae and with a number of hair-like setae ventrally. Hind femur as in
<taxonomicName id="EDA04D6DBF3AFFF0FEF1FA24889AB26F" baseAuthorityName="MEIGEN" baseAuthorityYear="1830" box="[344,432,1521,1543]" class="Insecta" family="Sphaeroceridae" genus="Crumomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="77" phylum="Arthropoda" rank="species" species="fimetaria">
<emphasis id="18D4EAFCBF3AFFF0FEF1FA24889AB26F" box="[344,432,1521,1543]" italics="true" pageId="6" pageNumber="77">fimetaria</emphasis>
</taxonomicName>
. Fore tibia with 1 short dorsal preapical seta and ventroapical brush ofdense, darker brown setulae. Mid tibia with usual subapical crown ofsetae, 6 rather thick anterodorsal setae, 1 strong anteroventral seta at apical 2/7, and 1 much weaker posteroventral seta opposite or slightly distal to anteroventral. Hind tibia with the usual dorsal preapical seta, without an anteroventral seta but with 2 apical spurs (the anterior short, the ventral strong, curved, about as long as tibial width). Fore and hind basitarsi with a well developed, small, black ventroapical hook, hind basitarsus and 2nd hind tarsal segment with dense ventral brush ofdark brown hairs.
</paragraph>
<paragraph id="2A1F36EEBF3AFFF1FED7F96F8DF2B58F" blockId="6.[317,1271,542,1802]" lastBlockId="7.[317,1271,436,890]" lastPageId="7" lastPageNumber="78" pageId="6" pageNumber="77">
<materialsCitation id="9AC83CB3BF3AFFF1FED7F96F8DF9B58F" collectionCode="R" lastPageId="7" lastPageNumber="78" pageId="6" pageNumber="77" specimenCode="R1" specimenCount="1" typeStatus="paratype">
Wing normal with normal venation; membrane and veins unicolorous light brownish, but both cross-veins brown infuscated, R-M more strongly so. Costal vein dark brown distally to
<specimenCode id="7A069E95BF3AFFF0FBDEF9028DA4B298" box="[1143,1166,1751,1776]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="77">
R
<subScript id="B62434ABBF3AFFF0FB2EF9318DA4B298" attach="left" box="[1159,1166,1764,1776]" fontSize="5" pageId="6" pageNumber="77">1</subScript>
</specimenCode>
, veins otherwise lighter brown. Wing vein ratios: C-index (
<typeStatus id="F51B884CBF3AFFF0FCBFF9218A46B362" box="[790,876,1780,1802]" pageId="6" pageNumber="77" type="paratype">paratype</typeStatus>
) =
<quantity id="ED589B0BBF3AFFF0FC25F9218AC7B362" box="[908,1005,1780,1802]" metricMagnitude="-3" metricUnit="m" metricValue="1.692" pageId="6" pageNumber="77" unit="mm" value="1.692">1.692 mm</quantity>
/
<quantity id="ED589B0BBF3AFFF0FBA9F9218D4BB362" box="[1024,1121,1780,1802]" metricMagnitude="-4" metricUnit="m" metricValue="5.77" pageId="6" pageNumber="77" unit="mm" value="0.577">0.577 mm</quantity>
= 2.93, M vein index =
<quantity id="ED589B0BBF3BFFF1FE23FE6188C1B5A2" box="[394,491,436,458]" metricMagnitude="-4" metricUnit="m" metricValue="9.23" pageId="7" pageNumber="78" unit="mm" value="0.923">0.923 mm</quantity>
/
<quantity id="ED589B0BBF3BFFF1FE5BFE618B6EB5A2" box="[498,580,436,458]" metricMagnitude="-4" metricUnit="m" metricValue="8.8" pageId="7" pageNumber="78" unit="mm" value="0.88">0.88 mm</quantity>
= 1.05, dM-Cu
<quantity id="ED589B0BBF3BFFF1FD72FE618A14B5A2" box="[731,830,436,458]" metricMagnitude="-4" metricUnit="m" metricValue="2.9499999999999997" pageId="7" pageNumber="78" unit="mm" value="0.295">0.295 mm</quantity>
, R-M-dM-Cu / dM-Cu ratio 3.13. Vein stub of Cu posterior to discal cell
<quantity id="ED589B0BBF3BFFF1FDE8FE048BB2B58F" box="[577,664,465,487]" metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="7" pageNumber="78" unit="mm" value="0.07">0.07 mm</quantity>
long. Haltere dirty yellow, knob usually darker than stem
</materialsCitation>
.
</paragraph>
<paragraph id="2A1F36EEBF3BFFF1FED7FE3B8D08B61F" blockId="7.[317,1271,436,890]" pageId="7" pageNumber="78">
Abdomen with well-sclerotized preabdominal sclerites. Terga black, broad, rather shiny, with fine setulae longest at lateral margins. Preabdominal sterna comparatively narrow, like in
<taxonomicName id="EDA04D6DBF3BFFF1FB3EFDDF8DC5B648" baseAuthorityName="MEIGEN" baseAuthorityYear="1830" box="[1175,1263,522,544]" class="Insecta" family="Sphaeroceridae" genus="Crumomyia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="78" phylum="Arthropoda" rank="species" species="fimetaria">
<emphasis id="18D4EAFCBF3BFFF1FB3EFDDF8DC5B648" box="[1175,1263,522,544]" italics="true" pageId="7" pageNumber="78">fimetaria</emphasis>
</taxonomicName>
, sterna 2–5 ofalmost equal breadth, 5th sternum unmodified. Postabdominal sclerites well-sclerot - ized, dark; 6th and 7th sterna strongly asymmetrical and fused as in congeners, 8th sternum also asymmetrical and relatively short, dorsally strongly convex and partly bare.
</paragraph>
<paragraph id="2A1F36EEBF3BFFFEFED7FDAB8D51B632" blockId="7.[317,1271,436,890]" lastBlockId="8.[317,1270,436,631]" lastPageId="8" lastPageNumber="79" pageId="7" pageNumber="78">
Male genital morphology is similar to that of
<taxonomicName id="EDA04D6DBF3BFFF1FC86FDAB8A88B6FC" baseAuthorityName="MEIGEN" baseAuthorityYear="1830" box="[815,930,638,660]" class="Insecta" family="Sphaeroceridae" genus="Crumomyia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="78" phylum="Arthropoda" rank="species" species="fimetaria">
<emphasis id="18D4EAFCBF3BFFF1FC86FDAB8A88B6FC" box="[815,930,638,660]" italics="true" pageId="7" pageNumber="78">C. fimetaria</emphasis>
</taxonomicName>
and
<taxonomicName id="EDA04D6DBF3BFFF1FC79FDAB8D01B6FB" baseAuthorityName="RONDANI" baseAuthorityYear="1880" box="[976,1067,638,660]" class="Insecta" family="Sphaeroceridae" genus="Crumomyia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="78" phylum="Arthropoda" rank="species" species="roserii">
<emphasis id="18D4EAFCBF3BFFF1FC79FDAB8D01B6FB" box="[976,1067,638,660]" italics="true" pageId="7" pageNumber="78">C. roserii</emphasis>
</taxonomicName>
. Epandrium strongly convex, with the usual ventral lateral cleft and densely uniformly haired, without extremely long setae. Cerci distinct but fused with epandrium (
<figureCitation id="B29B2A6BBF3BFFF1FCA6FD628A7DB6A5" box="[783,855,695,717]" captionStart="Figs 10–13" captionStartId="7.[317,360,1745,1767]" captionTargetBox="[326,1260,962,1711]" captionTargetId="figure-265@7.[323,1263,950,1716]" captionTargetPageId="7" captionText="Figs 10–13. Crumomyia topali sp. n., paratype male, genitalia: 10 = right halfofterminalia caudally, 11 = surstylus, broadest extension (setae on inner (medial) surface not depicted), 12 = phallus and" figureDoi="http://doi.org/10.5281/zenodo.12587214" httpUri="https://zenodo.org/record/12587214/files/figure.png" pageId="7" pageNumber="78">Fig. 10</figureCitation>
) and medially well separated, each with a number oflong and almost straight hair-like setae and a number offine setulae. Subepandrial sclerite well developed (
<figureCitation id="B29B2A6BBF3BFFF1FE70FD248B35B76F" box="[473,543,753,775]" captionStart="Figs 10–13" captionStartId="7.[317,360,1745,1767]" captionTargetBox="[326,1260,962,1711]" captionTargetId="figure-265@7.[323,1263,950,1716]" captionTargetPageId="7" captionText="Figs 10–13. Crumomyia topali sp. n., paratype male, genitalia: 10 = right halfofterminalia caudally, 11 = surstylus, broadest extension (setae on inner (medial) surface not depicted), 12 = phallus and" figureDoi="http://doi.org/10.5281/zenodo.12587214" httpUri="https://zenodo.org/record/12587214/files/figure.png" pageId="7" pageNumber="78">Fig. 10</figureCitation>
). Surstylus (
<figureCitation id="B29B2A6BBF3BFFF1FD3AFD248BFDB76F" box="[659,727,753,775]" captionStart="Figs 10–13" captionStartId="7.[317,360,1745,1767]" captionTargetBox="[326,1260,962,1711]" captionTargetId="figure-265@7.[323,1263,950,1716]" captionTargetPageId="7" captionText="Figs 10–13. Crumomyia topali sp. n., paratype male, genitalia: 10 = right halfofterminalia caudally, 11 = surstylus, broadest extension (setae on inner (medial) surface not depicted), 12 = phallus and" figureDoi="http://doi.org/10.5281/zenodo.12587214" httpUri="https://zenodo.org/record/12587214/files/figure.png" pageId="7" pageNumber="78">Fig. 11</figureCitation>
) broad and round in broadest extension, in this view anterior border not broken (as in
<taxonomicName id="EDA04D6DBF3BFFF1FDE1FCDB8B88B74B" baseAuthorityName="RONDANI" baseAuthorityYear="1880" box="[584,674,782,804]" class="Insecta" family="Sphaeroceridae" genus="Crumomyia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="78" phylum="Arthropoda" rank="species" species="roserii">
<emphasis id="18D4EAFCBF3BFFF1FDE1FCDB8B88B74B" box="[584,674,782,804]" italics="true" pageId="7" pageNumber="78">C. roserii</emphasis>
</taxonomicName>
), practically no apex on surstylus (cf. fig. 105 of
<taxonomicName id="EDA04D6DBF3BFFF1FBC2FCDB8DF7B74C" baseAuthorityName="MEIGEN" baseAuthorityYear="1830" box="[1131,1245,782,804]" class="Insecta" family="Sphaeroceridae" genus="Crumomyia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="78" phylum="Arthropoda" rank="species" species="fimetaria">
<emphasis id="18D4EAFCBF3BFFF1FBC2FCDB8DF7B74C" box="[1131,1245,782,804]" italics="true" pageId="7" pageNumber="78">C. fimetaria</emphasis>
</taxonomicName>
of
<bibRefCitation id="4E314B1FBF3BFFF1FE94FCFF8B00B728" author="NORRBOM, A. L. &amp; KIM, K. C." box="[317,554,810,832]" pageId="7" pageNumber="78" pagination="167 - 225" refId="ref7257" refString="NORRBOM, A. L. &amp; KIM, K. C. (1985 a) Systematics ofCrumomyia Macquart and Alloborborus (Diptera: Sphaeroceridae). Syst. Ent. 10: 167 - 225." type="journal article" year="1985">
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&amp;
<smallCapsWord id="2CF9A032BF3BFFF1FE16FCFF88CDB757" baselines="826,827" box="[447,487,810,832]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Kim" pageId="7" pageNumber="78">KIM</smallCapsWord>
1985
<emphasis id="18D4EAFCBF3BFFF1FDB7FCFE8B00B728" box="[542,554,811,832]" italics="true" pageId="7" pageNumber="78">a</emphasis>
</bibRefCitation>
); lateral surface in apical half covered by medium long setae, i.e. no such setae in basal half (cf. figs 100 and 105 of
<bibRefCitation id="4E314B1FBF3BFFF1FD73FC928AECB735" author="NORRBOM, A. L. &amp; KIM, K. C." box="[730,966,839,861]" pageId="7" pageNumber="78" pagination="167 - 225" refId="ref7257" refString="NORRBOM, A. L. &amp; KIM, K. C. (1985 a) Systematics ofCrumomyia Macquart and Alloborborus (Diptera: Sphaeroceridae). Syst. Ent. 10: 167 - 225." type="journal article" year="1985">
<smallCapsWord id="2CF9A032BF3BFFF1FD73FC928A16B734" baselines="855,856" box="[730,828,839,861]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Norrbom" pageId="7" pageNumber="78">NORRBOM</smallCapsWord>
&amp;
<smallCapsWord id="2CF9A032BF3BFFF1FCF5FC928AAEB734" baselines="855,856" box="[860,900,839,861]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Kim" pageId="7" pageNumber="78">KIM</smallCapsWord>
1985
<emphasis id="18D4EAFCBF3BFFF1FC13FC9D8AECB735" box="[954,966,840,861]" italics="true" pageId="7" pageNumber="78">a</emphasis>
</bibRefCitation>
). Hypandrium as in congeners. Aedeagal complex (
<figureCitation id="B29B2A6BBF3BFFF1FE56FCB18B6FB712" box="[511,581,868,890]" captionStart="Figs 10–13" captionStartId="7.[317,360,1745,1767]" captionTargetBox="[326,1260,962,1711]" captionTargetId="figure-265@7.[323,1263,950,1716]" captionTargetPageId="7" captionText="Figs 10–13. Crumomyia topali sp. n., paratype male, genitalia: 10 = right halfofterminalia caudally, 11 = surstylus, broadest extension (setae on inner (medial) surface not depicted), 12 = phallus and" figureDoi="http://doi.org/10.5281/zenodo.12587214" httpUri="https://zenodo.org/record/12587214/files/figure.png" pageId="7" pageNumber="78">Fig. 12</figureCitation>
) ofthe
<taxonomicName id="EDA04D6DBF3BFFF1FD3BFCB18A2DB712" baseAuthorityName="MEIGEN" baseAuthorityYear="1830" box="[658,775,868,890]" class="Insecta" family="Sphaeroceridae" genus="Crumomyia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="78" phylum="Arthropoda" rank="species" species="fimetaria">
<emphasis id="18D4EAFCBF3BFFF1FD3BFCB18A2DB712" box="[658,775,868,890]" italics="true" pageId="7" pageNumber="78">C. fimetaria</emphasis>
</taxonomicName>
-subgroup
<typeStatus id="F51B884CBF3BFFF1FCC6FCB18AB1B712" box="[879,923,868,890]" pageId="7" pageNumber="78">type</typeStatus>
, resembling that of
<taxonomicName id="EDA04D6DBF3BFFF1FBF2FCB18DFAB712" baseAuthorityName="MEIGEN" baseAuthorityYear="1830" box="[1115,1232,868,890]" class="Insecta" family="Sphaeroceridae" genus="Crumomyia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="78" phylum="Arthropoda" rank="species" species="fimetaria">
<emphasis id="18D4EAFCBF3BFFF1FBF2FCB18DFAB712" box="[1115,1232,868,890]" italics="true" pageId="7" pageNumber="78">C. fimetaria</emphasis>
</taxonomicName>
but postgonite, lateral view, 13 = postgonite laterally. Scale:
<quantity id="ED589B0BBF3BFFF1FC4FF8DF8D05B348" box="[998,1071,1802,1824]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="78" unit="mm" value="0.2">0.2 mm</quantity>
for all differing as follows: basiphallus somewhat more slender with thicker and straight epiphallus (resembling more to that of
<taxonomicName id="EDA04D6DBF34FFFEFDA8FE048B76B58E" baseAuthorityName="RONDANI" baseAuthorityYear="1880" box="[513,604,465,487]" class="Insecta" family="Sphaeroceridae" genus="Crumomyia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="79" phylum="Arthropoda" rank="species" species="roserii">
<emphasis id="18D4EAFCBF34FFFEFDA8FE048B76B58E" box="[513,604,465,487]" italics="true" pageId="8" pageNumber="79">C. roserii</emphasis>
</taxonomicName>
), pre-epiphallus arising rather distally (similar to
<taxonomicName id="EDA04D6DBF34FFFEFB99FE048D89B58F" baseAuthorityName="MEIGEN" baseAuthorityYear="1830" box="[1072,1187,465,487]" class="Insecta" family="Sphaeroceridae" genus="Crumomyia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="79" phylum="Arthropoda" rank="species" species="fimetaria">
<emphasis id="18D4EAFCBF34FFFEFB99FE048D89B58F" box="[1072,1187,465,487]" italics="true" pageId="8" pageNumber="79">C. fimetaria</emphasis>
</taxonomicName>
but contrary to
<taxonomicName id="EDA04D6DBF34FFFEFE27FE3B88C7B66B" baseAuthorityName="RONDANI" baseAuthorityYear="1880" box="[398,493,494,516]" class="Insecta" family="Sphaeroceridae" genus="Crumomyia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="79" phylum="Arthropoda" rank="species" species="roserii">
<emphasis id="18D4EAFCBF34FFFEFE27FE3B88C7B66B" box="[398,493,494,516]" italics="true" pageId="8" pageNumber="79">C. roserii</emphasis>
</taxonomicName>
) but stronger than in
<taxonomicName id="EDA04D6DBF34FFFEFD60FE3B8A15B66C" baseAuthorityName="MEIGEN" baseAuthorityYear="1830" box="[713,831,494,516]" class="Insecta" family="Sphaeroceridae" genus="Crumomyia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="79" phylum="Arthropoda" rank="species" species="fimetaria">
<emphasis id="18D4EAFCBF34FFFEFD60FE3B8A15B66C" box="[713,831,494,516]" italics="true" pageId="8" pageNumber="79">C. fimetaria</emphasis>
</taxonomicName>
. Distiphallus with differently formed paired dorsal hooks. Postgonite with broader apical part in profile, i.e. the lobe dorsal to it projects only slightly more anteriorly (
<figureCitation id="B29B2A6BBF34FFFEFD87FDF28B5FB655" box="[558,629,551,573]" captionStart="Figs 10–13" captionStartId="7.[317,360,1745,1767]" captionTargetBox="[326,1260,962,1711]" captionTargetId="figure-265@7.[323,1263,950,1716]" captionTargetPageId="7" captionText="Figs 10–13. Crumomyia topali sp. n., paratype male, genitalia: 10 = right halfofterminalia caudally, 11 = surstylus, broadest extension (setae on inner (medial) surface not depicted), 12 = phallus and" figureDoi="http://doi.org/10.5281/zenodo.12587214" httpUri="https://zenodo.org/record/12587214/files/figure.png" pageId="8" pageNumber="79">Fig. 13</figureCitation>
), seta on lateromedial edge seems longer than that of
<taxonomicName id="EDA04D6DBF34FFFEFBD2FDF28DC5B655" baseAuthorityName="MEIGEN" baseAuthorityYear="1830" box="[1147,1263,551,573]" class="Insecta" family="Sphaeroceridae" genus="Crumomyia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="79" phylum="Arthropoda" rank="species" species="fimetaria">
<emphasis id="18D4EAFCBF34FFFEFBD2FDF28DC5B655" box="[1147,1263,551,573]" italics="true" pageId="8" pageNumber="79">C. fimetaria</emphasis>
</taxonomicName>
. Ejaculatory apodeme rather slender, situated in posterodorsal opening ofphallophore.
</paragraph>
<caption id="7EDF6666BF3BFFF1FE94F9048DDCB36C" ID-DOI="http://doi.org/10.5281/zenodo.12587214" ID-Zenodo-Dep="12587214" httpUri="https://zenodo.org/record/12587214/files/figure.png" pageId="7" pageNumber="78" startId="7.[317,360,1745,1767]" targetBox="[326,1260,962,1711]" targetPageId="7" targetType="figure">
<paragraph id="2A1F36EEBF3BFFF1FE94F9048DDCB36C" blockId="7.[317,1270,1745,1824]" pageId="7" pageNumber="78">
<emphasis id="18D4EAFCBF3BFFF1FE94F9048884B28F" bold="true" box="[317,430,1745,1767]" pageId="7" pageNumber="78">Figs 10–13.</emphasis>
<taxonomicName id="EDA04D6DBF3BFFF1FE1DF9048B48B28F" authorityName="Papp" authorityYear="2003" box="[436,610,1745,1767]" class="Insecta" family="Sphaeroceridae" genus="Crumomyia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="78" phylum="Arthropoda" rank="species" species="topali" status="sp. nov.">
<emphasis id="18D4EAFCBF3BFFF1FE1DF9048B48B28F" box="[436,610,1745,1767]" italics="true" pageId="7" pageNumber="78">Crumomyia topali</emphasis>
</taxonomicName>
<taxonomicNameLabel id="03E75787BF3BFFF1FDCEF9048BB1B28F" box="[615,667,1745,1767]" pageId="7" pageNumber="78" rank="species">sp. n.</taxonomicNameLabel>
, paratype male, genitalia: 10 = right halfofterminalia caudally, 11 = surstylus, broadest extension (setae on inner (medial) surface not depicted), 12 = phallus and
</paragraph>
</caption>
<paragraph id="2A1F36EEBF34FFFEFED7FDB48B07B61F" blockId="8.[317,1270,436,631]" box="[382,557,609,631]" pageId="8" pageNumber="79">Female unknown.</paragraph>
<paragraph id="2A1F36EEBF34FFFEFED7FD488DCCB052" blockId="8.[317,1270,669,1188]" pageId="8" pageNumber="79">
Discussion. This new species is a member ofthe
<taxonomicName id="EDA04D6DBF34FFFEFC75FD488DC1B6DF" baseAuthorityName="Spuler" baseAuthorityYear="1925" box="[988,1259,669,695]" class="Insecta" family="Sphaeroceridae" genus="Crumomyia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="79" phylum="Arthropoda" rank="species" species="setitibialis">
<emphasis id="18D4EAFCBF34FFFEFC75FD488DC1B6DF" box="[988,1259,669,695]" italics="true" pageId="8" pageNumber="79">Crumomyia setitibialis</emphasis>
</taxonomicName>
- group,
<taxonomicName id="EDA04D6DBF34FFFEFE26FD1588D1B6B2" baseAuthorityName="MEIGEN" baseAuthorityYear="1830" box="[399,507,704,730]" class="Insecta" family="Sphaeroceridae" genus="Crumomyia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="79" phylum="Arthropoda" rank="species" species="fimetaria">
<emphasis id="18D4EAFCBF34FFFEFE26FD1588D1B6B2" box="[399,507,704,730]" italics="true" pageId="8" pageNumber="79">fimetaria</emphasis>
</taxonomicName>
subgroup sensu
<bibRefCitation id="4E314B1FBF34FFFEFD13FD158D15B6B2" author="NORRBOM, A. L. &amp; KIM, K. C." box="[698,1087,704,730]" pageId="8" pageNumber="79" pagination="167 - 225" refId="ref7257" refString="NORRBOM, A. L. &amp; KIM, K. C. (1985 a) Systematics ofCrumomyia Macquart and Alloborborus (Diptera: Sphaeroceridae). Syst. Ent. 10: 167 - 225." type="journal article" year="1985">
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and
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(1985
<emphasis id="18D4EAFCBF34FFFEFC40FD148AD2B6B2" box="[1001,1016,705,730]" italics="true" pageId="8" pageNumber="79">a</emphasis>
: 184)
</bibRefCitation>
. Based on similarities in the male genitalia, it appears intermediate between
<taxonomicName id="EDA04D6DBF34FFFEFBBCFD318B24B749" authority="(RONDANI, 1880)" baseAuthorityName="RONDANI" baseAuthorityYear="1880" class="Insecta" family="Sphaeroceridae" genus="Crumomyia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="79" phylum="Arthropoda" rank="species" species="roserii">
<emphasis id="18D4EAFCBF34FFFEFBBCFD318DDFB695" box="[1045,1269,740,766]" italics="true" pageId="8" pageNumber="79">Crumomyia roserii</emphasis>
(
<smallCapsWord id="2CF9A032BF34FFFEFEECFCD28893B748" baselines="794,794" box="[325,441,775,801]" lowerCaseFontSize="9" mainFontSize="11" normCase="title" normString="Rondani" pageId="8" pageNumber="79">RONDANI</smallCapsWord>
, 1880)
</taxonomicName>
and
<taxonomicName id="EDA04D6DBF34FFFEFDE5FCD28A83B749" authority="(MEIGEN, 1830)" baseAuthorityName="MEIGEN" baseAuthorityYear="1830" box="[588,937,775,801]" class="Insecta" family="Sphaeroceridae" genus="Crumomyia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="79" phylum="Arthropoda" rank="species" species="fimetaria">
<emphasis id="18D4EAFCBF34FFFEFDE5FCD28BF6B749" box="[588,732,775,801]" italics="true" pageId="8" pageNumber="79">C. fimetaria</emphasis>
(
<smallCapsWord id="2CF9A032BF34FFFEFD44FCD28A7BB748" baselines="794,794" box="[749,849,775,801]" lowerCaseFontSize="9" mainFontSize="11" normCase="title" normString="Meigen" pageId="8" pageNumber="79">MEIGEN</smallCapsWord>
, 1830)
</taxonomicName>
, which are consequently its sister-species. All these species share similar, very small (ifany) microtomentose pattern on anepisternum, only a few setae on dorsal part of katepisternum, no additional marginal setae on scutellum, male femora, tibiae and abdominal membrane with rather short setae, hind tibiae without an anteroventral seta, and similar structure ofinner male genitalia. However, with its bare scutellum, similar colouration oflegs, and details ofgenital structures, the new species seems closer to
<taxonomicName id="EDA04D6DBF34FFFEFB3EFC0F885CB07F" baseAuthorityName="MEIGEN" baseAuthorityYear="1830" class="Insecta" family="Sphaeroceridae" genus="Crumomyia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="79" phylum="Arthropoda" rank="species" species="fimetaria">
<emphasis id="18D4EAFCBF34FFFEFB3EFC0F885CB07F" italics="true" pageId="8" pageNumber="79">C. fimetaria</emphasis>
</taxonomicName>
. Indeed, only details of genital structures, described above, differentiate this species from
<taxonomicName id="EDA04D6DBF34FFFEFE7DFBF58B76B052" baseAuthorityName="MEIGEN" baseAuthorityYear="1830" box="[468,604,1056,1082]" class="Insecta" family="Sphaeroceridae" genus="Crumomyia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="79" phylum="Arthropoda" rank="species" species="fimetaria">
<emphasis id="18D4EAFCBF34FFFEFE7DFBF58B76B052" box="[468,604,1056,1082]" italics="true" pageId="8" pageNumber="79">C. fimetaria</emphasis>
</taxonomicName>
(
<figureCitation id="B29B2A6BBF34FFFEFDC0FBF58BC6B052" box="[617,748,1056,1082]" captionStart="Figs 10–13" captionStartId="7.[317,360,1745,1767]" captionTargetBox="[326,1260,962,1711]" captionTargetId="figure-265@7.[323,1263,950,1716]" captionTargetPageId="7" captionText="Figs 10–13. Crumomyia topali sp. n., paratype male, genitalia: 10 = right halfofterminalia caudally, 11 = surstylus, broadest extension (setae on inner (medial) surface not depicted), 12 = phallus and" figureDoi="http://doi.org/10.5281/zenodo.12587214" httpUri="https://zenodo.org/record/12587214/files/figure.png" pageId="8" pageNumber="79">Figs 10–13</figureCitation>
, cf.
<bibRefCitation id="4E314B1FBF34FFFEFCB1FBF58D18B052" author="NORRBOM, A. L. &amp; KIM, K. C." box="[792,1074,1056,1082]" pageId="8" pageNumber="79" pagination="167 - 225" refId="ref7257" refString="NORRBOM, A. L. &amp; KIM, K. C. (1985 a) Systematics ofCrumomyia Macquart and Alloborborus (Diptera: Sphaeroceridae). Syst. Ent. 10: 167 - 225." type="journal article" year="1985">
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&amp;
<smallCapsWord id="2CF9A032BF34FFFEFC1DFBF58ACCB052" baselines="1075,1076" box="[948,998,1056,1082]" lowerCaseFontSize="9" mainFontSize="11" normCase="title" normString="Kim" pageId="8" pageNumber="79">KIM</smallCapsWord>
1985
<emphasis id="18D4EAFCBF34FFFEFB8DFBF48D18B052" box="[1060,1074,1057,1082]" italics="true" pageId="8" pageNumber="79">a</emphasis>
</bibRefCitation>
: figs 105–106).
</paragraph>
<paragraph id="2A1F36EEBF34FFFEFED7FB918886B0E9" blockId="8.[317,1270,669,1188]" pageId="8" pageNumber="79">
The new species, like
<taxonomicName id="EDA04D6DBF34FFFEFD3AFB918A0FB036" baseAuthorityName="MEIGEN" baseAuthorityYear="1830" box="[659,805,1092,1118]" class="Insecta" family="Sphaeroceridae" genus="Crumomyia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="79" phylum="Arthropoda" rank="species" species="fimetaria">
<emphasis id="18D4EAFCBF34FFFEFD3AFB918A0FB036" box="[659,805,1092,1118]" italics="true" pageId="8" pageNumber="79">C. fimetaria</emphasis>
</taxonomicName>
, is terricolous but probably not cavedwelling.
</paragraph>
<paragraph id="2A1F36EEBF34FFFEFED7FB5F8D57B0CC" blockId="8.[317,1270,669,1188]" box="[382,1149,1162,1188]" pageId="8" pageNumber="79">
The genus
<taxonomicName id="EDA04D6DBF34FFFEFE56FB5F8A35B0CC" authority="MACQUART" authorityName="MACQUART" box="[511,799,1162,1188]" class="Insecta" family="Sphaeroceridae" genus="Crumomyia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis id="18D4EAFCBF34FFFEFE56FB5F8BA0B0CC" box="[511,650,1162,1188]" italics="true" pageId="8" pageNumber="79">Crumomyia</emphasis>
<smallCapsWord id="2CF9A032BF34FFFEFD38FB5F8A35B0CC" baselines="1181,1182" box="[657,799,1162,1188]" lowerCaseFontSize="9" mainFontSize="11" normCase="title" normString="Macquart" pageId="8" pageNumber="79">MACQUART</smallCapsWord>
</taxonomicName>
is new to the Oriental region.
</paragraph>
</subSubSection>
</treatment>
</document>
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