plazi/treatments-xml
2
0
Fork 0
Code Issues Pull requests Releases Activity
treatments-xml/data/A2/09/49/A209493BFD1BFFC4AFA61BADFE7BBCB4.xml
maintenance 618489d4ad added A2
2024-06-21 12:46:14 +02:00

283 lines
36 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="E975066D64707BFC9F17BF5BDBA0EEE7" ID-CLB-Dataset="202083" ID-DOI="10.11646/zootaxa.5211.1.1" ID-GBIF-Dataset="bb97b9dd-2acc-4244-8f30-72bef07b9ed9" ID-ISSN="1175-5326" ID-Zenodo-Dep="7345007" ID-ZooBank="5754769C-B747-4714-BDD9-7D5509D48BEB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1669101340847" checkinUser="plazi" docAuthor="López-García, Margarita M. &amp; Deloya, Cuauhtémoc" docDate="2022" docId="A209493BFD1BFFC4AFA61BADFE7BBCB4" docLanguage="en" docName="zootaxa.5211.1.1.pdf" docOrigin="Zootaxa 5211 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5211.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Tomarus roigjunenti Neita &amp; Ratcliffe 2017" docType="treatment" docVersion="9" lastPageNumber="70" masterDocId="5E303143FD5FFF81AF311956FFBEBC61" masterDocTitle="Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision" masterLastPageNumber="119" masterPageNumber="1" pageNumber="69" updateTime="1699396337306" updateUser="plazi">
<mods:mods id="431D16254F1C259F5669556551AE3AF2" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B1B5D547F4AC2798886208D9F35B4A8D">
<mods:title id="366EFE6A1288FA98550EDF2FCD9FCF22">Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision</mods:title>
</mods:titleInfo>
<mods:name id="78315A64D07CC19EDC27D555C7A0FB6A" type="personal">
<mods:role id="6BCFB587C832195EC7DAF9E141F4DD84">
<mods:roleTerm id="1C58846EE7BFDC0185C92D6BD3434907">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D867CAD7F6DFFE69BD326B4F90E95EEF">López-García, Margarita M.</mods:namePart>
<mods:nameIdentifier id="1F53D07D269D675ECB64CA37B697F05A" type="ORCID">0000-0003-2796-2931</mods:nameIdentifier>
<mods:nameIdentifier id="479710F07B96845151D819442C1AD06B" type="email">margaralopezg@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="FDE89CE5130A6AA62B981BBD5CC9A8DB" type="personal">
<mods:role id="6AC5D4763B68879771D4EA88103B0BFC">
<mods:roleTerm id="027AFB4D19133C4091C4EFDE78464E46">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C1DAA8BA187E4659D83C4703746828A1">Deloya, Cuauhtémoc</mods:namePart>
<mods:nameIdentifier id="E0367F1FBEED819C419906E51F234414" type="ORCID">0000-0002-4774-140X</mods:nameIdentifier>
<mods:nameIdentifier id="CD54E73D83696103CEDA23F615E69685" type="email">cuauhtemoc.deloya@inecol.mx</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="A85AC54C39C57A371B1B22A79088E7A9">text</mods:typeOfResource>
<mods:relatedItem id="3B093EF141B6A0F06EC0C9444977EC66" type="host">
<mods:titleInfo id="EAA9B2611C5A5B1A18025D366C1E2630">
<mods:title id="FE89F06242C5E9FE62B8BCCD3B251B2F">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="11FBA3F11854BFC8237DD42AE4858B33">
<mods:date id="862193AA9BEBFC1B6C9C80E305A0FCC7">2022</mods:date>
<mods:detail id="4DFB12580B6675A1AD5CF57BB5CC5CE3" type="pubDate">
<mods:number id="7D89853604CEE9BA2E9212CD8A0E7313">2022-11-22</mods:number>
</mods:detail>
<mods:detail id="9AFC5FC9D12D26B9999DFAF4E8E89B12" type="volume">
<mods:number id="CF2F56656D08684FF929566331728C2C">5211</mods:number>
</mods:detail>
<mods:detail id="B8FFD1B792D9A9246D5F6131AD39F5E7" type="issue">
<mods:number id="D52ADEF984155FAC9F60DBBD008560C9">1</mods:number>
</mods:detail>
<mods:extent id="22D4F1918D98F7B7DA33673FD10A113D" unit="page">
<mods:start id="EAD2CC5B4C7D5D5F9186ABEFA722A1DF">1</mods:start>
<mods:end id="5662D1CF88C3193808D9E3BFA21AA9C1">119</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="CFF5CBB81342D0FBC00D09E6742C0A70">
<mods:url id="FF16F45374D07B06445BFCAE3F2970BC">http://dx.doi.org/10.11646/zootaxa.5211.1.1</mods:url>
</mods:location>
<mods:classification id="8EBBABE5649936600734544D56FD2A96">journal article</mods:classification>
<mods:identifier id="D36ECF26EC0B8437E68F8EB2A384B390" type="CLB-Dataset">202083</mods:identifier>
<mods:identifier id="D5241428D6DF92EA249C758E575840B9" type="DOI">10.11646/zootaxa.5211.1.1</mods:identifier>
<mods:identifier id="36B22C7D8E0F6624A6F95107F59A23F4" type="GBIF-Dataset">bb97b9dd-2acc-4244-8f30-72bef07b9ed9</mods:identifier>
<mods:identifier id="F331ACCAC4D8EA9B77BF8EE02CDB1601" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="EFC9E3196F056F6EAD1B02F7E19655D3" type="Zenodo-Dep">7345007</mods:identifier>
<mods:identifier id="EEABD67F502D923D0E4ACB70DEFE12F2" type="ZooBank">5754769C-B747-4714-BDD9-7D5509D48BEB</mods:identifier>
</mods:mods>
<treatment id="A209493BFD1BFFC4AFA61BADFE7BBCB4" ID-DOI="http://doi.org/10.5281/zenodo.7383787" ID-GBIF-Taxon="205794515" ID-Zenodo-Dep="7383787" LSID="urn:lsid:plazi:treatment:A209493BFD1BFFC4AFA61BADFE7BBCB4" httpUri="http://treatment.plazi.org/id/A209493BFD1BFFC4AFA61BADFE7BBCB4" lastPageId="69" lastPageNumber="70" pageId="68" pageNumber="69">
<subSubSection id="62BAABA6FD1BFFC5AFA61BADFD09BF75" box="[151,695,762,789]" pageId="68" pageNumber="69" type="nomenclature">
<paragraph id="2A1FF82DFD1BFFC5AFA61BADFD09BF75" blockId="68.[151,695,762,825]" box="[151,695,762,789]" pageId="68" pageNumber="69">
<heading id="71574F41FD1BFFC5AFA61BADFD09BF75" bold="true" box="[151,695,762,789]" fontSize="11" level="2" pageId="68" pageNumber="69" reason="4">
<taxonomicName id="EDA083AEFD1BFFC5AFA61BADFD09BF75" authority="Neita &amp; Ratcliffe, 2017" authorityName="Neita &amp; Ratcliffe" authorityYear="2017" box="[151,695,762,789]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="roigjunenti">
<emphasis id="18D4243FFD1BFFC5AFA61BADFD09BF75" bold="true" box="[151,695,762,789]" pageId="68" pageNumber="69">
<emphasis id="18D4243FFD1BFFC5AFA61BADFE2EBF74" bold="true" box="[151,400,763,789]" italics="true" pageId="68" pageNumber="69">Tomarus roigjunenti</emphasis>
Neita &amp;
<bibRefCitation id="4E3185DCFD1BFFC5AD331BACFD09BF75" author="Ratcliffe, B. C. &amp; Cave, R. D." box="[514,695,762,788]" pageId="68" pageNumber="69" pagination="1 - 298" refId="ref92923" refString="Ratcliffe, B. C. &amp; Cave, R. D. (2017) The dynastine scarab beetles of the United States and Canada (Coleoptera: Scarabaeidae). Bulletin of the University of Nebraska State Museum, 30, 1 - 298." type="journal article" year="2017">Ratcliffe, 2017</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="62BAABA6FD1BFFC5AFA61A49FE5ABF58" box="[151,484,799,825]" pageId="68" pageNumber="69" type="description">
<paragraph id="2A1FF82DFD1BFFC5AFA61A49FE5ABF58" blockId="68.[151,695,762,825]" box="[151,484,799,825]" pageId="68" pageNumber="69">
(
<figureCitation id="B29BE4A8FD1BFFC5AFAE1A49FEBABF58" box="[159,260,799,825]" captionStart="FIGURE 9" captionStartId="87.[151,250,1074,1098]" captionTargetBox="[151,1436,181,1050]" captionTargetId="figure-17@87.[151,1436,181,1050]" captionTargetPageId="87" captionText="FIGURE 9. Pygidium of male. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) Ligyrus (Anagrylius) cuniculus, D) L. (Ligyrus) fossor, E) L. (Ligyrus) paranaensis, F) Tomarus bituberculatus, G) T. colombianus, H) T. maracaiboensis, I) T. pilcopataensis, J) T. pullus, K) T. roigjunenti, L) T. rostratus." figureDoi="http://doi.org/10.5281/zenodo.7345041" httpUri="https://zenodo.org/record/7345041/files/figure.png" pageId="68" pageNumber="69">Figs. 9K</figureCitation>
,
<figureCitation id="B29BE4A8FD1BFFC5AE3E1A49FEFCBF58" box="[271,322,799,825]" captionStart="FIGURE 16" captionStartId="91.[151,250,1633,1657]" captionTargetBox="[151,1426,189,1602]" captionTargetId="figure-17@91.[151,1436,181,1610]" captionTargetPageId="91" captionText="FIGURE 16. Parameres in frontal view. A) Tomarus adoceteus, B) T. amazonicus, C) T. bituberculatus, D) T. colombianus, E) T. discrepans, F) T. gyas, G) T. laevicollis, H) T. laticaudus, I) T. maimon, J) T. maracaiboensis, K) T. maternus, L) T. pilcopataensis, M) T. pullus, N) T. pumilus, O) T. roigjunenti, P) T. rostratus, Q) T. selanderi, R) T. subtropicus." figureDoi="http://doi.org/10.5281/zenodo.7345065" httpUri="https://zenodo.org/record/7345065/files/figure.png" pageId="68" pageNumber="69">16O</figureCitation>
,
<figureCitation id="B29BE4A8FD1BFFC5AE7C1A49FEC7BF58" box="[333,377,799,825]" captionStart="FIGURE 20" captionStartId="95.[151,250,1225,1249]" captionTargetBox="[159,1435,191,1201]" captionTargetId="figure-17@95.[151,1436,181,1202]" captionTargetPageId="95" captionText="FIGURE 20. Parameres in lateral view. A) Tomarus maracaiboensis, B) T. maternus, C) T. pilcopataensis, D) T. pullus, E) T. pumilus, F) T. roigjunenti, G) T. rostratus, H) T. selanderi, I) T. subtropicus." figureDoi="http://doi.org/10.5281/zenodo.7345083" httpUri="https://zenodo.org/record/7345083/files/figure.png" pageId="68" pageNumber="69">20F</figureCitation>
,
<figureCitation id="B29BE4A8FD1BFFC5AEB41A49FE0DBF58" box="[389,435,799,825]" captionStart="FIGURE 28" captionStartId="103.[151,250,1868,1892]" captionTargetBox="[159,1426,182,1839]" captionTargetId="figure-17@103.[151,1436,181,1844]" captionTargetPageId="103" captionText="FIGURE 28. Habitus of the species of Tomarus: A) Tomarus maracaiboensis, B) T. maternus, C) T. pilcopataensis, D) T. pullus, E) T. pumilus, F) T. roigjunenti, G) T. rostratus, H) T. selanderi, I) T. subtropicus. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7345107" httpUri="https://zenodo.org/record/7345107/files/figure.png" pageId="68" pageNumber="69">28F</figureCitation>
;
<figureCitation id="B29BE4A8FD1BFFC5AE8E1A49FE62BF58" box="[447,476,799,825]" captionStart="FIGURE 47" captionStartId="114.[151,250,1084,1108]" captionTargetBox="[203,1393,181,1051]" captionTargetId="figure-24@114.[195,1393,181,1061]" captionTargetPageId="114" captionText="FIGURE 47. Distributional records of Tomarus pilcopataensis, T. roigjunenti, and T. pumilus." figureDoi="http://doi.org/10.5281/zenodo.7345176" httpUri="https://zenodo.org/record/7345176/files/figure.png" pageId="68" pageNumber="69">47</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="62BAABA6FD1BFFC5AFA61A32FCD2BF1A" box="[151,876,867,891]" pageId="68" pageNumber="69" type="reference_group">
<paragraph id="2A1FF82DFD1BFFC5AFA61A32FCD2BF1A" blockId="68.[151,1437,867,1014]" box="[151,876,867,891]" pageId="68" pageNumber="69">
<treatmentCitationGroup id="0AB0DF03FD1BFFC5AFA61A32FCD2BF1A" box="[151,876,867,891]" pageId="68" pageNumber="69">
<treatmentCitation id="AB01DE3CFD1BFFC5AFA61A32FDC0BF1A" author="Ratcliffe, B. C. &amp; Cave, R. D." box="[151,638,867,891]" page="8" pageId="68" pageNumber="69" year="2017">
<taxonomicName id="EDA083AEFD1BFFC5AFA61A32FDC0BF1A" authority="Neita &amp; Ratcliffe, 2017: 8" authorityName="Neita &amp; Ratcliffe" authorityPageNumber="8" authorityYear="2017" box="[151,638,867,891]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="roigjunenti">
<emphasis id="18D4243FFD1BFFC5AFA61A32FED9BF1A" box="[151,359,868,891]" italics="true" pageId="68" pageNumber="69">Tomarus roigjunenti</emphasis>
Neita &amp;
<bibRefCitation id="4E3185DCFD1BFFC5AEF91A35FDC0BF1A" author="Ratcliffe, B. C. &amp; Cave, R. D." box="[456,638,867,891]" pageId="68" pageNumber="69" pagination="1 - 298" refId="ref92923" refString="Ratcliffe, B. C. &amp; Cave, R. D. (2017) The dynastine scarab beetles of the United States and Canada (Coleoptera: Scarabaeidae). Bulletin of the University of Nebraska State Museum, 30, 1 - 298." type="journal article" year="2017">Ratcliffe, 2017: 8</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. Original combination.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="62BAABA6FD1BFFC5AC421A35FC9DBF97" pageId="68" pageNumber="69" type="materials_examined">
<paragraph id="2A1FF82DFD1BFFC5AC421A35FC9DBF97" blockId="68.[151,1437,867,1014]" pageId="68" pageNumber="69">
Male
<typeStatus id="F51B468FFD1BFFC5AC9E1A35FBB6BF1A" box="[943,1032,867,891]" pageId="68" pageNumber="69" type="holotype">holotype</typeStatus>
<materialsCitation id="9AC8F270FD1BFFC5AB3E1A32FCA1BF97" ID-GBIF-Occurrence="3986696632" collectingDate="1961-03-01" collectionCode="USNM" collectorName="C. S. Reed" country="Argentina" location="Neuquen" pageId="68" pageNumber="69" specimenCount="6" stateProvince="Neuquen" typeStatus="paratype">
not examined. Male
<typeStatus id="F51B468FFD1BFFC5ABD21A32FA84BF1D" box="[1251,1338,868,892]" pageId="68" pageNumber="69" type="paratype">paratype</typeStatus>
(
<collectionCode id="4CB160E8FD1BFFC5AA781A35FA2BBF1A" box="[1353,1429,867,891]" country="USA" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="68" pageNumber="69" type="Museum">USNM</collectionCode>
) “
<collectingRegion id="E86436CFFD1BFFC5AFE11AD4FE90BFFB" box="[208,302,898,922]" country="Argentina" name="Neuquen" pageId="68" pageNumber="69">Neuquen</collectingRegion>
/ Capital Federal /
<date id="5E1EDEEDFD1BFFC5AEDD1AD4FDF3BFFB" box="[492,589,898,922]" pageId="68" pageNumber="69" value="1961-03-01">
<collectingDate id="4E5A2705FD1BFFC5AEDD1AD4FDF3BFFB" box="[492,589,898,922]" pageId="68" pageNumber="69" value="1961-03-01">1.iii.1961</collectingDate>
</date>
/ Dafuern //
<taxonomicName id="EDA083AEFD1BFFC5ADFB1AD4FCFBBFFB" baseAuthorityName="sensu Moron &amp; Grossi" baseAuthorityYear="2015" box="[714,837,898,922]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="genus">TOMARUS</taxonomicName>
/ ROIGJUNENTI / NEITA &amp; RATCLIFFE /
<typeStatus id="F51B468FFD1BFFC5AA3C1AD4FA2FBFFB" box="[1293,1425,898,922]" pageId="68" pageNumber="69" type="paratype">PARATYPE</typeStatus>
”. Female
<typeStatus id="F51B468FFD1BFFC5AE2A1AF7FECCBFD8" box="[283,370,929,953]" pageId="68" pageNumber="69" type="paratype">paratype</typeStatus>
(
<collectionCode id="4CB160E8FD1BFFC5AEB21AF7FE71BFD8" box="[387,463,929,953]" country="USA" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="68" pageNumber="69" type="Museum">USNM</collectionCode>
) “
<collectingRegion id="E86436CFFD1BFFC5AED91AF7FDF8BFD8" box="[488,582,929,953]" country="Argentina" name="Neuquen" pageId="68" pageNumber="69">Neuquen</collectingRegion>
/ Capital /
<date id="5E1EDEEDFD1BFFC5ADF31AF6FC9DBFD9" box="[706,803,928,952]" pageId="68" pageNumber="69" value="1961-03-01">
<collectingDate id="4E5A2705FD1BFFC5ADF31AF6FC9DBFD9" box="[706,803,928,952]" pageId="68" pageNumber="69" value="1961-03-01">1.iii.1961</collectingDate>
</date>
//
<taxonomicName id="EDA083AEFD1BFFC5AC751AF7FC01BFD8" baseAuthorityName="sensu Moron &amp; Grossi" baseAuthorityYear="2015" box="[836,959,929,953]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="genus">TOMARUS</taxonomicName>
/ ROIGJUNENTI / NEITA &amp; RATCLIFFE /
<typeStatus id="F51B468FFD1BFFC5AFF61AE9FEF5BFB6" box="[199,331,959,983]" pageId="68" pageNumber="69" type="paratype">PARATYPE</typeStatus>
”. Two female
<typeStatus id="F51B468FFD1BFFC5AEEF1A96FD81BFB9" box="[478,575,960,984]" pageId="68" pageNumber="69" type="paratype">paratypes</typeStatus>
(
<collectionCode id="4CB160E8FD1BFFC5AD7F1AE9FD24BFB6" box="[590,666,959,983]" country="USA" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="68" pageNumber="69" type="Museum">USNM</collectionCode>
) “
<collectingRegion id="E86436CFFD1BFFC5AD831AE9FCADBFB6" box="[690,787,959,983]" country="Argentina" name="Mendoza" pageId="68" pageNumber="69">Mendoza</collectingRegion>
/
<collectorName id="87559DFBFD1BFFC5AC181AE9FC2EBFB6" box="[809,912,959,983]" pageId="68" pageNumber="69">C.S. Reed</collectorName>
//
<taxonomicName id="EDA083AEFD1BFFC5AC9C1AE9FB96BFB6" baseAuthorityName="sensu Moron &amp; Grossi" baseAuthorityYear="2015" box="[941,1064,959,983]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="genus">TOMARUS</taxonomicName>
/ ROIGJUNENTI / NEITA &amp; RATCLIFFE /
<typeStatus id="F51B468FFD1BFFC5AE1E1A88FE0EBF97" box="[303,432,990,1014]" pageId="68" pageNumber="69" type="paratype">PARATYPE</typeStatus>
” Type locality:
<collectingRegion id="E86436CFFD1BFFC5AD611A88FD10BF97" box="[592,686,990,1014]" country="Argentina" name="Neuquen" pageId="68" pageNumber="69">Neuquén</collectingRegion>
,
<collectingCountry id="52B7B8BDFD1BFFC5AD861A88FCA1BF97" box="[695,799,990,1014]" name="Argentina" pageId="68" pageNumber="69">Argentina</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="62BAABA6FD1BFFC5AFA61D72FC46BA3B" pageId="68" pageNumber="69" type="description">
<paragraph id="2A1FF82DFD1BFFC5AFA61D72FC46BA3B" blockId="68.[151,1437,1060,1987]" pageId="68" pageNumber="69">
<emphasis id="18D4243FFD1BFFC5AFA61D72FE97B85F" bold="true" box="[151,297,1060,1086]" pageId="68" pageNumber="69">Description.</emphasis>
Habitus as in
<figureCitation id="B29BE4A8FD1BFFC5AEE61D72FD85B85F" box="[471,571,1060,1086]" captionStart="FIGURE 28" captionStartId="103.[151,250,1868,1892]" captionTargetBox="[159,1426,182,1839]" captionTargetId="figure-17@103.[151,1436,181,1844]" captionTargetPageId="103" captionText="FIGURE 28. Habitus of the species of Tomarus: A) Tomarus maracaiboensis, B) T. maternus, C) T. pilcopataensis, D) T. pullus, E) T. pumilus, F) T. roigjunenti, G) T. rostratus, H) T. selanderi, I) T. subtropicus. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7345107" httpUri="https://zenodo.org/record/7345107/files/figure.png" pageId="68" pageNumber="69">
Fig.
<specimenCount id="3CA633A4FD1BFFC5AD3F1D72FD85B85F" box="[526,571,1060,1086]" pageId="68" pageNumber="69" type="generic">28F</specimenCount>
</figureCitation>
. Length
<quantity id="ED5855C8FD1BFFC5AD921D72FCF2B85E" box="[675,844,1060,1087]" metricMagnitude="-2" metricUnit="m" metricValue="2.695" metricValueMax="2.76" metricValueMin="2.63" pageId="68" pageNumber="69" unit="mm" value="26.95" valueMax="27.6" valueMin="26.3">26.3–27.6 mm</quantity>
; humeral width
<quantity id="ED5855C8FD1BFFC5AB3D1D72FB0BB85E" box="[1036,1205,1060,1087]" metricMagnitude="-2" metricUnit="m" metricValue="1.325" metricValueMax="1.39" metricValueMin="1.26" pageId="68" pageNumber="69" unit="mm" value="13.25" valueMax="13.9" valueMin="12.6">12.6–13.9 mm</quantity>
. Color dark reddish brown to black.
<emphasis id="18D4243FFD1BFFC5AE7F1D1EFE29B803" bold="true" box="[334,407,1096,1122]" pageId="68" pageNumber="69">Head:</emphasis>
Frons coarsely rugose. Frontoclypeal region with 2 transverse tubercles separated by about 3 tubercle diameters. Clypeus narrowed towards apex, base 3.8 wider than apex. Clypeal teeth triangular, separated by less than a tooth diameter. Mandible with 2 apical teeth and a lateral, narrowly rounded tooth. Mentum abruptly constricted at apical 3rd. Galea of maxilla with teeth 5 and 6 well developed. Interocular distance 4 times an eye width.
<emphasis id="18D4243FFD1BFFC5AE051D8EFE09B893" bold="true" box="[308,439,1240,1266]" pageId="68" pageNumber="69">Pronotum:</emphasis>
Surface with dense, large punctures. Apical tubercle small, rounded. Fovea shallow, striate, narrow (1/5 as wide as interocular distance).
<emphasis id="18D4243FFD1BFFC5ADF41DAAFCF8B977" bold="true" box="[709,838,1276,1302]" pageId="68" pageNumber="69">Scutellum:</emphasis>
Surface with minute, sparse punctures.
<emphasis id="18D4243FFD1BFFC5AA3A1DAAFADFB977" bold="true" box="[1291,1377,1276,1302]" pageId="68" pageNumber="69">Elytra:</emphasis>
First interval punctate, punctures similar in size to other intervals; sutural stria complete. Inner surface of apex with rounded, small tubercles forming 38–40 parallel lines.
<emphasis id="18D4243FFD1BFFC5AC381C12FC39B93F" bold="true" box="[777,903,1348,1374]" pageId="68" pageNumber="69">Abdomen:</emphasis>
Apex of tergite IV with a triangular wide area with large, elongate tubercles forming about 18 diagonal, parallel lines. Pygidium with dense rugosity on basal 3rd (
<figureCitation id="B29BE4A8FD1BFFC5AFAE1CDAFF44B9C7" box="[159,250,1420,1446]" captionStart="FIGURE 9" captionStartId="87.[151,250,1074,1098]" captionTargetBox="[151,1436,181,1050]" captionTargetId="figure-17@87.[151,1436,181,1050]" captionTargetPageId="87" captionText="FIGURE 9. Pygidium of male. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) Ligyrus (Anagrylius) cuniculus, D) L. (Ligyrus) fossor, E) L. (Ligyrus) paranaensis, F) Tomarus bituberculatus, G) T. colombianus, H) T. maracaiboensis, I) T. pilcopataensis, J) T. pullus, K) T. roigjunenti, L) T. rostratus." figureDoi="http://doi.org/10.5281/zenodo.7345041" httpUri="https://zenodo.org/record/7345041/files/figure.png" pageId="68" pageNumber="69">Fig. 9K</figureCitation>
), apex regularly rounded.
<emphasis id="18D4243FFD1BFFC5AD131CDAFDC5B9C7" bold="true" box="[546,635,1420,1446]" pageId="68" pageNumber="69">Venter:</emphasis>
Apex of prosternal process flat, longitudinally oval. Metasternum with short, sparse setae on anterior angles.
<emphasis id="18D4243FFD1BFFC5AD761CE6FD36B9AB" bold="true" box="[583,648,1456,1482]" pageId="68" pageNumber="69">Legs:</emphasis>
Protibia tridentate with an additional inconspicuous, basal convexity. Metatibia triangular; apex slightly crenulate, with 21–25 spinules.
<emphasis id="18D4243FFD1BFFC5AC4C1C82FB8EB98F" bold="true" box="[893,1072,1492,1518]" pageId="68" pageNumber="69">Male genitalia:</emphasis>
Spiculum gastrale without basal plates. Phallobase 1.1 times longer than parameres. Parameres with lateral teeth large, acute; apical half narrowed, apices outwards, rounded (
<figureCitation id="B29BE4A8FD1BFFC5AEF51F4AFD88BA57" box="[452,566,1564,1590]" captionStart="FIGURE 16" captionStartId="91.[151,250,1633,1657]" captionTargetBox="[151,1426,189,1602]" captionTargetId="figure-17@91.[151,1436,181,1610]" captionTargetPageId="91" captionText="FIGURE 16. Parameres in frontal view. A) Tomarus adoceteus, B) T. amazonicus, C) T. bituberculatus, D) T. colombianus, E) T. discrepans, F) T. gyas, G) T. laevicollis, H) T. laticaudus, I) T. maimon, J) T. maracaiboensis, K) T. maternus, L) T. pilcopataensis, M) T. pullus, N) T. pumilus, O) T. roigjunenti, P) T. rostratus, Q) T. selanderi, R) T. subtropicus." figureDoi="http://doi.org/10.5281/zenodo.7345065" httpUri="https://zenodo.org/record/7345065/files/figure.png" pageId="68" pageNumber="69">Figs. 16O</figureCitation>
,
<figureCitation id="B29BE4A8FD1BFFC5AD701F4AFDCFBA57" box="[577,625,1564,1590]" captionStart="FIGURE 20" captionStartId="95.[151,250,1225,1249]" captionTargetBox="[159,1435,191,1201]" captionTargetId="figure-17@95.[151,1436,181,1202]" captionTargetPageId="95" captionText="FIGURE 20. Parameres in lateral view. A) Tomarus maracaiboensis, B) T. maternus, C) T. pilcopataensis, D) T. pullus, E) T. pumilus, F) T. roigjunenti, G) T. rostratus, H) T. selanderi, I) T. subtropicus." figureDoi="http://doi.org/10.5281/zenodo.7345083" httpUri="https://zenodo.org/record/7345083/files/figure.png" pageId="68" pageNumber="69">20F</figureCitation>
). Internal sac with copulatory lamella and short lamellar spiny belt; with a complex of 1–3 spine-like, accessory lamellae, with dense granules at base.
</paragraph>
</subSubSection>
<subSubSection id="62BAABA6FD1BFFC5AFF61F32FF6DBB6F" pageId="68" pageNumber="69" type="diagnosis">
<paragraph id="2A1FF82DFD1BFFC5AFF61F32FF6DBB6F" blockId="68.[151,1437,1060,1987]" pageId="68" pageNumber="69">
<emphasis id="18D4243FFD1BFFC5AFF61F32FEFCBA1F" bold="true" box="[199,322,1636,1662]" pageId="68" pageNumber="69">Diagnosis.</emphasis>
<taxonomicName id="EDA083AEFD1BFFC5AE7B1F33FD8EBA1F" authorityName="Neita &amp; Ratcliffe" authorityYear="2017" box="[330,560,1637,1662]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="roigjunenti">
<emphasis id="18D4243FFD1BFFC5AE7B1F33FD8EBA1F" box="[330,560,1637,1662]" italics="true" pageId="68" pageNumber="69">Tomarus roigjunenti</emphasis>
</taxonomicName>
can be recognized by the following character combination: frontal tubercles transverse; pronotal fovea and tubercle inconspicuous (
<figureCitation id="B29BE4A8FD1BFFC5ADCE1FDEFCDABAC3" box="[767,868,1672,1698]" captionStart="FIGURE 28" captionStartId="103.[151,250,1868,1892]" captionTargetBox="[159,1426,182,1839]" captionTargetId="figure-17@103.[151,1436,181,1844]" captionTargetPageId="103" captionText="FIGURE 28. Habitus of the species of Tomarus: A) Tomarus maracaiboensis, B) T. maternus, C) T. pilcopataensis, D) T. pullus, E) T. pumilus, F) T. roigjunenti, G) T. rostratus, H) T. selanderi, I) T. subtropicus. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7345107" httpUri="https://zenodo.org/record/7345107/files/figure.png" pageId="68" pageNumber="69">Fig. 28F</figureCitation>
); pronotum deeply and densely punctate; metasternum with sparse, minute setae on anterior angles; internal sac with a complex of 1–3 spine-like, accessory lamellae; and parameres with lateral teeth large and long, arcuate; apical half narrowed, apices outwards, rounded (
<figureCitation id="B29BE4A8FD1BFFC5AA171F86FA26BA8B" box="[1318,1432,1744,1770]" captionStart="FIGURE 16" captionStartId="91.[151,250,1633,1657]" captionTargetBox="[151,1426,189,1602]" captionTargetId="figure-17@91.[151,1436,181,1610]" captionTargetPageId="91" captionText="FIGURE 16. Parameres in frontal view. A) Tomarus adoceteus, B) T. amazonicus, C) T. bituberculatus, D) T. colombianus, E) T. discrepans, F) T. gyas, G) T. laevicollis, H) T. laticaudus, I) T. maimon, J) T. maracaiboensis, K) T. maternus, L) T. pilcopataensis, M) T. pullus, N) T. pumilus, O) T. roigjunenti, P) T. rostratus, Q) T. selanderi, R) T. subtropicus." figureDoi="http://doi.org/10.5281/zenodo.7345065" httpUri="https://zenodo.org/record/7345065/files/figure.png" pageId="68" pageNumber="69">Figs. 16O</figureCitation>
,
<figureCitation id="B29BE4A8FD1BFFC5AFA61FA2FF79BB6F" box="[151,199,1780,1806]" captionStart="FIGURE 20" captionStartId="95.[151,250,1225,1249]" captionTargetBox="[159,1435,191,1201]" captionTargetId="figure-17@95.[151,1436,181,1202]" captionTargetPageId="95" captionText="FIGURE 20. Parameres in lateral view. A) Tomarus maracaiboensis, B) T. maternus, C) T. pilcopataensis, D) T. pullus, E) T. pumilus, F) T. roigjunenti, G) T. rostratus, H) T. selanderi, I) T. subtropicus." figureDoi="http://doi.org/10.5281/zenodo.7345083" httpUri="https://zenodo.org/record/7345083/files/figure.png" pageId="68" pageNumber="69">20F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="62BAABA6FD1BFFC4AFF61E4EFE7BBCB4" lastPageId="69" lastPageNumber="70" pageId="68" pageNumber="69" type="distribution">
<paragraph id="2A1FF82DFD1BFFC5AFF61E4EFCE7BB53" blockId="68.[151,1437,1060,1987]" box="[199,857,1816,1842]" pageId="68" pageNumber="69">
<emphasis id="18D4243FFD1BFFC5AFF61E4EFEDFBB53" bold="true" box="[199,353,1816,1842]" pageId="68" pageNumber="69">Distribution.</emphasis>
<collectingCountry id="52B7B8BDFD1BFFC5AE571E4EFE6BBB53" box="[358,469,1816,1842]" name="Argentina" pageId="68" pageNumber="69">Argentina</collectingCountry>
(
<bibRefCitation id="4E3185DCFD1BFFC5AED51E4EFCF3BB53" author="Neita-Moreno, J. C. &amp; Ratcliffe, B. C." box="[484,845,1816,1842]" pageId="68" pageNumber="69" pagination="1 - 36" refId="ref91798" refString="Neita-Moreno, J. C. &amp; Ratcliffe, B. C. (2017) The genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) in Argentina, Chile, and Uruguay. Insecta Mundi, 547, 1 - 36." type="journal article" year="2017">Neita-Moreno &amp; Ratcliffe 2017</bibRefCitation>
).
</paragraph>
<paragraph id="2A1FF82DFD1BFFC5AFF61E6AFC74BBA3" blockId="68.[151,1437,1060,1987]" pageId="68" pageNumber="69">
<emphasis id="18D4243FFD1BFFC5AFF61E6AFDB9BB37" bold="true" box="[199,519,1852,1878]" pageId="68" pageNumber="69">
Locality records (
<figureCitation id="B29BE4A8FD1BFFC5AE901E6AFE44BB37" box="[417,506,1852,1878]" captionStart="FIGURE 47" captionStartId="114.[151,250,1084,1108]" captionTargetBox="[203,1393,181,1051]" captionTargetId="figure-24@114.[195,1393,181,1061]" captionTargetPageId="114" captionText="FIGURE 47. Distributional records of Tomarus pilcopataensis, T. roigjunenti, and T. pumilus." figureDoi="http://doi.org/10.5281/zenodo.7345176" httpUri="https://zenodo.org/record/7345176/files/figure.png" pageId="68" pageNumber="69">Fig. 47</figureCitation>
).
</emphasis>
<materialsCitation id="9AC8F270FD1BFFC5AD221E6AFC78BBA3" ID-GBIF-Occurrence="3986696853" collectionCode="USNM" country="Argentina" county="Records" location="Capital Federal" municipality="Tunuyan" pageId="68" pageNumber="69" specimenCount="1" stateProvince="Chubut">
4 examined specimens from
<collectionCode id="4CB160E8FD1BFFC5AC541E6AFC04BB37" box="[869,954,1852,1878]" country="USA" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="68" pageNumber="69" type="Museum">USNM</collectionCode>
.
<collectingCounty id="C37E80A1FD1BFFC5ACF91E6AFB9ABB37" box="[968,1060,1852,1878]" pageId="68" pageNumber="69">Records</collectingCounty>
from
<bibRefCitation id="4E3185DCFD1BFFC5AB411E6AFF5BBB1B" author="Neita-Moreno, J. C. &amp; Ratcliffe, B. C." pageId="68" pageNumber="69" pagination="1 - 36" refId="ref91798" refString="Neita-Moreno, J. C. &amp; Ratcliffe, B. C. (2017) The genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) in Argentina, Chile, and Uruguay. Insecta Mundi, 547, 1 - 36." type="journal article" year="2017">Neita-Moreno &amp; Ratcliffe (2017)</bibRefCitation>
.
<emphasis id="18D4243FFD1BFFC5AFDE1E36FDD9BB1B" bold="true" box="[239,615,1888,1914]" pageId="68" pageNumber="69">
<collectingCountry id="52B7B8BDFD1BFFC5AFDE1E36FE27BB1B" box="[239,409,1888,1914]" name="Argentina" pageId="68" pageNumber="69">ARGENTINA</collectingCountry>
(20).
<collectingRegion id="E86436CFFD1BFFC5AEEA1E36FD89BB1B" box="[475,567,1888,1914]" country="Argentina" name="Chubut" pageId="68" pageNumber="69">Chubut</collectingRegion>
(1):
</emphasis>
Chubut.
<emphasis id="18D4243FFD1BFFC5ADFC1E36FC28BB1B" bold="true" box="[717,918,1888,1914]" pageId="68" pageNumber="69">
<collectingRegion id="E86436CFFD1BFFC5ADFC1E36FCD6BB1B" box="[717,872,1888,1914]" country="Argentina" name="Ciudad Autonoma de Buenos Aires" pageId="68" pageNumber="69">Buenos Aires</collectingRegion>
(1).
</emphasis>
Buenos Aires.
<emphasis id="18D4243FFD1BFFC5AB0E1E36FB65BB1B" bold="true" box="[1087,1243,1888,1914]" pageId="68" pageNumber="69">
<collectingRegion id="E86436CFFD1BFFC5AB0E1E36FB13BB1B" box="[1087,1197,1888,1914]" country="Argentina" name="Mendoza" pageId="68" pageNumber="69">Mendoza</collectingRegion>
(4).
</emphasis>
<collectingMunicipality id="CA7B6257FD1BFFC5ABD01E37FAF9BB1A" box="[1249,1351,1889,1915]" pageId="68" pageNumber="69">Tunuyan</collectingMunicipality>
; Uspallata.
<emphasis id="18D4243FFD1BFFC5AFFD1ED5FEDBBBFF" bold="true" box="[204,357,1923,1950]" pageId="68" pageNumber="69">
<collectingRegion id="E86436CFFD1BFFC5AFFD1ED5FE89BBFF" box="[204,311,1923,1950]" country="Argentina" name="Neuquen" pageId="68" pageNumber="69">Neuquén</collectingRegion>
(9).
</emphasis>
<location id="2F7FAEF6FD1BFFC5AE5A1ED2FDA7BBFF" LSID="urn:lsid:plazi:treatment:A209493BFD1BFFC4AFA61BADFE7BBCB4:2F7FAEF6FD1BFFC5AE5A1ED2FDA7BBFF" box="[363,537,1924,1950]" country="Argentina" county="Records" municipality="Tunuyan" name="Capital Federal" pageId="68" pageNumber="69" stateProvince="Chubut">Capital Federal</location>
;
<location id="2F7FAEF6FD1BFFC5AD121ED3FD29BBFE" LSID="urn:lsid:plazi:treatment:A209493BFD1BFFC4AFA61BADFE7BBCB4:2F7FAEF6FD1BFFC5AD121ED3FD29BBFE" box="[547,663,1925,1951]" country="Argentina" county="Records" municipality="Tunuyan" name="Barrancas" pageId="68" pageNumber="69" stateProvince="Chubut">Barrancas</location>
;
<location id="2F7FAEF6FD1BFFC5AD911ED2FC92BBFF" LSID="urn:lsid:plazi:treatment:A209493BFD1BFFC4AFA61BADFE7BBCB4:2F7FAEF6FD1BFFC5AD911ED2FC92BBFF" box="[672,812,1924,1950]" country="Argentina" county="Records" municipality="Tunuyan" name="Collon Cura" pageId="68" pageNumber="69" stateProvince="Chubut">Collón Cura</location>
;
<location id="2F7FAEF6FD1BFFC5AC071ED2FC21BBFF" LSID="urn:lsid:plazi:treatment:A209493BFD1BFFC4AFA61BADFE7BBCB4:2F7FAEF6FD1BFFC5AC071ED2FC21BBFF" box="[822,927,1924,1950]" country="Argentina" county="Records" municipality="Tunuyan" name="Covunco" pageId="68" pageNumber="69" stateProvince="Chubut">Covunco</location>
;
<location id="2F7FAEF6FD1BFFC5AC981ED2FB8ABBFF" LSID="urn:lsid:plazi:treatment:A209493BFD1BFFC4AFA61BADFE7BBCB4:2F7FAEF6FD1BFFC5AC981ED2FB8ABBFF" box="[937,1076,1924,1950]" country="Argentina" county="Records" municipality="Tunuyan" name="Isla Victoria" pageId="68" pageNumber="69" stateProvince="Chubut">Isla Victoria</location>
;
<location id="2F7FAEF6FD1BFFC5AB0F1ED3FB57BBFF" LSID="urn:lsid:plazi:treatment:A209493BFD1BFFC4AFA61BADFE7BBCB4:2F7FAEF6FD1BFFC5AB0F1ED3FB57BBFF" box="[1086,1257,1924,1951]" country="Argentina" county="Records" municipality="Tunuyan" name="Laguna Blanca" pageId="68" pageNumber="69" stateProvince="Chubut">Laguna Blanca</location>
; Vatra Lauquen.
<emphasis id="18D4243FFD1BFFC5AFA61EF1FEFEBBA3" bold="true" box="[151,320,1959,1986]" pageId="68" pageNumber="69">
<collectingRegion id="E86436CFFD1BFFC5AFA61EF1FEAFBBA3" box="[151,273,1959,1986]" country="Argentina" name="Rio Negro" pageId="68" pageNumber="69">Río Negro</collectingRegion>
(5).
</emphasis>
<location id="2F7FAEF6FD1BFFC5AE741EFEFE39BBA3" LSID="urn:lsid:plazi:treatment:A209493BFD1BFFC4AFA61BADFE7BBCB4:2F7FAEF6FD1BFFC5AE741EFEFE39BBA3" box="[325,391,1960,1986]" country="Argentina" county="Records" municipality="Tunuyan" name="Allen" pageId="68" pageNumber="69" stateProvince="Chubut">Allen</location>
;
<location id="2F7FAEF6FD1BFFC5AEA31EFEFE41BBA3" LSID="urn:lsid:plazi:treatment:A209493BFD1BFFC4AFA61BADFE7BBCB4:2F7FAEF6FD1BFFC5AEA31EFEFE41BBA3" box="[402,511,1960,1986]" country="Argentina" county="Records" municipality="Tunuyan" name="Cipolletti" pageId="68" pageNumber="69" stateProvince="Chubut">Cipolletti</location>
;
<location id="2F7FAEF6FD1BFFC5AD3B1EFEFD19BBA2" LSID="urn:lsid:plazi:treatment:A209493BFD1BFFC4AFA61BADFE7BBCB4:2F7FAEF6FD1BFFC5AD3B1EFEFD19BBA2" box="[522,679,1960,1987]" country="Argentina" county="Records" municipality="Tunuyan" name="General Roca" pageId="68" pageNumber="69" stateProvince="Chubut">General Roca</location>
;
<location id="2F7FAEF6FD1BFFC5AD831EFEFC92BBA2" LSID="urn:lsid:plazi:treatment:A209493BFD1BFFC4AFA61BADFE7BBCB4:2F7FAEF6FD1BFFC5AD831EFEFC92BBA2" box="[690,812,1960,1987]" country="Argentina" county="Records" municipality="Tunuyan" name="Rio Negro" pageId="68" pageNumber="69" stateProvince="Chubut">Río Negro</location>
;
<location id="2F7FAEF6FD1BFFC5AC071EFEFC78BBA3" LSID="urn:lsid:plazi:treatment:A209493BFD1BFFC4AFA61BADFE7BBCB4:2F7FAEF6FD1BFFC5AC071EFEFC78BBA3" box="[822,966,1960,1986]" country="Argentina" county="Records" municipality="Tunuyan" name="Villa Regina" pageId="68" pageNumber="69" stateProvince="Chubut">Villa Regina</location>
</materialsCitation>
.
</paragraph>
<paragraph id="2A1FF82DFD1AFFC4AFF619C1FE7BBCB4" blockId="69.[151,1436,150,213]" pageId="69" pageNumber="70">
<emphasis id="18D4243FFD1AFFC4AFF619C1FE38BCD1" bold="true" box="[199,390,150,177]" pageId="69" pageNumber="70">Natural history.</emphasis>
Some specimens were found in sand dunes and adults are attracted to lights at night (NeitaMoreno &amp;
<bibRefCitation id="4E3185DCFD1AFFC4AE2519EDFE07BCB4" author="Ratcliffe, B. C. &amp; Cave, R. D." box="[276,441,187,213]" pageId="69" pageNumber="70" pagination="1 - 298" refId="ref92923" refString="Ratcliffe, B. C. &amp; Cave, R. D. (2017) The dynastine scarab beetles of the United States and Canada (Coleoptera: Scarabaeidae). Bulletin of the University of Nebraska State Museum, 30, 1 - 298." type="journal article" year="2017">Ratcliffe 2017</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
Reference in a new issue View git blame Copy permalink