451 lines
60 KiB
XML
451 lines
60 KiB
XML
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<mods:title id="87D2C1F62E371817078ED50CF9F29EBD">Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision</mods:title>
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<mods:namePart id="3D6625B98EB7FB616C3409C33FEC0A3F">López-García, Margarita M.</mods:namePart>
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<mods:namePart id="0718A8920F2C0CFF26D9FAC472B574D8">Deloya, Cuauhtémoc</mods:namePart>
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<treatment id="A209493BFD4BFF94AFA61F28FB4BBA30" ID-DOI="http://doi.org/10.5281/zenodo.7399788" ID-GBIF-Taxon="205794542" ID-Zenodo-Dep="7399788" LSID="urn:lsid:plazi:treatment:A209493BFD4BFF94AFA61F28FB4BBA30" httpUri="http://treatment.plazi.org/id/A209493BFD4BFF94AFA61F28FB4BBA30" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
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<heading id="71574F41FD4BFF95AFA61F28FCF6BAF9" bold="true" box="[151,840,1662,1689]" fontSize="11" level="2" pageId="20" pageNumber="21" reason="4">
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<taxonomicName id="EDA083AEFD4BFF95AFA61F28FDD4BAF9" ID-CoL="9WJ2M" authority="(Prell, 1937)" baseAuthorityName="Prell" baseAuthorityYear="1937" box="[151,618,1662,1689]" class="Insecta" family="Scarabaeidae" genus="Proculigyrus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="cicatricosus" status="comb. nov.">
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<emphasis id="18D4243FFD4BFF95AFA61F28FE74BAF8" bold="true" box="[151,458,1662,1689]" italics="true" pageId="20" pageNumber="21">Proculigyrus cicatricosus</emphasis>
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(
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<bibRefCitation id="4E3185DCFD4BFF95AEEB1F28FDDCBAF9" author="Prell, H." box="[474,610,1662,1688]" pageId="20" pageNumber="21" pagination="89 - 90" refId="ref92338" refString="Prell, H. (1937) Beitrage zur Kenntnis der Dynastiden. XV. Neue Arten und Rassen. Entomologische Zeitschrift, 51, 89 - 90." type="journal article" year="1937">Prell, 1937</bibRefCitation>
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<taxonomicNameLabel id="03E79944FD4BFF95AD431F29FCF6BAF9" box="[626,840,1662,1689]" pageId="20" pageNumber="21" rank="species">new combination</taxonomicNameLabel>
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<paragraph id="2A1FF82DFD4BFF95AFA61FF5FBE0BADC" blockId="20.[151,1118,1662,1725]" box="[151,1118,1698,1725]" pageId="20" pageNumber="21">
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(
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<figureCitation id="B29BE4A8FD4BFF95AFAE1FF5FEBFBADC" box="[159,257,1699,1725]" captionStart="FIGURE 1" captionStartId="81.[151,250,1535,1559]" captionTargetBox="[151,1436,181,1511]" captionTargetId="figure-17@81.[151,1436,181,1511]" captionTargetPageId="81" captionText="FIGURE 1. Head. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) Ligyrus (Ligyrodes) peruvianus, D) L. (Anagrylius) moroni, E) L. (Ligyrus) ruginasus, F) L. (Ligyrus) nasutus, G) L. (Ligyrus) bidentulus, H) Tomarus adoceteus, I) T. maimon, J) T. pullus, K) T. pumilus." figureDoi="http://doi.org/10.5281/zenodo.7345011" httpUri="https://zenodo.org/record/7345011/files/figure.png" pageId="20" pageNumber="21">Figs. 1B</figureCitation>
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,
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<figureCitation id="B29BE4A8FD4BFF95AE3C1FF5FE95BADC" box="[269,299,1699,1725]" captionStart="FIGURE 2" captionStartId="82.[151,250,1584,1608]" captionTargetBox="[151,1436,181,1548]" captionTargetId="figure-24@82.[151,1436,181,1560]" captionTargetPageId="82" captionText="FIGURE 2. Labium. A) Ligyrus (Anagrylius) moroni, B) Ligyrus (Ligyrus) allonasutus, C) L. (Ligyrus) gibbosus, D) L. (Ligyrus) villosus, E) L. (Ligyrodes) relictus, F) Proculigyrus cicatricosus, G) Euligyrus ebenus, H) E. similis, I) Tomarus gyas, J) Tomarus selanderi, K) Tomarus subtropicus. Abbreviations: LL: Ligular lobes, LP: Labial palpus, M: Mentum, S: Suture." figureDoi="http://doi.org/10.5281/zenodo.7345015" httpUri="https://zenodo.org/record/7345015/files/figure.png" pageId="20" pageNumber="21">2F</figureCitation>
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<figureCitation id="B29BE4A8FD4BFF95AE061FF5FEE7BADC" box="[311,345,1699,1725]" captionStart="FIGURE 4" captionStartId="83.[151,250,1872,1896]" captionTargetBox="[151,1437,1118,1848]" captionTargetId="figure-81@83.[151,1437,1118,1848]" captionTargetPageId="83" captionText="FIGURE 4. Maxilla. A) Euligyrus similis, B) Proculigyrus cicatricosus, C) Ligyrus (Ligyrodes) relictus, D) L. (Ligyrus) burmeisteri, E) L. (Ligyrus) villosus, F) Tomarus gyas, G) Tomarus maternus, H) T. selanderi, I) T. subtropicus." figureDoi="http://doi.org/10.5281/zenodo.7345023" httpUri="https://zenodo.org/record/7345023/files/figure.png" pageId="20" pageNumber="21">4B</figureCitation>
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<figureCitation id="B29BE4A8FD4BFF95AE541FF4FE36BADC" box="[357,392,1698,1725]" captionStart="FIGURE 5" captionStartId="84.[151,250,931,955]" captionTargetBox="[188,1400,181,907]" captionTargetId="figure-24@84.[188,1400,181,907]" captionTargetPageId="84" captionText="FIGURE 5. Mandible. A) Euligyrus similis, B) Proculigyrus cicatricosus, C) Ligyrus (Ligyrodes) relictus, D) L. (Ligyrus) burmeisteri, E) L. (Ligyrus) villosus, F) Tomarus rostratus, G) T. gyas, H) T. maternus." figureDoi="http://doi.org/10.5281/zenodo.7345025" httpUri="https://zenodo.org/record/7345025/files/figure.png" pageId="20" pageNumber="21">5B</figureCitation>
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<figureCitation id="B29BE4A8FD4BFF95AEA51FF5FE65BADC" box="[404,475,1699,1725]" captionStart="FIGURE 8" captionStartId="86.[151,250,1712,1736]" captionTargetBox="[151,1436,181,1688]" captionTargetId="figure-24@86.[151,1436,181,1688]" captionTargetPageId="86" captionText="FIGURE 8. Stridulatory apparatus. A, D) Proculigyrus cicatricosus, B, E) Ligyrus (Ligyrus) burmeisteri, C, F) L. (Ligyrodes) relictus, G, K) Tomarus amazonicus, H, L) T. maimon, I, M) T. maternus, J, N) T. gyas." figureDoi="http://doi.org/10.5281/zenodo.7345035" httpUri="https://zenodo.org/record/7345035/files/figure.png" pageId="20" pageNumber="21">8A, D</figureCitation>
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<figureCitation id="B29BE4A8FD4BFF95AED61FF5FDB7BADC" box="[487,521,1699,1725]" captionStart="FIGURE 9" captionStartId="87.[151,250,1074,1098]" captionTargetBox="[151,1436,181,1050]" captionTargetId="figure-17@87.[151,1436,181,1050]" captionTargetPageId="87" captionText="FIGURE 9. Pygidium of male. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) Ligyrus (Anagrylius) cuniculus, D) L. (Ligyrus) fossor, E) L. (Ligyrus) paranaensis, F) Tomarus bituberculatus, G) T. colombianus, H) T. maracaiboensis, I) T. pilcopataensis, J) T. pullus, K) T. roigjunenti, L) T. rostratus." figureDoi="http://doi.org/10.5281/zenodo.7345041" httpUri="https://zenodo.org/record/7345041/files/figure.png" pageId="20" pageNumber="21">9B</figureCitation>
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<figureCitation id="B29BE4A8FD4BFF95AD241FF5FDFBBADC" box="[533,581,1699,1725]" captionStart="FIGURE 11" captionStartId="88.[151,250,931,955]" captionTargetBox="[234,1354,186,899]" captionTargetId="figure-24@88.[231,1357,181,907]" captionTargetPageId="88" captionText="FIGURE 11. Protibia in frontal view.A) Euligyrus similis, B) Proculigyrus cicatricosus, C) Ligyrus (Anagrylius) moroni, D) L. (Ligyrodes) relictus, E) L. (Ligyrodes) sallaei, F) L. (Ligyrus) allonasutus, G) L. (Ligyrus) villosus, H) Tomarus bituberculatus, I) T. colombianus, J) T. pullus." figureDoi="http://doi.org/10.5281/zenodo.7345051" httpUri="https://zenodo.org/record/7345051/files/figure.png" pageId="20" pageNumber="21">11B</figureCitation>
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<figureCitation id="B29BE4A8FD4BFF95AD601FF5FD7EBADC" box="[593,704,1699,1725]" captionStart="FIGURE 12" captionStartId="88.[151,250,1921,1945]" captionTargetBox="[237,1351,1055,1897]" captionTargetId="figure-89@88.[233,1355,1052,1897]" captionTargetPageId="88" captionText="FIGURE 12. Mesotibia (A–F). A) Euligyrus similis, B) Proculigyrus cicatricosus, C) Ligyrus (Anagrylius) moroni, D) L. (Ligyrodes) sallaei; E) L. (Ligyrus) allonasutus, F) L. (Ligyrus) villosus. Metatibia (G–L). G) Euligyrus similis, H) Proculigyrus cicatricosus, I) Ligyrus (Anagrylius) moroni, J) L. (Ligyrodes) sallaei, K) L. (Ligyrus) allonasutus, L) L. (Ligyrus) villosus." figureDoi="http://doi.org/10.5281/zenodo.7345053" httpUri="https://zenodo.org/record/7345053/files/figure.png" pageId="20" pageNumber="21">12B, 12H</figureCitation>
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<figureCitation id="B29BE4A8FD4BFF95ADFD1FF5FD42BADC" box="[716,764,1699,1725]" captionStart="FIGURE 13" captionStartId="89.[151,250,1041,1065]" captionTargetBox="[154,1432,183,1016]" captionTargetId="figure-17@89.[151,1436,181,1017]" captionTargetPageId="89" captionText="FIGURE 13. Female genital plates. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) Ligyrus (Anagrylius) moroni, D) L. (Ligyrodes) sallaei, E) L. (Ligyrus) allonasutus, F) L. (Ligyrus) bidentulus, G) L. (Ligyrus) neglectus, H) Tomarus laticaudus, I) T. maternus, J) T. rostratus, K) T. subtropicus. Abbreviations: SCx: Subcoxite, Cx: Coxite." figureDoi="http://doi.org/10.5281/zenodo.7345057" httpUri="https://zenodo.org/record/7345057/files/figure.png" pageId="20" pageNumber="21">13B</figureCitation>
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<figureCitation id="B29BE4A8FD4BFF95AC391FF5FC84BADC" box="[776,826,1699,1725]" captionStart="FIGURE 14" captionStartId="89.[151,250,1677,1701]" captionTargetBox="[152,1432,1161,1649]" captionTargetId="figure-90@89.[152,1436,1161,1653]" captionTargetPageId="89" captionText="FIGURE 14. Spiculum gastrale. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) L. (Ligyrodes) relictus, D) L. (Ligyrus) villosus, E) Tomarus pullus, F) T. rostratus, G) T. selanderi, H) T. subtropicus." figureDoi="http://doi.org/10.5281/zenodo.7345059" httpUri="https://zenodo.org/record/7345059/files/figure.png" pageId="20" pageNumber="21">14B</figureCitation>
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<figureCitation id="B29BE4A8FD4BFF95AC741FF4FCC8BADC" box="[837,886,1698,1725]" captionStart="FIGURE 15" captionStartId="90.[151,250,1535,1559]" captionTargetBox="[154,1430,183,1509]" captionTargetId="figure-24@90.[151,1436,181,1512]" captionTargetPageId="90" captionText="FIGURE 15. Parameres in frontal view. A) Euligyrus ebenus, B) E. similis, C) Proculigyrus cicatricosus, D) Ligyrus (Anagrylius) cuniculus, E) L. (Anagrylius) moroni, F) L. (Ligyrodes) peruvianus, G) L. (Ligyrodes) relictus, H) L. (Ligyrodes) sallaei, I) L. (Ligyrus) allonasutus, J) L. (Ligyrus) bidentulus, K) L. (Ligyrus) burmeisteri, L) Ligyrus (Ligyrus) fossor, M) L. (Ligyrus) gibbosus, N) L. (Ligyrus) nasutus, O) L. (Ligyrus) neglectus, P) L. (Ligyrus) paranaensis, Q) L. (Ligyrus) peninsularis, R) L. (Ligyrus) rubripes, S) L. (Ligyrus) ruginasus, T) L. (Ligyrus) villosus." figureDoi="http://doi.org/10.5281/zenodo.7345063" httpUri="https://zenodo.org/record/7345063/files/figure.png" pageId="20" pageNumber="21">15C</figureCitation>
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<figureCitation id="B29BE4A8FD4BFF95ACB31FF5FC0DBADC" box="[898,947,1699,1725]" captionStart="FIGURE 17" captionStartId="92.[151,250,1215,1239]" captionTargetBox="[155,1430,188,1184]" captionTargetId="figure-24@92.[151,1436,181,1191]" captionTargetPageId="92" captionText="FIGURE 17. Parameres in lateral view. A) Euligyrus ebenus, B) E. similis, C) Proculigyrus cicatricosus, D) Ligyrus (Anagrylius) cuniculus; E) L. (Anagrylius) moroni, F) L. (Ligyrodes) relictus, G) L. (Ligyrodes) sallaei." figureDoi="http://doi.org/10.5281/zenodo.7345073" httpUri="https://zenodo.org/record/7345073/files/figure.png" pageId="20" pageNumber="21">17C</figureCitation>
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<figureCitation id="B29BE4A8FD4BFF95AC8F1FF5FC4EBADC" box="[958,1008,1699,1725]" captionStart="FIGURE 21" captionStartId="96.[151,250,1739,1763]" captionTargetBox="[161,1418,194,1706]" captionTargetId="figure-24@96.[151,1435,181,1715]" captionTargetPageId="96" captionText="FIGURE 21. Internal sac. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) Ligyrus (Anagrylius) moroni, D) L. (Ligyrodes) sallaei, E) L. (Ligyrus) allonasutus. Abbreviations:AL:Accessory lamellae, Br: Brush of setae, CL: Copulatory lamella, LSB: Lamellar spiny belt, Ra: Raspula." figureDoi="http://doi.org/10.5281/zenodo.7345085" httpUri="https://zenodo.org/record/7345085/files/figure.png" pageId="20" pageNumber="21">21B</figureCitation>
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<figureCitation id="B29BE4A8FD4BFF95ACCA1FF5FB90BADC" box="[1019,1070,1699,1725]" captionStart="FIGURE 24" captionStartId="99.[151,250,883,907]" captionTargetBox="[163,1419,185,844]" captionTargetId="figure-17@99.[151,1436,181,859]" captionTargetPageId="99" captionText="FIGURE 24. Habitus. A) Euligyrus ebenus, B) E. similis, C) Proculigyrus cicatricosus. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7345095" httpUri="https://zenodo.org/record/7345095/files/figure.png" pageId="20" pageNumber="21">24C</figureCitation>
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;
|
||
<figureCitation id="B29BE4A8FD4BFF95AB081FF5FBE9BADC" box="[1081,1111,1699,1725]" captionStart="FIGURE 40" captionStartId="110.[151,250,1952,1976]" captionTargetBox="[163,1434,994,1923]" captionTargetId="figure-41@110.[153,1434,994,1929]" captionTargetPageId="110" captionText="FIGURE 40. Distributional records of Proculigyrus cicatricosus." figureDoi="http://doi.org/10.5281/zenodo.7345150" httpUri="https://zenodo.org/record/7345150/files/figure.png" pageId="20" pageNumber="21">40</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="62BAABA6FD4BFF95AFA61FBEFD4DBA9E" box="[151,755,1767,1791]" pageId="20" pageNumber="21" type="reference_group">
|
||
<paragraph id="2A1FF82DFD4BFF95AFA61FBEFD4DBA9E" blockId="20.[151,1437,1767,1884]" box="[151,755,1767,1791]" pageId="20" pageNumber="21">
|
||
<treatmentCitationGroup id="0AB0DF03FD4BFF95AFA61FBEFD4DBA9E" box="[151,755,1767,1791]" pageId="20" pageNumber="21">
|
||
<treatmentCitation id="AB01DE3CFD4BFF95AFA61FBEFDB9BA9E" author="Prell, H." box="[151,519,1767,1791]" page="90" pageId="20" pageNumber="21" year="1937">
|
||
<taxonomicName id="EDA083AEFD4BFF95AFA61FBEFDB9BA9E" authority="Prell, 1937: 90" authorityName="Prell" authorityPageNumber="90" authorityYear="1937" box="[151,519,1767,1791]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="cicatricosus">
|
||
<emphasis id="18D4243FFD4BFF95AFA61FBEFED8BA9E" box="[151,358,1768,1791]" italics="true" pageId="20" pageNumber="21">Ligyrus cicatricosus</emphasis>
|
||
<bibRefCitation id="4E3185DCFD4BFF95AE5C1FB1FDB9BA9E" author="Prell, H." box="[365,519,1767,1791]" pageId="20" pageNumber="21" pagination="89 - 90" refId="ref92338" refString="Prell, H. (1937) Beitrage zur Kenntnis der Dynastiden. XV. Neue Arten und Rassen. Entomologische Zeitschrift, 51, 89 - 90." type="journal article" year="1937">Prell, 1937: 90</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
. Original combination.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="62BAABA6FD4BFF95ADC91FB1FDBDBB3A" pageId="20" pageNumber="21" type="materials_examined">
|
||
<paragraph id="2A1FF82DFD4BFF95ADC91FB1FDBDBB3A" blockId="20.[151,1437,1767,1884]" pageId="20" pageNumber="21">
|
||
Male
|
||
<typeStatus id="F51B468FFD4BFF95AC021FB1FC2EBA9E" box="[819,912,1767,1791]" pageId="20" pageNumber="21" type="lectotype">lectotype</typeStatus>
|
||
designated by
|
||
<bibRefCitation id="4E3185DCFD4BFF95AB1A1FB1FB78BA9E" author="Endrodi, S." box="[1067,1222,1767,1791]" pageId="20" pageNumber="21" pagination="1 - 145" refId="ref89768" refString="Endrodi, S. (1969) Monographie der Dynastinae. 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). Entomologische Abhandlungen, 37, 1 - 145." type="journal article" year="1969">Endrödi (1969)</bibRefCitation>
|
||
(ZMHB) “
|
||
<collectingCountry id="52B7B8BDFD4BFF95AA061FB1FA39BA9E" box="[1335,1415,1767,1791]" name="Mexico" pageId="20" pageNumber="21">Mexico</collectingCountry>
|
||
//
|
||
<taxonomicName id="EDA083AEFD4BFF95AFF61E50FEA8BB7F" authorityName="Burmeister" authorityYear="1847" box="[199,278,1798,1822]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">Ligyrus</taxonomicName>
|
||
/ (
|
||
<taxonomicName id="EDA083AEFD4BFF95AE1E1E50FE1DBB7F" authorityName="Casey" authorityYear="1915" box="[303,419,1798,1822]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="101" phylum="Arthropoda" rank="subGenus" subGenus="Anagrylius">Anagrylius</taxonomicName>
|
||
) /
|
||
<taxonomicName id="EDA083AEFD4BFF95AE8A1E50FDD6BB7F" authority="Prell" authorityName="Prell" authorityYear="1937" box="[443,616,1798,1822]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="cicatricosus">cicatricosus Prell</taxonomicName>
|
||
/
|
||
<specimenCount id="3CA633A4FD4BFF95AD481E50FD34BB7F" box="[633,650,1798,1822]" pageId="20" pageNumber="21" type="male">♂</specimenCount>
|
||
-Type //
|
||
<typeStatus id="F51B468FFD4BFF95ADEE1E50FCEEBB7F" box="[735,848,1798,1822]" pageId="20" pageNumber="21" type="lectotype">Lectotypus</typeStatus>
|
||
/
|
||
<taxonomicName id="EDA083AEFD4BFF95AC531E50FC0FBB7F" authorityName="Burmeister" authorityYear="1847" box="[866,945,1798,1822]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">Ligyrus</taxonomicName>
|
||
/ cicatricosus / Prell / Endrody”. Female
|
||
<typeStatus id="F51B468FFD4BFF95AA7A1E50FEB8BB5C" pageId="20" pageNumber="21" type="paralectotype">paralectotype</typeStatus>
|
||
(ZMHB) “
|
||
<collectingCountry id="52B7B8BDFD4BFF95AE451E72FE7BBB5D" box="[372,453,1828,1852]" name="Mexico" pageId="20" pageNumber="21">Mexiko</collectingCountry>
|
||
/ Coll / Dr. ITZINGER //
|
||
<taxonomicName id="EDA083AEFD4BFF95ADF91E72FCA9BB5D" authorityName="Burmeister" authorityYear="1847" box="[712,791,1828,1852]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">Ligyrus</taxonomicName>
|
||
/
|
||
<taxonomicName id="EDA083AEFD4BFF95AC181E72FC68BB5D" authority="Prell" authorityName="Prell" authorityYear="1937" box="[809,982,1828,1852]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="cicatricosus">cicatricosus Prell</taxonomicName>
|
||
/
|
||
<specimenCount id="3CA633A4FD4BFF95ACD61E73FC4ABB5C" box="[999,1012,1829,1853]" pageId="20" pageNumber="21" type="female">♀</specimenCount>
|
||
-Type //
|
||
<typeStatus id="F51B468FFD4BFF95AB7F1E73FB0CBB5C" box="[1102,1202,1829,1853]" pageId="20" pageNumber="21" type="paratype">Paratypus</typeStatus>
|
||
/
|
||
<taxonomicName id="EDA083AEFD4BFF95ABF51E72FAADBB5D" authorityName="Burmeister" authorityYear="1847" box="[1220,1299,1828,1852]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">Ligyrus</taxonomicName>
|
||
/ cicatricosus / Prell”. Type locality:
|
||
<collectingCountry id="52B7B8BDFD4BFF95AE9F1E15FE41BB3A" box="[430,511,1859,1883]" name="Mexico" pageId="20" pageNumber="21">Mexico</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="62BAABA6FD4BFF94AFA61EDFFC5BBEC8" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="description">
|
||
<paragraph id="2A1FF82DFD4BFF94AFA61EDFFC5BBEC8" blockId="20.[151,1437,1929,2028]" lastBlockId="21.[151,1437,150,1617]" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
|
||
<emphasis id="18D4243FFD4BFF95AFA61EDFFE97BBC2" bold="true" box="[151,297,1929,1955]" pageId="20" pageNumber="21">Description.</emphasis>
|
||
Habitus as in
|
||
<figureCitation id="B29BE4A8FD4BFF95AEF71EDFFD92BBC2" box="[454,556,1929,1955]" captionStart="FIGURE 24" captionStartId="99.[151,250,883,907]" captionTargetBox="[163,1419,185,844]" captionTargetId="figure-17@99.[151,1436,181,859]" captionTargetPageId="99" captionText="FIGURE 24. Habitus. A) Euligyrus ebenus, B) E. similis, C) Proculigyrus cicatricosus. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7345095" httpUri="https://zenodo.org/record/7345095/files/figure.png" pageId="20" pageNumber="21">Fig. 24C</figureCitation>
|
||
. Length
|
||
<quantity id="ED5855C8FD4BFF95ADBA1EDFFC8EBBC5" box="[651,816,1929,1956]" metricMagnitude="-2" metricUnit="m" metricValue="2.085" metricValueMax="2.52" metricValueMin="1.65" pageId="20" pageNumber="21" unit="mm" value="20.85" valueMax="25.2" valueMin="16.5">16.5–25.2 mm</quantity>
|
||
. Humeral width
|
||
<quantity id="ED5855C8FD4BFF95ACD81EDFFB3EBBC5" box="[1001,1152,1929,1956]" metricMagnitude="-2" metricUnit="m" metricValue="1.075" metricValueMax="1.33" metricValueMin="0.82" pageId="20" pageNumber="21" unit="mm" value="10.75" valueMax="13.3" valueMin="8.2">8.2–13.3 mm</quantity>
|
||
. Color black, legs lighter.
|
||
<emphasis id="18D4243FFD4BFF95AFA61EFBFF5EBBA6" bold="true" box="[151,224,1965,1991]" pageId="20" pageNumber="21">Head:</emphasis>
|
||
Frons slightly concave between eyes and behind frontoclypeal tubercles; surface with deep, large, confluent punctures, nearly smooth on vertex and only with sparser punctures. Clypeus trapezoidal (
|
||
<figureCitation id="B29BE4A8FD4BFF95ABBF1E87FB5BBB8A" box="[1166,1253,2001,2027]" captionStart="FIGURE 1" captionStartId="81.[151,250,1535,1559]" captionTargetBox="[151,1436,181,1511]" captionTargetId="figure-17@81.[151,1436,181,1511]" captionTargetPageId="81" captionText="FIGURE 1. Head. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) Ligyrus (Ligyrodes) peruvianus, D) L. (Anagrylius) moroni, E) L. (Ligyrus) ruginasus, F) L. (Ligyrus) nasutus, G) L. (Ligyrus) bidentulus, H) Tomarus adoceteus, I) T. maimon, J) T. pullus, K) T. pumilus." figureDoi="http://doi.org/10.5281/zenodo.7345011" httpUri="https://zenodo.org/record/7345011/files/figure.png" pageId="20" pageNumber="21">Fig. 1B</figureCitation>
|
||
). Apical clypeal teeth transverse, small, separated by 5 tooth diameters. Ocular canthus acute. Frons with 2 tumescences. Interocular distance 3.7 times the transverse ocular diameter. Antennal club short.
|
||
<emphasis id="18D4243FFD4AFF94AC8B19EDFB83BCB4" bold="true" box="[954,1085,187,213]" pageId="21" pageNumber="22">Pronotum:</emphasis>
|
||
Surface with punctures evenly sparse, deep, large, weakly umbilicate. Apex without defined tubercle, with an inconspicuous swelling and small fovea.
|
||
<emphasis id="18D4243FFD4AFF94AFD21855FE86BD7C" bold="true" box="[227,312,259,285]" pageId="21" pageNumber="22">Elytra:</emphasis>
|
||
Inner surface of apex with transverse tubercles forming about 85 parallel lines (
|
||
<figureCitation id="B29BE4A8FD4AFF94AB801855FAB5BD7C" box="[1201,1291,259,285]" captionStart="FIGURE 8" captionStartId="86.[151,250,1712,1736]" captionTargetBox="[151,1436,181,1688]" captionTargetId="figure-24@86.[151,1436,181,1688]" captionTargetPageId="86" captionText="FIGURE 8. Stridulatory apparatus. A, D) Proculigyrus cicatricosus, B, E) Ligyrus (Ligyrus) burmeisteri, C, F) L. (Ligyrodes) relictus, G, K) Tomarus amazonicus, H, L) T. maimon, I, M) T. maternus, J, N) T. gyas." figureDoi="http://doi.org/10.5281/zenodo.7345035" httpUri="https://zenodo.org/record/7345035/files/figure.png" pageId="21" pageNumber="22">Fig. 8D</figureCitation>
|
||
).
|
||
<emphasis id="18D4243FFD4AFF94AA2F1854FA22BD7D" bold="true" box="[1310,1436,258,284]" pageId="21" pageNumber="22">Abdomen:</emphasis>
|
||
Tergite IV with 6 transverse, inconspicuous striae (
|
||
<figureCitation id="B29BE4A8FD4AFF94ADE01871FC95BD20" box="[721,811,295,321]" captionStart="FIGURE 8" captionStartId="86.[151,250,1712,1736]" captionTargetBox="[151,1436,181,1688]" captionTargetId="figure-24@86.[151,1436,181,1688]" captionTargetPageId="86" captionText="FIGURE 8. Stridulatory apparatus. A, D) Proculigyrus cicatricosus, B, E) Ligyrus (Ligyrus) burmeisteri, C, F) L. (Ligyrodes) relictus, G, K) Tomarus amazonicus, H, L) T. maimon, I, M) T. maternus, J, N) T. gyas." figureDoi="http://doi.org/10.5281/zenodo.7345035" httpUri="https://zenodo.org/record/7345035/files/figure.png" pageId="21" pageNumber="22">Fig. 8A</figureCitation>
|
||
). Sternite VIII of female with narrow emargination. Pygidial surface completely punctate (
|
||
<figureCitation id="B29BE4A8FD4AFF94AD19181DFD3EBD04" box="[552,640,331,357]" captionStart="FIGURE 9" captionStartId="87.[151,250,1074,1098]" captionTargetBox="[151,1436,181,1050]" captionTargetId="figure-17@87.[151,1436,181,1050]" captionTargetPageId="87" captionText="FIGURE 9. Pygidium of male. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) Ligyrus (Anagrylius) cuniculus, D) L. (Ligyrus) fossor, E) L. (Ligyrus) paranaensis, F) Tomarus bituberculatus, G) T. colombianus, H) T. maracaiboensis, I) T. pilcopataensis, J) T. pullus, K) T. roigjunenti, L) T. rostratus." figureDoi="http://doi.org/10.5281/zenodo.7345041" httpUri="https://zenodo.org/record/7345041/files/figure.png" pageId="21" pageNumber="22">Fig. 9B</figureCitation>
|
||
); punctures subequal in size to those of elytral apex, ocellate, minutely setigerous. Surface evenly convex in male, slightly convex in female.
|
||
<emphasis id="18D4243FFD4AFF94AC901839FC44BDE8" bold="true" box="[929,1018,367,393]" pageId="21" pageNumber="22">Venter:</emphasis>
|
||
Apex of prosternal process spherical, with dense setae. Metespisternum without setae. Metasternum with small, deep, confluent punctures; with long setae, denser on anterior corners.
|
||
<emphasis id="18D4243FFD4AFF94AD2718E1FDE6BDB0" bold="true" box="[534,600,439,465]" pageId="21" pageNumber="22">Legs:</emphasis>
|
||
Male protarsus simple; inner claw simple. Protibia tridentate, basal tooth distant from the others (
|
||
<figureCitation id="B29BE4A8FD4AFF94AE95188DFDB4BD94" box="[420,522,475,501]" captionStart="FIGURE 11" captionStartId="88.[151,250,931,955]" captionTargetBox="[234,1354,186,899]" captionTargetId="figure-24@88.[231,1357,181,907]" captionTargetPageId="88" captionText="FIGURE 11. Protibia in frontal view.A) Euligyrus similis, B) Proculigyrus cicatricosus, C) Ligyrus (Anagrylius) moroni, D) L. (Ligyrodes) relictus, E) L. (Ligyrodes) sallaei, F) L. (Ligyrus) allonasutus, G) L. (Ligyrus) villosus, H) Tomarus bituberculatus, I) T. colombianus, J) T. pullus." figureDoi="http://doi.org/10.5281/zenodo.7345051" httpUri="https://zenodo.org/record/7345051/files/figure.png" pageId="21" pageNumber="22">Fig. 11B</figureCitation>
|
||
). Apical margin of mesotibia and metatibia dentate (
|
||
<figureCitation id="B29BE4A8FD4AFF94AB67188DFB5EBD94" box="[1110,1248,475,501]" captionStart="FIGURE 12" captionStartId="88.[151,250,1921,1945]" captionTargetBox="[237,1351,1055,1897]" captionTargetId="figure-89@88.[233,1355,1052,1897]" captionTargetPageId="88" captionText="FIGURE 12. Mesotibia (A–F). A) Euligyrus similis, B) Proculigyrus cicatricosus, C) Ligyrus (Anagrylius) moroni, D) L. (Ligyrodes) sallaei; E) L. (Ligyrus) allonasutus, F) L. (Ligyrus) villosus. Metatibia (G–L). G) Euligyrus similis, H) Proculigyrus cicatricosus, I) Ligyrus (Anagrylius) moroni, J) L. (Ligyrodes) sallaei, K) L. (Ligyrus) allonasutus, L) L. (Ligyrus) villosus." figureDoi="http://doi.org/10.5281/zenodo.7345053" httpUri="https://zenodo.org/record/7345053/files/figure.png" pageId="21" pageNumber="22">Fig. 12B, H</figureCitation>
|
||
). Apex of metatibia with 4–7 spinules.
|
||
<emphasis id="18D4243FFD4AFF94AEAC18A9FDD5BE79" bold="true" box="[413,619,510,537]" pageId="21" pageNumber="22">Female genitalia:</emphasis>
|
||
Subcoxite 1.8 times wider than long, outer sides narrowed towards apex (basal margin 1.5 times wider than apical margin). Coxite subrounded (as long as wide); surface strongly concave. Subcoxite as wide as coxite; shorter than coxite (
|
||
<figureCitation id="B29BE4A8FD4AFF94AD881B11FCA1BE00" box="[697,799,583,609]" captionStart="FIGURE 13" captionStartId="89.[151,250,1041,1065]" captionTargetBox="[154,1432,183,1016]" captionTargetId="figure-17@89.[151,1436,181,1017]" captionTargetPageId="89" captionText="FIGURE 13. Female genital plates. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) Ligyrus (Anagrylius) moroni, D) L. (Ligyrodes) sallaei, E) L. (Ligyrus) allonasutus, F) L. (Ligyrus) bidentulus, G) L. (Ligyrus) neglectus, H) Tomarus laticaudus, I) T. maternus, J) T. rostratus, K) T. subtropicus. Abbreviations: SCx: Subcoxite, Cx: Coxite." figureDoi="http://doi.org/10.5281/zenodo.7345057" httpUri="https://zenodo.org/record/7345057/files/figure.png" pageId="21" pageNumber="22">Fig. 13B</figureCitation>
|
||
).
|
||
<emphasis id="18D4243FFD4AFF94AC021B11FC58BE01" bold="true" box="[819,998,582,609]" pageId="21" pageNumber="22">Male genitalia:</emphasis>
|
||
Spiculum gastrale T-shaped, thickened (
|
||
<figureCitation id="B29BE4A8FD4AFF94AFAE1B3DFEBBBEE4" box="[159,261,619,645]" captionStart="FIGURE 14" captionStartId="89.[151,250,1677,1701]" captionTargetBox="[152,1432,1161,1649]" captionTargetId="figure-90@89.[152,1436,1161,1653]" captionTargetPageId="89" captionText="FIGURE 14. Spiculum gastrale. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) L. (Ligyrodes) relictus, D) L. (Ligyrus) villosus, E) Tomarus pullus, F) T. rostratus, G) T. selanderi, H) T. subtropicus." figureDoi="http://doi.org/10.5281/zenodo.7345059" httpUri="https://zenodo.org/record/7345059/files/figure.png" pageId="21" pageNumber="22">Fig. 14B</figureCitation>
|
||
). Parameres with a ventral, truncate tooth on each side; without dorsal teeth (
|
||
<figureCitation id="B29BE4A8FD4AFF94AB6A1B3DFB75BEE4" box="[1115,1227,618,645]" captionStart="FIGURE 15" captionStartId="90.[151,250,1535,1559]" captionTargetBox="[154,1430,183,1509]" captionTargetId="figure-24@90.[151,1436,181,1512]" captionTargetPageId="90" captionText="FIGURE 15. Parameres in frontal view. A) Euligyrus ebenus, B) E. similis, C) Proculigyrus cicatricosus, D) Ligyrus (Anagrylius) cuniculus, E) L. (Anagrylius) moroni, F) L. (Ligyrodes) peruvianus, G) L. (Ligyrodes) relictus, H) L. (Ligyrodes) sallaei, I) L. (Ligyrus) allonasutus, J) L. (Ligyrus) bidentulus, K) L. (Ligyrus) burmeisteri, L) Ligyrus (Ligyrus) fossor, M) L. (Ligyrus) gibbosus, N) L. (Ligyrus) nasutus, O) L. (Ligyrus) neglectus, P) L. (Ligyrus) paranaensis, Q) L. (Ligyrus) peninsularis, R) L. (Ligyrus) rubripes, S) L. (Ligyrus) ruginasus, T) L. (Ligyrus) villosus." figureDoi="http://doi.org/10.5281/zenodo.7345063" httpUri="https://zenodo.org/record/7345063/files/figure.png" pageId="21" pageNumber="22">Figs. 15C</figureCitation>
|
||
,
|
||
<figureCitation id="B29BE4A8FD4AFF94ABE71B3DFAB6BEE4" box="[1238,1288,619,645]" captionStart="FIGURE 17" captionStartId="92.[151,250,1215,1239]" captionTargetBox="[155,1430,188,1184]" captionTargetId="figure-24@92.[151,1436,181,1191]" captionTargetPageId="92" captionText="FIGURE 17. Parameres in lateral view. A) Euligyrus ebenus, B) E. similis, C) Proculigyrus cicatricosus, D) Ligyrus (Anagrylius) cuniculus; E) L. (Anagrylius) moroni, F) L. (Ligyrodes) relictus, G) L. (Ligyrodes) sallaei." figureDoi="http://doi.org/10.5281/zenodo.7345073" httpUri="https://zenodo.org/record/7345073/files/figure.png" pageId="21" pageNumber="22">17C</figureCitation>
|
||
). Internal sac with 12 accessory lamellae like spines and with a flat, oval base (
|
||
<figureCitation id="B29BE4A8FD4AFF94AC431BD9FC66BEC8" box="[882,984,655,681]" captionStart="FIGURE 21" captionStartId="96.[151,250,1739,1763]" captionTargetBox="[161,1418,194,1706]" captionTargetId="figure-24@96.[151,1435,181,1715]" captionTargetPageId="96" captionText="FIGURE 21. Internal sac. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) Ligyrus (Anagrylius) moroni, D) L. (Ligyrodes) sallaei, E) L. (Ligyrus) allonasutus. Abbreviations:AL:Accessory lamellae, Br: Brush of setae, CL: Copulatory lamella, LSB: Lamellar spiny belt, Ra: Raspula." figureDoi="http://doi.org/10.5281/zenodo.7345085" httpUri="https://zenodo.org/record/7345085/files/figure.png" pageId="21" pageNumber="22">Fig. 21B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="62BAABA6FD4AFF94AFF61BE4FC08BF3C" pageId="21" pageNumber="22" type="diagnosis">
|
||
<paragraph id="2A1FF82DFD4AFF94AFF61BE4FC08BF3C" blockId="21.[151,1437,150,1617]" pageId="21" pageNumber="22">
|
||
<emphasis id="18D4243FFD4AFF94AFF61BE4FEFCBEAD" bold="true" box="[199,322,690,716]" pageId="21" pageNumber="22">Diagnosis.</emphasis>
|
||
It is the only known species in the genus and can be differentiated from other
|
||
<taxonomicName id="EDA083AEFD4AFF94AB8E1BE5FAF0BEAC" authorityName="Endrodi" authorityYear="1966" box="[1215,1358,691,717]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="79" phylum="Arthropoda" rank="tribe" tribe="Pentodontini">Pentodontini</taxonomicName>
|
||
by the combination of the following characters: clypeus trapezoidal, clypeal teeth small, frontoclypeus with tumescences (
|
||
<figureCitation id="B29BE4A8FD4AFF94AFAE1BADFF48BF74" box="[159,246,763,789]" captionStart="FIGURE 1" captionStartId="81.[151,250,1535,1559]" captionTargetBox="[151,1436,181,1511]" captionTargetId="figure-17@81.[151,1436,181,1511]" captionTargetPageId="81" captionText="FIGURE 1. Head. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) Ligyrus (Ligyrodes) peruvianus, D) L. (Anagrylius) moroni, E) L. (Ligyrus) ruginasus, F) L. (Ligyrus) nasutus, G) L. (Ligyrus) bidentulus, H) Tomarus adoceteus, I) T. maimon, J) T. pullus, K) T. pumilus." figureDoi="http://doi.org/10.5281/zenodo.7345011" httpUri="https://zenodo.org/record/7345011/files/figure.png" pageId="21" pageNumber="22">Fig. 1B</figureCitation>
|
||
); apex of mesotibia and metatibia dentate (
|
||
<figureCitation id="B29BE4A8FD4AFF94ADE81BADFCDCBF74" box="[729,866,763,789]" captionStart="FIGURE 12" captionStartId="88.[151,250,1921,1945]" captionTargetBox="[237,1351,1055,1897]" captionTargetId="figure-89@88.[233,1355,1052,1897]" captionTargetPageId="88" captionText="FIGURE 12. Mesotibia (A–F). A) Euligyrus similis, B) Proculigyrus cicatricosus, C) Ligyrus (Anagrylius) moroni, D) L. (Ligyrodes) sallaei; E) L. (Ligyrus) allonasutus, F) L. (Ligyrus) villosus. Metatibia (G–L). G) Euligyrus similis, H) Proculigyrus cicatricosus, I) Ligyrus (Anagrylius) moroni, J) L. (Ligyrodes) sallaei, K) L. (Ligyrus) allonasutus, L) L. (Ligyrus) villosus." figureDoi="http://doi.org/10.5281/zenodo.7345053" httpUri="https://zenodo.org/record/7345053/files/figure.png" pageId="21" pageNumber="22">Fig. 12B, H</figureCitation>
|
||
); mentum subquadrate (
|
||
<figureCitation id="B29BE4A8FD4AFF94AB411BADFB7BBF74" box="[1136,1221,763,789]" captionStart="FIGURE 2" captionStartId="82.[151,250,1584,1608]" captionTargetBox="[151,1436,181,1548]" captionTargetId="figure-24@82.[151,1436,181,1560]" captionTargetPageId="82" captionText="FIGURE 2. Labium. A) Ligyrus (Anagrylius) moroni, B) Ligyrus (Ligyrus) allonasutus, C) L. (Ligyrus) gibbosus, D) L. (Ligyrus) villosus, E) L. (Ligyrodes) relictus, F) Proculigyrus cicatricosus, G) Euligyrus ebenus, H) E. similis, I) Tomarus gyas, J) Tomarus selanderi, K) Tomarus subtropicus. Abbreviations: LL: Ligular lobes, LP: Labial palpus, M: Mentum, S: Suture." figureDoi="http://doi.org/10.5281/zenodo.7345015" httpUri="https://zenodo.org/record/7345015/files/figure.png" pageId="21" pageNumber="22">Fig. 2F</figureCitation>
|
||
); protarsus of male simple; pronotum and pygidium with deep, large punctures; and parameres with wide, acute medial teeth and an apical long projection directed outwards on each side (
|
||
<figureCitation id="B29BE4A8FD4AFF94ADCA1A15FCD5BF3C" box="[763,875,834,861]" captionStart="FIGURE 15" captionStartId="90.[151,250,1535,1559]" captionTargetBox="[154,1430,183,1509]" captionTargetId="figure-24@90.[151,1436,181,1512]" captionTargetPageId="90" captionText="FIGURE 15. Parameres in frontal view. A) Euligyrus ebenus, B) E. similis, C) Proculigyrus cicatricosus, D) Ligyrus (Anagrylius) cuniculus, E) L. (Anagrylius) moroni, F) L. (Ligyrodes) peruvianus, G) L. (Ligyrodes) relictus, H) L. (Ligyrodes) sallaei, I) L. (Ligyrus) allonasutus, J) L. (Ligyrus) bidentulus, K) L. (Ligyrus) burmeisteri, L) Ligyrus (Ligyrus) fossor, M) L. (Ligyrus) gibbosus, N) L. (Ligyrus) nasutus, O) L. (Ligyrus) neglectus, P) L. (Ligyrus) paranaensis, Q) L. (Ligyrus) peninsularis, R) L. (Ligyrus) rubripes, S) L. (Ligyrus) ruginasus, T) L. (Ligyrus) villosus." figureDoi="http://doi.org/10.5281/zenodo.7345063" httpUri="https://zenodo.org/record/7345063/files/figure.png" pageId="21" pageNumber="22">Figs. 15C</figureCitation>
|
||
,
|
||
<figureCitation id="B29BE4A8FD4AFF94AC461A15FC17BF3C" box="[887,937,835,861]" captionStart="FIGURE 17" captionStartId="92.[151,250,1215,1239]" captionTargetBox="[155,1430,188,1184]" captionTargetId="figure-24@92.[151,1436,181,1191]" captionTargetPageId="92" captionText="FIGURE 17. Parameres in lateral view. A) Euligyrus ebenus, B) E. similis, C) Proculigyrus cicatricosus, D) Ligyrus (Anagrylius) cuniculus; E) L. (Anagrylius) moroni, F) L. (Ligyrodes) relictus, G) L. (Ligyrodes) sallaei." figureDoi="http://doi.org/10.5281/zenodo.7345073" httpUri="https://zenodo.org/record/7345073/files/figure.png" pageId="21" pageNumber="22">17C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="62BAABA6FD4AFF94AFF61A30FB4BBA30" pageId="21" pageNumber="22" type="distribution">
|
||
<paragraph id="2A1FF82DFD4AFF94AFF61A30FCD7BFC4" blockId="21.[151,1437,150,1617]" pageId="21" pageNumber="22">
|
||
<emphasis id="18D4243FFD4AFF94AFF61A30FEDFBFE1" bold="true" box="[199,353,870,896]" pageId="21" pageNumber="22">Distribution.</emphasis>
|
||
<collectingCountry id="52B7B8BDFD4AFF94AE571A31FE00BFE0" box="[358,446,871,897]" name="Mexico" pageId="21" pageNumber="22">Mexico</collectingCountry>
|
||
,
|
||
<collectingCountry id="52B7B8BDFD4AFF94AEF91A31FDFFBFE0" box="[456,577,871,897]" name="Guatemala" pageId="21" pageNumber="22">Guatemala</collectingCountry>
|
||
(
|
||
<bibRefCitation id="4E3185DCFD4AFF94AD7E1A31FC8FBFE0" author="Ratcliffe, B. C. & Cave, R. D. & Cano, E. B." box="[591,817,871,897]" pageId="21" pageNumber="22" pagination="1 - 666" refId="ref93157" refString="Ratcliffe, B. C., Cave, R. D. & Cano, E. B. (2013) The dynastine scarab beetles of Mexico, Guatemala, and Belize (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum, 27, 1 - 666." type="journal article" year="2013">
|
||
Ratcliffe
|
||
<emphasis id="18D4243FFD4AFF94AD891A31FD56BFE0" box="[696,744,871,897]" italics="true" pageId="21" pageNumber="22">et al</emphasis>
|
||
. 2013
|
||
</bibRefCitation>
|
||
),
|
||
<collectingCountry id="52B7B8BDFD4AFF94AC721A31FC78BFE0" box="[835,966,871,897]" name="El Salvador" pageId="21" pageNumber="22">El Salvador</collectingCountry>
|
||
,
|
||
<collectingCountry id="52B7B8BDFD4AFF94ACE11A31FB83BFE0" box="[976,1085,871,897]" name="Honduras" pageId="21" pageNumber="22">Honduras</collectingCountry>
|
||
(
|
||
<bibRefCitation id="4E3185DCFD4AFF94AB7B1A31FAF4BFE0" author="Ratcliffe, B. C. & Cave, R. D." box="[1098,1354,871,897]" pageId="21" pageNumber="22" pagination="1 - 424" refId="ref92768" refString="Ratcliffe, B. C. & Cave, R. D. (2006) The dynastine scarab beetles of Honduras, Nicaragua, and El Salvador. Bulletin of the University of Nebraska State Museum, 21, 1 - 424." type="journal article" year="2006">Ratcliffe & Cave 2006</bibRefCitation>
|
||
),
|
||
<collectingCountry id="52B7B8BDFD4AFF94AA6D1A31FF74BFC4" name="Costa Rica" pageId="21" pageNumber="22">Costa Rica</collectingCountry>
|
||
(
|
||
<bibRefCitation id="4E3185DCFD4AFF94AFE81ADDFEC1BFC4" author="Ratcliffe, B. C." box="[217,383,907,933]" pageId="21" pageNumber="22" pagination="1 - 506" refId="ref92614" refString="Ratcliffe, B. C. (2003) The dynastine scarab beetles of Costa Rica and Panama. Bulletin of the University of Nebraska State Museum, 16, 1 - 506." type="journal article" year="2003">Ratcliffe 2003</bibRefCitation>
|
||
), and
|
||
<collectingCountry id="52B7B8BDFD4AFF94AEF31ADDFD8EBFC4" box="[450,560,907,933]" name="Colombia" pageId="21" pageNumber="22">Colombia</collectingCountry>
|
||
(
|
||
<bibRefCitation id="4E3185DCFD4AFF94AD0E1ADDFCE0BFC4" author="Lopez-Garcia, M. M. & Gasca-Alvarez, H. J. & Amat-Garcia, G." box="[575,862,906,933]" pageId="21" pageNumber="22" pagination="451 - 492" refId="ref91458" refString="Lopez-Garcia, M. M., Gasca-Alvarez, H. J. & Amat-Garcia, G. (2015) The scarab beetle tribe Pentodontini (Coleoptera: Scarabaeidae: Dynastinae) of Colombia: taxonomy, natural history, and distribution. Zootaxa, 4048 (4), 451 - 492. https: // doi. org / 10.11646 / zootaxa. 4048.4.1" type="journal article" year="2015">
|
||
López-García
|
||
<emphasis id="18D4243FFD4AFF94ADD31ADDFCADBFC4" box="[738,787,907,933]" italics="true" pageId="21" pageNumber="22">et al</emphasis>
|
||
. 2015
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="2A1FF82DFD4AFF94AFF61AF8FE45BA68" blockId="21.[151,1437,150,1617]" pageId="21" pageNumber="22">
|
||
<emphasis id="18D4243FFD4AFF94AFF61AF8FE43BFA9" bold="true" box="[199,509,942,969]" pageId="21" pageNumber="22">
|
||
Locality records (
|
||
<figureCitation id="B29BE4A8FD4AFF94AEAB1AF8FE4EBFA9" box="[410,496,942,968]" captionStart="FIGURE 40" captionStartId="110.[151,250,1952,1976]" captionTargetBox="[163,1434,994,1923]" captionTargetId="figure-41@110.[153,1434,994,1929]" captionTargetPageId="110" captionText="FIGURE 40. Distributional records of Proculigyrus cicatricosus." figureDoi="http://doi.org/10.5281/zenodo.7345150" httpUri="https://zenodo.org/record/7345150/files/figure.png" pageId="21" pageNumber="22">Fig. 40</figureCitation>
|
||
).
|
||
</emphasis>
|
||
33 examined specimens from FSCA, ICN, IEXA, UNSM, USNM, ZMHB. Some records from
|
||
<bibRefCitation id="4E3185DCFD4AFF94AE021A85FE55BF8C" author="Ratcliffe, B. C." box="[307,491,979,1005]" pageId="21" pageNumber="22" pagination="1 - 506" refId="ref92614" refString="Ratcliffe, B. C. (2003) The dynastine scarab beetles of Costa Rica and Panama. Bulletin of the University of Nebraska State Museum, 16, 1 - 506." type="journal article" year="2003">Ratcliffe (2003)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="4E3185DCFD4AFF94AEC81A85FCADBF8C" author="Ratcliffe, B. C. & Cave, R. D." box="[505,787,979,1005]" pageId="21" pageNumber="22" pagination="1 - 424" refId="ref92768" refString="Ratcliffe, B. C. & Cave, R. D. (2006) The dynastine scarab beetles of Honduras, Nicaragua, and El Salvador. Bulletin of the University of Nebraska State Museum, 21, 1 - 424." type="journal article" year="2006">Ratcliffe & Cave (2006)</bibRefCitation>
|
||
, and
|
||
<bibRefCitation id="4E3185DCFD4AFF94AC631A85FBF1BF8C" author="Ratcliffe, B. C. & Cave, R. D. & Cano, E. B." box="[850,1103,979,1005]" pageId="21" pageNumber="22" pagination="1 - 666" refId="ref93157" refString="Ratcliffe, B. C., Cave, R. D. & Cano, E. B. (2013) The dynastine scarab beetles of Mexico, Guatemala, and Belize (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum, 27, 1 - 666." type="journal article" year="2013">
|
||
Ratcliffe
|
||
<emphasis id="18D4243FFD4AFF94AC8C1A85FC46BF8C" box="[957,1016,979,1005]" italics="true" pageId="21" pageNumber="22">et al.</emphasis>
|
||
(2013)
|
||
</bibRefCitation>
|
||
.
|
||
<emphasis id="18D4243FFD4AFF94AB6D1A84FEAAB871" bold="true" pageId="21" pageNumber="22">
|
||
<collectingCountry id="52B7B8BDFD4AFF94AB6D1A84FB47BF8D" box="[1116,1273,978,1004]" name="Colombia" pageId="21" pageNumber="22">COLOMBIA</collectingCountry>
|
||
(1).
|
||
<collectingRegion id="E86436CFFD4AFF94AA021A85FF5DB871" country="Colombia" name="Valle del Cauca" pageId="21" pageNumber="22">Valle del Cauca</collectingRegion>
|
||
(1):
|
||
</emphasis>
|
||
Buenaventura, Río San Juan.
|
||
<emphasis id="18D4243FFD4AFF94AD541AA0FC43B871" bold="true" box="[613,1021,1014,1041]" pageId="21" pageNumber="22">
|
||
<collectingCountry id="52B7B8BDFD4AFF94AD541AA0FCB1B871" box="[613,783,1014,1040]" name="Costa Rica" pageId="21" pageNumber="22">COSTA RICA</collectingCountry>
|
||
(1).
|
||
<collectingRegion id="E86436CFFD4AFF94AC751AA1FC72B870" box="[836,972,1015,1041]" country="Costa Rica" name="Puntarenas" pageId="21" pageNumber="22">Puntarenas</collectingRegion>
|
||
(1):
|
||
</emphasis>
|
||
Rancho Quemado.
|
||
<emphasis id="18D4243FFD4AFF94ABEA1AA1FE1AB855" bold="true" pageId="21" pageNumber="22">
|
||
<collectingCountry id="52B7B8BDFD4AFF94ABEA1AA1FA22B871" box="[1243,1436,1014,1041]" name="El Salvador" pageId="21" pageNumber="22">EL SALVADOR</collectingCountry>
|
||
(23).
|
||
<collectingRegion id="E86436CFFD4AFF94AFE61D4CFED1B855" box="[215,367,1050,1076]" country="El Salvador" name="Ahuachapan" pageId="21" pageNumber="22">Ahuachapan</collectingRegion>
|
||
(3):
|
||
</emphasis>
|
||
Caserío San Miguelito; Parque Nacional El Imposible.
|
||
<emphasis id="18D4243FFD4AFF94AB1E1D4DFAB9B855" bold="true" box="[1071,1287,1050,1077]" pageId="21" pageNumber="22">
|
||
<collectingRegion id="E86436CFFD4AFF94AB1E1D4DFB7AB855" box="[1071,1220,1050,1077]" country="El Salvador" name="La Libertad" pageId="21" pageNumber="22">La Libertad</collectingRegion>
|
||
(16):
|
||
</emphasis>
|
||
San Andrés; Santa Tecla.
|
||
<emphasis id="18D4243FFD4AFF94AE171D68FE6DB839" bold="true" box="[294,467,1086,1112]" pageId="21" pageNumber="22">
|
||
<collectingRegion id="E86436CFFD4AFF94AE171D68FE1DB839" box="[294,419,1086,1112]" country="El Salvador" name="Santa Ana" pageId="21" pageNumber="22">Santa Ana</collectingRegion>
|
||
(3):
|
||
</emphasis>
|
||
Hacienda Montecristo, Cerro Miramundo, Metapán.
|
||
<emphasis id="18D4243FFD4AFF94AB1A1D68FB45B839" bold="true" box="[1067,1275,1086,1112]" pageId="21" pageNumber="22">
|
||
<collectingRegion id="E86436CFFD4AFF94AB1A1D68FB74B839" box="[1067,1226,1086,1112]" country="El Salvador" name="San Salvador" pageId="21" pageNumber="22">San Salvador</collectingRegion>
|
||
(1):
|
||
</emphasis>
|
||
<collectingRegion id="E86436CFFD4AFF94AA331D69FA27B838" box="[1282,1433,1087,1113]" inferredBySuffix="true" pageId="21" pageNumber="22">San Salvador</collectingRegion>
|
||
.
|
||
<emphasis id="18D4243FFD4AFF94AFA61D34FDE3B81D" bold="true" box="[151,605,1122,1149]" pageId="21" pageNumber="22">
|
||
<collectingCountry id="52B7B8BDFD4AFF94AFA61D34FEF2B81D" box="[151,332,1122,1148]" name="Guatemala" pageId="21" pageNumber="22">GUATEMALA</collectingCountry>
|
||
(31).
|
||
<collectingRegion id="E86436CFFD4AFF94AEA11D34FD93B81C" box="[400,557,1122,1149]" country="Guatemala" name="Alta Verapaz" pageId="21" pageNumber="22">Alta Verapaz</collectingRegion>
|
||
(1):
|
||
</emphasis>
|
||
Finca Chulac, Mayanaquel Mining Company.
|
||
<emphasis id="18D4243FFD4AFF94AB581D34FA85B81D" bold="true" box="[1129,1339,1122,1149]" pageId="21" pageNumber="22">
|
||
<collectingRegion id="E86436CFFD4AFF94AB581D34FAB5B81C" box="[1129,1291,1122,1149]" country="Guatemala" name="Baja Verapaz" pageId="21" pageNumber="22">Baja Verapaz</collectingRegion>
|
||
(7):
|
||
</emphasis>
|
||
Biotopo de Quetzal; Carretera a Chilasco; Purulha.
|
||
<emphasis id="18D4243FFD4AFF94AD4B1DD0FC82B8C1" bold="true" box="[634,828,1158,1184]" pageId="21" pageNumber="22">
|
||
<collectingRegion id="E86436CFFD4AFF94AD4B1DD0FCB7B8C1" box="[634,777,1158,1184]" country="Guatemala" name="Chiquimula" pageId="21" pageNumber="22">Chiquimula</collectingRegion>
|
||
(1):
|
||
</emphasis>
|
||
Las Presas, arriba de Concepción Las Minas.
|
||
<emphasis id="18D4243FFD4AFF94AA771DD1FEB6B8A5" bold="true" pageId="21" pageNumber="22">
|
||
<collectingRegion id="E86436CFFD4AFF94AA771DD1FF68B8A4" country="Guatemala" name="El Progreso" pageId="21" pageNumber="22">El Progreso</collectingRegion>
|
||
(2):
|
||
</emphasis>
|
||
El Rancho; Finca El Riscosa (
|
||
<quantity id="ED5855C8FD4AFF94AD501DFCFD25B8A4" box="[609,667,1194,1221]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="21" pageNumber="22" unit="km" value="5.0">5 km</quantity>
|
||
N El Pacayal).
|
||
<emphasis id="18D4243FFD4AFF94AC7C1DFCFC50B8A5" bold="true" box="[845,1006,1194,1220]" pageId="21" pageNumber="22">
|
||
<collectingRegion id="E86436CFFD4AFF94AC7C1DFCFC03B8A5" box="[845,957,1194,1220]" country="Guatemala" name="Escuintla" pageId="21" pageNumber="22">Escuintla</collectingRegion>
|
||
(3):
|
||
</emphasis>
|
||
Peñón de Siquinalá; Santa Lucía Cotzumalguapa.
|
||
<emphasis id="18D4243FFD4AFF94AE1D1D99FDA4B889" bold="true" box="[300,538,1230,1257]" pageId="21" pageNumber="22">
|
||
<collectingRegion id="E86436CFFD4AFF94AE1D1D99FE56B888" box="[300,488,1231,1257]" country="Guatemala" name="Huehuetenango" pageId="21" pageNumber="22">Huehuetenango</collectingRegion>
|
||
(1):
|
||
</emphasis>
|
||
Barillas, Malpaís.
|
||
<emphasis id="18D4243FFD4AFF94ADDF1D99FCD7B889" bold="true" box="[750,873,1230,1257]" pageId="21" pageNumber="22">
|
||
<collectingRegion id="E86436CFFD4AFF94ADDF1D99FC87B888" box="[750,825,1231,1257]" country="Guatemala" name="Izabal" pageId="21" pageNumber="22">Izabal</collectingRegion>
|
||
(5):
|
||
</emphasis>
|
||
Cerro Negro Norte (
|
||
<quantity id="ED5855C8FD4AFF94AB671D99FB33B888" box="[1110,1165,1231,1257]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="21" pageNumber="22" unit="km" value="1.0">1 km</quantity>
|
||
W Honduras); Morales, Finca Firmeza.
|
||
<emphasis id="18D4243FFD4AFF94AE781DA4FD96B96D" bold="true" box="[329,552,1266,1292]" pageId="21" pageNumber="22">
|
||
<collectingRegion id="E86436CFFD4AFF94AE781DA4FE48B96D" box="[329,502,1266,1292]" country="Guatemala" name="Suchitepequez" pageId="21" pageNumber="22">Suchitepequez</collectingRegion>
|
||
(1):
|
||
</emphasis>
|
||
Finca El Vesubio (
|
||
<quantity id="ED5855C8FD4AFF94AC331DA5FCF5B96C" box="[770,843,1267,1293]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="21" pageNumber="22" unit="km" value="12.0">12 km</quantity>
|
||
N Patulul).
|
||
<emphasis id="18D4243FFD4AFF94ACE91DA5FBCFB96D" bold="true" box="[984,1137,1266,1293]" pageId="21" pageNumber="22">
|
||
<collectingRegion id="E86436CFFD4AFF94ACE91DA5FB8FB96C" box="[984,1073,1267,1293]" country="Guatemala" name="Zacapa" pageId="21" pageNumber="22">Zacapa</collectingRegion>
|
||
(10):
|
||
</emphasis>
|
||
Finca Santa Cruz, Marble Quarry road (NE Teculután); La Union (
|
||
<quantity id="ED5855C8FD4AFF94AD681C40FD17B950" box="[601,681,1302,1329]" metricMagnitude="3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="21" pageNumber="22" unit="km" value="5.5" valueMax="6.0" valueMin="5.0">5-6 km</quantity>
|
||
S); road to San Lorenzo Marmol Mine (NW Tuculután); Teculután.
|
||
<emphasis id="18D4243FFD4AFF94AFA61C6CFDA2B935" bold="true" box="[151,540,1338,1365]" pageId="21" pageNumber="22">
|
||
<collectingCountry id="52B7B8BDFD4AFF94AFA61C6CFE89B935" box="[151,311,1338,1364]" name="Honduras" pageId="21" pageNumber="22">HONDURAS</collectingCountry>
|
||
(28).
|
||
<collectingRegion id="E86436CFFD4AFF94AE4B1C6CFE52B934" box="[378,492,1338,1365]" country="Honduras" name="Atlantida" pageId="21" pageNumber="22">Atlántida</collectingRegion>
|
||
(2):
|
||
</emphasis>
|
||
La Ceiba.
|
||
<emphasis id="18D4243FFD4AFF94ADA41C6CFCEAB935" bold="true" box="[661,852,1338,1364]" pageId="21" pageNumber="22">
|
||
<collectingRegion id="E86436CFFD4AFF94ADA41C6CFC9AB935" box="[661,804,1338,1364]" country="Honduras" name="Comayagua" pageId="21" pageNumber="22">Comayagua</collectingRegion>
|
||
(3):
|
||
</emphasis>
|
||
El Taladro; Siguatepeque.
|
||
<emphasis id="18D4243FFD4AFF94ABB01C6CFABFB935" bold="true" box="[1153,1281,1338,1365]" pageId="21" pageNumber="22">
|
||
<collectingRegion id="E86436CFFD4AFF94ABB01C6CFB6EB934" box="[1153,1232,1338,1365]" country="Honduras" name="Cortes" pageId="21" pageNumber="22">Cortés</collectingRegion>
|
||
(6):
|
||
</emphasis>
|
||
Cofradia (
|
||
<quantity id="ED5855C8FD4AFF94AA491C6DFF05B918" metricMagnitude="3" metricUnit="m" metricValue="9.3" pageId="21" pageNumber="22" unit="km" value="9.3">9.3 km</quantity>
|
||
NW); Lago de Yojoa; Parque Nacional Cusuco.
|
||
<emphasis id="18D4243FFD4AFF94ADDC1C09FC14B919" bold="true" box="[749,938,1374,1401]" pageId="21" pageNumber="22">
|
||
<collectingRegion id="E86436CFFD4AFF94ADDC1C09FCD4B918" box="[749,874,1374,1401]" country="Honduras" name="El Paraiso" pageId="21" pageNumber="22">El Paraíso</collectingRegion>
|
||
(11):
|
||
</emphasis>
|
||
Capire (
|
||
<quantity id="ED5855C8FD4AFF94AB3E1C09FBDEB918" box="[1039,1120,1375,1401]" metricMagnitude="3" metricUnit="m" metricValue="8.3" pageId="21" pageNumber="22" unit="km" value="8.3">8.3 km</quantity>
|
||
SE);
|
||
<collectingRegion id="E86436CFFD4AFF94AB921C09FAA5B918" box="[1187,1307,1375,1401]" inferredBySuffix="true" pageId="21" pageNumber="22">El Paraíso</collectingRegion>
|
||
.
|
||
<emphasis id="18D4243FFD4AFF94AA191C08FE8BB9FD" bold="true" pageId="21" pageNumber="22">
|
||
<collectingRegion id="E86436CFFD4AFF94AA191C08FEBAB9FC" country="Honduras" name="Francisco Morazan" pageId="21" pageNumber="22">Francisco Morazán</collectingRegion>
|
||
(1):
|
||
</emphasis>
|
||
Cerro Uyuca.
|
||
<emphasis id="18D4243FFD4AFF94AEE91CD4FDD0B9FD" bold="true" box="[472,622,1410,1436]" pageId="21" pageNumber="22">
|
||
<collectingRegion id="E86436CFFD4AFF94AEE91CD4FD83B9FD" box="[472,573,1410,1436]" country="Honduras" name="Olancho" pageId="21" pageNumber="22">Olancho</collectingRegion>
|
||
(2):
|
||
</emphasis>
|
||
Montaña del Malacate.
|
||
<emphasis id="18D4243FFD4AFF94AC4A1CD5FC5BB9FD" bold="true" box="[891,997,1410,1437]" pageId="21" pageNumber="22">
|
||
<collectingRegion id="E86436CFFD4AFF94AC4A1CD5FC0AB9FC" box="[891,948,1411,1437]" country="Honduras" name="Yoro" pageId="21" pageNumber="22">Yoro</collectingRegion>
|
||
(3):
|
||
</emphasis>
|
||
Parque Nacional Pico Pijol.
|
||
<emphasis id="18D4243FFD4AFF94AA171CD4FEDBB9A1" bold="true" pageId="21" pageNumber="22">
|
||
<collectingCountry id="52B7B8BDFD4AFF94AA171CD4FA22B9FD" box="[1318,1436,1410,1436]" name="Mexico" pageId="21" pageNumber="22">MEXICO</collectingCountry>
|
||
(11).
|
||
<collectingRegion id="E86436CFFD4AFF94AFE31CF0FE8AB9A1" box="[210,308,1446,1472]" country="Mexico" name="Chiapas" pageId="21" pageNumber="22">Chiapas</collectingRegion>
|
||
(5):
|
||
</emphasis>
|
||
Cañón El Chorreadero (
|
||
<quantity id="ED5855C8FD4AFF94AD491CF0FD0CB9A0" box="[632,690,1446,1473]" metricMagnitude="3" metricUnit="m" metricValue="8.04672" pageId="21" pageNumber="22" unit="mi" value="5.0">5 mi.</quantity>
|
||
E Chiapa de Corzo); El Aguacero (
|
||
<quantity id="ED5855C8FD4AFF94AB711CF1FB36B9A0" box="[1088,1160,1447,1473]" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="21" pageNumber="22" unit="km" value="16.0">16 km</quantity>
|
||
W Ocozocautla); Suchiapa (
|
||
<quantity id="ED5855C8FD4AFF94AFFE1C9CFEB4B984" box="[207,266,1482,1509]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="21" pageNumber="22" unit="km" value="5.0">5 km</quantity>
|
||
SE).
|
||
<emphasis id="18D4243FFD4AFF94AE7A1C9CFE69B985" bold="true" box="[331,471,1482,1508]" pageId="21" pageNumber="22">
|
||
<collectingRegion id="E86436CFFD4AFF94AE7A1C9CFE1BB985" box="[331,421,1482,1508]" country="Mexico" name="Oaxaca" pageId="21" pageNumber="22">Oaxaca</collectingRegion>
|
||
(2):
|
||
</emphasis>
|
||
Cuicatlán (
|
||
<quantity id="ED5855C8FD4AFF94AD6D1C9CFD18B984" box="[604,678,1482,1509]" metricMagnitude="4" metricUnit="m" metricValue="2.414016" pageId="21" pageNumber="22" unit="mi" value="15.0">15 mi.</quantity>
|
||
N); Salina Cruz (
|
||
<quantity id="ED5855C8FD4AFF94AC401C9CFC12B984" box="[881,940,1482,1509]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="21" pageNumber="22" unit="km" value="5.0">5 km</quantity>
|
||
N).
|
||
<emphasis id="18D4243FFD4AFF94ACD01C9DFBCAB985" bold="true" box="[993,1140,1482,1509]" pageId="21" pageNumber="22">
|
||
<collectingRegion id="E86436CFFD4AFF94ACD01C9DFBFCB984" box="[993,1090,1483,1509]" country="Mexico" name="Tabasco" pageId="21" pageNumber="22">Tabasco</collectingRegion>
|
||
(1):
|
||
</emphasis>
|
||
Villa Hermosa.
|
||
<emphasis id="18D4243FFD4AFF94AA1E1C9DFF7FBA69" bold="true" pageId="21" pageNumber="22">
|
||
<collectingRegion id="E86436CFFD4AFF94AA1E1C9DFA22B984" box="[1327,1436,1483,1509]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="21" pageNumber="22">Veracruz</collectingRegion>
|
||
(3):
|
||
</emphasis>
|
||
Puente Nacional; Veracruz.
|
||
</paragraph>
|
||
<paragraph id="2A1FF82DFD4AFF94AFF61F45FB4BBA30" blockId="21.[151,1437,150,1617]" pageId="21" pageNumber="22">
|
||
<emphasis id="18D4243FFD4AFF94AFF61F45FE3BBA4D" bold="true" box="[199,389,1554,1581]" pageId="21" pageNumber="22">Natural history.</emphasis>
|
||
Specimens have been found mainly in cloud forests up to
|
||
<quantity id="ED5855C8FD4AFF94AB291F45FBCABA4C" box="[1048,1140,1555,1581]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="21" pageNumber="22" unit="m" value="2300.0">2,300 m</quantity>
|
||
but also in tropical humid forests near sea level. Most individuals have been collected with light traps between April and May.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |