327 lines
40 KiB
XML
327 lines
40 KiB
XML
<document id="45A2BFA4AE20B69798DE1A072FBF6BAD" ID-CLB-Dataset="202083" ID-DOI="10.11646/zootaxa.5211.1.1" ID-GBIF-Dataset="bb97b9dd-2acc-4244-8f30-72bef07b9ed9" ID-ISSN="1175-5326" ID-Zenodo-Dep="7345007" ID-ZooBank="5754769C-B747-4714-BDD9-7D5509D48BEB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1669101340847" checkinUser="plazi" docAuthor="López-García, Margarita M. & Deloya, Cuauhtémoc" docDate="2022" docId="A209493BFD6EFFB3AFA61AF9FE30B855" docLanguage="en" docName="zootaxa.5211.1.1.pdf" docOrigin="Zootaxa 5211 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5211.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Ligyrus (Ligyrus) ruginasus LeConte 1856" docType="treatment" docVersion="9" lastPageNumber="51" masterDocId="5E303143FD5FFF81AF311956FFBEBC61" masterDocTitle="Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision" masterLastPageNumber="119" masterPageNumber="1" pageNumber="50" updateTime="1699396337306" updateUser="plazi">
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<mods:title id="C88D16D1ECDB16DB026B5DB9643B0B8A">Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision</mods:title>
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<mods:namePart id="C8CE2091D203119181B835634343C54C">López-García, Margarita M.</mods:namePart>
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<mods:nameIdentifier id="9D818D2957BE61E0B5F5A4A4C48F9267" type="email">margaralopezg@gmail.com</mods:nameIdentifier>
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<mods:namePart id="B239F5DFDA01038699856867B9FCA9CD">Deloya, Cuauhtémoc</mods:namePart>
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<mods:nameIdentifier id="9BFE1545D5064F5C09B83D2EE0140B3B" type="email">cuauhtemoc.deloya@inecol.mx</mods:nameIdentifier>
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<mods:title id="86ABC73E87C9D3661565CE9567B97D24">Zootaxa</mods:title>
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<mods:date id="E92E5CAE3F355C856ADBFC26D2DFF81A">2022</mods:date>
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<mods:number id="7FE3C8860258525A6BE31DAE3388F3E1">2022-11-22</mods:number>
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<treatment id="A209493BFD6EFFB3AFA61AF9FE30B855" ID-DOI="http://doi.org/10.5281/zenodo.7383767" ID-GBIF-Taxon="205794496" ID-Zenodo-Dep="7383767" LSID="urn:lsid:plazi:treatment:A209493BFD6EFFB3AFA61AF9FE30B855" httpUri="http://treatment.plazi.org/id/A209493BFD6EFFB3AFA61AF9FE30B855" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50">
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<subSubSection id="62BAABA6FD6EFFB0AFA61AF9FCE7BFA8" box="[151,857,942,969]" pageId="49" pageNumber="50" type="nomenclature">
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<paragraph id="2A1FF82DFD6EFFB0AFA61AF9FCE7BFA8" blockId="49.[151,857,942,1005]" box="[151,857,942,969]" pageId="49" pageNumber="50">
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<heading id="71574F41FD6EFFB0AFA61AF9FCE7BFA8" bold="true" box="[151,857,942,969]" fontSize="11" level="2" pageId="49" pageNumber="50" reason="4">
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<emphasis id="18D4243FFD6EFFB0AFA61AF9FCE7BFA8" bold="true" box="[151,857,942,969]" pageId="49" pageNumber="50">
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<taxonomicName id="EDA083AEFD6EFFB0AFA61AF9FD16BFA9" authority="LeConte, 1856" authorityName="LeConte" authorityYear="1856" box="[151,680,942,969]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="ruginasus" subGenus="Ligyrus">
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<emphasis id="18D4243FFD6EFFB0AFA61AF9FF4ABFA8" bold="true" box="[151,244,943,969]" italics="true" pageId="49" pageNumber="50">Ligyrus</emphasis>
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||
(
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<emphasis id="18D4243FFD6EFFB0AE341AF9FEDCBFA8" bold="true" box="[261,354,943,969]" italics="true" pageId="49" pageNumber="50">Ligyrus</emphasis>
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||
)
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<emphasis id="18D4243FFD6EFFB0AE421AF9FE52BFA8" bold="true" box="[371,492,943,969]" italics="true" pageId="49" pageNumber="50">ruginasus</emphasis>
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||
<bibRefCitation id="4E3185DCFD6EFFB0AEC21AF8FD16BFA9" author="LeConte, J. L." box="[499,680,942,968]" pageId="49" pageNumber="50" pagination="19 - 25" refId="ref91224" refString="LeConte, J. L. (1856) Notice of three genera of Scarabaeidae found in the United States. Proceedings of the Academy of Natural Sciences of Philadelphia, 8, 19 - 25." type="journal article" year="1856">LeConte, 1856</bibRefCitation>
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||
</taxonomicName>
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revised status
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="62BAABA6FD6EFFB0AFA61A85FE56BF8C" box="[151,488,978,1005]" pageId="49" pageNumber="50" type="description">
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<paragraph id="2A1FF82DFD6EFFB0AFA61A85FE56BF8C" blockId="49.[151,857,942,1005]" box="[151,488,978,1005]" pageId="49" pageNumber="50">
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(
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||
<figureCitation id="B29BE4A8FD6EFFB0AFAE1A85FF41BF8C" box="[159,255,979,1005]" captionStart="FIGURE 1" captionStartId="81.[151,250,1535,1559]" captionTargetBox="[151,1436,181,1511]" captionTargetId="figure-17@81.[151,1436,181,1511]" captionTargetPageId="81" captionText="FIGURE 1. Head. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) Ligyrus (Ligyrodes) peruvianus, D) L. (Anagrylius) moroni, E) L. (Ligyrus) ruginasus, F) L. (Ligyrus) nasutus, G) L. (Ligyrus) bidentulus, H) Tomarus adoceteus, I) T. maimon, J) T. pullus, K) T. pumilus." figureDoi="http://doi.org/10.5281/zenodo.7345011" httpUri="https://zenodo.org/record/7345011/files/figure.png" pageId="49" pageNumber="50">Figs. 1E</figureCitation>
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||
,
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||
<figureCitation id="B29BE4A8FD6EFFB0AE3D1A84FE87BF8C" box="[268,313,978,1005]" captionStart="FIGURE 15" captionStartId="90.[151,250,1535,1559]" captionTargetBox="[154,1430,183,1509]" captionTargetId="figure-24@90.[151,1436,181,1512]" captionTargetPageId="90" captionText="FIGURE 15. Parameres in frontal view. A) Euligyrus ebenus, B) E. similis, C) Proculigyrus cicatricosus, D) Ligyrus (Anagrylius) cuniculus, E) L. (Anagrylius) moroni, F) L. (Ligyrodes) peruvianus, G) L. (Ligyrodes) relictus, H) L. (Ligyrodes) sallaei, I) L. (Ligyrus) allonasutus, J) L. (Ligyrus) bidentulus, K) L. (Ligyrus) burmeisteri, L) Ligyrus (Ligyrus) fossor, M) L. (Ligyrus) gibbosus, N) L. (Ligyrus) nasutus, O) L. (Ligyrus) neglectus, P) L. (Ligyrus) paranaensis, Q) L. (Ligyrus) peninsularis, R) L. (Ligyrus) rubripes, S) L. (Ligyrus) ruginasus, T) L. (Ligyrus) villosus." figureDoi="http://doi.org/10.5281/zenodo.7345063" httpUri="https://zenodo.org/record/7345063/files/figure.png" pageId="49" pageNumber="50">15S</figureCitation>
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||
,
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||
<figureCitation id="B29BE4A8FD6EFFB0AE741A85FEC6BF8C" box="[325,376,979,1005]" captionStart="FIGURE 18" captionStartId="93.[151,250,1597,1621]" captionTargetBox="[152,1436,184,1567]" captionTargetId="figure-17@93.[151,1436,181,1573]" captionTargetPageId="93" captionText="FIGURE 18. Parameres in lateral view.A) L. (Ligyrus.) allonasutus, B) L. (Ligyrus) bidentulus, C) L. (Ligyrus) burmeisteri, D) Ligyrus (Ligyrus) fossor, E) L. (Ligyrus) gibbosus, F) L. (Ligyrus) nasutus, G) L. (Ligyrus) neglectus, H) L. (Ligyrus) paranaensis, I) L. (Ligyrus) peninsularis, J) L. (Ligyrus) rubripes, K) L. (Ligyrus) ruginasus, L) L. (Ligyrus) villosus." figureDoi="http://doi.org/10.5281/zenodo.7345077" httpUri="https://zenodo.org/record/7345077/files/figure.png" pageId="49" pageNumber="50">18K</figureCitation>
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,
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<figureCitation id="B29BE4A8FD6EFFB0AEB51A85FE09BF8C" box="[388,439,979,1005]" captionStart="FIGURE 26" captionStartId="101.[151,250,1827,1851]" captionTargetBox="[159,1430,189,1791]" captionTargetId="figure-17@101.[151,1436,181,1803]" captionTargetPageId="101" captionText="FIGURE 26. Habitus of the species of Ligyrus. Subgenus Ligyrus: A) Ligyrus fossor, B) L. gibbosus, C) L. nasutus, D) L. neglectus, E) L. paranaensis, F) L. peninsularis, G) L. rubripes, H) L. ruginasus, I) L. villosus. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7345101" httpUri="https://zenodo.org/record/7345101/files/figure.png" pageId="49" pageNumber="50">26H</figureCitation>
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||
;
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||
<figureCitation id="B29BE4A8FD6EFFB0AEF21A85FE5EBF8C" box="[451,480,979,1005]" captionStart="FIGURE 39" captionStartId="110.[151,250,945,969]" captionTargetBox="[162,1434,181,912]" captionTargetId="figure-24@110.[153,1434,181,921]" captionTargetPageId="110" captionText="FIGURE 39. Distributional records of Ligyrus (Ligyrus) ruginasus and L. peninsularis." figureDoi="http://doi.org/10.5281/zenodo.7345147" httpUri="https://zenodo.org/record/7345147/files/figure.png" pageId="49" pageNumber="50">39</figureCitation>
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||
)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="62BAABA6FD6EFFB0AFA61D4EFCB8B84E" box="[151,774,1047,1071]" pageId="49" pageNumber="50" type="reference_group">
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||
<paragraph id="2A1FF82DFD6EFFB0AFA61D4EFCB8B84E" blockId="49.[151,1437,1047,1133]" box="[151,774,1047,1071]" pageId="49" pageNumber="50">
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||
<treatmentCitationGroup id="0AB0DF03FD6EFFB0AFA61D4EFCB8B84E" box="[151,774,1047,1071]" pageId="49" pageNumber="50">
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<treatmentCitation id="AB01DE3CFD6EFFB0AFA61D4EFDA7B84E" author="LeConte, J. L." box="[151,537,1047,1071]" page="20" pageId="49" pageNumber="50" year="1856">
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<taxonomicName id="EDA083AEFD6EFFB0AFA61D4EFDA7B84E" authority="LeConte, 1856: 20" authorityName="LeConte" authorityPageNumber="20" authorityYear="1856" box="[151,537,1047,1071]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="ruginasus">
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||
<emphasis id="18D4243FFD6EFFB0AFA61D4EFEEEB84E" box="[151,336,1048,1071]" italics="true" pageId="49" pageNumber="50">Ligyrus ruginasus</emphasis>
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||
<bibRefCitation id="4E3185DCFD6EFFB0AE661D41FDA7B84E" author="LeConte, J. L." box="[343,537,1047,1071]" pageId="49" pageNumber="50" pagination="19 - 25" refId="ref91224" refString="LeConte, J. L. (1856) Notice of three genera of Scarabaeidae found in the United States. Proceedings of the Academy of Natural Sciences of Philadelphia, 8, 19 - 25." type="journal article" year="1856">LeConte, 1856: 20</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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||
. Original combination.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="62BAABA6FD6EFFB0AC3D1D41FC59B80D" pageId="49" pageNumber="50" type="materials_examined">
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||
<paragraph id="2A1FF82DFD6EFFB0AC3D1D41FB11B82F" blockId="49.[151,1437,1047,1133]" pageId="49" pageNumber="50">
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Female
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<typeStatus id="F51B468FFD6EFFB0AC6F1D41FC09B84E" box="[862,951,1047,1071]" pageId="49" pageNumber="50" type="holotype">holotype</typeStatus>
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||
(MCZ) “Type / 3717 //
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<taxonomicName id="EDA083AEFD6EFFB0AB811D4EFA91B84E" authorityName="LeConte" authorityYear="1856" box="[1200,1327,1047,1071]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="ruginasus">L. ruginasus</taxonomicName>
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/
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<collectorName id="87559DFBFD6EFFB0AA751D4EFF42B82F" pageId="49" pageNumber="50">Rg. Barracks</collectorName>
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||
/ Hald. / Lec.” Type locality: Ringgold Barracks,
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<collectingRegion id="E86436CFFD6EFFB0ADC71D60FC8FB82C" box="[758,817,1078,1101]" country="United States of America" name="Texas" pageId="49" pageNumber="50">Texas</collectingRegion>
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(near
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<collectingRegion id="E86436CFFD6EFFB0AC401D60FC59B82F" box="[881,999,1078,1102]" country="Puerto Rico" name="Rio Grande" pageId="49" pageNumber="50">Rio Grande</collectingRegion>
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City,
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<collectingCounty id="C37E80A1FD6EFFB0AB151D60FB19B82F" box="[1060,1191,1078,1102]" pageId="49" pageNumber="50">Starr County</collectingCounty>
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||
)
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</paragraph>
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<paragraph id="2A1FF82DFD6EFFB0AFA61D02FC59B80D" blockId="49.[151,1437,1047,1133]" box="[151,999,1108,1133]" pageId="49" pageNumber="50">
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||
<treatmentCitationGroup id="0AB0DF03FD6EFFB0AFA61D02FC59B80D" box="[151,999,1108,1133]" pageId="49" pageNumber="50">
|
||
<treatmentCitation id="AB01DE3CFD6EFFB0AFA61D02FDAEB80C" author="Casey, T. L." box="[151,528,1108,1133]" page="209" pageId="49" pageNumber="50" year="1915">
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||
<taxonomicName id="EDA083AEFD6EFFB0AFA61D02FDAEB80C" authority="Casey, 1915: 209" authorityName="Casey" authorityPageNumber="209" authorityYear="1915" box="[151,528,1108,1133]" class="Insecta" family="Dynastidae" genus="Oxygrylius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="pimalis">
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||
<emphasis id="18D4243FFD6EFFB0AFA61D02FEE6B80D" box="[151,344,1108,1132]" italics="true" pageId="49" pageNumber="50">Oxygrylius pimalis</emphasis>
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||
<bibRefCitation id="4E3185DCFD6EFFB0AE6F1D03FDAEB80C" author="Casey, T. L." box="[350,528,1108,1133]" pageId="49" pageNumber="50" pagination="1 - 394" refId="ref88969" refString="Casey, T. L. (1915) A review of the American species of Rutelinae, Dynastinae, and Cetoniinae. Memoirs on the Coleoptera, 6, 1 - 394." type="journal article" year="1915">Casey, 1915: 209</bibRefCitation>
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||
</taxonomicName>
|
||
</treatmentCitation>
|
||
. Synonym.
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||
<typeStatus id="F51B468FFD6EFFB0ADB61D03FD5AB80C" box="[647,740,1109,1133]" pageId="49" pageNumber="50" type="syntype">Syntypes</typeStatus>
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||
(USNM). Not examined.
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||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="62BAABA6FD6EFFB0AFA61DCCFCCDBA94" pageId="49" pageNumber="50" type="description">
|
||
<paragraph id="2A1FF82DFD6EFFB0AFA61DCCFCCDBA94" blockId="49.[151,1437,1178,2033]" pageId="49" pageNumber="50">
|
||
<emphasis id="18D4243FFD6EFFB0AFA61DCCFE97B8D5" bold="true" box="[151,297,1178,1204]" pageId="49" pageNumber="50">Description.</emphasis>
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||
Habitus as in
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<figureCitation id="B29BE4A8FD6EFFB0AEE81DCDFDFAB8D4" box="[473,580,1179,1205]" captionStart="FIGURE 26" captionStartId="101.[151,250,1827,1851]" captionTargetBox="[159,1430,189,1791]" captionTargetId="figure-17@101.[151,1436,181,1803]" captionTargetPageId="101" captionText="FIGURE 26. Habitus of the species of Ligyrus. Subgenus Ligyrus: A) Ligyrus fossor, B) L. gibbosus, C) L. nasutus, D) L. neglectus, E) L. paranaensis, F) L. peninsularis, G) L. rubripes, H) L. ruginasus, I) L. villosus. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7345101" httpUri="https://zenodo.org/record/7345101/files/figure.png" pageId="49" pageNumber="50">Fig. 26H</figureCitation>
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. Length
|
||
<quantity id="ED5855C8FD6EFFB0AD9C1DCDFCE9B8D4" box="[685,855,1179,1205]" metricMagnitude="-2" metricUnit="m" metricValue="1.645" metricValueMax="1.8699999999999999" metricValueMin="1.42" pageId="49" pageNumber="50" unit="mm" value="16.45" valueMax="18.7" valueMin="14.2">14.2–18.7 mm</quantity>
|
||
; humeral width 7.2–10.0 mm. Color dark reddish brown.
|
||
<emphasis id="18D4243FFD6EFFB0AFC11DE9FE87B8B8" bold="true" box="[240,313,1215,1241]" pageId="49" pageNumber="50">Head:</emphasis>
|
||
Frons deeply rugopunctate, with large and sparse punctures, nearly smooth on vertex, only with sparse, small punctures. Frontoclypeal region with a straight, complete carina (
|
||
<figureCitation id="B29BE4A8FD6EFFB0AB391DB5FBE3B89C" box="[1032,1117,1251,1277]" captionStart="FIGURE 1" captionStartId="81.[151,250,1535,1559]" captionTargetBox="[151,1436,181,1511]" captionTargetId="figure-17@81.[151,1436,181,1511]" captionTargetPageId="81" captionText="FIGURE 1. Head. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) Ligyrus (Ligyrodes) peruvianus, D) L. (Anagrylius) moroni, E) L. (Ligyrus) ruginasus, F) L. (Ligyrus) nasutus, G) L. (Ligyrus) bidentulus, H) Tomarus adoceteus, I) T. maimon, J) T. pullus, K) T. pumilus." figureDoi="http://doi.org/10.5281/zenodo.7345011" httpUri="https://zenodo.org/record/7345011/files/figure.png" pageId="49" pageNumber="50">Fig. 1E</figureCitation>
|
||
). Ocular canthus acute, with 10 ventral setae. Clypeal surface rugopunctate to transversely rugose, striae denser than those on frons. Clypeus triangular, ending in an acute tooth (
|
||
<figureCitation id="B29BE4A8FD6EFFB0AD001C7DFD39B924" box="[561,647,1323,1349]" captionStart="FIGURE 1" captionStartId="81.[151,250,1535,1559]" captionTargetBox="[151,1436,181,1511]" captionTargetId="figure-17@81.[151,1436,181,1511]" captionTargetPageId="81" captionText="FIGURE 1. Head. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) Ligyrus (Ligyrodes) peruvianus, D) L. (Anagrylius) moroni, E) L. (Ligyrus) ruginasus, F) L. (Ligyrus) nasutus, G) L. (Ligyrus) bidentulus, H) Tomarus adoceteus, I) T. maimon, J) T. pullus, K) T. pumilus." figureDoi="http://doi.org/10.5281/zenodo.7345011" httpUri="https://zenodo.org/record/7345011/files/figure.png" pageId="49" pageNumber="50">Fig. 1E</figureCitation>
|
||
). Mandible with 2 apical and 1 lateral, acute tooth, tooth 1 longer and narrower than tooth 2. Maxilla rectangular; galea with 3 dorsal teeth. Apex of labrum straight. Interocular distance 3.5 times an eye width. Antennal club very long.
|
||
<emphasis id="18D4243FFD6EFFB0ADE91C25FCE4B9EC" bold="true" box="[728,858,1395,1421]" pageId="49" pageNumber="50">Pronotum:</emphasis>
|
||
Surface with large, dense punctures. Apex with a prominent tubercle and a deep, rounded fovea.
|
||
<emphasis id="18D4243FFD6EFFB0AD9A1CC1FCBEB9D0" bold="true" box="[683,768,1431,1457]" pageId="49" pageNumber="50">Elytra:</emphasis>
|
||
First interval punctate, punctures similar to other intervals. Inner surface of apex with small, transverse tubercles forming about 66 parallel lines.
|
||
<emphasis id="18D4243FFD6EFFB0AB5D1CECFB54B9B5" bold="true" box="[1132,1258,1466,1492]" pageId="49" pageNumber="50">Abdomen:</emphasis>
|
||
Apex of tergite IV with a diagonal line of 7 transverse, parallel, short carinae. Pygidial surface with small punctures, denser on base and anterior corners. Surface in lateral view strongly convex (male) to flat (female).
|
||
<emphasis id="18D4243FFD6EFFB0AB751F55FB3BBA7C" bold="true" box="[1092,1157,1539,1565]" pageId="49" pageNumber="50">Legs:</emphasis>
|
||
Protibia tridentate; basal tooth slightly distant from others. Protibial surface sparsely punctate. Protarsus of male simple, inner claw simple. Metatibia strongly contracted at apex. Apex of metatibia entire, with 25 spinules.
|
||
<emphasis id="18D4243FFD6EFFB0AB1C1F1DFB45BA05" bold="true" box="[1069,1275,1610,1637]" pageId="49" pageNumber="50">Female genitalia:</emphasis>
|
||
Subcoxite subrectangular (2 times wider than long). Coxite subquadrate (as long as wide); surface strongly concave. Subcoxite wider and subequal in length to coxite.
|
||
<emphasis id="18D4243FFD6EFFB0AD681FC5FCB0BACD" bold="true" box="[601,782,1682,1709]" pageId="49" pageNumber="50">Male genitalia:</emphasis>
|
||
Spiculum gastrale thin, apex truncate. Parameres without ventral teeth; apex widely expanded (
|
||
<figureCitation id="B29BE4A8FD6EFFB0AD0F1FE1FD15BAB0" box="[574,683,1718,1745]" captionStart="FIGURE 15" captionStartId="90.[151,250,1535,1559]" captionTargetBox="[154,1430,183,1509]" captionTargetId="figure-24@90.[151,1436,181,1512]" captionTargetPageId="90" captionText="FIGURE 15. Parameres in frontal view. A) Euligyrus ebenus, B) E. similis, C) Proculigyrus cicatricosus, D) Ligyrus (Anagrylius) cuniculus, E) L. (Anagrylius) moroni, F) L. (Ligyrodes) peruvianus, G) L. (Ligyrodes) relictus, H) L. (Ligyrodes) sallaei, I) L. (Ligyrus) allonasutus, J) L. (Ligyrus) bidentulus, K) L. (Ligyrus) burmeisteri, L) Ligyrus (Ligyrus) fossor, M) L. (Ligyrus) gibbosus, N) L. (Ligyrus) nasutus, O) L. (Ligyrus) neglectus, P) L. (Ligyrus) paranaensis, Q) L. (Ligyrus) peninsularis, R) L. (Ligyrus) rubripes, S) L. (Ligyrus) ruginasus, T) L. (Ligyrus) villosus." figureDoi="http://doi.org/10.5281/zenodo.7345063" httpUri="https://zenodo.org/record/7345063/files/figure.png" pageId="49" pageNumber="50">Figs. 15S</figureCitation>
|
||
,
|
||
<figureCitation id="B29BE4A8FD6EFFB0AD891FE1FD52BAB0" box="[696,748,1719,1745]" captionStart="FIGURE 18" captionStartId="93.[151,250,1597,1621]" captionTargetBox="[152,1436,184,1567]" captionTargetId="figure-17@93.[151,1436,181,1573]" captionTargetPageId="93" captionText="FIGURE 18. Parameres in lateral view.A) L. (Ligyrus.) allonasutus, B) L. (Ligyrus) bidentulus, C) L. (Ligyrus) burmeisteri, D) Ligyrus (Ligyrus) fossor, E) L. (Ligyrus) gibbosus, F) L. (Ligyrus) nasutus, G) L. (Ligyrus) neglectus, H) L. (Ligyrus) paranaensis, I) L. (Ligyrus) peninsularis, J) L. (Ligyrus) rubripes, K) L. (Ligyrus) ruginasus, L) L. (Ligyrus) villosus." figureDoi="http://doi.org/10.5281/zenodo.7345077" httpUri="https://zenodo.org/record/7345077/files/figure.png" pageId="49" pageNumber="50">18K</figureCitation>
|
||
). Internal sac with lamellar spiny belt short (2.5 times longer than copulatory lamellae); accessory lamella thickened, elongate.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="62BAABA6FD6EFFB0AFF61FA8FB4BBB90" pageId="49" pageNumber="50" type="diagnosis">
|
||
<paragraph id="2A1FF82DFD6EFFB0AFF61FA8FF45BBC8" blockId="49.[151,1437,1178,2033]" pageId="49" pageNumber="50">
|
||
<emphasis id="18D4243FFD6EFFB0AFF61FA8FEFCBB79" bold="true" box="[199,322,1790,1816]" pageId="49" pageNumber="50">Diagnosis.</emphasis>
|
||
<taxonomicName id="EDA083AEFD6EFFB0AE781FA9FDABBB79" authorityName="LeConte" authorityYear="1856" box="[329,533,1791,1816]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="ruginasus">
|
||
<emphasis id="18D4243FFD6EFFB0AE781FA9FDABBB79" box="[329,533,1791,1816]" italics="true" pageId="49" pageNumber="50">Ligyrus ruginasus</emphasis>
|
||
</taxonomicName>
|
||
can be recognized by the triangular clypeus, with 1 central tooth; frontal carina straight, not interrupted (
|
||
<figureCitation id="B29BE4A8FD6EFFB0AE9F1E75FDBDBB5C" box="[430,515,1827,1853]" captionStart="FIGURE 1" captionStartId="81.[151,250,1535,1559]" captionTargetBox="[151,1436,181,1511]" captionTargetId="figure-17@81.[151,1436,181,1511]" captionTargetPageId="81" captionText="FIGURE 1. Head. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) Ligyrus (Ligyrodes) peruvianus, D) L. (Anagrylius) moroni, E) L. (Ligyrus) ruginasus, F) L. (Ligyrus) nasutus, G) L. (Ligyrus) bidentulus, H) Tomarus adoceteus, I) T. maimon, J) T. pullus, K) T. pumilus." figureDoi="http://doi.org/10.5281/zenodo.7345011" httpUri="https://zenodo.org/record/7345011/files/figure.png" pageId="49" pageNumber="50">Fig. 1E</figureCitation>
|
||
); pronotum with prominent tubercle and deep, rounded fovea (
|
||
<figureCitation id="B29BE4A8FD6EFFB0ABF31E75FA95BB5C" box="[1218,1323,1827,1853]" captionStart="FIGURE 26" captionStartId="101.[151,250,1827,1851]" captionTargetBox="[159,1430,189,1791]" captionTargetId="figure-17@101.[151,1436,181,1803]" captionTargetPageId="101" captionText="FIGURE 26. Habitus of the species of Ligyrus. Subgenus Ligyrus: A) Ligyrus fossor, B) L. gibbosus, C) L. nasutus, D) L. neglectus, E) L. paranaensis, F) L. peninsularis, G) L. rubripes, H) L. ruginasus, I) L. villosus. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7345101" httpUri="https://zenodo.org/record/7345101/files/figure.png" pageId="49" pageNumber="50">Fig. 26H</figureCitation>
|
||
); pronotal punctation deep and large; metatibia elongate with apex strongly narrowed; parameres without ventral teeth; apex widely expanded (
|
||
<figureCitation id="B29BE4A8FD6EFFB0AE521E3DFE6EBBE4" box="[355,464,1898,1925]" captionStart="FIGURE 15" captionStartId="90.[151,250,1535,1559]" captionTargetBox="[154,1430,183,1509]" captionTargetId="figure-24@90.[151,1436,181,1512]" captionTargetPageId="90" captionText="FIGURE 15. Parameres in frontal view. A) Euligyrus ebenus, B) E. similis, C) Proculigyrus cicatricosus, D) Ligyrus (Anagrylius) cuniculus, E) L. (Anagrylius) moroni, F) L. (Ligyrodes) peruvianus, G) L. (Ligyrodes) relictus, H) L. (Ligyrodes) sallaei, I) L. (Ligyrus) allonasutus, J) L. (Ligyrus) bidentulus, K) L. (Ligyrus) burmeisteri, L) Ligyrus (Ligyrus) fossor, M) L. (Ligyrus) gibbosus, N) L. (Ligyrus) nasutus, O) L. (Ligyrus) neglectus, P) L. (Ligyrus) paranaensis, Q) L. (Ligyrus) peninsularis, R) L. (Ligyrus) rubripes, S) L. (Ligyrus) ruginasus, T) L. (Ligyrus) villosus." figureDoi="http://doi.org/10.5281/zenodo.7345063" httpUri="https://zenodo.org/record/7345063/files/figure.png" pageId="49" pageNumber="50">Figs. 15S</figureCitation>
|
||
,
|
||
<figureCitation id="B29BE4A8FD6EFFB0AEEA1E3DFDB0BBE4" box="[475,526,1899,1925]" captionStart="FIGURE 18" captionStartId="93.[151,250,1597,1621]" captionTargetBox="[152,1436,184,1567]" captionTargetId="figure-17@93.[151,1436,181,1573]" captionTargetPageId="93" captionText="FIGURE 18. Parameres in lateral view.A) L. (Ligyrus.) allonasutus, B) L. (Ligyrus) bidentulus, C) L. (Ligyrus) burmeisteri, D) Ligyrus (Ligyrus) fossor, E) L. (Ligyrus) gibbosus, F) L. (Ligyrus) nasutus, G) L. (Ligyrus) neglectus, H) L. (Ligyrus) paranaensis, I) L. (Ligyrus) peninsularis, J) L. (Ligyrus) rubripes, K) L. (Ligyrus) ruginasus, L) L. (Ligyrus) villosus." figureDoi="http://doi.org/10.5281/zenodo.7345077" httpUri="https://zenodo.org/record/7345077/files/figure.png" pageId="49" pageNumber="50">18K</figureCitation>
|
||
); internal sac with accessory lamella thick and elongate; coxite of female strongly concave.
|
||
</paragraph>
|
||
<paragraph id="2A1FF82DFD6EFFB0AFF61EE4FB4BBB90" blockId="49.[151,1437,1178,2033]" pageId="49" pageNumber="50">
|
||
<emphasis id="18D4243FFD6EFFB0AFF61EE4FE7EBBAC" bold="true" box="[199,448,1970,1997]" pageId="49" pageNumber="50">Taxonomic remarks.</emphasis>
|
||
The species was originally described as
|
||
<taxonomicName id="EDA083AEFD6EFFB0ACA81EE5FBD9BBAD" authorityName="LeConte" authorityYear="1856" box="[921,1127,1971,1996]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="ruginasus">
|
||
<emphasis id="18D4243FFD6EFFB0ACA81EE5FBD9BBAD" box="[921,1127,1971,1996]" italics="true" pageId="49" pageNumber="50">Ligyrus ruginasus</emphasis>
|
||
</taxonomicName>
|
||
but then transferred in the genus
|
||
<taxonomicName id="EDA083AEFD6EFFB0AFD11E81FE44BB90" authority="(Casey 1915)" baseAuthorityName="Casey" baseAuthorityYear="1915" box="[224,506,2006,2033]" class="Insecta" family="Dynastidae" genus="Oxygrylius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="18D4243FFD6EFFB0AFD11E81FEE5BB90" box="[224,347,2007,2033]" italics="true" pageId="49" pageNumber="50">Oxygrylius</emphasis>
|
||
(
|
||
<bibRefCitation id="4E3185DCFD6EFFB0AE581E81FE4CBB90" author="Casey, T. L." box="[361,498,2006,2033]" pageId="49" pageNumber="50" pagination="1 - 394" refId="ref88969" refString="Casey, T. L. (1915) A review of the American species of Rutelinae, Dynastinae, and Cetoniinae. Memoirs on the Coleoptera, 6, 1 - 394." type="journal article" year="1915">Casey 1915</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. More details are provided in taxonomic remarks of
|
||
<taxonomicName id="EDA083AEFD6EFFB0AB771E81FB50BB90" authorityName="López-García & Deloya" authorityYear="2022" baseAuthorityName="Casey" baseAuthorityYear="1915" box="[1094,1262,2007,2033]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="peninsularis">
|
||
<emphasis id="18D4243FFD6EFFB0AB771E81FB50BB90" box="[1094,1262,2007,2033]" italics="true" pageId="49" pageNumber="50">L. peninsularis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="62BAABA6FD6DFFB3AFF619C0FE30B855" pageId="50" pageNumber="51" type="distribution">
|
||
<paragraph id="2A1FF82DFD6DFFB3AFF619C0FC77BCB5" blockId="50.[151,1437,150,1076]" pageId="50" pageNumber="51">
|
||
<emphasis id="18D4243FFD6DFFB3AFF619C0FEDFBCD1" bold="true" box="[199,353,150,176]" pageId="50" pageNumber="51">Distribution.</emphasis>
|
||
This species inhabits the Sonoran Desert and drier areas of the southwestern
|
||
<collectingCountry id="52B7B8BDFD6DFFB3ABE119C0FAD9BCD1" box="[1232,1383,150,176]" name="United States of America" pageId="50" pageNumber="51">United States</collectingCountry>
|
||
(except most of
|
||
<collectingRegion id="E86436CFFD6DFFB3AE1819ECFE23BCB5" box="[297,413,186,212]" country="United States of America" name="California" pageId="50" pageNumber="51">California</collectingRegion>
|
||
) and northern and western
|
||
<collectingCountry id="52B7B8BDFD6DFFB3ADFC19ECFC9DBCB5" box="[717,803,186,212]" name="Mexico" pageId="50" pageNumber="51">Mexico</collectingCountry>
|
||
(
|
||
<bibRefCitation id="4E3185DCFD6DFFB3AC0019ECFC02BCB5" author="Hardy, A." box="[817,956,186,212]" pageId="50" pageNumber="51" pagination="122 - 127" refId="ref90534" refString="Hardy, A. (1964) Studies on Oxygrylius Casey (Coleoptera: Scarabaeidae: Dynastinae). The Coleopterists Bulletin, 18, 122 - 127." type="journal article" year="1964">Hardy 1964</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="2A1FF82DFD6DFFB3AFF61988FBDFBF8C" blockId="50.[151,1437,150,1076]" pageId="50" pageNumber="51">
|
||
<emphasis id="18D4243FFD6DFFB3AFF61988FE2FBC99" bold="true" box="[199,401,222,248]" pageId="50" pageNumber="51">Locality records.</emphasis>
|
||
<specimenCount id="3CA633A4FD6DFFB3AEA61988FD91BC99" box="[407,559,222,248]" pageId="50" pageNumber="51" type="generic">21 specimens</specimenCount>
|
||
examined from CERPE, FSCA, IEXA, MN/UFRJ, MZUSP, UNSM. Mexican localities from
|
||
<bibRefCitation id="4E3185DCFD6DFFB3AE731854FD8BBD7D" author="Ratcliffe, B. C. & Cave, R. D. & Cano, E. B." box="[322,565,258,284]" pageId="50" pageNumber="51" pagination="1 - 666" refId="ref93157" refString="Ratcliffe, B. C., Cave, R. D. & Cano, E. B. (2013) The dynastine scarab beetles of Mexico, Guatemala, and Belize (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum, 27, 1 - 666." type="journal article" year="2013">
|
||
Ratcliffe
|
||
<emphasis id="18D4243FFD6DFFB3AE9B1855FE5DBD7D" box="[426,483,258,284]" italics="true" pageId="50" pageNumber="51">et al.</emphasis>
|
||
(2013)
|
||
</bibRefCitation>
|
||
and United States of America counties from
|
||
<bibRefCitation id="4E3185DCFD6DFFB3AB001854FAF8BD7D" author="Ratcliffe, B. C. & Cave, R. D." box="[1073,1350,258,284]" pageId="50" pageNumber="51" pagination="1 - 298" refId="ref92923" refString="Ratcliffe, B. C. & Cave, R. D. (2017) The dynastine scarab beetles of the United States and Canada (Coleoptera: Scarabaeidae). Bulletin of the University of Nebraska State Museum, 30, 1 - 298." type="journal article" year="2017">Ratcliffe & Cave (2017)</bibRefCitation>
|
||
.
|
||
<emphasis id="18D4243FFD6DFFB3AA631854FE9BBD21" bold="true" pageId="50" pageNumber="51">MEXICO (1,801)</emphasis>
|
||
.
|
||
<emphasis id="18D4243FFD6DFFB3AE021870FE46BD21" bold="true" box="[307,504,294,320]" pageId="50" pageNumber="51">
|
||
<collectingRegion id="E86436CFFD6DFFB3AE021870FE07BD21" box="[307,441,294,320]" name="Chihuahua" pageId="50" pageNumber="51">Chihuahua</collectingRegion>
|
||
(81):
|
||
</emphasis>
|
||
Chihuahua; Ciudad Camargo; Ciudad Jiménez; Ciudad Juárez; Crell; Nueva Casas Grandes; Ojo del Lucero; San Pedro; Santa Eulalia.
|
||
<emphasis id="18D4243FFD6DFFB3ADEA181CFC39BD05" bold="true" box="[731,903,330,356]" pageId="50" pageNumber="51">
|
||
<collectingRegion id="E86436CFFD6DFFB3ADEA181CFCF6BD05" box="[731,840,330,356]" name="Coahuila" pageId="50" pageNumber="51">Coahuila</collectingRegion>
|
||
(26):
|
||
</emphasis>
|
||
Cuatro Ciénagas; Dolores; Saltillo; San José de la Niña; San Pedro de las Colonias; Torreón.
|
||
<emphasis id="18D4243FFD6DFFB3ADA01838FC84BDE9" bold="true" box="[657,826,366,392]" pageId="50" pageNumber="51">
|
||
<collectingRegion id="E86436CFFD6DFFB3ADA01838FD45BDE9" box="[657,763,366,392]" name="Durango" pageId="50" pageNumber="51">Durango</collectingRegion>
|
||
(85):
|
||
</emphasis>
|
||
Ciudad Lerdo; Cuencamé; El Entroque; Reserva de la Biosfera Mapimí; San Ignacio.
|
||
<emphasis id="18D4243FFD6DFFB3AEC318C4FD4DBDCD" bold="true" box="[498,755,401,428]" pageId="50" pageNumber="51">
|
||
<collectingRegion id="E86436CFFD6DFFB3AEC318C4FD7DBDCD" box="[498,707,401,428]" name="Estado de Mexico" pageId="50" pageNumber="51">Estado de México</collectingRegion>
|
||
(4):
|
||
</emphasis>
|
||
San José Villa de Allende.
|
||
<emphasis id="18D4243FFD6DFFB3AB2F18C4FB12BDCD" bold="true" box="[1054,1196,402,428]" pageId="50" pageNumber="51">
|
||
<collectingRegion id="E86436CFFD6DFFB3AB2F18C4FBD1BDCD" box="[1054,1135,402,428]" name="Jalisco" pageId="50" pageNumber="51">Jalisco</collectingRegion>
|
||
(16):
|
||
</emphasis>
|
||
Ameca; Guadalajara; Hostotipaquillo; Zapopan.
|
||
<emphasis id="18D4243FFD6DFFB3AEF918E0FDDABDB1" bold="true" box="[456,612,438,464]" pageId="50" pageNumber="51">
|
||
<collectingRegion id="E86436CFFD6DFFB3AEF918E0FD9ABDB1" box="[456,548,438,464]" name="Nayarit" pageId="50" pageNumber="51">Nayarit</collectingRegion>
|
||
(73):
|
||
</emphasis>
|
||
Jesús María; Los Sabinos; Pochotitlan; Tepic; Volcán Ceboruco.
|
||
<emphasis id="18D4243FFD6DFFB3AA6118E0FEAABD95" bold="true" pageId="50" pageNumber="51">
|
||
<collectingRegion id="E86436CFFD6DFFB3AA6118E0FF6ABD95" name="Nuevo Leon" pageId="50" pageNumber="51">Nuevo León</collectingRegion>
|
||
(89):
|
||
</emphasis>
|
||
Apodaca; Cañón de la Huasteca; El Álamo; Linares; Monterrey; Punta de la Loma; San Nicolás de los Garza; Parque Ecoturístico Cola de Caballo.
|
||
<emphasis id="18D4243FFD6DFFB3ADBA18A8FCCDBE79" bold="true" box="[651,883,509,536]" pageId="50" pageNumber="51">
|
||
<collectingRegion id="E86436CFFD6DFFB3ADBA18A8FCFCBE79" box="[651,834,509,536]" name="San Luis Potosi" pageId="50" pageNumber="51">San Luís Potosí</collectingRegion>
|
||
(1):
|
||
</emphasis>
|
||
Ciudad Valles.
|
||
<emphasis id="18D4243FFD6DFFB3AB1318A8FB79BE79" bold="true" box="[1058,1223,510,536]" pageId="50" pageNumber="51">
|
||
<collectingRegion id="E86436CFFD6DFFB3AB1318A8FBC5BE79" box="[1058,1147,510,536]" name="Sinaloa" pageId="50" pageNumber="51">Sinaloa</collectingRegion>
|
||
(405):
|
||
</emphasis>
|
||
Choix; Concordia; Culiacán; El Dorado; El Zapotillo; Elota; Escuinapa; Guamúchil; Isla La Chiva; Isla Macapule; Isla Mazocahui; Isla Niscoco; Isla Pájaros II; Isla San Ignacio; Isla Tesobiare; Isla Vinorama; Los Mochis; Mazatlán; Mesa El Carrizal; Río Piaxta; Rosario; San Blas; Tierra Blanca; Vado Hondo; Venados; Villa Unión.
|
||
<emphasis id="18D4243FFD6DFFB3AB771B3CFB55BEE5" bold="true" box="[1094,1259,618,644]" pageId="50" pageNumber="51">
|
||
<collectingRegion id="E86436CFFD6DFFB3AB771B3CFB22BEE5" box="[1094,1180,618,644]" name="Sonora" pageId="50" pageNumber="51">Sonora</collectingRegion>
|
||
(963):
|
||
</emphasis>
|
||
Agua Caliente; Agua Pietra; Álamos; Bahía Kino; Benjamin Hill; Caborca; Cananea; Carbó; Cerro Basura; Cerro Masiaca; Ciudad Obregón; Esperanza; Estación Don; Estación Llano; Estación Luís; Guaymas; Hermosillo; Imuris; Isla Tiburón; La Sandía; Magdalena; Moctezuma; Navajoa; Nogales; Pitiquito; Poza; Puerto Peñasco; Rosario del Tesopaco; Santa Ana; Saric; Sonoita; Tónichi; Valle de Yaqui; Yécora.
|
||
<emphasis id="18D4243FFD6DFFB3ADC61BACFC7FBF75" bold="true" box="[759,961,762,788]" pageId="50" pageNumber="51">
|
||
<collectingRegion id="E86436CFFD6DFFB3ADC61BACFC3CBF75" box="[759,898,762,788]" name="Tamaulipas" pageId="50" pageNumber="51">Tamaulipas</collectingRegion>
|
||
(53):
|
||
</emphasis>
|
||
Ciudad Mante; Ciudad Victoria; Güemez; Llera; Nuevo Laredo; Rancho San Felipe; Santander Jiménez.
|
||
<emphasis id="18D4243FFD6DFFB3AC6A1A48FBAFBF59" bold="true" box="[859,1041,798,824]" pageId="50" pageNumber="51">
|
||
<collectingRegion id="E86436CFFD6DFFB3AC6A1A48FC6FBF59" box="[859,977,798,824]" name="Zacatecas" pageId="50" pageNumber="51">Zacatecas</collectingRegion>
|
||
(10):
|
||
</emphasis>
|
||
Apazol; Jalpa.
|
||
<emphasis id="18D4243FFD6DFFB3ABF11A48FDD0BF3D" bold="true" pageId="50" pageNumber="51">
|
||
<collectingCountry id="52B7B8BDFD6DFFB3ABF11A48FEF6BF3D" name="United States of America" pageId="50" pageNumber="51">UNITED STATES OF AMERICA</collectingCountry>
|
||
(3,807).
|
||
<collectingRegion id="E86436CFFD6DFFB3AE9C1A14FDB3BF3D" box="[429,525,834,860]" country="United States of America" name="Arizona" pageId="50" pageNumber="51">Arizona</collectingRegion>
|
||
(3,017):
|
||
</emphasis>
|
||
Apache; Cochise; Coconino; Gila; Graham; Greenlee; La Paz; Maricopa; Mohave; Pima; Pinal; Santa Cruz; Yavapi; Yuma.
|
||
<emphasis id="18D4243FFD6DFFB3ADE01A30FC25BFE1" bold="true" box="[721,923,870,896]" pageId="50" pageNumber="51">
|
||
<collectingRegion id="E86436CFFD6DFFB3ADE01A30FCF2BFE1" box="[721,844,870,896]" country="United States of America" name="California" pageId="50" pageNumber="51">California</collectingRegion>
|
||
(105):
|
||
</emphasis>
|
||
Imperial; Riverside.
|
||
<emphasis id="18D4243FFD6DFFB3ABBD1A30FA91BFE1" bold="true" box="[1164,1327,870,896]" pageId="50" pageNumber="51">
|
||
<collectingRegion id="E86436CFFD6DFFB3ABBD1A30FB42BFE1" box="[1164,1276,870,896]" country="United States of America" name="Colorado" pageId="50" pageNumber="51">Colorado</collectingRegion>
|
||
(8):
|
||
</emphasis>
|
||
La Plata.
|
||
<emphasis id="18D4243FFD6DFFB3AFA61ADCFE9FBFC5" bold="true" box="[151,289,906,932]" pageId="50" pageNumber="51">
|
||
<collectingRegion id="E86436CFFD6DFFB3AFA61ADCFF50BFC5" box="[151,238,906,932]" country="United States of America" name="Kansas" pageId="50" pageNumber="51">Kansas</collectingRegion>
|
||
(2):
|
||
</emphasis>
|
||
Morton.
|
||
<emphasis id="18D4243FFD6DFFB3AEBC1ADCFDCFBFC5" bold="true" box="[397,625,906,932]" pageId="50" pageNumber="51">
|
||
<collectingRegion id="E86436CFFD6DFFB3AEBC1ADCFD9CBFC5" box="[397,546,906,932]" country="United States of America" name="New Mexico" pageId="50" pageNumber="51">New Mexico</collectingRegion>
|
||
(108):
|
||
</emphasis>
|
||
Doña Ana; Eddy; Grant; Hidalgo; Luna; Otero; Sierra; Socorro.
|
||
<emphasis id="18D4243FFD6DFFB3AA681ADCFF60BFA9" bold="true" pageId="50" pageNumber="51">
|
||
<collectingRegion id="E86436CFFD6DFFB3AA681ADCFA22BFC5" box="[1369,1436,906,932]" country="United States of America" name="Texas" pageId="50" pageNumber="51">Texas</collectingRegion>
|
||
(554):
|
||
</emphasis>
|
||
Bexar; Brewster; Cameron; Culberson; Dimmit; Duval; El Paso; Hidalgo; Hudspeth; Jeff Davis; Kimble; Kleberg; Maverick; Nueces; Presidio; Starr; Terrell; Uvalde; Val Verde; Webb; Zapata.
|
||
</paragraph>
|
||
<paragraph id="2A1FF82DFD6DFFB3AFF61AA0FE30B855" blockId="50.[151,1437,150,1076]" pageId="50" pageNumber="51">
|
||
<emphasis id="18D4243FFD6DFFB3AFF61AA0FE3BB871" bold="true" box="[199,389,1014,1040]" pageId="50" pageNumber="51">Natural history.</emphasis>
|
||
The adults are attracted to lights and found from May to December, but are most abundant in August (
|
||
<bibRefCitation id="4E3185DCFD6DFFB3AFC61D4CFE3FB855" author="Hardy, A." box="[247,385,1050,1076]" pageId="50" pageNumber="51" pagination="122 - 127" refId="ref90534" refString="Hardy, A. (1964) Studies on Oxygrylius Casey (Coleoptera: Scarabaeidae: Dynastinae). The Coleopterists Bulletin, 18, 122 - 127." type="journal article" year="1964">Hardy 1964</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |