244 lines
36 KiB
XML
244 lines
36 KiB
XML
<document id="1E5325B0DC5C69B28B24B85AA230DCD3" ID-CLB-Dataset="202083" ID-DOI="10.11646/zootaxa.5211.1.1" ID-GBIF-Dataset="bb97b9dd-2acc-4244-8f30-72bef07b9ed9" ID-ISSN="1175-5326" ID-Zenodo-Dep="7345007" ID-ZooBank="5754769C-B747-4714-BDD9-7D5509D48BEB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1669101340847" checkinUser="plazi" docAuthor="López-García, Margarita M. & Deloya, Cuauhtémoc" docDate="2022" docId="A209493BFD1AFFC4AFA61871FA3EBAE8" docLanguage="en" docName="zootaxa.5211.1.1.pdf" docOrigin="Zootaxa 5211 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5211.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Tomarus rostratus Dupuis 2014" docType="treatment" docVersion="7" lastPageNumber="70" masterDocId="5E303143FD5FFF81AF311956FFBEBC61" masterDocTitle="Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision" masterLastPageNumber="119" masterPageNumber="1" pageNumber="70" updateTime="1699396337306" updateUser="plazi">
|
||
<mods:mods id="CC559406BC98D323CFC6568244B91F1E" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="DE8D1262F6FDA12D9CC9ABB3E7C2742F">
|
||
<mods:title id="276EF973FF5AFB431C283D1BA39348F0">Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="26324902B1DD495B87CAAF4B7BE1CC96" type="personal">
|
||
<mods:role id="2B453EA7784A2C572EF5961BACDDA93B">
|
||
<mods:roleTerm id="4AEA66857692F24C310394F27C45BCDD">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="54E2752808B23E9DBCF6ADAA45C08EB1">López-García, Margarita M.</mods:namePart>
|
||
<mods:nameIdentifier id="556586AE64FDF0B51C4CC1B274C24C0F" type="ORCID">0000-0003-2796-2931</mods:nameIdentifier>
|
||
<mods:nameIdentifier id="A9E0088FC6193F2C79BA467F72687784" type="email">margaralopezg@gmail.com</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name id="C23F249FD843109306AC937829F97BD3" type="personal">
|
||
<mods:role id="3DCAFD1E45AAD64EDCA7FCD2AB0303EA">
|
||
<mods:roleTerm id="30D06AFD25F8911EB9FD24309C3E5023">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="06A138A4C556324E836684DCA6BAC048">Deloya, Cuauhtémoc</mods:namePart>
|
||
<mods:nameIdentifier id="A1A304DCAF833FAB466B46F3BCF07135" type="ORCID">0000-0002-4774-140X</mods:nameIdentifier>
|
||
<mods:nameIdentifier id="117812B82BC47BB052BBD1EF25BC1C18" type="email">cuauhtemoc.deloya@inecol.mx</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:typeOfResource id="75844C885ECF5689E2DB6C477F675C9A">text</mods:typeOfResource>
|
||
<mods:relatedItem id="6842A94917F917EBB6BE7327833E2282" type="host">
|
||
<mods:titleInfo id="9E41F517E73804CDF53A5A762E585A90">
|
||
<mods:title id="39BAB68179D13F8DA375B2A3188DC9A8">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="B2058BDF63C1C41003A7E6F72EC6AC89">
|
||
<mods:date id="1476E4FB6C32ACD736DB445EBA1BEDA6">2022</mods:date>
|
||
<mods:detail id="D2596EAF2B8EE12B8F8B1DED9973A9E4" type="pubDate">
|
||
<mods:number id="4619DF4D4A0667CE1D3F9193D7914C46">2022-11-22</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="97CD13E9F59CDCDB1B8B84B853D1BFDB" type="volume">
|
||
<mods:number id="AFAE97E836FDF4BB1910EDBD07E91669">5211</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="733FC166A4C8895FDD67649B918389AA" type="issue">
|
||
<mods:number id="A01F864B950B6A244F0C42210A88EC24">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="35CD0F37550C0A0CDCB94A76E75675DE" unit="page">
|
||
<mods:start id="85DCD68A1FB1333837CBFCF8865E588C">1</mods:start>
|
||
<mods:end id="B062D335831119CF7F94289458C3ADE5">119</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="8897C827A60A83AF57BB5124100B2243">
|
||
<mods:url id="9BF0526566F03DA9C610C78A08F534C9">http://dx.doi.org/10.11646/zootaxa.5211.1.1</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="5C33B9429C95C2191E74137A400405A5">journal article</mods:classification>
|
||
<mods:identifier id="3B49D52FDB4FC81AB5B2733378537225" type="CLB-Dataset">202083</mods:identifier>
|
||
<mods:identifier id="DDCC9B1CFFC96DAB8E78C2A4E1C0ADC0" type="DOI">10.11646/zootaxa.5211.1.1</mods:identifier>
|
||
<mods:identifier id="679CFB6E69ABD285B5BD10B3778D9E49" type="GBIF-Dataset">bb97b9dd-2acc-4244-8f30-72bef07b9ed9</mods:identifier>
|
||
<mods:identifier id="4BEACCEF406F44778E52869784EA00E9" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="F8986D5054F79183DD620A99392CC495" type="Zenodo-Dep">7345007</mods:identifier>
|
||
<mods:identifier id="2AD29A1632B6AF1B041D3495F98D161B" type="ZooBank">5754769C-B747-4714-BDD9-7D5509D48BEB</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="A209493BFD1AFFC4AFA61871FA3EBAE8" ID-DOI="http://doi.org/10.5281/zenodo.7345121" ID-GBIF-Taxon="205794534" ID-Zenodo-Dep="7345121" LSID="urn:lsid:plazi:treatment:A209493BFD1AFFC4AFA61871FA3EBAE8" httpUri="http://treatment.plazi.org/id/A209493BFD1AFFC4AFA61871FA3EBAE8" lastPageNumber="70" pageId="69" pageNumber="70">
|
||
<subSubSection id="62BAABA6FD1AFFC4AFA61871FDA1BD21" box="[151,543,294,321]" pageId="69" pageNumber="70" type="nomenclature">
|
||
<paragraph id="2A1FF82DFD1AFFC4AFA61871FDA1BD21" blockId="69.[151,743,294,357]" box="[151,543,294,321]" pageId="69" pageNumber="70">
|
||
<heading id="71574F41FD1AFFC4AFA61871FDA1BD21" bold="true" box="[151,543,294,321]" fontSize="11" level="2" pageId="69" pageNumber="70" reason="4">
|
||
<taxonomicName id="EDA083AEFD1AFFC4AFA61871FDA1BD21" authority="Dupuis, 2014" authorityName="Dupuis" authorityYear="2014" box="[151,543,294,321]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="rostratus">
|
||
<emphasis id="18D4243FFD1AFFC4AFA61871FDA1BD21" bold="true" box="[151,543,294,321]" pageId="69" pageNumber="70">
|
||
<emphasis id="18D4243FFD1AFFC4AFA61871FECBBD20" bold="true" box="[151,373,295,321]" italics="true" pageId="69" pageNumber="70">Tomarus rostratus</emphasis>
|
||
<bibRefCitation id="4E3185DCFD1AFFC4AE4D1870FDA1BD21" author="Dupuis, F." box="[380,543,294,320]" pageId="69" pageNumber="70" pagination="1 - 4" refId="ref89615" refString="Dupuis, F. (2014) Tomarus rostratus nouvelle espece du Perou (Coleoptera, Dynastidae). Coleopteres, 20, 1 - 4." type="journal article" year="2014">Dupuis, 2014</bibRefCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="62BAABA6FD1AFFC4AFA6181DFD59BD04" box="[151,743,330,357]" pageId="69" pageNumber="70" type="description">
|
||
<paragraph id="2A1FF82DFD1AFFC4AFA6181DFD59BD04" blockId="69.[151,743,294,357]" box="[151,743,330,357]" pageId="69" pageNumber="70">
|
||
(
|
||
<figureCitation id="B29BE4A8FD1AFFC4AFAE181DFF43BD04" box="[159,253,330,357]" captionStart="FIGURE 5" captionStartId="84.[151,250,931,955]" captionTargetBox="[188,1400,181,907]" captionTargetId="figure-24@84.[188,1400,181,907]" captionTargetPageId="84" captionText="FIGURE 5. Mandible. A) Euligyrus similis, B) Proculigyrus cicatricosus, C) Ligyrus (Ligyrodes) relictus, D) L. (Ligyrus) burmeisteri, E) L. (Ligyrus) villosus, F) Tomarus rostratus, G) T. gyas, H) T. maternus." figureDoi="http://doi.org/10.5281/zenodo.7345025" httpUri="https://zenodo.org/record/7345025/files/figure.png" pageId="69" pageNumber="70">Figs. 5F</figureCitation>
|
||
,
|
||
<figureCitation id="B29BE4A8FD1AFFC4AE39181DFE92BD04" box="[264,300,331,357]" captionStart="FIGURE 7" captionStartId="85.[151,250,1391,1415]" captionTargetBox="[151,1436,184,1368]" captionTargetId="figure-17@85.[151,1436,181,1368]" captionTargetPageId="85" captionText="FIGURE 7. Left hindwing in ventral view. A, C) Euligyrus similis, B, D) Ligyrus (Anagrylius) cuniculus, E) L. (Ligyrus) burmeisteri, F) Tomarus gyas, G) T. maternus, H) T. rostratus." figureDoi="http://doi.org/10.5281/zenodo.7345031" httpUri="https://zenodo.org/record/7345031/files/figure.png" pageId="69" pageNumber="70">7H</figureCitation>
|
||
,
|
||
<figureCitation id="B29BE4A8FD1AFFC4AE09181DFEE7BD04" box="[312,345,331,357]" captionStart="FIGURE 9" captionStartId="87.[151,250,1074,1098]" captionTargetBox="[151,1436,181,1050]" captionTargetId="figure-17@87.[151,1436,181,1050]" captionTargetPageId="87" captionText="FIGURE 9. Pygidium of male. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) Ligyrus (Anagrylius) cuniculus, D) L. (Ligyrus) fossor, E) L. (Ligyrus) paranaensis, F) Tomarus bituberculatus, G) T. colombianus, H) T. maracaiboensis, I) T. pilcopataensis, J) T. pullus, K) T. roigjunenti, L) T. rostratus." figureDoi="http://doi.org/10.5281/zenodo.7345041" httpUri="https://zenodo.org/record/7345041/files/figure.png" pageId="69" pageNumber="70">9L</figureCitation>
|
||
,
|
||
<figureCitation id="B29BE4A8FD1AFFC4AE54181DFE30BD04" box="[357,398,331,357]" captionStart="FIGURE 13" captionStartId="89.[151,250,1041,1065]" captionTargetBox="[154,1432,183,1016]" captionTargetId="figure-17@89.[151,1436,181,1017]" captionTargetPageId="89" captionText="FIGURE 13. Female genital plates. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) Ligyrus (Anagrylius) moroni, D) L. (Ligyrodes) sallaei, E) L. (Ligyrus) allonasutus, F) L. (Ligyrus) bidentulus, G) L. (Ligyrus) neglectus, H) Tomarus laticaudus, I) T. maternus, J) T. rostratus, K) T. subtropicus. Abbreviations: SCx: Subcoxite, Cx: Coxite." figureDoi="http://doi.org/10.5281/zenodo.7345057" httpUri="https://zenodo.org/record/7345057/files/figure.png" pageId="69" pageNumber="70">13J</figureCitation>
|
||
,
|
||
<figureCitation id="B29BE4A8FD1AFFC4AEAB181DFE78BD04" box="[410,454,331,357]" captionStart="FIGURE 14" captionStartId="89.[151,250,1677,1701]" captionTargetBox="[152,1432,1161,1649]" captionTargetId="figure-90@89.[152,1436,1161,1653]" captionTargetPageId="89" captionText="FIGURE 14. Spiculum gastrale. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) L. (Ligyrodes) relictus, D) L. (Ligyrus) villosus, E) Tomarus pullus, F) T. rostratus, G) T. selanderi, H) T. subtropicus." figureDoi="http://doi.org/10.5281/zenodo.7345059" httpUri="https://zenodo.org/record/7345059/files/figure.png" pageId="69" pageNumber="70">14F</figureCitation>
|
||
,
|
||
<figureCitation id="B29BE4A8FD1AFFC4AEE3181DFE42BD04" box="[466,508,331,357]" captionStart="FIGURE 16" captionStartId="91.[151,250,1633,1657]" captionTargetBox="[151,1426,189,1602]" captionTargetId="figure-17@91.[151,1436,181,1610]" captionTargetPageId="91" captionText="FIGURE 16. Parameres in frontal view. A) Tomarus adoceteus, B) T. amazonicus, C) T. bituberculatus, D) T. colombianus, E) T. discrepans, F) T. gyas, G) T. laevicollis, H) T. laticaudus, I) T. maimon, J) T. maracaiboensis, K) T. maternus, L) T. pilcopataensis, M) T. pullus, N) T. pumilus, O) T. roigjunenti, P) T. rostratus, Q) T. selanderi, R) T. subtropicus." figureDoi="http://doi.org/10.5281/zenodo.7345065" httpUri="https://zenodo.org/record/7345065/files/figure.png" pageId="69" pageNumber="70">16P</figureCitation>
|
||
,
|
||
<figureCitation id="B29BE4A8FD1AFFC4AD39181DFD85BD04" box="[520,571,331,357]" captionStart="FIGURE 20" captionStartId="95.[151,250,1225,1249]" captionTargetBox="[159,1435,191,1201]" captionTargetId="figure-17@95.[151,1436,181,1202]" captionTargetPageId="95" captionText="FIGURE 20. Parameres in lateral view. A) Tomarus maracaiboensis, B) T. maternus, C) T. pilcopataensis, D) T. pullus, E) T. pumilus, F) T. roigjunenti, G) T. rostratus, H) T. selanderi, I) T. subtropicus." figureDoi="http://doi.org/10.5281/zenodo.7345083" httpUri="https://zenodo.org/record/7345083/files/figure.png" pageId="69" pageNumber="70">20G</figureCitation>
|
||
,
|
||
<figureCitation id="B29BE4A8FD1AFFC4AD77181DFDC6BD04" box="[582,632,331,357]" captionStart="FIGURE 22" captionStartId="97.[151,250,1544,1568]" captionTargetBox="[159,1424,185,1501]" captionTargetId="figure-17@97.[151,1436,181,1520]" captionTargetPageId="97" captionText="FIGURE 22. Internal sac. A) Tomarus pullus, B) T. rostratus, C) T. maternus. Abbreviations: AL: Accessory lamellae, CL: Copulatory lamella, LSB: Lamellar spiny belt, Ra: Raspula, SD: Sclerotized diverticulum." figureDoi="http://doi.org/10.5281/zenodo.7345087" httpUri="https://zenodo.org/record/7345087/files/figure.png" pageId="69" pageNumber="70">22B</figureCitation>
|
||
,
|
||
<figureCitation id="B29BE4A8FD1AFFC4ADB2181DFD09BD04" box="[643,695,331,357]" captionStart="FIGURE 28" captionStartId="103.[151,250,1868,1892]" captionTargetBox="[159,1426,182,1839]" captionTargetId="figure-17@103.[151,1436,181,1844]" captionTargetPageId="103" captionText="FIGURE 28. Habitus of the species of Tomarus: A) Tomarus maracaiboensis, B) T. maternus, C) T. pilcopataensis, D) T. pullus, E) T. pumilus, F) T. roigjunenti, G) T. rostratus, H) T. selanderi, I) T. subtropicus. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7345107" httpUri="https://zenodo.org/record/7345107/files/figure.png" pageId="69" pageNumber="70">28G</figureCitation>
|
||
;
|
||
<figureCitation id="B29BE4A8FD1AFFC4ADF3181DFD61BD04" box="[706,735,331,357]" captionStart="FIGURE 49" captionStartId="115.[151,250,1113,1137]" captionTargetBox="[162,1433,182,1078]" captionTargetId="figure-17@115.[154,1433,181,1089]" captionTargetPageId="115" captionText="FIGURE 49. Distributional records of Tomarus rostratus." figureDoi="http://doi.org/10.5281/zenodo.7345184" httpUri="https://zenodo.org/record/7345184/files/figure.png" pageId="69" pageNumber="70">49</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="62BAABA6FD1AFFC4AFA618C6FD63BDC6" box="[151,733,399,423]" pageId="69" pageNumber="70" type="reference_group">
|
||
<paragraph id="2A1FF82DFD1AFFC4AFA618C6FD63BDC6" blockId="69.[151,1436,399,485]" box="[151,733,399,423]" pageId="69" pageNumber="70">
|
||
<treatmentCitationGroup id="0AB0DF03FD1AFFC4AFA618C6FD63BDC6" box="[151,733,399,423]" pageId="69" pageNumber="70">
|
||
<treatmentCitation id="AB01DE3CFD1AFFC4AFA618C6FE4ABDC6" author="Dupuis, F." box="[151,500,399,423]" page="1" pageId="69" pageNumber="70" year="2014">
|
||
<taxonomicName id="EDA083AEFD1AFFC4AFA618C6FE4ABDC6" authority="Dupuis, 2014: 1" authorityName="Dupuis" authorityPageNumber="1" authorityYear="2014" box="[151,500,399,423]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="rostratus">
|
||
<emphasis id="18D4243FFD1AFFC4AFA618C6FEF1BDC6" box="[151,335,400,423]" italics="true" pageId="69" pageNumber="70">Tomarus rostratus</emphasis>
|
||
<bibRefCitation id="4E3185DCFD1AFFC4AE6218D9FE4ABDC6" author="Dupuis, F." box="[339,500,399,423]" pageId="69" pageNumber="70" pagination="1 - 4" refId="ref89615" refString="Dupuis, F. (2014) Tomarus rostratus nouvelle espece du Perou (Coleoptera, Dynastidae). Coleopteres, 20, 1 - 4." type="journal article" year="2014">Dupuis, 2014: 1</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
. Original combination.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="62BAABA6FD1AFFC4ADD118D9FDB9BD85" pageId="69" pageNumber="70" type="materials_examined">
|
||
<paragraph id="2A1FF82DFD1AFFC4ADD118D9FDB9BD85" blockId="69.[151,1436,399,485]" pageId="69" pageNumber="70">
|
||
Male
|
||
<typeStatus id="F51B468FFD1AFFC4AC2818D9FCCCBDC6" box="[793,882,399,423]" pageId="69" pageNumber="70" type="holotype">holotype</typeStatus>
|
||
(MNHN) “
|
||
<collectingCountry id="52B7B8BDFD1AFFC4ACD218D9FBA9BDC6" box="[995,1047,399,423]" name="Peru" pageId="69" pageNumber="70">Peru</collectingCountry>
|
||
. /
|
||
<taxonomicName id="EDA083AEFD1AFFC4AB1B18C6FB3CBDC6" baseAuthorityName="sensu Moron & Grossi" baseAuthorityYear="2015" box="[1066,1154,400,423]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="genus">Tomarus</taxonomicName>
|
||
/ Er. / Maimon // MUSÉUM PARIS / 1952 / coll. R. OBERTHÜR //
|
||
<taxonomicName id="EDA083AEFD1AFFC4AD6818F8FCB0BDA4" authority="López-García & Deloya, 2022" authorityName="López-García & Deloya" authorityYear="2022" box="[601,782,430,453]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="rostratus" status="sp. nov.">Tomarus rostratus</taxonomicName>
|
||
<taxonomicNameLabel id="03E79944FD1AFFC4AC2418F8FCF2BDA4" box="[789,844,430,453]" pageId="69" pageNumber="70" rank="species">n. sp.</taxonomicNameLabel>
|
||
/
|
||
<typeStatus id="F51B468FFD1AFFC4AC6E18F8FC5BBDA7" box="[863,997,430,454]" pageId="69" pageNumber="70" type="holotype">HOLOTYPE</typeStatus>
|
||
/ F.
|
||
<bibRefCitation id="4E3185DCFD1AFFC4AB2018F8FB2ABDA7" author="Dupuis, F." box="[1041,1172,430,454]" pageId="69" pageNumber="70" pagination="1 - 4" refId="ref89615" refString="Dupuis, F. (2014) Tomarus rostratus nouvelle espece du Perou (Coleoptera, Dynastidae). Coleopteres, 20, 1 - 4." type="journal article" year="2014">Dupuis 2014</bibRefCitation>
|
||
//
|
||
<typeStatus id="F51B468FFD1AFFC4AB9F18F8FA8ABDA7" box="[1198,1332,430,454]" pageId="69" pageNumber="70" type="holotype">HOLOTYPE</typeStatus>
|
||
// MNHN / EC4187”. Type locality:
|
||
<collectingCountry id="52B7B8BDFD1AFFC4AEE0189BFDBDBD85" box="[465,515,461,484]" name="Peru" pageId="69" pageNumber="70">Peru</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="62BAABA6FD1AFFC4AFA61B44FBB7B8D4" pageId="69" pageNumber="70" type="description">
|
||
<paragraph id="2A1FF82DFD1AFFC4AFA61B44FBB7B8D4" blockId="69.[151,1437,530,1673]" pageId="69" pageNumber="70">
|
||
<emphasis id="18D4243FFD1AFFC4AFA61B44FE97BE4D" bold="true" box="[151,297,530,556]" pageId="69" pageNumber="70">Description.</emphasis>
|
||
Habitus as in
|
||
<figureCitation id="B29BE4A8FD1AFFC4AEE81B45FDFABE4C" box="[473,580,531,557]" captionStart="FIGURE 28" captionStartId="103.[151,250,1868,1892]" captionTargetBox="[159,1426,182,1839]" captionTargetId="figure-17@103.[151,1436,181,1844]" captionTargetPageId="103" captionText="FIGURE 28. Habitus of the species of Tomarus: A) Tomarus maracaiboensis, B) T. maternus, C) T. pilcopataensis, D) T. pullus, E) T. pumilus, F) T. roigjunenti, G) T. rostratus, H) T. selanderi, I) T. subtropicus. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7345107" httpUri="https://zenodo.org/record/7345107/files/figure.png" pageId="69" pageNumber="70">Fig. 28G</figureCitation>
|
||
. Length
|
||
<quantity id="ED5855C8FD1AFFC4AD9C1B44FCE9BE4C" box="[685,855,530,557]" metricMagnitude="-2" metricUnit="m" metricValue="2.245" metricValueMax="2.75" metricValueMin="1.7399999999999998" pageId="69" pageNumber="70" unit="mm" value="22.45" valueMax="27.5" valueMin="17.4">17.4–27.5 mm</quantity>
|
||
; humeral width 8.3–10.0 mm. Color dark reddish brown.
|
||
<emphasis id="18D4243FFD1AFFC4AFDC1B61FE88BE30" bold="true" box="[237,310,567,593]" pageId="69" pageNumber="70">Head:</emphasis>
|
||
Frons coarsely and irregularly rugose. Frontoclypeal region with 2 transverse tubercles separated by about 3 tubercle diameters. Clypeus trapezoidal, not strongly narrowed toward apex; apex with 2 triangular teeth separated by about 1.5 tooth diameters. Mandible with 2 apical teeth and a lateral, widely rounded tooth (
|
||
<figureCitation id="B29BE4A8FD1AFFC4AA0B1B29FA2EBEF8" box="[1338,1424,638,665]" captionStart="FIGURE 5" captionStartId="84.[151,250,931,955]" captionTargetBox="[188,1400,181,907]" captionTargetId="figure-24@84.[188,1400,181,907]" captionTargetPageId="84" captionText="FIGURE 5. Mandible. A) Euligyrus similis, B) Proculigyrus cicatricosus, C) Ligyrus (Ligyrodes) relictus, D) L. (Ligyrus) burmeisteri, E) L. (Ligyrus) villosus, F) Tomarus rostratus, G) T. gyas, H) T. maternus." figureDoi="http://doi.org/10.5281/zenodo.7345025" httpUri="https://zenodo.org/record/7345025/files/figure.png" pageId="69" pageNumber="70">Fig. 5F</figureCitation>
|
||
). Mentum abruptly constricted at apical 3rd. Galea of maxilla with teeth 5 and 6 well developed.
|
||
<emphasis id="18D4243FFD1AFFC4AB8F1BF5FAFEBEDC" bold="true" box="[1214,1344,675,701]" pageId="69" pageNumber="70">Pronotum:</emphasis>
|
||
Surface punctate, punctures rounded, smaller than punctures on elytra; densely distributed on apical and lateral margins. Apical margin with a small tubercle, visible in lateral view. Pronotum without rugose fovea, with small to large punctures on each side of tubercle.
|
||
<emphasis id="18D4243FFD1AFFC4AD1B1A58FD15BF49" bold="true" box="[554,683,782,808]" pageId="69" pageNumber="70">Scutellum:</emphasis>
|
||
Surface without punctures.
|
||
<emphasis id="18D4243FFD1AFFC4ACDA1A59FBFEBF48" bold="true" box="[1003,1088,783,809]" pageId="69" pageNumber="70">Elytra:</emphasis>
|
||
Sutural stria complete. Elytral punctures dense and deep, punctures mainly ocellate, 3 pairs of distinct double rows.
|
||
<emphasis id="18D4243FFD1AFFC4AB621A64FB6FBF2D" bold="true" box="[1107,1233,818,844]" pageId="69" pageNumber="70">Abdomen:</emphasis>
|
||
Pygidium convex (male) to flat (female) in lateral view; surface deeply rugopunctate on basal 3rd with disc smooth (male) (
|
||
<figureCitation id="B29BE4A8FD1AFFC4AA091A01FA31BF10" box="[1336,1423,855,881]" captionStart="FIGURE 9" captionStartId="87.[151,250,1074,1098]" captionTargetBox="[151,1436,181,1050]" captionTargetId="figure-17@87.[151,1436,181,1050]" captionTargetPageId="87" captionText="FIGURE 9. Pygidium of male. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) Ligyrus (Anagrylius) cuniculus, D) L. (Ligyrus) fossor, E) L. (Ligyrus) paranaensis, F) Tomarus bituberculatus, G) T. colombianus, H) T. maracaiboensis, I) T. pilcopataensis, J) T. pullus, K) T. roigjunenti, L) T. rostratus." figureDoi="http://doi.org/10.5281/zenodo.7345041" httpUri="https://zenodo.org/record/7345041/files/figure.png" pageId="69" pageNumber="70">Fig. 9L</figureCitation>
|
||
), or completely rugopunctate (female). Apical margin of sternite VII straight in male, slightly emarginate with notch broad and shallow in female. Apex of sternite VIII emarginate, notch wide in male and narrow and deep in female.
|
||
<emphasis id="18D4243FFD1AFFC4AFA61A95FF4EBFBC" bold="true" box="[151,240,963,989]" pageId="69" pageNumber="70">Venter:</emphasis>
|
||
Prosternal process long, thick, apex transversely oval to rounded, and bordered with long, pale yellow setae. Metasternum with short, sparse setae on anterior angles.
|
||
<emphasis id="18D4243FFD1AFFC4AC571AB1FC19B860" bold="true" box="[870,935,999,1025]" pageId="69" pageNumber="70">Legs:</emphasis>
|
||
Protibia tridentate with an additional small, basal denticle. Apex of metatibia slightly crenulate, with 8–15 spinules.
|
||
<emphasis id="18D4243FFD1AFFC4ACF01D5DFB31B845" bold="true" box="[961,1167,1034,1061]" pageId="69" pageNumber="70">Female genitalia:</emphasis>
|
||
Coxite surface strongly concave (
|
||
<figureCitation id="B29BE4A8FD1AFFC4AE351D79FEDBB828" box="[260,357,1071,1097]" captionStart="FIGURE 13" captionStartId="89.[151,250,1041,1065]" captionTargetBox="[154,1432,183,1016]" captionTargetId="figure-17@89.[151,1436,181,1017]" captionTargetPageId="89" captionText="FIGURE 13. Female genital plates. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) Ligyrus (Anagrylius) moroni, D) L. (Ligyrodes) sallaei, E) L. (Ligyrus) allonasutus, F) L. (Ligyrus) bidentulus, G) L. (Ligyrus) neglectus, H) Tomarus laticaudus, I) T. maternus, J) T. rostratus, K) T. subtropicus. Abbreviations: SCx: Subcoxite, Cx: Coxite." figureDoi="http://doi.org/10.5281/zenodo.7345057" httpUri="https://zenodo.org/record/7345057/files/figure.png" pageId="69" pageNumber="70">Fig. 13J</figureCitation>
|
||
).
|
||
<emphasis id="18D4243FFD1AFFC4AE4B1D79FD8EB829" bold="true" box="[378,560,1070,1097]" pageId="69" pageNumber="70">Male genitalia:</emphasis>
|
||
Spiculum gastrale with basal plates (
|
||
<figureCitation id="B29BE4A8FD1AFFC4ACED1D79FBFCB828" box="[988,1090,1071,1097]" captionStart="FIGURE 14" captionStartId="89.[151,250,1677,1701]" captionTargetBox="[152,1432,1161,1649]" captionTargetId="figure-90@89.[152,1436,1161,1653]" captionTargetPageId="89" captionText="FIGURE 14. Spiculum gastrale. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) L. (Ligyrodes) relictus, D) L. (Ligyrus) villosus, E) Tomarus pullus, F) T. rostratus, G) T. selanderi, H) T. subtropicus." figureDoi="http://doi.org/10.5281/zenodo.7345059" httpUri="https://zenodo.org/record/7345059/files/figure.png" pageId="69" pageNumber="70">Fig. 14F</figureCitation>
|
||
). Parameres widened at apical 4th, elongate from basal 4th to apex; sides parallel, without teeth or lateral projections (
|
||
<figureCitation id="B29BE4A8FD1AFFC4AB5D1D05FB6EB80C" box="[1132,1232,1107,1133]" captionStart="FIGURE 16" captionStartId="91.[151,250,1633,1657]" captionTargetBox="[151,1426,189,1602]" captionTargetId="figure-17@91.[151,1436,181,1610]" captionTargetPageId="91" captionText="FIGURE 16. Parameres in frontal view. A) Tomarus adoceteus, B) T. amazonicus, C) T. bituberculatus, D) T. colombianus, E) T. discrepans, F) T. gyas, G) T. laevicollis, H) T. laticaudus, I) T. maimon, J) T. maracaiboensis, K) T. maternus, L) T. pilcopataensis, M) T. pullus, N) T. pumilus, O) T. roigjunenti, P) T. rostratus, Q) T. selanderi, R) T. subtropicus." figureDoi="http://doi.org/10.5281/zenodo.7345065" httpUri="https://zenodo.org/record/7345065/files/figure.png" pageId="69" pageNumber="70">Fig. 16P</figureCitation>
|
||
), regularly curved at lateral view (
|
||
<figureCitation id="B29BE4A8FD1AFFC4AE791D21FE0CB8F0" box="[328,434,1143,1169]" captionStart="FIGURE 20" captionStartId="95.[151,250,1225,1249]" captionTargetBox="[159,1435,191,1201]" captionTargetId="figure-17@95.[151,1436,181,1202]" captionTargetPageId="95" captionText="FIGURE 20. Parameres in lateral view. A) Tomarus maracaiboensis, B) T. maternus, C) T. pilcopataensis, D) T. pullus, E) T. pumilus, F) T. roigjunenti, G) T. rostratus, H) T. selanderi, I) T. subtropicus." figureDoi="http://doi.org/10.5281/zenodo.7345083" httpUri="https://zenodo.org/record/7345083/files/figure.png" pageId="69" pageNumber="70">Fig. 20G</figureCitation>
|
||
). Internal sac with small copulatory lamella, lamellar spiny belt reduced, complex of accessory lamellae with 6 spines joined at the base by dense granules (
|
||
<figureCitation id="B29BE4A8FD1AFFC4ACA41DCDFC42B8D4" box="[917,1020,1179,1205]" captionStart="FIGURE 22" captionStartId="97.[151,250,1544,1568]" captionTargetBox="[159,1424,185,1501]" captionTargetId="figure-17@97.[151,1436,181,1520]" captionTargetPageId="97" captionText="FIGURE 22. Internal sac. A) Tomarus pullus, B) T. rostratus, C) T. maternus. Abbreviations: AL: Accessory lamellae, CL: Copulatory lamella, LSB: Lamellar spiny belt, Ra: Raspula, SD: Sclerotized diverticulum." figureDoi="http://doi.org/10.5281/zenodo.7345087" httpUri="https://zenodo.org/record/7345087/files/figure.png" pageId="69" pageNumber="70">Fig. 22B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="62BAABA6FD1AFFC4AFF61DE8FB5CB9EC" pageId="69" pageNumber="70" type="diagnosis">
|
||
<paragraph id="2A1FF82DFD1AFFC4AFF61DE8FB5CB9EC" blockId="69.[151,1437,530,1673]" pageId="69" pageNumber="70">
|
||
<emphasis id="18D4243FFD1AFFC4AFF61DE8FEFCB8B9" bold="true" box="[199,322,1214,1240]" pageId="69" pageNumber="70">Diagnosis.</emphasis>
|
||
<taxonomicName id="EDA083AEFD1AFFC4AE761DE9FDACB8B9" authorityName="López-García & Deloya" authorityYear="2022" box="[327,530,1215,1240]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="rostratus">
|
||
<emphasis id="18D4243FFD1AFFC4AE761DE9FDACB8B9" box="[327,530,1215,1240]" italics="true" pageId="69" pageNumber="70">Tomarus rostratus</emphasis>
|
||
</taxonomicName>
|
||
is unique in the genus because of its extremely elongate parameres, without lateral teeth. It can be also differentiated by the following combination of characters: clypeus trapezoidal; pronotum with small tubercle and without rugose fovea, with small to large punctures on each side of tubercle (
|
||
<figureCitation id="B29BE4A8FD1AFFC4ABF81C51FA8FB940" box="[1225,1329,1287,1313]" captionStart="FIGURE 28" captionStartId="103.[151,250,1868,1892]" captionTargetBox="[159,1426,182,1839]" captionTargetId="figure-17@103.[151,1436,181,1844]" captionTargetPageId="103" captionText="FIGURE 28. Habitus of the species of Tomarus: A) Tomarus maracaiboensis, B) T. maternus, C) T. pilcopataensis, D) T. pullus, E) T. pumilus, F) T. roigjunenti, G) T. rostratus, H) T. selanderi, I) T. subtropicus. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7345107" httpUri="https://zenodo.org/record/7345107/files/figure.png" pageId="69" pageNumber="70">Fig. 28G</figureCitation>
|
||
); protibia tridentate with a small denticle; apical border of pygidium strongly truncate in male; apical margin of sternite VIII emarginated in female; coxite surface strongly concave (
|
||
<figureCitation id="B29BE4A8FD1AFFC4AC281C19FCC7B908" box="[793,889,1359,1385]" captionStart="FIGURE 13" captionStartId="89.[151,250,1041,1065]" captionTargetBox="[154,1432,183,1016]" captionTargetId="figure-17@89.[151,1436,181,1017]" captionTargetPageId="89" captionText="FIGURE 13. Female genital plates. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) Ligyrus (Anagrylius) moroni, D) L. (Ligyrodes) sallaei, E) L. (Ligyrus) allonasutus, F) L. (Ligyrus) bidentulus, G) L. (Ligyrus) neglectus, H) Tomarus laticaudus, I) T. maternus, J) T. rostratus, K) T. subtropicus. Abbreviations: SCx: Subcoxite, Cx: Coxite." figureDoi="http://doi.org/10.5281/zenodo.7345057" httpUri="https://zenodo.org/record/7345057/files/figure.png" pageId="69" pageNumber="70">Fig. 13J</figureCitation>
|
||
); spiculum gastrale with basal plates (
|
||
<figureCitation id="B29BE4A8FD1AFFC4AA1B1C19FA31B908" box="[1322,1423,1359,1385]" captionStart="FIGURE 14" captionStartId="89.[151,250,1677,1701]" captionTargetBox="[152,1432,1161,1649]" captionTargetId="figure-90@89.[152,1436,1161,1653]" captionTargetPageId="89" captionText="FIGURE 14. Spiculum gastrale. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) L. (Ligyrodes) relictus, D) L. (Ligyrus) villosus, E) Tomarus pullus, F) T. rostratus, G) T. selanderi, H) T. subtropicus." figureDoi="http://doi.org/10.5281/zenodo.7345059" httpUri="https://zenodo.org/record/7345059/files/figure.png" pageId="69" pageNumber="70">Fig. 14F</figureCitation>
|
||
); internal sac of aedeagus with small copulatory lamella and reduced lamellar spiny belt (
|
||
<figureCitation id="B29BE4A8FD1AFFC4AB5F1C25FB6BB9EC" box="[1134,1237,1395,1421]" captionStart="FIGURE 22" captionStartId="97.[151,250,1544,1568]" captionTargetBox="[159,1424,185,1501]" captionTargetId="figure-17@97.[151,1436,181,1520]" captionTargetPageId="97" captionText="FIGURE 22. Internal sac. A) Tomarus pullus, B) T. rostratus, C) T. maternus. Abbreviations: AL: Accessory lamellae, CL: Copulatory lamella, LSB: Lamellar spiny belt, Ra: Raspula, SD: Sclerotized diverticulum." figureDoi="http://doi.org/10.5281/zenodo.7345087" httpUri="https://zenodo.org/record/7345087/files/figure.png" pageId="69" pageNumber="70">Fig. 22B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="62BAABA6FD1AFFC4AFF61CC0FA3EBAE8" pageId="69" pageNumber="70" type="distribution">
|
||
<paragraph id="2A1FF82DFD1AFFC4AFF61CC0FE41B9B4" blockId="69.[151,1437,530,1673]" pageId="69" pageNumber="70">
|
||
<emphasis id="18D4243FFD1AFFC4AFF61CC0FEDFB9D1" bold="true" box="[199,353,1430,1456]" pageId="69" pageNumber="70">Distribution.</emphasis>
|
||
<taxonomicName id="EDA083AEFD1AFFC4AE561CC1FD8CB9D1" authorityName="López-García & Deloya" authorityYear="2022" box="[359,562,1431,1456]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="rostratus">
|
||
<emphasis id="18D4243FFD1AFFC4AE561CC1FD8CB9D1" box="[359,562,1431,1456]" italics="true" pageId="69" pageNumber="70">Tomarus rostratus</emphasis>
|
||
</taxonomicName>
|
||
is an endemic species from the coastal desert of
|
||
<collectingCountry id="52B7B8BDFD1AFFC4AB7E1CC1FB3CB9D0" box="[1103,1154,1431,1457]" name="Peru" pageId="69" pageNumber="70">Peru</collectingCountry>
|
||
(
|
||
<bibRefCitation id="4E3185DCFD1AFFC4ABA11CC1FA9DB9D0" author="Dupuis, F." box="[1168,1315,1431,1457]" pageId="69" pageNumber="70" pagination="1 - 4" refId="ref89615" refString="Dupuis, F. (2014) Tomarus rostratus nouvelle espece du Perou (Coleoptera, Dynastidae). Coleopteres, 20, 1 - 4." type="journal article" year="2014">Dupuis 2014</bibRefCitation>
|
||
) and
|
||
<collectingCountry id="52B7B8BDFD1AFFC4AA511CC1FA23B9D0" box="[1376,1437,1431,1457]" name="Chile" pageId="69" pageNumber="70">Chile</collectingCountry>
|
||
(
|
||
<bibRefCitation id="4E3185DCFD1AFFC4AFAE1CEDFE4DB9B4" author="Lopez-Garcia, M. M. & Deloya, C." box="[159,499,1467,1493]" pageId="69" pageNumber="70" pagination="331 - 338" refId="ref91261" refString="Lopez-Garcia, M. M. & Deloya, C. (2018) Description of the female of Tomarus rostratus Dupuis and new synonyms and records for the genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae). The Coleopterists Bulletin, 72 (2), 331 - 338. https: // doi. org / 10.1649 / 0010 - 065 X- 72.2.331" type="journal article" year="2018">López-García & Deloya 2018</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="2A1FF82DFD1AFFC4AFF61C88FC34BA04" blockId="69.[151,1437,530,1673]" pageId="69" pageNumber="70">
|
||
<emphasis id="18D4243FFD1AFFC4AFF61C88FE46B999" bold="true" box="[199,504,1502,1529]" pageId="69" pageNumber="70">
|
||
Locality records (
|
||
<figureCitation id="B29BE4A8FD1AFFC4AEA61C88FE55B999" box="[407,491,1502,1528]" captionStart="FIGURE 49" captionStartId="115.[151,250,1113,1137]" captionTargetBox="[162,1433,182,1078]" captionTargetId="figure-17@115.[154,1433,181,1089]" captionTargetPageId="115" captionText="FIGURE 49. Distributional records of Tomarus rostratus." figureDoi="http://doi.org/10.5281/zenodo.7345184" httpUri="https://zenodo.org/record/7345184/files/figure.png" pageId="69" pageNumber="70">Fig. 49</figureCitation>
|
||
).
|
||
</emphasis>
|
||
39 examined specimens from FSCA, MEKRB, MNHN, USNM, SENASA, UGBC.
|
||
<emphasis id="18D4243FFD1AFFC4AFA61F54FD9BBA7D" bold="true" box="[151,549,1538,1565]" pageId="69" pageNumber="70">
|
||
<collectingCountry id="52B7B8BDFD1AFFC4AFA61F54FF4FBA7D" box="[151,241,1538,1564]" name="Chile" pageId="69" pageNumber="70">CHILE</collectingCountry>
|
||
(12).
|
||
<collectingRegion id="E86436CFFD1AFFC4AE091F54FD9EBA7D" box="[312,544,1538,1565]" country="Chile" name="Arica y Parinacota" pageId="69" pageNumber="70">Arica y Parinacota</collectingRegion>
|
||
:
|
||
</emphasis>
|
||
Cuya; Puente Chapa (
|
||
<quantity id="ED5855C8FD1AFFC4AC151F54FCD0BA7C" box="[804,878,1538,1565]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="69" pageNumber="70" unit="km" value="50.0">50 km</quantity>
|
||
S Arica); Valle de Azapa (
|
||
<quantity id="ED5855C8FD1AFFC4ABAE1F55FB67BA7C" box="[1183,1241,1539,1565]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="69" pageNumber="70" unit="km" value="6.0">6 km</quantity>
|
||
E Arica).
|
||
<emphasis id="18D4243FFD1AFFC4AA611F55FEC8BA21" bold="true" pageId="69" pageNumber="70">
|
||
<collectingCountry id="52B7B8BDFD1AFFC4AA611F55FA22BA7C" box="[1360,1436,1539,1565]" name="Peru" pageId="69" pageNumber="70">PERU</collectingCountry>
|
||
(27).
|
||
<collectingRegion id="E86436CFFD1AFFC4AFE51F70FEFBBA21" box="[212,325,1574,1600]" country="Peru" name="Arequipa" pageId="69" pageNumber="70">Arequipa</collectingRegion>
|
||
(3):
|
||
</emphasis>
|
||
Camaná, Quilca; Quiroz.
|
||
<emphasis id="18D4243FFD1AFFC4ADAF1F70FCA3BA21" bold="true" box="[670,797,1574,1600]" pageId="69" pageNumber="70">
|
||
<collectingRegion id="E86436CFFD1AFFC4ADAF1F70FD60BA21" box="[670,734,1574,1600]" country="Peru" name="Lima" pageId="69" pageNumber="70">Lima</collectingRegion>
|
||
(19):
|
||
</emphasis>
|
||
Canta, Santa Rosa de Quives; Cañete; Huarochirí, Santa Eulalia; Huaura, Végueta, Medio Mundo; Lima.
|
||
<emphasis id="18D4243FFD1AFFC4AD881F1DFCF6BA05" bold="true" box="[697,840,1610,1637]" pageId="69" pageNumber="70">No data (5):</emphasis>
|
||
Peru.
|
||
</paragraph>
|
||
<paragraph id="2A1FF82DFD1AFFC4AFF61F39FA3EBAE8" blockId="69.[151,1437,530,1673]" box="[199,1408,1646,1673]" pageId="69" pageNumber="70">
|
||
<emphasis id="18D4243FFD1AFFC4AFF61F39FE3BBAE9" bold="true" box="[199,389,1646,1673]" pageId="69" pageNumber="70">Natural history.</emphasis>
|
||
Specimens have been found between
|
||
<quantity id="ED5855C8FD1AFFC4AC001F39FC09BAE8" box="[817,951,1647,1673]" metricMagnitude="2" metricUnit="m" metricValue="5.24" metricValueMax="10.36" metricValueMin="0.12" pageId="69" pageNumber="70" unit="m" value="524.0" valueMax="1036.0" valueMin="12.0">12–1,036 m</quantity>
|
||
elevation between August and February.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |