155 lines
16 KiB
XML
155 lines
16 KiB
XML
<document id="61AF5AE11E5C9CB7D5643CBD43DFF1CC" ID-CLB-Dataset="202083" ID-DOI="10.11646/zootaxa.5211.1.1" ID-GBIF-Dataset="bb97b9dd-2acc-4244-8f30-72bef07b9ed9" ID-ISSN="1175-5326" ID-Zenodo-Dep="7345007" ID-ZooBank="5754769C-B747-4714-BDD9-7D5509D48BEB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1669101340847" checkinUser="plazi" docAuthor="López-García, Margarita M. & Deloya, Cuauhtémoc" docDate="2022" docId="A209493BFD6DFFB3AFA61DD0FBF2BA27" docLanguage="en" docName="zootaxa.5211.1.1.pdf" docOrigin="Zootaxa 5211 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5211.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Ligyrus (Ligyrus) spinipenis López-García & Deloya 2022, new combination" docType="treatment" docVersion="7" lastPageNumber="51" masterDocId="5E303143FD5FFF81AF311956FFBEBC61" masterDocTitle="Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision" masterLastPageNumber="119" masterPageNumber="1" pageNumber="51" updateTime="1699396337306" updateUser="plazi">
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<mods:mods id="5C3F993FE1161F554F1814D8B7F5A7C5" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="E7E2DDC215AAC206D5B8B351BA931B4A">
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<mods:title id="B8DCA187B4F0C231AFB210602C946990">Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision</mods:title>
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<mods:namePart id="0B697F82D16A8C8B71A1367B3F81D667">López-García, Margarita M.</mods:namePart>
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<mods:nameIdentifier id="D2416775331EBA6A33627741E7629427" type="ORCID">0000-0003-2796-2931</mods:nameIdentifier>
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<mods:nameIdentifier id="5749994E2BC53FFB15447BFD97FB953F" type="email">margaralopezg@gmail.com</mods:nameIdentifier>
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<mods:namePart id="6E6CE46889973B85888AA50022D08F77">Deloya, Cuauhtémoc</mods:namePart>
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<mods:nameIdentifier id="A6F722347781B9545F55F43DD1C36A49" type="ORCID">0000-0002-4774-140X</mods:nameIdentifier>
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<mods:nameIdentifier id="595E52BAF73A2F5FC183000378AD639C" type="email">cuauhtemoc.deloya@inecol.mx</mods:nameIdentifier>
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<mods:typeOfResource id="9F99CCC09C870B9EB3C118D16DAF9FEC">text</mods:typeOfResource>
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<mods:title id="81B5AFA42E9EC39397DEFD582CF3B39D">Zootaxa</mods:title>
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<mods:part id="612390DC36E18602CA32050BB3CD9373">
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<mods:date id="48B378700717290D2B399589D4152865">2022</mods:date>
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<mods:number id="9FC8B95F4D25BF9D86C813312E4DE114">2022-11-22</mods:number>
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<mods:number id="2526525284C7EE1A663816C1E6D0B93F">5211</mods:number>
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<mods:number id="79E11EC47AB8D838C32C790C8B106B8A">1</mods:number>
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<mods:classification id="9289BD0CCD1294F9074EADAEC73F195E">journal article</mods:classification>
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<treatment id="A209493BFD6DFFB3AFA61DD0FBF2BA27" ID-DOI="http://doi.org/10.5281/zenodo.7345081" ID-GBIF-Taxon="205794478" ID-Zenodo-Dep="7345081" LSID="urn:lsid:plazi:treatment:A209493BFD6DFFB3AFA61DD0FBF2BA27" httpUri="http://treatment.plazi.org/id/A209493BFD6DFFB3AFA61DD0FBF2BA27" lastPageNumber="51" pageId="50" pageNumber="51">
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<subSubSection id="62BAABA6FD6DFFB3AFA61DD0FBBAB8C1" box="[151,1028,1158,1184]" pageId="50" pageNumber="51" type="nomenclature">
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<paragraph id="2A1FF82DFD6DFFB3AFA61DD0FBBAB8C1" blockId="50.[151,1028,1158,1184]" box="[151,1028,1158,1184]" pageId="50" pageNumber="51">
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<heading id="71574F41FD6DFFB3AFA61DD0FBBAB8C1" bold="true" box="[151,1028,1158,1184]" fontSize="11" level="2" pageId="50" pageNumber="51" reason="3">
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<emphasis id="18D4243FFD6DFFB3AFA61DD0FBBAB8C1" bold="true" box="[151,1028,1158,1184]" pageId="50" pageNumber="51">
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<taxonomicName id="EDA083AEFD6DFFB3AFA61DD0FC99B8C1" authority="(Neita & Ratcliffe, 2017)" authorityName="López-García & Deloya" authorityYear="2022" baseAuthorityName="Neita & Ratcliffe" baseAuthorityYear="2017" box="[151,807,1158,1184]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="spinipenis" status="comb. nov." subGenus="Ligyrus">
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<emphasis id="18D4243FFD6DFFB3AFA61DD0FF4AB8C1" bold="true" box="[151,244,1158,1184]" italics="true" pageId="50" pageNumber="51">Ligyrus</emphasis>
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(
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<emphasis id="18D4243FFD6DFFB3AE341DD0FEDCB8C1" bold="true" box="[261,354,1158,1184]" italics="true" pageId="50" pageNumber="51">Ligyrus</emphasis>
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)
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<emphasis id="18D4243FFD6DFFB3AE421DD0FE53B8C1" bold="true" box="[371,493,1158,1184]" italics="true" pageId="50" pageNumber="51">spinipenis</emphasis>
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(Neita &
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<bibRefCitation id="4E3185DCFD6DFFB3AD591DD0FCA1B8C1" author="Ratcliffe, B. C. & Cave, R. D." box="[616,799,1158,1184]" pageId="50" pageNumber="51" pagination="1 - 298" refId="ref92923" refString="Ratcliffe, B. C. & Cave, R. D. (2017) The dynastine scarab beetles of the United States and Canada (Coleoptera: Scarabaeidae). Bulletin of the University of Nebraska State Museum, 30, 1 - 298." type="journal article" year="2017">Ratcliffe, 2017</bibRefCitation>
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)
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</taxonomicName>
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<taxonomicNameLabel id="03E79944FD6DFFB3AC1F1DD0FBBAB8C1" box="[814,1028,1158,1184]" pageId="50" pageNumber="51" rank="species">new combination</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="62BAABA6FD6DFFB3AFA61D9DFD4EB960" pageId="50" pageNumber="51" type="reference_group">
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<paragraph id="2A1FF82DFD6DFFB3AFA61D9DFD4EB960" blockId="50.[151,1437,1227,1284]" pageId="50" pageNumber="51">
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<treatmentCitationGroup id="0AB0DF03FD6DFFB3AFA61D9DFD4EB960" pageId="50" pageNumber="51">
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<treatmentCitation id="AB01DE3CFD6DFFB3AFA61D9DFDC2B882" author="Ratcliffe, B. C. & Cave, R. D." box="[151,636,1227,1251]" page="11" pageId="50" pageNumber="51" year="2017">
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<taxonomicName id="EDA083AEFD6DFFB3AFA61D9DFDC2B882" authority="Neita & Ratcliffe, 2017: 11" authorityName="Neita & Ratcliffe" authorityPageNumber="11" authorityYear="2017" box="[151,636,1227,1251]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="spinipenis">
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<emphasis id="18D4243FFD6DFFB3AFA61D9DFEE3B883" box="[151,349,1227,1250]" italics="true" pageId="50" pageNumber="51">Tomarus spinipenis</emphasis>
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Neita &
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<bibRefCitation id="4E3185DCFD6DFFB3AE8A1D9DFDC2B882" author="Ratcliffe, B. C. & Cave, R. D." box="[443,636,1227,1251]" pageId="50" pageNumber="51" pagination="1 - 298" refId="ref92923" refString="Ratcliffe, B. C. & Cave, R. D. (2017) The dynastine scarab beetles of the United States and Canada (Coleoptera: Scarabaeidae). Bulletin of the University of Nebraska State Museum, 30, 1 - 298." type="journal article" year="2017">Ratcliffe, 2017: 11</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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. Original combination.
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<typeStatus id="F51B468FFD6DFFB3AC5C1D9DFC21B882" box="[877,927,1227,1251]" pageId="50" pageNumber="51">Type</typeStatus>
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material at
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<location id="2F7FAEF6FD6DFFB3AB281D9DFECEB965" LSID="urn:lsid:plazi:treatment:A209493BFD6DFFB3AFA61DD0FBF2BA27:2F7FAEF6FD6DFFB3AB281D9DFECEB965" country="Argentina" name="Instituto Argentino de Investigaciones de Zonas Aridas" pageId="50" pageNumber="51" stateProvince="Mendoza">Instituto Argentino de Investigaciones de Zonas Áridas</location>
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,
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<collectingRegion id="E86436CFFD6DFFB3AE4B1DBFFE65B960" box="[378,475,1257,1281]" country="Argentina" name="Mendoza" pageId="50" pageNumber="51">Mendoza</collectingRegion>
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,
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<collectingCountry id="52B7B8BDFD6DFFB3AED21DBFFDF6B960" box="[483,584,1257,1281]" name="Argentina" pageId="50" pageNumber="51">Argentina</collectingCountry>
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(not examined).
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="62BAABA6FD6DFFB3AFA61C79FBF2BA27" pageId="50" pageNumber="51" type="diagnosis">
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<paragraph id="2A1FF82DFD6DFFB3AFA61C79FF45B9D4" blockId="50.[151,1437,1327,1606]" pageId="50" pageNumber="51">
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<emphasis id="18D4243FFD6DFFB3AFA61C79FEACB928" bold="true" box="[151,274,1327,1353]" pageId="50" pageNumber="51">Diagnosis.</emphasis>
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For complete description see
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<bibRefCitation id="4E3185DCFD6DFFB3AD5B1C79FC59B92B" author="Neita-Moreno, J. C. & Ratcliffe, B. C." box="[618,999,1327,1354]" pageId="50" pageNumber="51" pagination="1 - 36" refId="ref91798" refString="Neita-Moreno, J. C. & Ratcliffe, B. C. (2017) The genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) in Argentina, Chile, and Uruguay. Insecta Mundi, 547, 1 - 36." type="journal article" year="2017">Neita-Moreno & Ratcliffe (2017)</bibRefCitation>
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. The species can be recognized by the subtriangular clypeus with 2 apical teeth; frontoclypeal carina incomplete at middle; pronotum without fovea or tubercle; apex of metatibia with about 29 spinules; coxite surface flat; and apical half of parameres with small spinules.
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</paragraph>
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<paragraph id="2A1FF82DFD6DFFB3AFF61CE9FAD9B9BB" blockId="50.[151,1437,1327,1606]" box="[199,1383,1471,1498]" pageId="50" pageNumber="51">
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<emphasis id="18D4243FFD6DFFB3AFF61CE9FE51B9B8" bold="true" box="[199,495,1471,1497]" pageId="50" pageNumber="51">Geographic distribution.</emphasis>
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<taxonomicName id="EDA083AEFD6DFFB3AEC71C96FD7AB9B8" authorityName="López-García & Deloya" authorityYear="2022" baseAuthorityName="Neita & Ratcliffe" baseAuthorityYear="2017" box="[502,708,1472,1497]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="spinipenis">
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<emphasis id="18D4243FFD6DFFB3AEC71C96FD7AB9B8" box="[502,708,1472,1497]" italics="true" pageId="50" pageNumber="51">Ligyrus spinipenis</emphasis>
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</taxonomicName>
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is known from
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<collectingCountry id="52B7B8BDFD6DFFB3AC451CE9FC5DB9B8" box="[884,995,1471,1497]" name="Argentina" pageId="50" pageNumber="51">Argentina</collectingCountry>
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(
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<bibRefCitation id="4E3185DCFD6DFFB3ACC01CE9FAE5B9BB" author="Neita-Moreno, J. C. & Ratcliffe, B. C." box="[1009,1371,1471,1498]" pageId="50" pageNumber="51" pagination="1 - 36" refId="ref91798" refString="Neita-Moreno, J. C. & Ratcliffe, B. C. (2017) The genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) in Argentina, Chile, and Uruguay. Insecta Mundi, 547, 1 - 36." type="journal article" year="2017">Neita-Moreno & Ratcliffe 2017</bibRefCitation>
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).
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</paragraph>
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<paragraph id="2A1FF82DFD6DFFB3AFF61CB5FAC3BA43" blockId="50.[151,1437,1327,1606]" pageId="50" pageNumber="51">
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<emphasis id="18D4243FFD6DFFB3AFF61CB5FE44B99C" bold="true" box="[199,506,1507,1534]" pageId="50" pageNumber="51">
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Locality records (
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<figureCitation id="B29BE4A8FD6DFFB3AEA91CB5FE53B99C" box="[408,493,1507,1533]" captionStart="FIGURE 36" captionStartId="108.[151,250,1989,2013]" captionTargetBox="[219,1377,1115,1961]" captionTargetId="figure-43@108.[211,1377,1115,1965]" captionTargetPageId="108" captionText="FIGURE 36. Distributional records of Ligyrus (Ligyrus) paranaensis, L. spinipenis, and L. burmeisteri." figureDoi="http://doi.org/10.5281/zenodo.7345135" httpUri="https://zenodo.org/record/7345135/files/figure.png" pageId="50" pageNumber="51">Fig. 36</figureCitation>
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).
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</emphasis>
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<materialsCitation id="9AC8F270FD6DFFB3AD331CB5FAC7BA43" ID-GBIF-Occurrence="3986696624" country="Argentina" location="Santiago del Estero" municipality="Parque Nacional Del Copo" pageId="50" pageNumber="51" specimenCount="1" stateProvince="La Rioja">
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Records from
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<bibRefCitation id="4E3185DCFD6DFFB3AD901CB5FBA7B99F" author="Neita-Moreno, J. C. & Ratcliffe, B. C." box="[673,1049,1507,1534]" pageId="50" pageNumber="51" pagination="1 - 36" refId="ref91798" refString="Neita-Moreno, J. C. & Ratcliffe, B. C. (2017) The genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) in Argentina, Chile, and Uruguay. Insecta Mundi, 547, 1 - 36." type="journal article" year="2017">Neita-Moreno & Ratcliffe (2017)</bibRefCitation>
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.
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<emphasis id="18D4243FFD6DFFB3AB151CB5FA22B99C" bold="true" box="[1060,1436,1507,1534]" pageId="50" pageNumber="51">
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<collectingCountry id="52B7B8BDFD6DFFB3AB151CB5FB70B99C" box="[1060,1230,1507,1533]" name="Argentina" pageId="50" pageNumber="51">ARGENTINA</collectingCountry>
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(9).
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<collectingRegion id="E86436CFFD6DFFB3AA321CB2FAD5B99C" box="[1283,1387,1507,1534]" country="Argentina" name="La Rioja" pageId="50" pageNumber="51">La Rioja</collectingRegion>
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(2):
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</emphasis>
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La Rioja.
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<emphasis id="18D4243FFD6DFFB3AE371F51FE1FBA40" bold="true" box="[262,417,1543,1570]" pageId="50" pageNumber="51">
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<collectingRegion id="E86436CFFD6DFFB3AE371F51FECEBA43" box="[262,368,1543,1570]" country="Argentina" name="San Luis" pageId="50" pageNumber="51">San Luís</collectingRegion>
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(1):
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</emphasis>
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San Antonio.
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<emphasis id="18D4243FFD6DFFB3AD701F51FCE7BA40" bold="true" box="[577,857,1543,1570]" pageId="50" pageNumber="51">
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<collectingRegion id="E86436CFFD6DFFB3AD701F51FC96BA43" box="[577,808,1543,1570]" country="Argentina" name="Santiago del Estero" pageId="50" pageNumber="51">Santiago del Estero</collectingRegion>
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(6):
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</emphasis>
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<collectingMunicipality id="CA7B6257FD6DFFB3AC511F5EFB2EBA43" box="[864,1168,1543,1570]" pageId="50" pageNumber="51">Parque Nacional Del Copo</collectingMunicipality>
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;
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<location id="2F7FAEF6FD6DFFB3ABAD1F51FAC7BA43" LSID="urn:lsid:plazi:treatment:A209493BFD6DFFB3AFA61DD0FBF2BA27:2F7FAEF6FD6DFFB3ABAD1F51FAC7BA43" box="[1180,1401,1543,1570]" country="Argentina" municipality="Parque Nacional Del Copo" name="Santiago del Estero" pageId="50" pageNumber="51" stateProvince="La Rioja">Santiago del Estero</location>
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="2A1FF82DFD6DFFB3AFF61F7AFBF2BA27" blockId="50.[151,1437,1327,1606]" box="[199,1100,1579,1606]" pageId="50" pageNumber="51">
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<emphasis id="18D4243FFD6DFFB3AFF61F7AFE3BBA24" bold="true" box="[199,389,1579,1606]" pageId="50" pageNumber="51">Natural history.</emphasis>
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Adults are attracted to lights (
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<bibRefCitation id="4E3185DCFD6DFFB3ADE71F7DFBFEBA27" author="Neita-Moreno, J. C. & Ratcliffe, B. C." box="[726,1088,1579,1606]" pageId="50" pageNumber="51" pagination="1 - 36" refId="ref91798" refString="Neita-Moreno, J. C. & Ratcliffe, B. C. (2017) The genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) in Argentina, Chile, and Uruguay. Insecta Mundi, 547, 1 - 36." type="journal article" year="2017">Neita-Moreno & Ratcliffe 2017</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |