534 lines
60 KiB
XML
534 lines
60 KiB
XML
<document id="CD18E5FEB8CE6B6765A49D7DBBA9DDA2" ID-CLB-Dataset="202083" ID-DOI="10.11646/zootaxa.5211.1.1" ID-GBIF-Dataset="bb97b9dd-2acc-4244-8f30-72bef07b9ed9" ID-ISSN="1175-5326" ID-Zenodo-Dep="7345007" ID-ZooBank="5754769C-B747-4714-BDD9-7D5509D48BEB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1669101340847" checkinUser="plazi" docAuthor="López-García, Margarita M. & Deloya, Cuauhtémoc" docDate="2022" docId="A209493BFD65FFBAAFA61F3FFA24BADC" docLanguage="en" docName="zootaxa.5211.1.1.pdf" docOrigin="Zootaxa 5211 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5211.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Tomarus discrepans" docType="treatment" docVersion="9" lastPageNumber="60" masterDocId="5E303143FD5FFF81AF311956FFBEBC61" masterDocTitle="Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision" masterLastPageNumber="119" masterPageNumber="1" pageNumber="59" updateTime="1699396337306" updateUser="plazi">
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<mods:title id="BD669946C37A22070DA299C1B227CE9B">Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision</mods:title>
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<mods:namePart id="02ECCC0ADC79ECB3098B310D239121CC">López-García, Margarita M.</mods:namePart>
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<mods:nameIdentifier id="8DABAF57109F3134C41BF9A314BB8799" type="email">margaralopezg@gmail.com</mods:nameIdentifier>
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<mods:namePart id="87A2C1EDC23646FA40D4C94F160D6D2A">Deloya, Cuauhtémoc</mods:namePart>
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<mods:nameIdentifier id="ED918C38A420AE0BB3124DA043E79BAD" type="email">cuauhtemoc.deloya@inecol.mx</mods:nameIdentifier>
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<mods:title id="96596DB4D5D4415D65919F187CC4BA3F">Zootaxa</mods:title>
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<mods:part id="2F039F9B8EE38B6AE3223154B6C6CC7F">
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<mods:date id="89E8B407D941CFA4FEEE3CDB621C209D">2022</mods:date>
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<mods:number id="C6C7C7BB02339407BD0124EFCC716019">2022-11-22</mods:number>
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<mods:number id="E2E82869269EED4BCB8614C03074442B">5211</mods:number>
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<mods:url id="BFB7120F17ED6C4752C6E497C864B7F2">http://dx.doi.org/10.11646/zootaxa.5211.1.1</mods:url>
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<mods:classification id="48177738BE915F24880514BF88E14921">journal article</mods:classification>
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<treatment id="A209493BFD65FFBAAFA61F3FFA24BADC" ID-DOI="http://doi.org/10.5281/zenodo.7383775" ID-GBIF-Taxon="205794518" ID-Zenodo-Dep="7383775" LSID="urn:lsid:plazi:treatment:A209493BFD65FFBAAFA61F3FFA24BADC" httpUri="http://treatment.plazi.org/id/A209493BFD65FFBAAFA61F3FFA24BADC" lastPageId="59" lastPageNumber="60" pageId="58" pageNumber="59">
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<subSubSection id="62BAABA6FD65FFBBAFA61F3FFD06BAE3" box="[151,696,1640,1667]" pageId="58" pageNumber="59" type="nomenclature">
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<paragraph id="2A1FF82DFD65FFBBAFA61F3FFD06BAE3" blockId="58.[151,696,1640,1703]" box="[151,696,1640,1667]" pageId="58" pageNumber="59">
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<heading id="71574F41FD65FFBBAFA61F3FFD06BAE3" bold="true" box="[151,696,1640,1667]" fontSize="11" level="2" pageId="58" pageNumber="59" reason="4">
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<taxonomicName id="EDA083AEFD65FFBBAFA61F3FFD06BAE3" authority="(Escalona & Joly, 2006)" baseAuthorityName="Escalona & Joly" baseAuthorityYear="2006" box="[151,696,1640,1667]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="discrepans">
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<emphasis id="18D4243FFD65FFBBAFA61F3FFD06BAE3" bold="true" box="[151,696,1640,1667]" pageId="58" pageNumber="59">
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<emphasis id="18D4243FFD65FFBBAFA61F3FFE34BAE2" bold="true" box="[151,394,1640,1667]" italics="true" pageId="58" pageNumber="59">Tomarus discrepans</emphasis>
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(
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<bibRefCitation id="4E3185DCFD65FFBBAEAA1F3FFD0EBAE3" author="Escalona, H. E. & Joly, L. J." box="[411,688,1640,1667]" pageId="58" pageNumber="59" pagination="111 - 137" refId="ref89956" refString="Escalona, H. E. & Joly, L. J. (2006) El genero Ligyrus Burmeister, 1847 en Venezuela (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini). Boletin Sociedad Entomologica Aragonesa, 39, 111 - 137." type="journal article" year="2006">Escalona & Joly, 2006</bibRefCitation>
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||
)
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||
</emphasis>
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||
</taxonomicName>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="62BAABA6FD65FFBBAFA61FDAFE4DBAC6" box="[151,499,1676,1703]" pageId="58" pageNumber="59" type="description">
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<paragraph id="2A1FF82DFD65FFBBAFA61FDAFE4DBAC6" blockId="58.[151,696,1640,1703]" box="[151,499,1676,1703]" pageId="58" pageNumber="59">
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(
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<figureCitation id="B29BE4A8FD65FFBBAFAE1FDAFEB0BAC7" box="[159,270,1676,1702]" captionStart="FIGURE 16" captionStartId="91.[151,250,1633,1657]" captionTargetBox="[151,1426,189,1602]" captionTargetId="figure-17@91.[151,1436,181,1610]" captionTargetPageId="91" captionText="FIGURE 16. Parameres in frontal view. A) Tomarus adoceteus, B) T. amazonicus, C) T. bituberculatus, D) T. colombianus, E) T. discrepans, F) T. gyas, G) T. laevicollis, H) T. laticaudus, I) T. maimon, J) T. maracaiboensis, K) T. maternus, L) T. pilcopataensis, M) T. pullus, N) T. pumilus, O) T. roigjunenti, P) T. rostratus, Q) T. selanderi, R) T. subtropicus." figureDoi="http://doi.org/10.5281/zenodo.7345065" httpUri="https://zenodo.org/record/7345065/files/figure.png" pageId="58" pageNumber="59">Figs. 16E</figureCitation>
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,
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<figureCitation id="B29BE4A8FD65FFBBAE2B1FDBFEF7BAC6" box="[282,329,1677,1703]" captionStart="FIGURE 19" captionStartId="94.[151,250,1155,1179]" captionTargetBox="[157,1432,188,1128]" captionTargetId="figure-24@94.[152,1436,181,1131]" captionTargetPageId="94" captionText="FIGURE 19. Parameres in lateral view. A) Tomarus adoceteus, B) T. amazonicus, C) T. bituberculatus, D) T. colombianus, E) T. discrepans, F) T. gyas, G) T. laevicollis, H) T. laticaudus, I) T. maimon." figureDoi="http://doi.org/10.5281/zenodo.7345079" httpUri="https://zenodo.org/record/7345079/files/figure.png" pageId="58" pageNumber="59">19E</figureCitation>
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,
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<figureCitation id="B29BE4A8FD65FFBBAE641FDBFE38BAC6" box="[341,390,1677,1703]" captionStart="FIGURE 23" captionStartId="98.[151,250,1728,1752]" captionTargetBox="[161,1421,189,1693]" captionTargetId="figure-24@98.[151,1436,181,1704]" captionTargetPageId="98" captionText="FIGURE 23. Internal sac. A) Tomarus amazonicus, B) T. discrepans, C) T. laticaudus, D) T. maimon. Abbreviations: AL: Accessory lamellae, CL: Copulatory lamella, LSB: Lamellar spiny belt, Ra: Raspula, SD: Sclerotized diverticulum." figureDoi="http://doi.org/10.5281/zenodo.7345091" httpUri="https://zenodo.org/record/7345091/files/figure.png" pageId="58" pageNumber="59">23B</figureCitation>
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,
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||
<figureCitation id="B29BE4A8FD65FFBBAEA31FDBFE7CBAC6" box="[402,450,1677,1703]" captionStart="FIGURE 27" captionStartId="102.[151,250,1868,1892]" captionTargetBox="[158,1409,184,1839]" captionTargetId="figure-24@102.[151,1436,181,1844]" captionTargetPageId="102" captionText="FIGURE 27. Habitus of the species of Tomarus: A) Tomarus adoceteus, B) T. amazonicus, C) T. bituberculatus, D) T. colombianus, E) T. discrepans, F) T. gyas, G) T. laevicollis, H) T. laticaudus, I) T. maimon. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7345103" httpUri="https://zenodo.org/record/7345103/files/figure.png" pageId="58" pageNumber="59">27E</figureCitation>
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;
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<figureCitation id="B29BE4A8FD65FFBBAEFF1FDAFE55BAC6" box="[462,491,1676,1703]" captionStart="FIGURE 45" captionStartId="113.[151,250,1077,1101]" captionTargetBox="[207,1391,181,1048]" captionTargetId="figure-17@113.[197,1391,181,1053]" captionTargetPageId="113" captionText="FIGURE 45. Distributional records of Tomarus discrepans and T. selanderi." figureDoi="http://doi.org/10.5281/zenodo.7345168" httpUri="https://zenodo.org/record/7345168/files/figure.png" pageId="58" pageNumber="59">45</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="62BAABA6FD65FFBBAFA61F84FEB1BB24" pageId="58" pageNumber="59" type="reference_group">
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||
<paragraph id="2A1FF82DFD65FFBBAFA61F84FEB1BB24" blockId="58.[151,1436,1745,1861]" pageId="58" pageNumber="59">
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<treatmentCitationGroup id="0AB0DF03FD65FFBBAFA61F84FEB1BB24" pageId="58" pageNumber="59">
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<treatmentCitation id="AB01DE3CFD65FFBBAFA61F84FD3CBA88" author="Escalona, H. E. & Joly, L. J." box="[151,642,1745,1769]" page="125" pageId="58" pageNumber="59" year="2006">
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<taxonomicName id="EDA083AEFD65FFBBAFA61F84FD3CBA88" authority="Escalona & Joly, 2006: 125" authorityName="Escalona & Joly" authorityPageNumber="125" authorityYear="2006" box="[151,642,1745,1769]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="discrepans">
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<emphasis id="18D4243FFD65FFBBAFA61F84FEE4BA88" box="[151,346,1745,1769]" italics="true" pageId="58" pageNumber="59">Ligyrus discrepans</emphasis>
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<materialsCitation id="9AC8F270FD65FFBBAE501F87FD3CBA88" ID-GBIF-Occurrence="3986696522" box="[353,642,1745,1769]" country="Venezuela" county="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" location="Estacion Experimental San Nicolas" municipality="Maracay" pageId="58" pageNumber="59" specimenCount="1" stateProvince="Portuguesa" typeStatus="holotype">
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<bibRefCitation id="4E3185DCFD65FFBBAE501F87FD3CBA88" author="Escalona, H. E. & Joly, L. J." box="[353,642,1745,1769]" pageId="58" pageNumber="59" pagination="111 - 137" refId="ref89956" refString="Escalona, H. E. & Joly, L. J. (2006) El genero Ligyrus Burmeister, 1847 en Venezuela (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini). Boletin Sociedad Entomologica Aragonesa, 39, 111 - 137." type="journal article" year="2006">Escalona & Joly, 2006: 125</bibRefCitation>
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</materialsCitation>
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</taxonomicName>
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</treatmentCitation>
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||
. Original combination.
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<typeStatus id="F51B468FFD65FFBBAC461F87FC6BBA88" box="[887,981,1745,1769]" pageId="58" pageNumber="59" type="holotype">Holotype</typeStatus>
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male not examined, which according to the original description is at
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<collectingCounty id="C37E80A1FD65FFBBAEF41FA6FB35BB69" box="[453,1163,1776,1800]" pageId="58" pageNumber="59">Museo del Instituto de Zoología Agrícola Francisco Fernández Yépez</collectingCounty>
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(
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<collectingMunicipality id="CA7B6257FD65FFBBABA81FA6FB4ABB69" box="[1177,1268,1776,1800]" pageId="58" pageNumber="59">Maracay</collectingMunicipality>
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,
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||
<collectingCountry id="52B7B8BDFD65FFBBABCF1FA6FAD4BB69" box="[1278,1386,1776,1800]" name="Venezuela" pageId="58" pageNumber="59">Venezuela</collectingCountry>
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), but it was not possible to get in contact with a curator. Type locality:
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<location id="2F7FAEF6FD65FFBBAC451E58FB5CBB47" LSID="urn:lsid:plazi:treatment:A209493BFD65FFBAAFA61F3FFA24BADC:2F7FAEF6FD65FFBBAC451E58FB5CBB47" box="[884,1250,1806,1830]" country="Venezuela" county="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" municipality="Maracay" name="Estacion Experimental San Nicolas" pageId="58" pageNumber="59" stateProvince="Portuguesa">Estación Experimental San Nicolás</location>
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,
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<collectingRegion id="E86436CFFD65FFBBABDF1E59FADFBB46" box="[1262,1377,1807,1831]" country="Venezuela" name="Portuguesa" pageId="58" pageNumber="59">Portuguesa</collectingRegion>
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,
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<collectingCountry id="52B7B8BDFD65FFBBAA5D1E59FEB5BB24" name="Venezuela" pageId="58" pageNumber="59">Venezuela</collectingCountry>
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.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="62BAABA6FD65FFBAAFA61E25FAB8BE5C" lastPageId="59" lastPageNumber="60" pageId="58" pageNumber="59" type="description">
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||
<paragraph id="2A1FF82DFD65FFBAAFA61E25FAB8BE5C" blockId="58.[151,1437,1907,2042]" lastBlockId="59.[151,1437,150,1725]" lastPageId="59" lastPageNumber="60" pageId="58" pageNumber="59">
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<emphasis id="18D4243FFD65FFBBAFA61E25FE97BBEC" bold="true" box="[151,297,1907,1933]" pageId="58" pageNumber="59">Description.</emphasis>
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Habitus as in
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<figureCitation id="B29BE4A8FD65FFBBAEE01E25FD89BBEC" box="[465,567,1907,1933]" captionStart="FIGURE 27" captionStartId="102.[151,250,1868,1892]" captionTargetBox="[158,1409,184,1839]" captionTargetId="figure-24@102.[151,1436,181,1844]" captionTargetPageId="102" captionText="FIGURE 27. Habitus of the species of Tomarus: A) Tomarus adoceteus, B) T. amazonicus, C) T. bituberculatus, D) T. colombianus, E) T. discrepans, F) T. gyas, G) T. laevicollis, H) T. laticaudus, I) T. maimon. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7345103" httpUri="https://zenodo.org/record/7345103/files/figure.png" pageId="58" pageNumber="59">Fig. 27E</figureCitation>
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. Length 21.0–
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||
<quantity id="ED5855C8FD65FFBBADED1E25FCFABBEF" box="[732,836,1907,1934]" metricMagnitude="-2" metricUnit="m" metricValue="2.95" pageId="58" pageNumber="59" unit="mm" value="29.5">29.5 mm</quantity>
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; humeral width 11.0–
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<quantity id="ED5855C8FD65FFBBAB0F1E25FB18BBEF" box="[1086,1190,1907,1934]" metricMagnitude="-2" metricUnit="m" metricValue="1.47" pageId="58" pageNumber="59" unit="mm" value="14.7">14.7 mm</quantity>
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. Color dark brown to black.
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<emphasis id="18D4243FFD65FFBBAFD21EC1FE92BBD0" bold="true" box="[227,300,1943,1969]" pageId="58" pageNumber="59">Head:</emphasis>
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Frons and clypeus coarsely and densely rugose. Frontoclypeal region with 2 conical tubercles separated by about 2.5 tubercle diameters. Clypeus narrowed towards apex, base 3.0–3.5 times as wide as apex. Clypeal teeth triangular, separated by a tooth diameter. Mandible with 2 apical teeth and a lateral, narrowly rounded tooth. Mentum abruptly constricted at apical 3rd. Galea of maxilla with teeth 5 and 6 well developed. Interocular distance 3.5 times an eye width.
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<emphasis id="18D4243FFD64FFBAAE8219EDFD88BCB4" bold="true" box="[435,566,187,213]" pageId="59" pageNumber="60">Pronotum:</emphasis>
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Surface with small punctures, denser and deeper on anterior and posterior angles. Apical tubercle prominent, acute; slightly recurved. Subapical fovea deep, rounded, very wide (as wide as the interocular distance), equal in both sexes. Foveal surface transversely rugose.
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<emphasis id="18D4243FFD64FFBAAB1B1854FB15BD7D" bold="true" box="[1066,1195,258,284]" pageId="59" pageNumber="60">Scutellum:</emphasis>
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Surface with minute, sparse punctures.
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<emphasis id="18D4243FFD64FFBAAE6E1871FE0BBD20" bold="true" box="[351,437,295,321]" pageId="59" pageNumber="60">Elytra:</emphasis>
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Punctures on first interval as large as those on other intervals. Inner surface of apex with rounded, large tubercles forming 25–30 parallel lines.
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<emphasis id="18D4243FFD64FFBAAC37181CFC3ABD05" bold="true" box="[774,900,330,356]" pageId="59" pageNumber="60">Abdomen:</emphasis>
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Apex of tergite IV with a triangular wide area with large, rounded tubercles forming 15–17 diagonal parallel lines. Pygidium with dense rugosity on basal 3rd, apex rounded.
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<emphasis id="18D4243FFD64FFBAAE7318C5FE22BDCC" bold="true" box="[322,412,403,429]" pageId="59" pageNumber="60">Venter:</emphasis>
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Apex of prosternal process flat, longitudinally oval to subrectangular. Metasternum with short, sparse setae.
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<emphasis id="18D4243FFD64FFBAAE4218E1FE0BBDB0" bold="true" box="[371,437,439,465]" pageId="59" pageNumber="60">Legs:</emphasis>
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Protibia tridentate, with a basal small denticle. Apex of metatibia crenulate; with 21–25 spinules.
|
||
<emphasis id="18D4243FFD64FFBAAE30188DFE0BBD95" bold="true" box="[257,437,474,501]" pageId="59" pageNumber="60">Male genitalia:</emphasis>
|
||
Spiculum gastrale without basal plates. Phallobase as long as parameres. Parameres with 1 dorsal tooth each side; tooth long, acute (
|
||
<figureCitation id="B29BE4A8FD64FFBAAD4818A9FD56BE78" box="[633,744,511,537]" captionStart="FIGURE 16" captionStartId="91.[151,250,1633,1657]" captionTargetBox="[151,1426,189,1602]" captionTargetId="figure-17@91.[151,1436,181,1610]" captionTargetPageId="91" captionText="FIGURE 16. Parameres in frontal view. A) Tomarus adoceteus, B) T. amazonicus, C) T. bituberculatus, D) T. colombianus, E) T. discrepans, F) T. gyas, G) T. laevicollis, H) T. laticaudus, I) T. maimon, J) T. maracaiboensis, K) T. maternus, L) T. pilcopataensis, M) T. pullus, N) T. pumilus, O) T. roigjunenti, P) T. rostratus, Q) T. selanderi, R) T. subtropicus." figureDoi="http://doi.org/10.5281/zenodo.7345065" httpUri="https://zenodo.org/record/7345065/files/figure.png" pageId="59" pageNumber="60">Figs. 16E</figureCitation>
|
||
,
|
||
<figureCitation id="B29BE4A8FD64FFBAADC518A9FC9BBE78" box="[756,805,511,537]" captionStart="FIGURE 19" captionStartId="94.[151,250,1155,1179]" captionTargetBox="[157,1432,188,1128]" captionTargetId="figure-24@94.[152,1436,181,1131]" captionTargetPageId="94" captionText="FIGURE 19. Parameres in lateral view. A) Tomarus adoceteus, B) T. amazonicus, C) T. bituberculatus, D) T. colombianus, E) T. discrepans, F) T. gyas, G) T. laevicollis, H) T. laticaudus, I) T. maimon." figureDoi="http://doi.org/10.5281/zenodo.7345079" httpUri="https://zenodo.org/record/7345079/files/figure.png" pageId="59" pageNumber="60">19E</figureCitation>
|
||
). Internal sac with copulatory lamella and short lamellar spiny belt; with a complex of 5 spine-like accessory lamellae, with dense granules at base (
|
||
<figureCitation id="B29BE4A8FD64FFBAABA31B75FB44BE5C" box="[1170,1274,547,573]" captionStart="FIGURE 23" captionStartId="98.[151,250,1728,1752]" captionTargetBox="[161,1421,189,1693]" captionTargetId="figure-24@98.[151,1436,181,1704]" captionTargetPageId="98" captionText="FIGURE 23. Internal sac. A) Tomarus amazonicus, B) T. discrepans, C) T. laticaudus, D) T. maimon. Abbreviations: AL: Accessory lamellae, CL: Copulatory lamella, LSB: Lamellar spiny belt, Ra: Raspula, SD: Sclerotized diverticulum." figureDoi="http://doi.org/10.5281/zenodo.7345091" httpUri="https://zenodo.org/record/7345091/files/figure.png" pageId="59" pageNumber="60">Fig. 23B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="62BAABA6FD64FFBAAFF61B10FD6DBFA8" pageId="59" pageNumber="60" type="diagnosis">
|
||
<paragraph id="2A1FF82DFD64FFBAAFF61B10FEE6BE90" blockId="59.[151,1437,150,1725]" pageId="59" pageNumber="60">
|
||
<emphasis id="18D4243FFD64FFBAAFF61B10FEFCBE01" bold="true" box="[199,322,582,608]" pageId="59" pageNumber="60">Diagnosis.</emphasis>
|
||
<taxonomicName id="EDA083AEFD64FFBAAE781B11FD94BE00" baseAuthorityName="Escalona & Joly" baseAuthorityYear="2006" box="[329,554,583,609]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="discrepans">
|
||
<emphasis id="18D4243FFD64FFBAAE781B11FD94BE00" box="[329,554,583,609]" italics="true" pageId="59" pageNumber="60">Tomarus discrepans</emphasis>
|
||
</taxonomicName>
|
||
is recognized by the following character combination: frontoclypeal tubercles conical; pronotal tubercle acute and prominent (
|
||
<figureCitation id="B29BE4A8FD64FFBAAD9D1B3DFCAFBEE4" box="[684,785,619,645]" captionStart="FIGURE 27" captionStartId="102.[151,250,1868,1892]" captionTargetBox="[158,1409,184,1839]" captionTargetId="figure-24@102.[151,1436,181,1844]" captionTargetPageId="102" captionText="FIGURE 27. Habitus of the species of Tomarus: A) Tomarus adoceteus, B) T. amazonicus, C) T. bituberculatus, D) T. colombianus, E) T. discrepans, F) T. gyas, G) T. laevicollis, H) T. laticaudus, I) T. maimon. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7345103" httpUri="https://zenodo.org/record/7345103/files/figure.png" pageId="59" pageNumber="60">Fig. 27E</figureCitation>
|
||
); pronotal fovea rounded (as wide as interocular distance); metasternum with short setae; protibia tridentate and with an additional basal denticle; apex of metatibia with 21–25 spinules; parameres with a long, acute tooth on each side, apical half strongly contracted, apices widely expanded (
|
||
<figureCitation id="B29BE4A8FD64FFBAAFAE1B81FEB0BE90" box="[159,270,727,753]" captionStart="FIGURE 16" captionStartId="91.[151,250,1633,1657]" captionTargetBox="[151,1426,189,1602]" captionTargetId="figure-17@91.[151,1436,181,1610]" captionTargetPageId="91" captionText="FIGURE 16. Parameres in frontal view. A) Tomarus adoceteus, B) T. amazonicus, C) T. bituberculatus, D) T. colombianus, E) T. discrepans, F) T. gyas, G) T. laevicollis, H) T. laticaudus, I) T. maimon, J) T. maracaiboensis, K) T. maternus, L) T. pilcopataensis, M) T. pullus, N) T. pumilus, O) T. roigjunenti, P) T. rostratus, Q) T. selanderi, R) T. subtropicus." figureDoi="http://doi.org/10.5281/zenodo.7345065" httpUri="https://zenodo.org/record/7345065/files/figure.png" pageId="59" pageNumber="60">Figs. 16E</figureCitation>
|
||
,
|
||
<figureCitation id="B29BE4A8FD64FFBAAE2B1B81FEF5BE90" box="[282,331,727,753]" captionStart="FIGURE 19" captionStartId="94.[151,250,1155,1179]" captionTargetBox="[157,1432,188,1128]" captionTargetId="figure-24@94.[152,1436,181,1131]" captionTargetPageId="94" captionText="FIGURE 19. Parameres in lateral view. A) Tomarus adoceteus, B) T. amazonicus, C) T. bituberculatus, D) T. colombianus, E) T. discrepans, F) T. gyas, G) T. laevicollis, H) T. laticaudus, I) T. maimon." figureDoi="http://doi.org/10.5281/zenodo.7345079" httpUri="https://zenodo.org/record/7345079/files/figure.png" pageId="59" pageNumber="60">19E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="2A1FF82DFD64FFBAAFF61BACFD6DBFA8" blockId="59.[151,1437,150,1725]" pageId="59" pageNumber="60">
|
||
<emphasis id="18D4243FFD64FFBAAFF61BACFE02BF74" bold="true" box="[199,444,762,789]" pageId="59" pageNumber="60">Taxonomic remarks.</emphasis>
|
||
<taxonomicName id="EDA083AEFD64FFBAAEF31BADFD1FBF74" baseAuthorityName="Escalona & Joly" baseAuthorityYear="2006" box="[450,673,763,789]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="discrepans">
|
||
<emphasis id="18D4243FFD64FFBAAEF31BADFD1FBF74" box="[450,673,763,789]" italics="true" pageId="59" pageNumber="60">Tomarus discrepans</emphasis>
|
||
</taxonomicName>
|
||
was described by
|
||
<bibRefCitation id="4E3185DCFD64FFBAAC5A1BADFBCCBF74" author="Escalona, H. E. & Joly, L. J." box="[875,1138,763,789]" pageId="59" pageNumber="60" pagination="111 - 137" refId="ref89956" refString="Escalona, H. E. & Joly, L. J. (2006) El genero Ligyrus Burmeister, 1847 en Venezuela (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini). Boletin Sociedad Entomologica Aragonesa, 39, 111 - 137." type="journal article" year="2006">Escalona & Joly (2006)</bibRefCitation>
|
||
to include specimens from
|
||
<collectingCountry id="52B7B8BDFD64FFBAAFA61A49FEB4BF58" box="[151,266,799,825]" name="Venezuela" pageId="59" pageNumber="60">Venezuela</collectingCountry>
|
||
similar in external morphology to
|
||
<taxonomicName id="EDA083AEFD64FFBAADA01A49FD5EBF59" authorityName="Erichson" authorityYear="1848" box="[657,736,799,824]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="gyas">
|
||
<emphasis id="18D4243FFD64FFBAADA01A49FD5EBF59" box="[657,736,799,824]" italics="true" pageId="59" pageNumber="60">T. gyas</emphasis>
|
||
</taxonomicName>
|
||
but with the characteristic parameres of the male. The authors said that the species was also in
|
||
<collectingCountry id="52B7B8BDFD64FFBAAD311A15FDE7BF3C" box="[512,601,835,861]" name="Panama" pageId="59" pageNumber="60">Panama</collectingCountry>
|
||
and
|
||
<collectingCountry id="52B7B8BDFD64FFBAADBE1A15FCB6BF3C" box="[655,776,835,861]" name="Costa Rica" pageId="59" pageNumber="60">Costa Rica</collectingCountry>
|
||
based on the illustrations of the parameres presented for
|
||
<taxonomicName id="EDA083AEFD64FFBAAAB91A15FF74BFE1" authorityName="Erichson" authorityYear="1848" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="gyas">
|
||
<emphasis id="18D4243FFD64FFBAAAB91A15FF74BFE1" italics="true" pageId="59" pageNumber="60">T. gyas</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<bibRefCitation id="4E3185DCFD64FFBAAFC11A31FE19BFE0" author="Ratcliffe, B. C." box="[240,423,871,897]" pageId="59" pageNumber="60" pagination="1 - 506" refId="ref92614" refString="Ratcliffe, B. C. (2003) The dynastine scarab beetles of Costa Rica and Panama. Bulletin of the University of Nebraska State Museum, 16, 1 - 506." type="journal article" year="2003">Ratcliffe (2003)</bibRefCitation>
|
||
. Although in the present revision the presence of the species is confirmed in the countries from
|
||
<collectingCountry id="52B7B8BDFD64FFBAAFE41ADDFE95BFC4" box="[213,299,907,933]" name="Mexico" pageId="59" pageNumber="60">Mexico</collectingCountry>
|
||
to
|
||
<collectingCountry id="52B7B8BDFD64FFBAAE7E1ADDFE15BFC4" box="[335,427,907,933]" name="Panama" pageId="59" pageNumber="60">Panama</collectingCountry>
|
||
,
|
||
<taxonomicName id="EDA083AEFD64FFBAAE871ADDFDBBBFC5" authorityName="Erichson" authorityYear="1848" box="[438,517,907,932]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="gyas">
|
||
<emphasis id="18D4243FFD64FFBAAE871ADDFDBBBFC5" box="[438,517,907,932]" italics="true" pageId="59" pageNumber="60">T. gyas</emphasis>
|
||
</taxonomicName>
|
||
is also distributed in
|
||
<collectingCountry id="52B7B8BDFD64FFBAADC61ADDFCCFBFC4" box="[759,881,907,933]" name="Costa Rica" pageId="59" pageNumber="60">Costa Rica</collectingCountry>
|
||
and
|
||
<collectingCountry id="52B7B8BDFD64FFBAAC991ADDFBBABFC4" box="[936,1028,907,933]" name="Panama" pageId="59" pageNumber="60">Panama</collectingCountry>
|
||
, and so examination of parameres is necessary for correct identification of both species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="62BAABA6FD64FFBAAFF61A84FA24BADC" pageId="59" pageNumber="60" type="distribution">
|
||
<paragraph id="2A1FF82DFD64FFBAAFF61A84FCB6B870" blockId="59.[151,1437,150,1725]" pageId="59" pageNumber="60">
|
||
<emphasis id="18D4243FFD64FFBAAFF61A84FEDFBF8D" bold="true" box="[199,353,978,1004]" pageId="59" pageNumber="60">Distribution.</emphasis>
|
||
<collectingCountry id="52B7B8BDFD64FFBAAE571A85FE62BF8C" box="[358,476,979,1005]" name="Venezuela" pageId="59" pageNumber="60">Venezuela</collectingCountry>
|
||
,
|
||
<collectingCountry id="52B7B8BDFD64FFBAAED71A85FDFCBF8C" box="[486,578,979,1005]" name="Panama" pageId="59" pageNumber="60">Panama</collectingCountry>
|
||
,
|
||
<collectingCountry id="52B7B8BDFD64FFBAAD7D1A85FD76BF8C" box="[588,712,979,1005]" name="Costa Rica" pageId="59" pageNumber="60">Costa Rica</collectingCountry>
|
||
,
|
||
<collectingCountry id="52B7B8BDFD64FFBAADE31A85FCE8BF8C" box="[722,854,979,1005]" name="El Salvador" pageId="59" pageNumber="60">El Salvador</collectingCountry>
|
||
,
|
||
<collectingCountry id="52B7B8BDFD64FFBAAC501A85FC6FBF8C" box="[865,977,979,1005]" name="Honduras" pageId="59" pageNumber="60">Honduras</collectingCountry>
|
||
,
|
||
<collectingCountry id="52B7B8BDFD64FFBAACEA1A85FBEFBF8C" box="[987,1105,979,1005]" name="Nicaragua" pageId="59" pageNumber="60">Nicaragua</collectingCountry>
|
||
,
|
||
<collectingCountry id="52B7B8BDFD64FFBAAB6A1A85FB69BF8C" box="[1115,1239,979,1005]" name="Guatemala" pageId="59" pageNumber="60">Guatemala</collectingCountry>
|
||
, and
|
||
<collectingCountry id="52B7B8BDFD64FFBAAA211A85FAD7BF8C" box="[1296,1385,979,1005]" name="Mexico" pageId="59" pageNumber="60">Mexico</collectingCountry>
|
||
. It is probably found along the Caribbean coast of
|
||
<collectingCountry id="52B7B8BDFD64FFBAADA01AA1FCBDB870" box="[657,771,1015,1041]" name="Colombia" pageId="59" pageNumber="60">Colombia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="2A1FF82DFD64FFBAAFF61D4CFA23B854" blockId="59.[151,1437,150,1725]" box="[199,1437,1050,1077]" pageId="59" pageNumber="60">
|
||
<emphasis id="18D4243FFD64FFBAAFF61D4CFE45B855" bold="true" box="[199,507,1050,1077]" pageId="59" pageNumber="60">
|
||
Locality records (
|
||
<figureCitation id="B29BE4A8FD64FFBAAEA81D4CFE50B855" box="[409,494,1050,1076]" captionStart="FIGURE 45" captionStartId="113.[151,250,1077,1101]" captionTargetBox="[207,1391,181,1048]" captionTargetId="figure-17@113.[197,1391,181,1053]" captionTargetPageId="113" captionText="FIGURE 45. Distributional records of Tomarus discrepans and T. selanderi." figureDoi="http://doi.org/10.5281/zenodo.7345168" httpUri="https://zenodo.org/record/7345168/files/figure.png" pageId="59" pageNumber="60">Fig. 45</figureCitation>
|
||
).
|
||
</emphasis>
|
||
<specimenCount id="3CA633A4FD64FFBAAD331D4DFD16B854" box="[514,680,1051,1077]" pageId="59" pageNumber="60" type="generic">184 specimens</specimenCount>
|
||
examined from CNIN, EMAC, FSCA, IEXA, UNSM, and USNM.
|
||
</paragraph>
|
||
<paragraph id="2A1FF82DFD64FFBAAFA61D69FEBCBA14" blockId="59.[151,1437,150,1725]" pageId="59" pageNumber="60">
|
||
Some records from
|
||
<bibRefCitation id="4E3185DCFD64FFBAAE4D1D69FD31B838" author="Escalona, H. E. & Joly, L. J." box="[380,655,1087,1113]" pageId="59" pageNumber="60" pagination="111 - 137" refId="ref89956" refString="Escalona, H. E. & Joly, L. J. (2006) El genero Ligyrus Burmeister, 1847 en Venezuela (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini). Boletin Sociedad Entomologica Aragonesa, 39, 111 - 137." type="journal article" year="2006">Escalona & Joly (2006)</bibRefCitation>
|
||
.
|
||
<materialsCitation id="9AC8F270FD64FFBAADAD1D68FF68B81C" ID-GBIF-Occurrence="3986696738" pageId="59" pageNumber="60">
|
||
<emphasis id="18D4243FFD64FFBAADAD1D68FB98B839" bold="true" box="[668,1062,1086,1112]" pageId="59" pageNumber="60">
|
||
<collectingCountry id="52B7B8BDFD64FFBAADAD1D68FCF6B839" box="[668,840,1086,1112]" name="Costa Rica" pageId="59" pageNumber="60">COSTA RICA</collectingCountry>
|
||
(15). Cartago (2):
|
||
</emphasis>
|
||
Turrialba.
|
||
<emphasis id="18D4243FFD64FFBAAB961D68FA97B839" bold="true" box="[1191,1321,1086,1113]" pageId="59" pageNumber="60">Limón (4):</emphasis>
|
||
Hamburg Farm
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="9AC8F270FD64FFBAAFD21D35FE22B81C" ID-GBIF-Occurrence="3986696433" box="[227,412,1123,1149]" pageId="59" pageNumber="60">Salvadora Farm</materialsCitation>
|
||
;
|
||
<materialsCitation id="9AC8F270FD64FFBAAE981D35FDBBB81C" ID-GBIF-Occurrence="3986696848" box="[425,517,1123,1149]" pageId="59" pageNumber="60">Sixaola.</materialsCitation>
|
||
<materialsCitation id="9AC8F270FD64FFBAAD3C1D34FC39B81C" ID-GBIF-Occurrence="3986696422" box="[525,903,1122,1149]" pageId="59" pageNumber="60">
|
||
<emphasis id="18D4243FFD64FFBAAD3C1D34FD14B81D" bold="true" box="[525,682,1122,1149]" pageId="59" pageNumber="60">San José (9):</emphasis>
|
||
<collectingCounty id="C37E80A1FD64FFBAAD851D35FC39B81C" box="[692,903,1123,1149]" pageId="59" pageNumber="60">Estación La Selva</collectingCounty>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="9AC8F270FD64FFBAACA51D35FC4EB81C" ID-GBIF-Occurrence="3986696567" box="[916,1008,1123,1149]" pageId="59" pageNumber="60">Heredia</materialsCitation>
|
||
;
|
||
<materialsCitation id="9AC8F270FD64FFBAACCF1D35FB3BB81C" ID-GBIF-Occurrence="3986696761" box="[1022,1157,1123,1149]" pageId="59" pageNumber="60">Santa Tecla</materialsCitation>
|
||
;
|
||
<materialsCitation id="9AC8F270FD64FFBAABA31D35FB71B81C" ID-GBIF-Occurrence="3986696473" box="[1170,1231,1123,1149]" pageId="59" pageNumber="60">Tena.</materialsCitation>
|
||
<materialsCitation id="9AC8F270FD64FFBAABE91D35FDA8B8C0" ID-GBIF-Occurrence="3986696561" pageId="59" pageNumber="60">
|
||
<emphasis id="18D4243FFD64FFBAABE91D35FE34B8C1" bold="true" pageId="59" pageNumber="60">
|
||
<collectingCountry id="52B7B8BDFD64FFBAABE91D35FA22B81D" box="[1240,1436,1122,1149]" name="El Salvador" pageId="59" pageNumber="60">EL SALVADOR</collectingCountry>
|
||
(7). La Libertad (2):
|
||
</emphasis>
|
||
San Andrés
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="9AC8F270FD64FFBAAD131DD1FD15B8C0" ID-GBIF-Occurrence="3986696844" box="[546,683,1159,1185]" pageId="59" pageNumber="60">Santa Tecla.</materialsCitation>
|
||
<materialsCitation id="9AC8F270FD64FFBAAD831DD0FB98B8C0" ID-GBIF-Occurrence="3986696444" box="[690,1062,1158,1185]" pageId="59" pageNumber="60">
|
||
<emphasis id="18D4243FFD64FFBAAD831DD0FC3DB8C1" bold="true" box="[690,899,1158,1184]" pageId="59" pageNumber="60">San Salvador (3):</emphasis>
|
||
San Salvador.
|
||
</materialsCitation>
|
||
<materialsCitation id="9AC8F270FD64FFBAAB1C1DD0FAE3B8C0" ID-GBIF-Occurrence="3986696401" box="[1069,1373,1158,1185]" pageId="59" pageNumber="60">
|
||
<emphasis id="18D4243FFD64FFBAAB1C1DD0FB65B8C1" bold="true" box="[1069,1243,1158,1184]" pageId="59" pageNumber="60">Santa Ana (1):</emphasis>
|
||
Santa Ana.
|
||
</materialsCitation>
|
||
<materialsCitation id="9AC8F270FD64FFBAAA551DD1FEC5B8A4" ID-GBIF-Occurrence="3986696551" pageId="59" pageNumber="60">
|
||
<emphasis id="18D4243FFD64FFBAAA551DD1FEB9B8A5" bold="true" pageId="59" pageNumber="60">Usulután (1):</emphasis>
|
||
Usulután.
|
||
</materialsCitation>
|
||
<materialsCitation id="9AC8F270FD64FFBAAEB31DFCFC1BB8A4" ID-GBIF-Occurrence="3986697021" box="[386,933,1194,1221]" pageId="59" pageNumber="60">
|
||
<emphasis id="18D4243FFD64FFBAAEB31DFCFCF4B8A5" bold="true" box="[386,842,1194,1221]" pageId="59" pageNumber="60">
|
||
<collectingCountry id="52B7B8BDFD64FFBAAEB31DFCFD89B8A5" box="[386,567,1194,1220]" name="Guatemala" pageId="59" pageNumber="60">GUATEMALA</collectingCountry>
|
||
(48). Alta Verapaz (6):
|
||
</emphasis>
|
||
Chulac
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="9AC8F270FD64FFBAAC811DFDFBADB8A4" ID-GBIF-Occurrence="3986696340" box="[944,1043,1195,1221]" pageId="59" pageNumber="60">
|
||
<collectorName id="87559DFBFD64FFBAAC811DFDFBADB8A4" box="[944,1043,1195,1221]" pageId="59" pageNumber="60">La Tinta</collectorName>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="9AC8F270FD64FFBAAB2F1DFDFBCFB8A4" ID-GBIF-Occurrence="3986696607" box="[1054,1137,1195,1221]" pageId="59" pageNumber="60">Panzos</materialsCitation>
|
||
;
|
||
<materialsCitation id="9AC8F270FD64FFBAAB4C1DFDFB69B8A4" ID-GBIF-Occurrence="3986696727" box="[1149,1239,1195,1221]" pageId="59" pageNumber="60">Senahú.</materialsCitation>
|
||
<materialsCitation id="9AC8F270FD64FFBAABEF1DFDFEBCB888" ID-GBIF-Occurrence="3986696876" pageId="59" pageNumber="60">
|
||
<emphasis id="18D4243FFD64FFBAABEF1DFDFA23B8A5" bold="true" box="[1246,1437,1194,1221]" pageId="59" pageNumber="60">El Progreso (1):</emphasis>
|
||
<collectorName id="87559DFBFD64FFBAAFA61D99FEBCB888" box="[151,258,1231,1257]" pageId="59" pageNumber="60">El Jicaro.</collectorName>
|
||
</materialsCitation>
|
||
<materialsCitation id="9AC8F270FD64FFBAAE381D99FDCCB888" ID-GBIF-Occurrence="3986696635" box="[265,626,1230,1257]" pageId="59" pageNumber="60">
|
||
<emphasis id="18D4243FFD64FFBAAE381D99FE2DB889" bold="true" box="[265,403,1230,1257]" pageId="59" pageNumber="60">Izabal (35):</emphasis>
|
||
Cerro Negro Norte
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="9AC8F270FD64FFBAAD4F1D99FC9DB888" ID-GBIF-Occurrence="3986696687" box="[638,803,1231,1257]" pageId="59" pageNumber="60">Finca Firmeza</materialsCitation>
|
||
;
|
||
<materialsCitation id="9AC8F270FD64FFBAAC1E1D99FC2CB888" ID-GBIF-Occurrence="3986696773" box="[815,914,1231,1257]" pageId="59" pageNumber="60">Morales.</materialsCitation>
|
||
<materialsCitation id="9AC8F270FD64FFBAACA81D99FB68B888" ID-GBIF-Occurrence="3986696692" box="[921,1238,1230,1257]" pageId="59" pageNumber="60">
|
||
<emphasis id="18D4243FFD64FFBAACA81D99FBF0B889" bold="true" box="[921,1102,1230,1257]" pageId="59" pageNumber="60">Retalhuleu (1):</emphasis>
|
||
Retalhuleu.
|
||
</materialsCitation>
|
||
<materialsCitation id="9AC8F270FD64FFBAABED1D98FEE9B96C" ID-GBIF-Occurrence="3986696606" pageId="59" pageNumber="60">
|
||
<emphasis id="18D4243FFD64FFBAABED1D98FA23B889" bold="true" box="[1244,1437,1230,1257]" pageId="59" pageNumber="60">San Marcos (1):</emphasis>
|
||
Puente Talismán.
|
||
</materialsCitation>
|
||
<materialsCitation id="9AC8F270FD64FFBAAE6C1DA4FD3CB96C" ID-GBIF-Occurrence="3986696458" box="[349,642,1266,1293]" pageId="59" pageNumber="60">
|
||
<emphasis id="18D4243FFD64FFBAAE6C1DA4FDADB96D" bold="true" box="[349,531,1266,1293]" pageId="59" pageNumber="60">Santa Rosa (1):</emphasis>
|
||
Mal País.
|
||
</materialsCitation>
|
||
<materialsCitation id="9AC8F270FD64FFBAADB91DA4FBB2B96C" ID-GBIF-Occurrence="3986696483" box="[648,1036,1266,1293]" pageId="59" pageNumber="60">
|
||
<emphasis id="18D4243FFD64FFBAADB91DA4FCDBB96D" bold="true" box="[648,869,1266,1293]" pageId="59" pageNumber="60">Suchitepéquez (3):</emphasis>
|
||
Santa Bárbara
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="9AC8F270FD64FFBAAB271DA5FB0AB96C" ID-GBIF-Occurrence="3986696760" box="[1046,1204,1267,1293]" pageId="59" pageNumber="60">Finca Parraxe</materialsCitation>
|
||
;
|
||
<materialsCitation id="9AC8F270FD64FFBAAB8F1DA5FF53B950" ID-GBIF-Occurrence="3986696316" pageId="59" pageNumber="60">Finca Moca, Volcán Atitlán.</materialsCitation>
|
||
<materialsCitation id="9AC8F270FD64FFBAAFC31C40FD51B950" ID-GBIF-Occurrence="3986696949" box="[242,751,1302,1329]" pageId="59" pageNumber="60">
|
||
<emphasis id="18D4243FFD64FFBAAFC31C40FD3FB951" bold="true" box="[242,641,1302,1329]" pageId="59" pageNumber="60">
|
||
<collectingCountry id="52B7B8BDFD64FFBAAFC31C40FE2CB951" box="[242,402,1302,1328]" name="Honduras" pageId="59" pageNumber="60">HONDURAS</collectingCountry>
|
||
(62). Atlántida (31):
|
||
</emphasis>
|
||
<collectorName id="87559DFBFD64FFBAADB61C41FD51B950" box="[647,751,1303,1329]" pageId="59" pageNumber="60">La Ceiba</collectorName>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="9AC8F270FD64FFBAADC81C41FC92B950" ID-GBIF-Occurrence="3986696350" box="[761,812,1303,1329]" pageId="59" pageNumber="60">Tela</materialsCitation>
|
||
;
|
||
<materialsCitation id="9AC8F270FD64FFBAAC071C41FB1CB950" ID-GBIF-Occurrence="3986696490" box="[822,1186,1303,1329]" pageId="59" pageNumber="60">Parque Nacional Cuero y Salado</materialsCitation>
|
||
;
|
||
<materialsCitation id="9AC8F270FD64FFBAAB9D1C41FF55B934" ID-GBIF-Occurrence="3986696918" pageId="59" pageNumber="60">Parque Nacional Pico Bonito.</materialsCitation>
|
||
<materialsCitation id="9AC8F270FD64FFBAAFC31C6CFDE4B934" ID-GBIF-Occurrence="3986696999" box="[242,602,1338,1365]" pageId="59" pageNumber="60">
|
||
<emphasis id="18D4243FFD64FFBAAFC31C6CFE0AB935" bold="true" box="[242,436,1338,1364]" pageId="59" pageNumber="60">Comayagua (1):</emphasis>
|
||
Siguatepeque.
|
||
</materialsCitation>
|
||
<materialsCitation id="9AC8F270FD64FFBAAD501C6CFCDCB934" ID-GBIF-Occurrence="3986696465" box="[609,866,1338,1365]" pageId="59" pageNumber="60">
|
||
<emphasis id="18D4243FFD64FFBAAD501C6CFD5DB935" bold="true" box="[609,739,1338,1365]" pageId="59" pageNumber="60">Copán (1):</emphasis>
|
||
Lancetilla.
|
||
</materialsCitation>
|
||
<materialsCitation id="9AC8F270FD64FFBAAC5B1C6CFBE6B934" ID-GBIF-Occurrence="3986696323" box="[874,1112,1338,1365]" pageId="59" pageNumber="60">
|
||
<emphasis id="18D4243FFD64FFBAAC5B1C6CFC55B935" bold="true" box="[874,1003,1338,1365]" pageId="59" pageNumber="60">Cortés (8):</emphasis>
|
||
<collectorName id="87559DFBFD64FFBAACC21C6DFBE6B934" box="[1011,1112,1339,1365]" pageId="59" pageNumber="60">La Lima</collectorName>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="9AC8F270FD64FFBAAB521C6DFAA6B934" ID-GBIF-Occurrence="3986696356" box="[1123,1304,1339,1365]" pageId="59" pageNumber="60">San Pedro Sula.</materialsCitation>
|
||
<materialsCitation id="9AC8F270FD64FFBAAA111C6DFE2DB918" ID-GBIF-Occurrence="3986696813" pageId="59" pageNumber="60">
|
||
<emphasis id="18D4243FFD64FFBAAA111C6DFF7FB919" bold="true" pageId="59" pageNumber="60">El Paraiso (2):</emphasis>
|
||
Capiré (
|
||
<quantity id="ED5855C8FD64FFBAAE131C09FEE5B918" box="[290,347,1375,1401]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="59" pageNumber="60" unit="km" value="8.0">8 km</quantity>
|
||
<collectionCode id="4CB160E8FD64FFBAAE531C09FE39B918" box="[354,391,1375,1401]" pageId="59" pageNumber="60">SE</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
<materialsCitation id="9AC8F270FD64FFBAAEA81C08FCD9B918" ID-GBIF-Occurrence="3986696440" box="[409,871,1374,1401]" pageId="59" pageNumber="60">
|
||
<emphasis id="18D4243FFD64FFBAAEA81C08FDC6B919" bold="true" box="[409,632,1374,1401]" pageId="59" pageNumber="60">Gracias a Dios (3):</emphasis>
|
||
<collectingMunicipality id="CA7B6257FD64FFBAAD4F1C09FCD9B918" box="[638,871,1375,1401]" pageId="59" pageNumber="60">
|
||
Reserva
|
||
<location id="2F7FAEF6FD64FFBAADEE1C09FCD9B918" LSID="urn:lsid:plazi:treatment:A209493BFD65FFBAAFA61F3FFA24BADC:2F7FAEF6FD64FFBAADEE1C09FCD9B918" box="[735,871,1375,1401]" country="Costa Rica" county="Estacion La Selva" municipality="Reserva Rio Platano" name="Rio Platano" pageId="59" pageNumber="60" stateProvince="Carabobo">Río Plátano</location>
|
||
</collectingMunicipality>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="9AC8F270FD64FFBAAC401C09FBBFB918" ID-GBIF-Occurrence="3986697047" box="[881,1025,1375,1401]" pageId="59" pageNumber="60">Wampusirpi.</materialsCitation>
|
||
<materialsCitation id="9AC8F270FD64FFBAAB371C08FA90B918" ID-GBIF-Occurrence="3986696816" box="[1030,1326,1374,1401]" pageId="59" pageNumber="60">
|
||
<emphasis id="18D4243FFD64FFBAAB371C08FB14B919" bold="true" box="[1030,1194,1374,1400]" pageId="59" pageNumber="60">Olancho (10):</emphasis>
|
||
<location id="2F7FAEF6FD64FFBAAB801C09FA90B918" LSID="urn:lsid:plazi:treatment:A209493BFD65FFBAAFA61F3FFA24BADC:2F7FAEF6FD64FFBAAB801C09FA90B918" box="[1201,1326,1375,1401]" country="Costa Rica" county="Estacion La Selva" municipality="Reserva Rio Platano" name="Catamacas" pageId="59" pageNumber="60" stateProvince="Carabobo">Catamacas</location>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="9AC8F270FD64FFBAAA081C09FE97B9FC" ID-GBIF-Occurrence="3986696709" pageId="59" pageNumber="60">
|
||
<location id="2F7FAEF6FD64FFBAAA081C09FE97B9FC" LSID="urn:lsid:plazi:treatment:A209493BFD65FFBAAFA61F3FFA24BADC:2F7FAEF6FD64FFBAAA081C09FE97B9FC" country="Costa Rica" county="Estacion La Selva" municipality="Reserva Rio Platano" name="Montana de Melacate." pageId="59" pageNumber="60" stateProvince="Carabobo">Montaña de Melacate.</location>
|
||
</materialsCitation>
|
||
<materialsCitation id="9AC8F270FD64FFBAAE031CD5FDB0B9FC" ID-GBIF-Occurrence="3986696894" box="[306,526,1410,1437]" pageId="59" pageNumber="60">
|
||
<emphasis id="18D4243FFD64FFBAAE031CD5FE20B9FD" bold="true" box="[306,414,1410,1437]" pageId="59" pageNumber="60">Yoro (6):</emphasis>
|
||
<location id="2F7FAEF6FD64FFBAAE991CD5FDB0B9FC" LSID="urn:lsid:plazi:treatment:A209493BFD65FFBAAFA61F3FFA24BADC:2F7FAEF6FD64FFBAAE991CD5FDB0B9FC" box="[424,526,1411,1437]" country="Costa Rica" county="Estacion La Selva" municipality="Reserva Rio Platano" name="Cuevitas" pageId="59" pageNumber="60" stateProvince="Carabobo">Cuevitas</location>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="9AC8F270FD64FFBAAD2A1CD5FCE7B9FC" ID-GBIF-Occurrence="3986696588" box="[539,857,1411,1437]" pageId="59" pageNumber="60">Parque Nacional Pico Pijol.</materialsCitation>
|
||
<materialsCitation id="9AC8F270FD64FFBAAC501CD4FAD8B9FC" ID-GBIF-Occurrence="3986696950" box="[865,1382,1410,1437]" pageId="59" pageNumber="60">
|
||
<emphasis id="18D4243FFD64FFBAAC501CD4FB19B9FD" bold="true" box="[865,1191,1410,1436]" pageId="59" pageNumber="60">
|
||
<collectingCountry id="52B7B8BDFD64FFBAAC501CD4FC69B9FD" box="[865,983,1410,1436]" name="Mexico" pageId="59" pageNumber="60">MEXICO</collectingCountry>
|
||
(9). Chiapas (8):
|
||
</emphasis>
|
||
<location id="2F7FAEF6FD64FFBAAB801CD5FAD8B9FC" LSID="urn:lsid:plazi:treatment:A209493BFD65FFBAAFA61F3FFA24BADC:2F7FAEF6FD64FFBAAB801CD5FAD8B9FC" box="[1201,1382,1411,1437]" country="Costa Rica" county="Estacion La Selva" municipality="Reserva Rio Platano" name="Boca de Chajul" pageId="59" pageNumber="60" stateProvince="Carabobo">
|
||
Boca
|
||
<collectorName id="87559DFBFD64FFBAABC51CD5FAD8B9FC" box="[1268,1382,1411,1437]" pageId="59" pageNumber="60">de Chajul</collectorName>
|
||
</location>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="9AC8F270FD64FFBAAA451CD5FE94B9A0" ID-GBIF-Occurrence="3986696436" pageId="59" pageNumber="60">
|
||
<location id="2F7FAEF6FD64FFBAAA451CD5FE94B9A0" LSID="urn:lsid:plazi:treatment:A209493BFD65FFBAAFA61F3FFA24BADC:2F7FAEF6FD64FFBAAA451CD5FE94B9A0" country="Costa Rica" county="Estacion La Selva" municipality="Reserva Rio Platano" name="Las Guacamayas" pageId="59" pageNumber="60" stateProvince="Carabobo">Las Guacamayas</location>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="9AC8F270FD64FFBAAE071CF1FE17B9A0" ID-GBIF-Occurrence="3986696312" box="[310,425,1447,1473]" pageId="59" pageNumber="60">Ocosingo.</materialsCitation>
|
||
<materialsCitation id="9AC8F270FD64FFBAAE801CF1FD0AB9A0" ID-GBIF-Occurrence="3986696543" box="[433,692,1446,1473]" pageId="59" pageNumber="60">
|
||
<emphasis id="18D4243FFD64FFBAAE801CF1FDEFB9A1" bold="true" box="[433,593,1446,1473]" pageId="59" pageNumber="60">Veracruz (1):</emphasis>
|
||
<collectorName id="87559DFBFD64FFBAAD6B1CF1FD0AB9A0" box="[602,692,1447,1473]" pageId="59" pageNumber="60">El Plan.</collectorName>
|
||
</materialsCitation>
|
||
<materialsCitation id="9AC8F270FD64FFBAAD8D1CF0FB4EB9A0" ID-GBIF-Occurrence="3986696554" box="[700,1264,1446,1473]" pageId="59" pageNumber="60">
|
||
<emphasis id="18D4243FFD64FFBAAD8D1CF0FBFEB9A1" bold="true" box="[700,1088,1446,1472]" pageId="59" pageNumber="60">
|
||
<collectingCountry id="52B7B8BDFD64FFBAAD8D1CF0FCD7B9A1" box="[700,873,1446,1472]" name="Nicaragua" pageId="59" pageNumber="60">NICARAGUA</collectingCountry>
|
||
(7). Granada (1):
|
||
</emphasis>
|
||
<collectorName id="87559DFBFD64FFBAAB791CF1FB01B9A0" box="[1096,1215,1447,1473]" pageId="59" pageNumber="60">La Calera.</collectorName>
|
||
<emphasis id="18D4243FFD64FFBAABF71CF0FB4EB9A0" bold="true" box="[1222,1264,1446,1473]" pageId="59" pageNumber="60">Río</emphasis>
|
||
</materialsCitation>
|
||
<materialsCitation id="9AC8F270FD64FFBAABC81CF0FF53B984" ID-GBIF-Occurrence="3986696780" pageId="59" pageNumber="60">
|
||
<emphasis id="18D4243FFD64FFBAABC81CF0FA22B9A1" bold="true" box="[1273,1436,1446,1473]" pageId="59" pageNumber="60">San Juan (4):</emphasis>
|
||
<location id="2F7FAEF6FD64FFBAAFA61C9DFF53B984" LSID="urn:lsid:plazi:treatment:A209493BFD65FFBAAFA61F3FFA24BADC:2F7FAEF6FD64FFBAAFA61C9DFF53B984" box="[151,237,1483,1509]" country="Costa Rica" county="Estacion La Selva" municipality="Reserva Rio Platano" name="Bartola" pageId="59" pageNumber="60" stateProvince="Carabobo">Bartola</location>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="9AC8F270FD64FFBAAFCB1C9DFE21B984" ID-GBIF-Occurrence="3986696707" box="[250,415,1483,1509]" pageId="59" pageNumber="60">
|
||
<location id="2F7FAEF6FD64FFBAAFCB1C9DFE21B984" LSID="urn:lsid:plazi:treatment:A209493BFD65FFBAAFA61F3FFA24BADC:2F7FAEF6FD64FFBAAFCB1C9DFE21B984" box="[250,415,1483,1509]" country="Costa Rica" county="Estacion La Selva" municipality="Reserva Rio Platano" name="Rio Papaturro." pageId="59" pageNumber="60" stateProvince="Carabobo">Río Papaturro.</location>
|
||
</materialsCitation>
|
||
<materialsCitation id="9AC8F270FD64FFBAAE971C9DFD29B984" ID-GBIF-Occurrence="3986696834" box="[422,663,1482,1509]" pageId="59" pageNumber="60">
|
||
<emphasis id="18D4243FFD64FFBAAE971C9DFD97B985" bold="true" box="[422,553,1482,1509]" pageId="59" pageNumber="60">Zelaya (2):</emphasis>
|
||
<location id="2F7FAEF6FD64FFBAAD001C9DFD29B984" LSID="urn:lsid:plazi:treatment:A209493BFD65FFBAAFA61F3FFA24BADC:2F7FAEF6FD64FFBAAD001C9DFD29B984" box="[561,663,1483,1509]" country="Costa Rica" county="Estacion La Selva" municipality="Reserva Rio Platano" name="Bonanza" pageId="59" pageNumber="60" stateProvince="Carabobo">Bonanza</location>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="9AC8F270FD64FFBAAD921C9DFCF6B984" ID-GBIF-Occurrence="3986696582" box="[675,840,1483,1509]" pageId="59" pageNumber="60">Cerro Saslaya.</materialsCitation>
|
||
<materialsCitation id="9AC8F270FD64FFBAAC611C9CFA8DB984" ID-GBIF-Occurrence="3986696911" box="[848,1331,1482,1509]" pageId="59" pageNumber="60">
|
||
<emphasis id="18D4243FFD64FFBAAC611C9CFB36B985" bold="true" box="[848,1160,1482,1509]" pageId="59" pageNumber="60">
|
||
<collectingCountry id="52B7B8BDFD64FFBAAC611C9CFC77B985" box="[848,969,1482,1508]" name="Panama" pageId="59" pageNumber="60">PANAMA</collectingCountry>
|
||
(28). Colón (3):
|
||
</emphasis>
|
||
Madden Dam.
|
||
</materialsCitation>
|
||
<materialsCitation id="9AC8F270FD64FFBAAA0B1C9CFEC6BA68" ID-GBIF-Occurrence="3986696672" pageId="59" pageNumber="60">
|
||
<emphasis id="18D4243FFD64FFBAAA0B1C9CFF71BA69" bold="true" pageId="59" pageNumber="60">Panamá (24):</emphasis>
|
||
<location id="2F7FAEF6FD64FFBAAFE71CB9FEC6BA68" LSID="urn:lsid:plazi:treatment:A209493BFD65FFBAAFA61F3FFA24BADC:2F7FAEF6FD64FFBAAFE71CB9FEC6BA68" box="[214,376,1519,1545]" country="Costa Rica" county="Estacion La Selva" municipality="Reserva Rio Platano" name="Altos de Maje" pageId="59" pageNumber="60" stateProvince="Carabobo">
|
||
Altos
|
||
<collectorName id="87559DFBFD64FFBAAE2B1CB9FEC6BA68" box="[282,376,1519,1545]" pageId="59" pageNumber="60">de Maje</collectorName>
|
||
</location>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="9AC8F270FD64FFBAAEB21CB9FD88BA68" ID-GBIF-Occurrence="3986696619" box="[387,566,1519,1545]" pageId="59" pageNumber="60">
|
||
<location id="2F7FAEF6FD64FFBAAEB21CB9FD88BA68" LSID="urn:lsid:plazi:treatment:A209493BFD65FFBAAFA61F3FFA24BADC:2F7FAEF6FD64FFBAAEB21CB9FD88BA68" box="[387,566,1519,1545]" country="Costa Rica" county="Estacion La Selva" municipality="Reserva Rio Platano" name="Barro Colorado" pageId="59" pageNumber="60" stateProvince="Carabobo">Barro Colorado</location>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="9AC8F270FD64FFBAAD701CB9FD72BA68" ID-GBIF-Occurrence="3986696674" box="[577,716,1519,1545]" pageId="59" pageNumber="60">
|
||
<collectorName id="87559DFBFD64FFBAAD701CB9FD72BA68" box="[577,716,1519,1545]" pageId="59" pageNumber="60">
|
||
<location id="2F7FAEF6FD64FFBAAD701CB9FD72BA68" LSID="urn:lsid:plazi:treatment:A209493BFD65FFBAAFA61F3FFA24BADC:2F7FAEF6FD64FFBAAD701CB9FD72BA68" box="[577,716,1519,1545]" country="Costa Rica" county="Estacion La Selva" municipality="Reserva Rio Platano" name="La Chorrera" pageId="59" pageNumber="60" stateProvince="Carabobo">La Chorrera</location>
|
||
</collectorName>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="9AC8F270FD64FFBAADE61CB9FC1EBA68" ID-GBIF-Occurrence="3986696540" box="[727,928,1519,1545]" pageId="59" pageNumber="60">
|
||
<location id="2F7FAEF6FD64FFBAADE61CB9FC1EBA68" LSID="urn:lsid:plazi:treatment:A209493BFD65FFBAAFA61F3FFA24BADC:2F7FAEF6FD64FFBAADE61CB9FC1EBA68" box="[727,928,1519,1545]" country="Costa Rica" county="Estacion La Selva" municipality="Reserva Rio Platano" name="Llano-Carti Road" pageId="59" pageNumber="60" stateProvince="Carabobo">Llano-Carti Road</location>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="9AC8F270FD64FFBAAC9A1CB9FBB5BA68" ID-GBIF-Occurrence="3986696627" box="[939,1035,1519,1545]" pageId="59" pageNumber="60">Panamá.</materialsCitation>
|
||
<materialsCitation id="9AC8F270FD64FFBAAB201CB8FF52BA4C" ID-GBIF-Occurrence="3986696402" pageId="59" pageNumber="60">
|
||
<emphasis id="18D4243FFD64FFBAAB201CB8FA23BA69" bold="true" box="[1041,1437,1518,1544]" pageId="59" pageNumber="60">
|
||
<collectingCountry id="52B7B8BDFD64FFBAAB201CB8FB00BA69" box="[1041,1214,1518,1544]" name="Venezuela" pageId="59" pageNumber="60">VENEZUELA</collectingCountry>
|
||
(6).
|
||
<collectingRegion id="E86436CFFD64FFBAABC51CB8FAD2BA69" box="[1268,1388,1518,1544]" country="Venezuela" name="Carabobo" pageId="59" pageNumber="60">Carabobo</collectingRegion>
|
||
(1):
|
||
</emphasis>
|
||
Nirgua.
|
||
</materialsCitation>
|
||
<materialsCitation id="9AC8F270FD64FFBAAFC31F44FDB7BA4C" ID-GBIF-Occurrence="3986696731" box="[242,521,1554,1581]" pageId="59" pageNumber="60">
|
||
<emphasis id="18D4243FFD64FFBAAFC31F44FE3FBA4D" bold="true" box="[242,385,1554,1580]" pageId="59" pageNumber="60">
|
||
<collectingRegion id="E86436CFFD64FFBAAFC31F44FEEFBA4D" box="[242,337,1554,1580]" country="Venezuela" name="Cojedes" pageId="59" pageNumber="60">Cojedes</collectingRegion>
|
||
(1):
|
||
</emphasis>
|
||
San Carlos.
|
||
</materialsCitation>
|
||
<materialsCitation id="9AC8F270FD64FFBAAD3F1F45FA86BA4C" ID-GBIF-Occurrence="3986696389" box="[526,1336,1554,1581]" pageId="59" pageNumber="60">
|
||
<emphasis id="18D4243FFD64FFBAAD3F1F45FD78BA4D" bold="true" box="[526,710,1554,1581]" pageId="59" pageNumber="60">
|
||
<collectingRegion id="E86436CFFD64FFBAAD3F1F45FD2BBA4C" box="[526,661,1555,1581]" country="Venezuela" name="Portuguesa" pageId="59" pageNumber="60">Portuguesa</collectingRegion>
|
||
(1):
|
||
</emphasis>
|
||
<location id="2F7FAEF6FD64FFBAADFD1F45FBE7BA4C" LSID="urn:lsid:plazi:treatment:A209493BFD65FFBAAFA61F3FFA24BADC:2F7FAEF6FD64FFBAADFD1F45FBE7BA4C" box="[716,1113,1555,1581]" country="Costa Rica" county="Estacion La Selva" municipality="Reserva Rio Platano" name="Estacion Experimental San Nicolas" pageId="59" pageNumber="60" stateProvince="Carabobo">Estación Experimental San Nicolás</location>
|
||
,
|
||
<quantity id="ED5855C8FD64FFBAAB551F44FB14BA4C" box="[1124,1194,1554,1581]" metricMagnitude="4" metricUnit="m" metricValue="5.6" pageId="59" pageNumber="60" unit="km" value="56.0">56 km</quantity>
|
||
<collectorName id="87559DFBFD64FFBAAB811F45FA86BA4C" box="[1200,1336,1555,1581]" pageId="59" pageNumber="60">de Guanare.</collectorName>
|
||
</materialsCitation>
|
||
<materialsCitation id="9AC8F270FD64FFBAAA0C1F44FC38BA30" ID-GBIF-Occurrence="3986696877" pageId="59" pageNumber="60">
|
||
<emphasis id="18D4243FFD64FFBAAA0C1F44FF7FBA31" bold="true" pageId="59" pageNumber="60">
|
||
<collectingRegion id="E86436CFFD64FFBAAA0C1F44FA22BA4C" box="[1341,1436,1554,1581]" country="Venezuela" name="Tachira" pageId="59" pageNumber="60">Táchira</collectingRegion>
|
||
(1):
|
||
</emphasis>
|
||
<location id="2F7FAEF6FD64FFBAAFFB1F61FD53BA30" LSID="urn:lsid:plazi:treatment:A209493BFD65FFBAAFA61F3FFA24BADC:2F7FAEF6FD64FFBAAFFB1F61FD53BA30" box="[202,749,1591,1617]" country="Costa Rica" county="Estacion La Selva" municipality="Reserva Rio Platano" name="Complejo Hidroelectrico Leonardo Ruiz Pineda" pageId="59" pageNumber="60" stateProvince="Carabobo">Complejo Hidroeléctrico Leonardo Ruíz Pineda</location>
|
||
, Las Cuevas.
|
||
</materialsCitation>
|
||
<materialsCitation id="9AC8F270FD64FFBAACBF1F60FB6ABA30" ID-GBIF-Occurrence="3986696595" box="[910,1236,1590,1617]" pageId="59" pageNumber="60">
|
||
<emphasis id="18D4243FFD64FFBAACBF1F60FBBFBA31" bold="true" box="[910,1025,1590,1616]" pageId="59" pageNumber="60">
|
||
<collectingRegion id="E86436CFFD64FFBAACBF1F60FC70BA31" box="[910,974,1590,1616]" country="Venezuela" name="Zulia" pageId="59" pageNumber="60">Zulia</collectingRegion>
|
||
(2):
|
||
</emphasis>
|
||
<location id="2F7FAEF6FD64FFBAAB3B1F61FBF0BA30" LSID="urn:lsid:plazi:treatment:A209493BFD65FFBAAFA61F3FFA24BADC:2F7FAEF6FD64FFBAAB3B1F61FBF0BA30" box="[1034,1102,1591,1617]" country="Costa Rica" county="Estacion La Selva" municipality="Reserva Rio Platano" name="Perija" pageId="59" pageNumber="60" stateProvince="Carabobo">Perijá</location>
|
||
,
|
||
<collectorName id="87559DFBFD64FFBAAB6A1F61FB6ABA30" box="[1115,1236,1591,1617]" pageId="59" pageNumber="60">
|
||
<location id="2F7FAEF6FD64FFBAAB6A1F61FB6ABA30" LSID="urn:lsid:plazi:treatment:A209493BFD65FFBAAFA61F3FFA24BADC:2F7FAEF6FD64FFBAAB6A1F61FB6ABA30" box="[1115,1236,1591,1617]" country="Costa Rica" county="Estacion La Selva" municipality="Reserva Rio Platano" name="El Tucuco" pageId="59" pageNumber="60" stateProvince="Carabobo">El Tucuco</location>
|
||
</collectorName>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="9AC8F270FD64FFBAABD11F61FEBCBA14" ID-GBIF-Occurrence="3986696517" pageId="59" pageNumber="60">
|
||
<location id="2F7FAEF6FD64FFBAABD11F61FA9ABA30" LSID="urn:lsid:plazi:treatment:A209493BFD65FFBAAFA61F3FFA24BADC:2F7FAEF6FD64FFBAABD11F61FA9ABA30" box="[1248,1316,1591,1617]" country="Costa Rica" county="Estacion La Selva" municipality="Reserva Rio Platano" name="Perija" pageId="59" pageNumber="60" stateProvince="Carabobo">Perijá</location>
|
||
,
|
||
<location id="2F7FAEF6FD64FFBAAA031F61FA26BA30" LSID="urn:lsid:plazi:treatment:A209493BFD65FFBAAFA61F3FFA24BADC:2F7FAEF6FD64FFBAAA031F61FA26BA30" box="[1330,1432,1591,1617]" country="Costa Rica" county="Estacion La Selva" municipality="Reserva Rio Platano" name="Kasmera" pageId="59" pageNumber="60" stateProvince="Carabobo">Kasmera</location>
|
||
,
|
||
<location id="2F7FAEF6FD64FFBAAFA61F0DFF41BA14" LSID="urn:lsid:plazi:treatment:A209493BFD65FFBAAFA61F3FFA24BADC:2F7FAEF6FD64FFBAAFA61F0DFF41BA14" box="[151,255,1627,1653]" country="Costa Rica" county="Estacion La Selva" municipality="Reserva Rio Platano" name="Rio Yasa" pageId="59" pageNumber="60" stateProvince="Carabobo">Río Yasa</location>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="2A1FF82DFD64FFBAAFF61F29FA24BADC" blockId="59.[151,1437,150,1725]" pageId="59" pageNumber="60">
|
||
<emphasis id="18D4243FFD64FFBAAFF61F29FE39BAF9" bold="true" box="[199,391,1662,1689]" pageId="59" pageNumber="60">Natural history.</emphasis>
|
||
Adults can be collected using lights at night. The species is found mainly in tropical dry to humid forests below
|
||
<quantity id="ED5855C8FD64FFBAAEB21FF4FE74BADC" box="[387,458,1698,1725]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="59" pageNumber="60" unit="m" value="650.0">650 m</quantity>
|
||
and infrequently in montane forest at
|
||
<quantity id="ED5855C8FD64FFBAAC471FF5FBA1BADC" box="[886,1055,1699,1725]" metricMagnitude="3" metricUnit="m" metricValue="1.3" metricValueMax="1.4" metricValueMin="1.2" pageId="59" pageNumber="60" unit="m" value="1300.0" valueMax="1400.0" valueMin="1200.0">
|
||
<geoCoordinate id="4F949EEAFD64FFBAAC471FF5FC0BBADC" box="[886,949,1699,1725]" degrees="1,200" direction="north" orientation="latitude" pageId="59" pageNumber="60" precision="55" value="1.2">1,200</geoCoordinate>
|
||
<geoCoordinate id="4F949EEAFD64FFBAAC841FF5FBBCBADC" box="[949,1026,1699,1725]" degrees="1,400" direction="west" orientation="longitude" pageId="59" pageNumber="60" precision="55" value="-1.4">–1,400</geoCoordinate>
|
||
m
|
||
</quantity>
|
||
(
|
||
<bibRefCitation id="4E3185DCFD64FFBAAB1C1FF5FAACBADC" author="Ratcliffe, B. C. & Cave, R. D. & Cano, E. B." box="[1069,1298,1699,1725]" pageId="59" pageNumber="60" pagination="1 - 666" refId="ref93157" refString="Ratcliffe, B. C., Cave, R. D. & Cano, E. B. (2013) The dynastine scarab beetles of Mexico, Guatemala, and Belize (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum, 27, 1 - 666." type="journal article" year="2013">
|
||
Ratcliffe
|
||
<emphasis id="18D4243FFD64FFBAABA91FF5FB6EBADC" box="[1176,1232,1699,1725]" italics="true" pageId="59" pageNumber="60">et al.</emphasis>
|
||
2013
|
||
</bibRefCitation>
|
||
, as
|
||
<taxonomicName id="EDA083AEFD64FFBAAA0A1FF5FA37BADD" authorityName="Erichson" authorityYear="1848" box="[1339,1417,1699,1724]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="gyas">
|
||
<emphasis id="18D4243FFD64FFBAAA0A1FF5FA37BADD" box="[1339,1417,1699,1724]" italics="true" pageId="59" pageNumber="60">T. gyas</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |