treatments-xml/data/82/3A/15/823A151BFFB7EB5A8DC9CA7839890361.xml
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<document ID-DOI="10.1590/S1679-62252012000200001" ID-ISSN="1982-0224" ID-Zenodo-Dep="4647117" approvalRequired="5" approvalRequired_for_textStreams="5" checkinTime="1617110711770" checkinUser="carolina" docAuthor="Dutra ¹, Guilherme M., Wosiacki ¹, Wolmar B. &amp; de Pinna ², Mario C. C." docDate="2012" docId="823A151BFFB7EB5A8DC9CA7839890361" docLanguage="en" docName="NeotropIchthyol.10.2.225-231.pdf" docOrigin="Neotropical Ichthyology 10 (2)" docSource="http://www.scielo.br/scielo.php?script=sci_arttext&amp;pid=S1679-62252012000200001&amp;lng=en&amp;tlng=en" docStyle="DocumentStyle:85A40B78B715C4497F837D9D78407CE5.3:NeotropIchthyol.2003-2013.journal_article" docStyleId="85A40B78B715C4497F837D9D78407CE5" docStyleName="NeotropIchthyol.2003-2013.journal_article" docStyleVersion="3" docTitle="Trichomycterus anhanga Dutra¹ &amp; Wosiacki¹ &amp; de Pinna² 2012, new species" docType="treatment" docVersion="2" lastPageId="4" lastPageNumber="229" masterDocId="7E036D63FFB6EB5E8D39C902382C0729" masterDocTitle="Trichomycterus anhanga, a new species of miniature catfish related to T. hasemani and T. johnsoni (Siluriformes: Trichomycteridae) from the Amazon basin, Brazil" masterLastPageNumber="231" masterPageNumber="225" pageId="1" pageNumber="226" updateTime="1617111146853" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Trichomycterus anhanga, a new species of miniature catfish related to T. hasemani and T. johnsoni (Siluriformes: Trichomycteridae) from the Amazon basin, Brazil</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dutra ¹, Guilherme M.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wosiacki ¹, Wolmar B.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>de Pinna ², Mario C. C.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Neotropical Ichthyology</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2012</mods:date>
<mods:detail type="pubDate">
<mods:number>2012-12-31</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>10</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>225</mods:start>
<mods:end>231</mods:end>
</mods:extent>
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<mods:location>
<mods:url>http://www.scielo.br/scielo.php?script=sci_arttext&amp;pid=S1679-62252012000200001&amp;lng=en&amp;tlng=en</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1590/S1679-62252012000200001</mods:identifier>
<mods:identifier type="ISSN">1982-0224</mods:identifier>
<mods:identifier type="Zenodo-Dep">4647117</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:823A151BFFB7EB5A8DC9CA7839890361" httpUri="http://treatment.plazi.org/id/823A151BFFB7EB5A8DC9CA7839890361" lastPageId="4" lastPageNumber="229" pageId="1" pageNumber="226">
<subSubSection box="[240,660,890,914]" pageId="1" pageNumber="226" type="nomenclature">
<paragraph blockId="1.[240,660,890,946]" box="[240,660,890,914]" pageId="1" pageNumber="226">
<heading box="[240,660,890,914]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="226" reason="2">
<emphasis bold="true" box="[240,660,890,914]" pageId="1" pageNumber="226">
<taxonomicName authority="Dutra¹ &amp; Wosiacki¹ &amp; de Pinna², 2012" authorityName="Dutra¹ &amp; Wosiacki¹ &amp; de Pinna²" authorityYear="2012" box="[240,518,890,914]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="226" phylum="Chordata" rank="species" species="anhanga" status="new species">
<emphasis bold="true" box="[240,518,890,914]" italics="true" pageId="1" pageNumber="226">Trichomycterus anhanga</emphasis>
</taxonomicName>
,
<taxonomicNameLabel box="[530,660,890,914]" pageId="1" pageNumber="226" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[367,532,922,946]" pageId="1" pageNumber="226" type="description">
<paragraph blockId="1.[240,660,890,946]" box="[367,532,922,946]" pageId="1" pageNumber="226">
<emphasis bold="true" box="[367,532,922,946]" pageId="1" pageNumber="226">
<figureCitation box="[367,441,922,946]" captionStart="Fig" captionStartId="2.[136,175,1593,1617]" captionTargetBox="[136,1459,174,1569]" captionTargetId="figure-149@2.[136,1459,174,1569]" captionTargetPageId="2" captionText="Fig. 1. Trichomycterus anhanga, holotype, INPA 25125, 10.0 mm SL. Brazil, Amazonas, igarapé Extrema, tributary of rio Aripuanã, rio Madeira drainage. a) lateral view; b) dorsal view; c) ventral view." figureDoi="http://doi.org/10.5281/zenodo.4647119" httpUri="https://zenodo.org/record/4647119/files/figure.png" pageId="1" pageNumber="226">Figs. 1</figureCitation>
,
<figureCitation box="[454,482,922,946]" captionStart="Fig" captionStartId="3.[129,168,666,690]" captionTargetBox="[141,759,201,618]" captionTargetId="figure-540@3.[129,772,178,631]" captionTargetPageId="3" captionText="Fig. 2. Palatine in Trichomycterus, dorsal view. a) T.anhanga, left side, INPA 36117, paratype; b) T. hasemani, inverted of right side, MPEG 4968. Scale bars = 0.1 mm. 1 = anterior articular facets; 2 = posterior articular facets." figureDoi="http://doi.org/10.5281/zenodo.4647121" httpUri="https://zenodo.org/record/4647121/files/figure.png" pageId="1" pageNumber="226">2a</figureCitation>
,
<figureCitation box="[493,507,922,946]" captionStart="Fig" captionStartId="3.[809,848,846,870]" captionTargetBox="[821,1392,185,792]" captionTargetId="figure-541@3.[809,1452,166,825]" captionTargetPageId="3" captionText="Fig. 3. Jaws of Trichomycterus anhanga, MZUSP 108822. Scale bars = 0.1 mm.a) upper jaw in ventral view; b) lower jaw in ventrolateral view, left side. PMX = premaxilar, MET = mesethmoid, DEN = dentary;AAR = anguloarticular." figureDoi="http://doi.org/10.5281/zenodo.4647123" httpUri="https://zenodo.org/record/4647123/files/figure.png" pageId="1" pageNumber="226">3</figureCitation>
,
<figureCitation box="[519,532,922,946]" captionStart="Fig" captionStartId="4.[136,175,798,822]" captionTargetBox="[151,771,189,759]" captionTargetId="figure-566@4.[136,779,174,770]" captionTargetPageId="4" captionText="Fig. 4. Opercle and Interopercle of Trichomycterus anhanga, lateral view, left side, INPA 36117, paratype. OP = opercle; IO = interopercle.Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4647125" httpUri="https://zenodo.org/record/4647125/files/figure.png" pageId="1" pageNumber="226">4</figureCitation>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="226" type="materials_examined">
<paragraph blockId="1.[129,771,986,1108]" pageId="1" pageNumber="226">
<materialsCitation collectingDate="2005-04-29" collectionCode="INPA" collectorName="L. Rapp &amp; Daniel, O. M &amp; Ribeiro, L. M &amp; A. Galuch." country="Brazil" county="Novo Aripuana" latitude="-5.120556" location="rio Aripuana" longLatPrecision="1" longitude="-60.379692" municipality="Extrema" pageId="1" pageNumber="226" specimenCode="INPA 25125" specimenCount="1" stateProvince="Amazonas" typeStatus="holotype">
<emphasis bold="true" box="[129,230,986,1007]" pageId="1" pageNumber="226">
<typeStatus box="[129,226,986,1007]" pageId="1" pageNumber="226">Holotype</typeStatus>
.
</emphasis>
<specimenCode box="[240,367,986,1008]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="1" pageNumber="226">INPA 25125</specimenCode>
, 10.0 mm SL,
<collectingCountry box="[523,586,986,1008]" name="Brazil" pageId="1" pageNumber="226">Brazil</collectingCountry>
,
<collectingRegion box="[597,703,986,1007]" country="Brazil" name="Amazonas" pageId="1" pageNumber="226">Amazonas</collectingRegion>
,
<collectingCounty pageId="1" pageNumber="226">Novo Aripuanã</collectingCounty>
, igarapé
<collectingMunicipality box="[310,395,1015,1036]" pageId="1" pageNumber="226">Extrema</collectingMunicipality>
, tributary of
<location LSID="urn:lsid:plazi:treatment:823A151BFFB7EB5A8DC9CA7839890361:0F4CF2D6FFB7EB5F8F30CAF53AAB0324" box="[521,647,1015,1037]" country="Brazil" county="Novo Aripuana" latitude="-5.120556" longLatPrecision="1" longitude="-60.379692" municipality="Extrema" name="rio Aripuana" pageId="1" pageNumber="226" stateProvince="Amazonas">rio Aripuanã</location>
,
<location LSID="urn:lsid:plazi:treatment:823A151BFFB7EB5A8DC9CA7839890361:0F4CF2D6FFB7EB5F8FA8CAF53B2F0324" box="[657,771,1015,1037]" country="Brazil" county="Novo Aripuana" latitude="-5.120556" longLatPrecision="1" longitude="-60.379692" municipality="Extrema" name="rio Madeira" pageId="1" pageNumber="226" stateProvince="Amazonas">rio Madeira</location>
drainage,
<geoCoordinate box="[225,359,1049,1071]" degrees="05" direction="south" minutes="07" orientation="latitude" pageId="1" pageNumber="226" precision="1" seconds="14.0" value="-5.120556">05°0714.0”S</geoCoordinate>
<geoCoordinate box="[365,508,1049,1071]" degrees="60" direction="west" minutes="22" orientation="longitude" pageId="1" pageNumber="226" precision="1" seconds="46.9" value="-60.379692">60°2246.9”W</geoCoordinate>
,
<date box="[518,642,1049,1071]" pageId="1" pageNumber="226" value="2005-04-29">
<collectingDate box="[518,642,1049,1071]" pageId="1" pageNumber="226" value="2005-04-29">29 Apr 2005</collectingDate>
</date>
,
<collectorName box="[652,731,1049,1070]" pageId="1" pageNumber="226">L. Rapp</collectorName>
Py-
<collectorName box="[129,259,1086,1108]" pageId="1" pageNumber="226">Daniel, O. M</collectorName>
.
<collectorName box="[269,405,1086,1108]" pageId="1" pageNumber="226">Ribeiro, L. M</collectorName>
. Sousa &amp;
<collectorName box="[503,610,1086,1108]" pageId="1" pageNumber="226">A. Galuch.</collectorName>
</materialsCitation>
</paragraph>
<paragraph blockId="1.[129,772,1142,1481]" pageId="1" pageNumber="226">
<materialsCitation collectingDate="2005-04-29" collectionCode="INPA" collectorName="L. Rapp &amp; Daniel, O. M &amp; Ribeiro, L. M &amp; A. Galuch." country="Brazil" county="Novo Aripuana" latitude="-5.120556" location="rio Aripuana" longLatPrecision="1" longitude="-60.379692" municipality="Extrema" pageId="1" pageNumber="226" specimenCode="INPA 36117, 3" specimenCount="1" stateProvince="Amazonas" typeStatus="paratype">
<emphasis bold="true" box="[129,424,1142,1164]" pageId="1" pageNumber="226">
<typeStatus box="[129,233,1143,1164]" pageId="1" pageNumber="226">Paratypes</typeStatus>
(all from
<collectingCountry box="[346,413,1142,1164]" name="Brazil" pageId="1" pageNumber="226">Brazil</collectingCountry>
).
</emphasis>
<specimenCode box="[433,559,1143,1165]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="1" pageNumber="226">INPA 36117</specimenCode>
,
<specimenCode box="[571,584,1143,1164]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="1" pageNumber="226">3</specimenCode>
,
<quantity box="[599,728,1143,1165]" metricMagnitude="-2" metricUnit="m" metricValue="1.02" metricValueMax="1.06" metricValueMin="0.98" pageId="1" pageNumber="226" unit="mm" value="10.2" valueMax="10.6" valueMin="9.8">9.8-10.6 mm</quantity>
SL, same data as holotype
</materialsCitation>
.
<materialsCitation collectingDate="2005-04-29" collectionCode="MZUSP" collectorName="L. Rapp &amp; Daniel, O. M &amp; Ribeiro, L. M &amp; A. Galuch." country="Brazil" county="Novo Aripuana" latitude="-5.120556" location="rio Aripuana" longLatPrecision="1" longitude="-60.379692" municipality="Extrema" pageId="1" pageNumber="226" specimenCode="MZUSP 108822, 2" specimenCount="1" stateProvince="Amazonas" typeStatus="paratype">
<specimenCode box="[366,530,1172,1193]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="226">MZUSP 108822</specimenCode>
,
<specimenCode box="[543,557,1172,1193]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="226">2</specimenCode>
+1 c&amp;s,
<quantity box="[644,772,1172,1193]" metricMagnitude="-3" metricUnit="m" metricValue="9.95" metricValueMax="10.2" metricValueMin="9.7" pageId="1" pageNumber="226" unit="mm" value="9.95" valueMax="10.2" valueMin="9.7">9.7-10.2 mm</quantity>
SL, same data as holotype
</materialsCitation>
.
<materialsCitation collectingDate="2007-09-06" collectionCode="INPA" country="Brazil" county="Novo Aripuana" latitude="-5.9922223" location="rio Aripuana" longLatPrecision="1" longitude="-60.209724" municipality="Palhalzinho" pageId="1" pageNumber="226" specimenCode="INPA 31402, 1" specimenCount="1" stateProvince="Amazonas" typeStatus="paratype">
<specimenCode box="[386,507,1200,1221]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="1" pageNumber="226">INPA 31402</specimenCode>
,
<specimenCode box="[515,527,1200,1221]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="1" pageNumber="226">1</specimenCode>
,
<quantity box="[537,621,1200,1222]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="1" pageNumber="226" unit="mm" value="10.5">10.5 mm</quantity>
SL,
<collectingRegion box="[663,767,1200,1221]" country="Brazil" name="Amazonas" pageId="1" pageNumber="226">Amazonas</collectingRegion>
,
<collectingCounty box="[129,280,1229,1251]" pageId="1" pageNumber="226">Novo Aripuanã</collectingCounty>
, igarapé
<collectingMunicipality box="[367,485,1229,1251]" pageId="1" pageNumber="226">Palhalzinho</collectingMunicipality>
, tributary of
<location LSID="urn:lsid:plazi:treatment:823A151BFFB7EB5A8DC9CA7839890361:0F4CF2D6FFB7EB5F8F5BCDCF3AF203CA" box="[610,734,1229,1251]" country="Brazil" county="Novo Aripuana" latitude="-5.9922223" longLatPrecision="1" longitude="-60.209724" municipality="Palhalzinho" name="rio Aripuana" pageId="1" pageNumber="226" stateProvince="Amazonas">rio Aripuanã</location>
,
<location LSID="urn:lsid:plazi:treatment:823A151BFFB7EB5A8DC9CA7839890361:0F4CF2D6FFB7EB5F8FD1CDCF38FD0229" country="Brazil" county="Novo Aripuana" latitude="-5.9922223" longLatPrecision="1" longitude="-60.209724" municipality="Palhalzinho" name="rio Madeira" pageId="1" pageNumber="226" stateProvince="Amazonas">rio Madeira</location>
drainage,
<geoCoordinate box="[308,440,1258,1280]" degrees="05" direction="south" minutes="59" orientation="latitude" pageId="1" pageNumber="226" precision="1" seconds="32.0" value="-5.9922223">05°5932.0”S</geoCoordinate>
<geoCoordinate box="[445,586,1258,1280]" degrees="60" direction="west" minutes="12" orientation="longitude" pageId="1" pageNumber="226" precision="1" seconds="35.0" value="-60.209724">60°1235.0”W</geoCoordinate>
,
<date box="[595,715,1258,1280]" pageId="1" pageNumber="226" value="2007-09-06">
<collectingDate box="[595,715,1258,1280]" pageId="1" pageNumber="226" value="2007-09-06">06 Sep 2007</collectingDate>
</date>
, L
</materialsCitation>
.
<materialsCitation collectionCode="R" collectorName="Py-Daniel, M. S &amp; Rocha" country="Brazil" pageId="1" pageNumber="226" specimenCount="1" typeStatus="paratype">
<collectionCode box="[749,768,1258,1279]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="226">R</collectionCode>
.
<collectorName box="[129,291,1287,1309]" pageId="1" pageNumber="226">Py-Daniel, M. S</collectorName>
.
<collectorName box="[302,364,1287,1309]" pageId="1" pageNumber="226">Rocha</collectorName>
</materialsCitation>
&amp;
<materialsCitation box="[397,569,1287,1309]" collectionCode="R" collectorName="Oliveira" country="Brazil" pageId="1" pageNumber="226" specimenCount="1" typeStatus="paratype">
<collectionCode box="[397,416,1287,1308]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="226">R</collectionCode>
.
<collectionCode box="[426,445,1287,1308]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="226">R</collectionCode>
. de
<collectorName box="[486,569,1287,1309]" pageId="1" pageNumber="226">Oliveira</collectorName>
</materialsCitation>
.
<materialsCitation collectingDate="2002-09-12" collectionCode="MPEG" collectorName="W. B. Wosiacki &amp; A. Bezerra" country="Brazil" latitude="-2.5774999" location="Juruti Grande" longLatPrecision="1" longitude="-56.399975" municipality="Juruti" pageId="1" pageNumber="226" specimenCode="MPEG 6973, 1" specimenCount="1" stateProvince="Para" typeStatus="paratype">
<specimenCode box="[579,704,1287,1309]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="1" pageNumber="226">MPEG 6973</specimenCode>
,
<specimenCode box="[715,727,1287,1308]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="1" pageNumber="226">1</specimenCode>
,
<quantity metricMagnitude="-3" metricUnit="m" metricValue="9.9" pageId="1" pageNumber="226" unit="mm" value="9.9">9.9 mm</quantity>
SL,
<collectingRegion box="[224,272,1316,1338]" country="Brazil" name="Para" pageId="1" pageNumber="226">Pará</collectingRegion>
,
<collectingMunicipality box="[287,349,1315,1337]" pageId="1" pageNumber="226">Juruti</collectingMunicipality>
, igarapé
<location LSID="urn:lsid:plazi:treatment:823A151BFFB7EB5A8DC9CA7839890361:0F4CF2D6FFB7EB5F8CF8CC213A790210" box="[449,597,1315,1337]" country="Brazil" latitude="-2.5774999" longLatPrecision="1" longitude="-56.399975" municipality="Juruti" name="Juruti Grande" pageId="1" pageNumber="226" stateProvince="Para">Juruti Grande</location>
, tributary of rio
<collectingRegion box="[129,238,1344,1365]" country="Brazil" name="Amazonas" pageId="1" pageNumber="226">Amazonas</collectingRegion>
,
<geoCoordinate box="[251,390,1344,1365]" degrees="02" direction="south" minutes="34" orientation="latitude" pageId="1" pageNumber="226" precision="1" seconds="39.0" value="-2.5774999">02°3439.0”S</geoCoordinate>
<geoCoordinate box="[400,548,1344,1366]" degrees="56" direction="west" minutes="23" orientation="longitude" pageId="1" pageNumber="226" precision="1" seconds="59.9" value="-56.399975">56°2359.9”W</geoCoordinate>
,
<date box="[562,698,1344,1365]" pageId="1" pageNumber="226" value="2002-09-12">
<collectingDate box="[562,698,1344,1365]" pageId="1" pageNumber="226" value="2002-09-12">12 Sep 2002</collectingDate>
</date>
,
<collectorName pageId="1" pageNumber="226">W. B. Wosiacki</collectorName>
&amp;
<collectorName box="[249,359,1373,1395]" pageId="1" pageNumber="226">A. Bezerra</collectorName>
</materialsCitation>
.
<materialsCitation collectingDate="2007-11-27" collectionCode="MPEG" collectorName="A. Hercos" country="Brazil" county="Juruti" latitude="-2.6116667" location="rio Arua" longLatPrecision="15" longitude="-56.193333" municipality="Mutum" pageId="1" pageNumber="226" specimenCode="MPEG 14356, 1" specimenCount="1" stateProvince="Para" typeStatus="paratype">
<specimenCode box="[368,505,1373,1395]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="1" pageNumber="226">MPEG 14356</specimenCode>
,
<specimenCode box="[516,528,1373,1394]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="1" pageNumber="226">1</specimenCode>
c&amp;s,
<quantity box="[587,674,1373,1395]" metricMagnitude="-2" metricUnit="m" metricValue="1.31" pageId="1" pageNumber="226" unit="mm" value="13.1">13.1 mm</quantity>
SL,
<collectingRegion box="[722,768,1373,1395]" country="Brazil" name="Para" pageId="1" pageNumber="226">Pará</collectingRegion>
,
<collectingCounty box="[129,186,1402,1424]" pageId="1" pageNumber="226">Juruti</collectingCounty>
, igarapé
<collectingMunicipality box="[270,343,1402,1423]" pageId="1" pageNumber="226">Mutum</collectingMunicipality>
, tributary of
<location LSID="urn:lsid:plazi:treatment:823A151BFFB7EB5A8DC9CA7839890361:0F4CF2D6FFB7EB5F8CE9CC783A0C02A6" box="[464,544,1402,1424]" country="Brazil" county="Juruti" latitude="-2.6116667" longLatPrecision="15" longitude="-56.193333" municipality="Mutum" name="rio Arua" pageId="1" pageNumber="226" stateProvince="Para">rio Aruã</location>
, tributary of
<location LSID="urn:lsid:plazi:treatment:823A151BFFB7EB5A8DC9CA7839890361:0F4CF2D6FFB7EB5F8FA1CC783B2C02A6" box="[664,768,1402,1424]" country="Brazil" county="Juruti" latitude="-2.6116667" longLatPrecision="15" longitude="-56.193333" municipality="Mutum" name="rio Branco" pageId="1" pageNumber="226" stateProvince="Para">rio Branco</location>
,
<location LSID="urn:lsid:plazi:treatment:823A151BFFB7EB5A8DC9CA7839890361:0F4CF2D6FFB7EB5F8DB8CC9538D90284" box="[129,245,1431,1453]" country="Brazil" county="Juruti" latitude="-2.6116667" longLatPrecision="15" longitude="-56.193333" municipality="Mutum" name="rio Arapiuns" pageId="1" pageNumber="226" stateProvince="Para">rio Arapiuns</location>
basin,
<location LSID="urn:lsid:plazi:treatment:823A151BFFB7EB5A8DC9CA7839890361:0F4CF2D6FFB7EB5F8C0ECC9539B30284" box="[311,415,1431,1453]" country="Brazil" county="Juruti" latitude="-2.6116667" longLatPrecision="15" longitude="-56.193333" municipality="Mutum" name="rio Tapajos" pageId="1" pageNumber="226" stateProvince="Para">rio Tapajós</location>
drainage,
<geoCoordinate box="[511,642,1431,1453]" degrees="02" direction="south" minutes="36" orientation="latitude" pageId="1" pageNumber="226" precision="1" seconds="42.0" value="-2.6116667">02°3642.0”S</geoCoordinate>
<geoCoordinate box="[647,768,1431,1453]" degrees="56" direction="west" minutes="11" orientation="longitude" pageId="1" pageNumber="226" precision="15" seconds="36" value="-56.193333">56°1136”W</geoCoordinate>
,
<date box="[129,258,1460,1481]" pageId="1" pageNumber="226" value="2007-11-27">
<collectingDate box="[129,258,1460,1481]" pageId="1" pageNumber="226" value="2007-11-27">27 Nov 2007</collectingDate>
</date>
,
<collectorName box="[267,369,1460,1481]" pageId="1" pageNumber="226">A. Hercos</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="226" type="diagnosis">
<paragraph blockId="1.[129,773,1520,1961]" pageId="1" pageNumber="226">
<emphasis bold="true" box="[129,247,1520,1544]" pageId="1" pageNumber="226">Diagnosis.</emphasis>
<taxonomicName authorityName="Dutra¹ &amp; Wosiacki¹ &amp; de Pinna²" authorityYear="2012" box="[256,527,1521,1545]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="226" phylum="Chordata" rank="species" species="anhanga">
<emphasis box="[256,527,1521,1545]" italics="true" pageId="1" pageNumber="226">Trichomycterus anhanga</emphasis>
</taxonomicName>
differs from all other congeners by its reduced latero-sensory canal system, which is restricted to a short section between LL1 and LL2, on the pterotic, and entirely absent on the rest of body and cranium. It also differs from all other species currently in
<taxonomicName authorityName="Valenciennes in Humboldt" authorityYear="1832" box="[610,771,1649,1673]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="226" phylum="Chordata" rank="genus">
<emphasis box="[610,771,1649,1673]" italics="true" pageId="1" pageNumber="226">Trichomycterus</emphasis>
</taxonomicName>
except
<taxonomicName baseAuthorityName="Eigenmann" baseAuthorityYear="1914" box="[222,374,1681,1705]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="226" phylum="Chordata" rank="species" species="hasemani">
<emphasis box="[222,374,1681,1705]" italics="true" pageId="1" pageNumber="226">T. hasemani</emphasis>
</taxonomicName>
and
<taxonomicName baseAuthorityName="Fowler" baseAuthorityYear="1932" box="[446,588,1681,1705]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="226" phylum="Chordata" rank="species" species="johnsoni">
<emphasis box="[446,588,1681,1705]" italics="true" pageId="1" pageNumber="226">T. johnsoni</emphasis>
</taxonomicName>
by the single hypertrophied cranial fontanel, which occupies nearly the entire skull roof (
<emphasis box="[310,332,1745,1768]" italics="true" pageId="1" pageNumber="226">vs</emphasis>
. fontanel restricted to one or more narrow spaces along the dorsal midline of skull).
</paragraph>
<paragraph blockId="1.[129,773,1520,1961]" lastBlockId="1.[809,1452,178,911]" pageId="1" pageNumber="226">
<taxonomicName authorityName="Dutra¹ &amp; Wosiacki¹ &amp; de Pinna²" authorityYear="2012" box="[166,430,1809,1833]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="226" phylum="Chordata" rank="species" species="anhanga">
<emphasis box="[166,430,1809,1833]" italics="true" pageId="1" pageNumber="226">Trichomycterus anhanga</emphasis>
</taxonomicName>
differs further from
<taxonomicName baseAuthorityName="Eigenmann" baseAuthorityYear="1914" box="[645,771,1809,1833]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="226" phylum="Chordata" rank="species" species="hasemani">
<emphasis box="[645,771,1809,1833]" italics="true" pageId="1" pageNumber="226">T. hasemani</emphasis>
</taxonomicName>
and
<taxonomicName baseAuthorityName="Fowler" baseAuthorityYear="1932" box="[172,288,1841,1865]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="226" phylum="Chordata" rank="species" species="johnsoni">
<emphasis box="[172,288,1841,1865]" italics="true" pageId="1" pageNumber="226">T. johnsoni</emphasis>
</taxonomicName>
by the presence of three (i,2) pectoral-fin rays (
<emphasis box="[138,160,1873,1896]" italics="true" pageId="1" pageNumber="226">vs</emphasis>
. i,3-5 rays); the absence of pelvic fins and girdle (
<emphasis box="[742,764,1873,1896]" italics="true" pageId="1" pageNumber="226">vs</emphasis>
. presence); the reduced oral dentition, with only two teeth each on premaxilla and dentary (
<emphasis box="[423,452,1937,1960]" italics="true" pageId="1" pageNumber="226">vs.</emphasis>
teeth numerous in both upper and lower jaws); the reduced pharyngeal dentition, with only a single tooth in the lower pharyngeal plate and none in the upper (
<emphasis box="[883,912,243,266]" italics="true" pageId="1" pageNumber="226">vs.</emphasis>
pharyngeal dentition well developed); the presence of five to seven odontodes in the interopercle (
<emphasis box="[1311,1333,275,298]" italics="true" pageId="1" pageNumber="226">vs</emphasis>
.
<quantity box="[1348,1425,274,298]" metricMagnitude="-1" metricUnit="m" metricValue="2.413" metricValueMax="2.54" metricValueMin="2.286" pageId="1" pageNumber="226" unit="in" value="9.5" valueMax="10.0" valueMin="9.0">9-10 in</quantity>
<taxonomicName baseAuthorityName="Eigenmann" baseAuthorityYear="1914" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="226" phylum="Chordata" rank="species" species="hasemani">
<emphasis italics="true" pageId="1" pageNumber="226">T. hasemani</emphasis>
</taxonomicName>
and
<quantity box="[962,1038,306,330]" metricMagnitude="-1" metricUnit="m" metricValue="2.667" metricValueMax="3.048" metricValueMin="2.286" pageId="1" pageNumber="226" unit="in" value="10.5" valueMax="12.0" valueMin="9.0">9-12 in</quantity>
<taxonomicName baseAuthorityName="Fowler" baseAuthorityYear="1932" box="[1044,1161,306,330]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="226" phylum="Chordata" rank="species" species="johnsoni">
<emphasis box="[1044,1161,306,330]" italics="true" pageId="1" pageNumber="226">T. johnsoni</emphasis>
</taxonomicName>
), the absence of procurrent rays anterior to dorsal and anal fins (
<emphasis box="[1171,1192,339,362]" italics="true" pageId="1" pageNumber="226">vs</emphasis>
. one vestigial ray anterior to the main ray series in each fin), the lack of a lateral series of dark spots (
<emphasis box="[930,952,403,426]" italics="true" pageId="1" pageNumber="226">vs</emphasis>
. presence); the presence of a small dark spot on the ventral surface of the lower lip (
<emphasis box="[1194,1216,438,461]" italics="true" pageId="1" pageNumber="226">vs</emphasis>
. absence); the narrow comma-shaped palatine, lacking a posterior process (
<emphasis box="[1339,1360,472,495]" italics="true" pageId="1" pageNumber="226">vs</emphasis>
. palatine broad, with large posterior process) (
<figureCitation box="[1218,1287,503,527]" captionStart="Fig" captionStartId="3.[129,168,666,690]" captionTargetBox="[141,759,201,618]" captionTargetId="figure-540@3.[129,772,178,631]" captionTargetPageId="3" captionText="Fig. 2. Palatine in Trichomycterus, dorsal view. a) T.anhanga, left side, INPA 36117, paratype; b) T. hasemani, inverted of right side, MPEG 4968. Scale bars = 0.1 mm. 1 = anterior articular facets; 2 = posterior articular facets." figureDoi="http://doi.org/10.5281/zenodo.4647121" httpUri="https://zenodo.org/record/4647121/files/figure.png" pageId="1" pageNumber="226">Fig. 2</figureCitation>
); the lack of a parasphenoid (
<emphasis box="[969,991,536,559]" italics="true" pageId="1" pageNumber="226">vs</emphasis>
. parasphenoid present); the presence of a single pair of pleural ribs (
<emphasis box="[1079,1101,568,591]" italics="true" pageId="1" pageNumber="226">vs</emphasis>
. two or three), the insertion of the first dorsal-fin pterygiophore anterior to the neural spine of the 16
<superScript attach="left" box="[835,847,629,643]" fontSize="6" pageId="1" pageNumber="226">th</superScript>
vertebra (
<emphasis box="[953,975,632,655]" italics="true" pageId="1" pageNumber="226">vs</emphasis>
. 17
<superScript attach="left" box="[1013,1025,629,643]" fontSize="6" pageId="1" pageNumber="226">th</superScript>
or 18
<superScript attach="left" box="[1083,1095,629,643]" fontSize="6" pageId="1" pageNumber="226">th</superScript>
vertebra in
<taxonomicName baseAuthorityName="Fowler" baseAuthorityYear="1932" box="[1218,1333,631,655]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="226" phylum="Chordata" rank="species" species="johnsoni">
<emphasis box="[1218,1333,631,655]" italics="true" pageId="1" pageNumber="226">T. johnsoni</emphasis>
</taxonomicName>
, and 18
<superScript attach="left" box="[1414,1426,629,643]" fontSize="6" pageId="1" pageNumber="226">th</superScript>
to 20
<superScript attach="left" box="[835,847,661,675]" fontSize="6" pageId="1" pageNumber="226">th</superScript>
in
<taxonomicName baseAuthorityName="Eigenmann" baseAuthorityYear="1914" box="[885,1014,663,687]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="226" phylum="Chordata" rank="species" species="hasemani">
<emphasis box="[885,1014,663,687]" italics="true" pageId="1" pageNumber="226">T. hasemani</emphasis>
</taxonomicName>
). It can be further distinguished from
<taxonomicName baseAuthorityName="Eigenmann" baseAuthorityYear="1914" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="226" phylum="Chordata" rank="species" species="hasemani">
<emphasis italics="true" pageId="1" pageNumber="226">T. hasemani</emphasis>
</taxonomicName>
by the insertion of the first anal-fin pterygiophore anterior to the hemal spine of 16
<superScript attach="right" box="[1134,1146,725,739]" fontSize="6" pageId="1" pageNumber="226">th</superScript>
vertebra (
<emphasis box="[1246,1275,728,751]" italics="true" pageId="1" pageNumber="226">vs.</emphasis>
anterior to hemal spine of 19
<superScript attach="none" box="[920,939,757,783]" fontSize="6" pageId="1" pageNumber="226">th-</superScript>
20
<superScript attach="left" box="[967,979,757,771]" fontSize="6" pageId="1" pageNumber="226">th</superScript>
vertebra); and from
<taxonomicName baseAuthorityName="Fowler" baseAuthorityYear="1932" box="[1185,1299,759,783]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="226" phylum="Chordata" rank="species" species="johnsoni">
<emphasis box="[1185,1299,759,783]" italics="true" pageId="1" pageNumber="226">T. johnsoni</emphasis>
</taxonomicName>
by the absence of irregular longitudinal dark lines on the sides of the body (
<emphasis box="[1423,1445,792,815]" italics="true" pageId="1" pageNumber="226">vs</emphasis>
. presence), the absence of a dark midventral line (
<emphasis box="[1311,1333,824,847]" italics="true" pageId="1" pageNumber="226">vs</emphasis>
. presence), and the smaller interorbital distance (17.8-23.2% SL
<emphasis box="[1362,1384,856,879]" italics="true" pageId="1" pageNumber="226">vs</emphasis>
. 25.5- 37.0% SL).
</paragraph>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="228" pageId="1" pageNumber="226" type="description">
<paragraph blockId="1.[809,1452,944,1961]" pageId="1" pageNumber="226">
<emphasis bold="true" box="[809,952,944,968]" pageId="1" pageNumber="226">Description.</emphasis>
Morphometric data given in
<tableCitation box="[1290,1376,945,969]" captionStart="Table 1" captionStartId="4.[816,879,177,201]" captionText="Table 1. Morphometric data for holotype and paratypes of Trichomycterus anhanga (n = 9 including holotype)." pageId="1" pageNumber="226">Table 1</tableCitation>
. Body elongate, approximately cylindrical near head, progressively more compressed posteriorly to strongly compressed caudal peduncle. Dorsal profile gently convex from tip of snout to posterior margin of head and straight from this point to end of caudal peduncle. Dorsal-fin base in recess on dorsal profile of body. Ventral profile straight from snout tip to pectoral-fin origin, gently convex from that point to anus, then straight to end of caudal peduncle. Anal opening on vertical line through origin of dorsal fin.
</paragraph>
<paragraph blockId="1.[809,1452,944,1961]" pageId="1" pageNumber="226">Head depressed, its greatest width across bases of interopercular odontodes; snout round in dorsal view; lateral surface of head with well-developed muscle protruding from posterior margin of eye to opercular patch of odontodes. Eye small and circular, with well-defined margin, located on anterior half of HL, dorsally oriented.</paragraph>
<paragraph blockId="1.[809,1452,944,1961]" pageId="1" pageNumber="226">Anterior nostril round, its diameter equal to eye lens, surrounded by short tube of integument continuous posterolaterally with nasal barbel. Posterior nostril occluded anteriorly by fleshy integument shaped like a half-shell, positioned closer to eye than to anterior nostril. Posterior nostril slightly displaced medially relative to line between anterior nostril and eye.</paragraph>
<paragraph blockId="1.[809,1452,944,1961]" pageId="1" pageNumber="226">
Mouth subterminal, with corners slightly posterolaterally oriented. Upper lip extending slightly further anteriorly than lower one, its anterior margin gently rounded in ventral view, continuous laterally with bases of maxillary and rictal barbels. Lower lip thinner than upper one, its anterior margin gently rounded in ventral view. Jaw dentition reduced, with two conical feeble teeth on each premaxilla and each dentary (
<figureCitation box="[1324,1387,1873,1897]" captionStart="Fig" captionStartId="3.[809,848,846,870]" captionTargetBox="[821,1392,185,792]" captionTargetId="figure-541@3.[809,1452,166,825]" captionTargetPageId="3" captionText="Fig. 3. Jaws of Trichomycterus anhanga, MZUSP 108822. Scale bars = 0.1 mm.a) upper jaw in ventral view; b) lower jaw in ventrolateral view, left side. PMX = premaxilar, MET = mesethmoid, DEN = dentary;AAR = anguloarticular." figureDoi="http://doi.org/10.5281/zenodo.4647123" httpUri="https://zenodo.org/record/4647123/files/figure.png" pageId="1" pageNumber="226">Fig. 3</figureCitation>
).
</paragraph>
<paragraph blockId="1.[809,1452,944,1961]" lastBlockId="2.[136,779,1713,1961]" lastPageId="2" lastPageNumber="227" pageId="1" pageNumber="226">Barbels elongate, wider at base, narrowing gradually distally, ending in sharp filament. Origin of nasal barbel on posterolateral portion of anterior nostril, reaching posterior margin of eye or slightly beyond that. Maxillary barbel variably reaching to point between base and posterior limit of interopercular patch of odontodes. Rictal barbel variably reaching to point between base and middle of interopercular patch of odontodes; its base lacking fleshy lobe.</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4647119" ID-Zenodo-Dep="4647119" httpUri="https://zenodo.org/record/4647119/files/figure.png" pageId="2" pageNumber="227" startId="2.[136,175,1593,1617]" targetBox="[136,1459,174,1569]" targetPageId="2">
<paragraph blockId="2.[136,1459,1593,1655]" pageId="2" pageNumber="227">
<emphasis bold="true" box="[136,207,1593,1618]" pageId="2" pageNumber="227">Fig. 1.</emphasis>
<taxonomicName authorityName="Dutra¹ &amp; Wosiacki¹ &amp; de Pinna²" authorityYear="2012" box="[215,484,1594,1618]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="227" phylum="Chordata" rank="species" species="anhanga">
<emphasis box="[215,484,1594,1618]" italics="true" pageId="2" pageNumber="227">Trichomycterus anhanga</emphasis>
</taxonomicName>
, holotype, INPA 25125, 10.0 mm SL. Brazil, Amazonas, igarapé Extrema, tributary of rio Aripuanã, rio Madeira drainage.
<emphasis bold="true" box="[481,494,1631,1655]" pageId="2" pageNumber="227">a</emphasis>
) lateral view;
<emphasis bold="true" box="[645,660,1631,1655]" pageId="2" pageNumber="227">b</emphasis>
) dorsal view;
<emphasis bold="true" box="[810,822,1631,1655]" pageId="2" pageNumber="227">c</emphasis>
) ventral view.
</paragraph>
</caption>
<paragraph blockId="2.[136,779,1713,1961]" lastBlockId="2.[816,1460,1713,1961]" pageId="2" pageNumber="227">
Interopercle with five to seven (six*) odontodes. Interopercular patch with odontodes arranged in two main series. Opercle with six to 10 (seven*) odontodes (
<figureCitation box="[1378,1446,1713,1737]" captionStart="Fig" captionStartId="4.[136,175,798,822]" captionTargetBox="[151,771,189,759]" captionTargetId="figure-566@4.[136,779,174,770]" captionTargetPageId="4" captionText="Fig. 4. Opercle and Interopercle of Trichomycterus anhanga, lateral view, left side, INPA 36117, paratype. OP = opercle; IO = interopercle.Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4647125" httpUri="https://zenodo.org/record/4647125/files/figure.png" pageId="2" pageNumber="227">Fig. 4</figureCitation>
). Odontodes on anterior series straight and those on posterior series slightly longer and curved medially. Opercle with small odontodes anterior to first series. Gill membrane united to isthmus only at anteriormost portion. Gill opening wide. Branchiostegal rays, four* or five.
</paragraph>
<paragraph blockId="2.[816,1460,1713,1961]" lastBlockId="3.[129,772,880,1481]" lastPageId="3" lastPageNumber="228" pageId="2" pageNumber="227">Pectoral fin i,2, with first ray prolonged as filament and branched rays approximately one third of length of first ray. Dorsal fin ii,5(5), iii,3(1), or iii,4*(2), its distal margin rounded; unbranched rays gradually increasing in size, last unbranched ray and first branched ray larger than others, branched rays gradually decreasing in size. Anal fin ii,4, its distal margin gently convex; first unbranched ray two-thirds of length of second ray; second unbranched ray and first branched ray longer than others; branched rays slightly decreasing in size, with their origin at vertical through second or third dorsal-fin ray. Caudal fin truncate, i,8,i(1), i,8,ii(1), ii,7,ii(2) or iii,7,ii*(5). Procurrent caudal-fin rays six or seven* dorsally (eight visible in c&amp;s), and six* or seven ventrally, visible in alcoholic specimens. Pelvic fin and girdle absent.</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4647121" ID-Zenodo-Dep="4647121" httpUri="https://zenodo.org/record/4647121/files/figure.png" pageId="3" pageNumber="228" startId="3.[129,168,666,690]" targetBox="[141,759,201,618]" targetPageId="3">
<paragraph blockId="3.[129,771,666,786]" pageId="3" pageNumber="228">
<emphasis bold="true" box="[129,196,666,690]" pageId="3" pageNumber="228">Fig. 2.</emphasis>
Palatine in
<taxonomicName authorityName="Valenciennes in Humboldt" authorityYear="1832" box="[316,480,666,690]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="228" phylum="Chordata" rank="genus">
<emphasis box="[316,480,666,690]" italics="true" pageId="3" pageNumber="228">Trichomycterus</emphasis>
</taxonomicName>
, dorsal view.
<emphasis bold="true" box="[622,635,666,690]" pageId="3" pageNumber="228">a</emphasis>
)
<taxonomicName authorityName="Dutra¹ &amp; Wosiacki¹ &amp; de Pinna²" authorityYear="2012" box="[649,764,666,690]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="228" phylum="Chordata" rank="species" species="anhanga">
<emphasis box="[649,764,666,690]" italics="true" pageId="3" pageNumber="228">T. anhanga</emphasis>
</taxonomicName>
, left side, INPA 36117, paratype;
<emphasis bold="true" box="[480,495,698,722]" pageId="3" pageNumber="228">b</emphasis>
)
<taxonomicName baseAuthorityName="Eigenmann" baseAuthorityYear="1914" box="[511,640,698,722]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="228" phylum="Chordata" rank="species" species="hasemani">
<emphasis box="[511,640,698,722]" italics="true" pageId="3" pageNumber="228">T. hasemani</emphasis>
</taxonomicName>
, inverted of right side, MPEG 4968. Scale bars = 0.1 mm. 1 = anterior articular facets; 2 = posterior articular facets.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4647123" ID-Zenodo-Dep="4647123" httpUri="https://zenodo.org/record/4647123/files/figure.png" pageId="3" pageNumber="228" startId="3.[809,848,846,870]" targetBox="[821,1392,185,792]" targetPageId="3">
<paragraph blockId="3.[809,1452,846,967]" pageId="3" pageNumber="228">
<emphasis bold="true" box="[809,881,846,870]" pageId="3" pageNumber="228">Fig. 3.</emphasis>
Jaws of
<taxonomicName authority=", MZUSP" authorityName="MZUSP" box="[982,1356,846,870]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="228" phylum="Chordata" rank="species" species="anhanga">
<emphasis box="[982,1252,846,870]" italics="true" pageId="3" pageNumber="228">Trichomycterus anhanga</emphasis>
, MZUSP
</taxonomicName>
108822. Scale bars = 0.1 mm.
<emphasis bold="true" box="[1025,1038,878,902]" pageId="3" pageNumber="228">a</emphasis>
) upper jaw in ventral view;
<emphasis bold="true" box="[1322,1337,878,902]" pageId="3" pageNumber="228">b</emphasis>
) lower jaw in ventrolateral view, left side. PMX = premaxilar, MET = mesethmoid, DEN = dentary; AAR = anguloarticular.
</paragraph>
</caption>
<paragraph blockId="3.[129,772,880,1481]" pageId="3" pageNumber="228">
Lateral line with two pores dorsal to pectoral-fin insertion, LL1 and LL2. Cephalic portion of latero-sensory canal system (supraorbital, infraorbital and preorbital) absent. Vertebrae 29(1), 31(3), or 32(4). One pair of pleural ribs. Dorsal fin with seven or eight pterygiophores, first one anterior to neural spine of 16
<superScript attach="none" box="[759,771,1391,1405]" fontSize="6" pageId="3" pageNumber="228">th</superScript>
or 17
<superScript attach="left" box="[181,193,1423,1437]" fontSize="6" pageId="3" pageNumber="228">th</superScript>
free vertebrae.Anal fin with six pterygiophores, first one anterior to hemal spine of 16
<superScript attach="left" box="[425,437,1455,1469]" fontSize="6" pageId="3" pageNumber="228">th</superScript>
vertebra.
</paragraph>
<paragraph blockId="3.[129,772,1520,1961]" pageId="3" pageNumber="228">
<emphasis bold="true" box="[129,323,1520,1545]" pageId="3" pageNumber="228">Color in ethanol.</emphasis>
Ground color cream. Flank, dorsum, and posteroventral region with tiny light brown spots, irregular in shape. Dorsum with series of four to eight transverse dark brown spots; when eight, first to fifth ones anterior to dorsal fin insertion, sixth over dorsal-fin origin, seventh on vertical line through posterior margin of dorsal fin and eighth on posterior portion of caudal peduncle. Series of three to five dark brown transverse spots on ventral surface of body; first one immediately anterior to anus, second at origin of anal-fin base, third immediately posterior to anal-fin base, fourth approximately in middle of caudal peduncle, and fifth near insertion of first ventral caudal-fin procurrent rays.
</paragraph>
<paragraph blockId="3.[129,772,1520,1961]" lastBlockId="3.[809,1452,1036,1609]" pageId="3" pageNumber="228">Ground color of head cream. Snout with spot extending from base of maxillary barbel to anterior margin of eye. Region of neurocranium with light brown spot extending lateroventrally to interopercular patch of odontodes, sometimes merging with first dorsal spot. Ventral surface of head with few light brown chromatophores concentrated between interopercular odontodes. Ventral surface of lower lip with conspicuous dark spot at symphysis. Opercular and interopercular patches of odontodes with few chromatophores more concentrated at odontode bases. Nasal, maxillary and rictal barbels with small brown spots on dorsal surface, more concentrated basally.</paragraph>
<paragraph blockId="3.[809,1452,1036,1609]" pageId="3" pageNumber="228">Pectoral fin hyaline. Bases of dorsal and anal fins with light brown spot on insertion of first three rays, individuals without spots sometimes with few brown chromatophores on dorsal and anal fins. Caudal fin with dark brown vertical bar at base, darker across central rays, gradually lighter toward dorsal and ventral ends. Middle part of fin with dark brown chromatophores, more concentrated near spot at base; edge hyaline.</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="228" type="distribution">
<paragraph blockId="3.[809,1452,1648,1865]" pageId="3" pageNumber="228">
<emphasis bold="true" box="[809,957,1648,1672]" pageId="3" pageNumber="228">Distribution.</emphasis>
Known from igarapé da Extrema and igarapé Palhalzinho, tributaries to the rio Aripuanã, itself a tributary to the rio Madeira; igarapé Juruti Grande, tributary to the right margin of the rio
<collectingRegion box="[1072,1189,1745,1769]" country="Brazil" name="Amazonas" pageId="3" pageNumber="228">Amazonas</collectingRegion>
; and igarapé Mutum, a tributary of the rio Aruã, a tributary of rio Branco, itself tributary of the rio Arapiuns, tributary of the rio Tapajós basin (
<figureCitation box="[878,941,1840,1865]" captionStart="Fig" captionStartId="5.[129,168,992,1016]" captionTargetBox="[212,1362,174,969]" captionTargetId="figure-505@5.[212,1370,174,969]" captionTargetPageId="5" captionText="Fig. 5. Map of northern South America showing the distribution" figureDoi="http://doi.org/10.5281/zenodo.4647127" httpUri="https://zenodo.org/record/4647127/files/figure.png" pageId="3" pageNumber="228">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="229" pageId="3" pageNumber="228" type="etymology">
<paragraph blockId="3.[809,1452,1905,1961]" pageId="3" pageNumber="228">
<emphasis bold="true" box="[809,940,1905,1929]" pageId="3" pageNumber="228">Etymology.</emphasis>
The specific name
<taxonomicName box="[1145,1240,1905,1929]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="228" phylum="Chordata" rank="species" species="anhanga">
<emphasis box="[1145,1240,1905,1929]" italics="true" pageId="3" pageNumber="228">anhanga</emphasis>
</taxonomicName>
is a reference to the Amazonian Anhangá legend, a spirit that lives in the woods,
</paragraph>
<caption pageId="4" pageNumber="229">
<paragraph blockId="4.[816,1458,177,233]" pageId="4" pageNumber="229">
<emphasis bold="true" box="[816,908,177,201]" pageId="4" pageNumber="229">Table 1.</emphasis>
Morphometric data for holotype and paratypes of
<taxonomicName box="[816,1084,209,233]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="229" phylum="Chordata" rank="species" species="anhanga">
<emphasis box="[816,1084,209,233]" italics="true" pageId="4" pageNumber="229">Trichomycterus anhanga</emphasis>
</taxonomicName>
(n = 9 including holotype).
</paragraph>
</caption>
<paragraph pageId="4" pageNumber="229">
<table box="[818,1449,256,842]" gridcols="6" gridrows="22" pageId="4" pageNumber="229">
<tr box="[818,1449,256,275]" gridrow="0" pageId="4" pageNumber="229" rowspan-0="1">
<td box="[1064,1180,256,275]" gridcol="1" gridrow="0" pageId="4" pageNumber="229">Holotype</td>
<td box="[1205,1253,256,275]" gridcol="2" gridrow="0" pageId="4" pageNumber="229">Min.</td>
<td box="[1277,1325,256,275]" gridcol="3" gridrow="0" pageId="4" pageNumber="229">Max.</td>
<td box="[1350,1399,256,275]" gridcol="4" gridrow="0" pageId="4" pageNumber="229">Mean</td>
<td box="[1422,1449,256,275]" gridcol="5" gridrow="0" pageId="4" pageNumber="229">SD</td>
</tr>
<tr box="[818,1449,306,326]" gridrow="1" pageId="4" pageNumber="229">
<td box="[818,1041,306,326]" gridcol="0" gridrow="1" pageId="4" pageNumber="229">Standard length (SL, mm)</td>
<td box="[1064,1180,306,326]" gridcol="1" gridrow="1" pageId="4" pageNumber="229">10.0</td>
<td box="[1205,1253,306,326]" gridcol="2" gridrow="1" pageId="4" pageNumber="229">9.7</td>
<td box="[1277,1325,306,326]" gridcol="3" gridrow="1" pageId="4" pageNumber="229">13.1</td>
<td box="[1350,1399,306,326]" gridcol="4" gridrow="1" pageId="4" pageNumber="229">-</td>
<td box="[1422,1449,306,326]" gridcol="5" gridrow="1" pageId="4" pageNumber="229">-</td>
</tr>
<tr box="[818,1449,355,375]" gridrow="2" pageId="4" pageNumber="229" rowspan-0="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<td box="[1064,1253,355,375]" colspan="2" colspanRight="1" gridcol="1" gridrow="2" pageId="4" pageNumber="229">Percentage of SL</td>
</tr>
<tr box="[818,1449,380,399]" gridrow="3" pageId="4" pageNumber="229">
<td box="[818,1041,380,399]" gridcol="0" gridrow="3" pageId="4" pageNumber="229">Total length</td>
<td box="[1064,1180,380,399]" gridcol="1" gridrow="3" pageId="4" pageNumber="229">120.7</td>
<td box="[1205,1253,380,399]" gridcol="2" gridrow="3" pageId="4" pageNumber="229">113.9</td>
<td box="[1277,1325,380,399]" gridcol="3" gridrow="3" pageId="4" pageNumber="229">126.3</td>
<td box="[1350,1399,380,399]" gridcol="4" gridrow="3" pageId="4" pageNumber="229">119.3</td>
<td box="[1422,1449,380,399]" gridcol="5" gridrow="3" pageId="4" pageNumber="229">3.5</td>
</tr>
<tr box="[818,1449,405,424]" gridrow="4" pageId="4" pageNumber="229">
<td box="[818,1041,405,424]" gridcol="0" gridrow="4" pageId="4" pageNumber="229">Body depth</td>
<td box="[1064,1180,405,424]" gridcol="1" gridrow="4" pageId="4" pageNumber="229">16.9</td>
<td box="[1205,1253,405,424]" gridcol="2" gridrow="4" pageId="4" pageNumber="229">11.6</td>
<td box="[1277,1325,405,424]" gridcol="3" gridrow="4" pageId="4" pageNumber="229">16.9</td>
<td box="[1350,1399,405,424]" gridcol="4" gridrow="4" pageId="4" pageNumber="229">14.0</td>
<td box="[1422,1449,405,424]" gridcol="5" gridrow="4" pageId="4" pageNumber="229">1.7</td>
</tr>
<tr box="[818,1449,429,448]" gridrow="5" pageId="4" pageNumber="229">
<td box="[818,1041,429,448]" gridcol="0" gridrow="5" pageId="4" pageNumber="229">Peduncle length</td>
<td box="[1064,1180,429,448]" gridcol="1" gridrow="5" pageId="4" pageNumber="229">22.3</td>
<td box="[1205,1253,429,448]" gridcol="2" gridrow="5" pageId="4" pageNumber="229">21.6</td>
<td box="[1277,1325,429,448]" gridcol="3" gridrow="5" pageId="4" pageNumber="229">24.9</td>
<td box="[1350,1399,429,448]" gridcol="4" gridrow="5" pageId="4" pageNumber="229">23.5</td>
<td box="[1422,1449,429,448]" gridcol="5" gridrow="5" pageId="4" pageNumber="229">1.1</td>
</tr>
<tr box="[818,1449,453,473]" gridrow="6" pageId="4" pageNumber="229">
<td box="[818,1041,453,473]" gridcol="0" gridrow="6" pageId="4" pageNumber="229">Peduncle depth</td>
<td box="[1064,1180,453,473]" gridcol="1" gridrow="6" pageId="4" pageNumber="229">15.2</td>
<td box="[1205,1253,453,473]" gridcol="2" gridrow="6" pageId="4" pageNumber="229">10.7</td>
<td box="[1277,1325,453,473]" gridcol="3" gridrow="6" pageId="4" pageNumber="229">15.2</td>
<td box="[1350,1399,453,473]" gridcol="4" gridrow="6" pageId="4" pageNumber="229">12.5</td>
<td box="[1422,1449,453,473]" gridcol="5" gridrow="6" pageId="4" pageNumber="229">1.7</td>
</tr>
<tr box="[818,1449,478,497]" gridrow="7" pageId="4" pageNumber="229">
<td box="[818,1041,478,497]" gridcol="0" gridrow="7" pageId="4" pageNumber="229">Predorsal length</td>
<td box="[1064,1180,478,497]" gridcol="1" gridrow="7" pageId="4" pageNumber="229">66.2</td>
<td box="[1205,1253,478,497]" gridcol="2" gridrow="7" pageId="4" pageNumber="229">65.5</td>
<td box="[1277,1325,478,497]" gridcol="3" gridrow="7" pageId="4" pageNumber="229">69.4</td>
<td box="[1350,1399,478,497]" gridcol="4" gridrow="7" pageId="4" pageNumber="229">67.8</td>
<td box="[1422,1449,478,497]" gridcol="5" gridrow="7" pageId="4" pageNumber="229">1.5</td>
</tr>
<tr box="[818,1449,503,522]" gridrow="8" pageId="4" pageNumber="229">
<td box="[818,1041,503,522]" gridcol="0" gridrow="8" pageId="4" pageNumber="229">Preanal length</td>
<td box="[1064,1180,503,522]" gridcol="1" gridrow="8" pageId="4" pageNumber="229">68.2</td>
<td box="[1205,1253,503,522]" gridcol="2" gridrow="8" pageId="4" pageNumber="229">63.5</td>
<td box="[1277,1325,503,522]" gridcol="3" gridrow="8" pageId="4" pageNumber="229">71.4</td>
<td box="[1350,1399,503,522]" gridcol="4" gridrow="8" pageId="4" pageNumber="229">67.4</td>
<td box="[1422,1449,503,522]" gridcol="5" gridrow="8" pageId="4" pageNumber="229">2.3</td>
</tr>
<tr box="[818,1449,527,546]" gridrow="9" pageId="4" pageNumber="229">
<td box="[818,1041,527,546]" gridcol="0" gridrow="9" pageId="4" pageNumber="229">Dorsal-fin base length</td>
<td box="[1064,1180,527,546]" gridcol="1" gridrow="9" pageId="4" pageNumber="229">8.0</td>
<td box="[1205,1253,527,546]" gridcol="2" gridrow="9" pageId="4" pageNumber="229">7.5</td>
<td box="[1277,1325,527,546]" gridcol="3" gridrow="9" pageId="4" pageNumber="229">10.1</td>
<td box="[1350,1399,527,546]" gridcol="4" gridrow="9" pageId="4" pageNumber="229">8.6</td>
<td box="[1422,1449,527,546]" gridcol="5" gridrow="9" pageId="4" pageNumber="229">0.8</td>
</tr>
<tr box="[818,1449,552,571]" gridrow="10" pageId="4" pageNumber="229">
<td box="[818,1041,552,571]" gridcol="0" gridrow="10" pageId="4" pageNumber="229">Anal-fin base length</td>
<td box="[1064,1180,552,571]" gridcol="1" gridrow="10" pageId="4" pageNumber="229">7.8</td>
<td box="[1205,1253,552,571]" gridcol="2" gridrow="10" pageId="4" pageNumber="229">5.5</td>
<td box="[1277,1325,552,571]" gridcol="3" gridrow="10" pageId="4" pageNumber="229">9.2</td>
<td box="[1350,1399,552,571]" gridcol="4" gridrow="10" pageId="4" pageNumber="229">7.7</td>
<td box="[1422,1449,552,571]" gridcol="5" gridrow="10" pageId="4" pageNumber="229">1.0</td>
</tr>
<tr box="[818,1449,576,595]" gridrow="11" pageId="4" pageNumber="229">
<td box="[818,1041,576,595]" gridcol="0" gridrow="11" pageId="4" pageNumber="229">Head length (HL)</td>
<td box="[1064,1180,576,595]" gridcol="1" gridrow="11" pageId="4" pageNumber="229">18.4</td>
<td box="[1205,1253,576,595]" gridcol="2" gridrow="11" pageId="4" pageNumber="229">17.2</td>
<td box="[1277,1325,576,595]" gridcol="3" gridrow="11" pageId="4" pageNumber="229">20.2</td>
<td box="[1350,1399,576,595]" gridcol="4" gridrow="11" pageId="4" pageNumber="229">18.4</td>
<td box="[1422,1449,576,595]" gridcol="5" gridrow="11" pageId="4" pageNumber="229">1.0</td>
</tr>
<tr box="[818,1449,601,621]" gridrow="12" pageId="4" pageNumber="229" rowspan-0="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<td box="[1064,1253,601,621]" colspan="2" colspanRight="1" gridcol="1" gridrow="12" pageId="4" pageNumber="229">Percentage of HL</td>
</tr>
<tr box="[818,1449,626,645]" gridrow="13" pageId="4" pageNumber="229">
<td box="[818,1041,626,645]" gridcol="0" gridrow="13" pageId="4" pageNumber="229">Head width</td>
<td box="[1064,1180,626,645]" gridcol="1" gridrow="13" pageId="4" pageNumber="229">93.5</td>
<td box="[1205,1253,626,645]" gridcol="2" gridrow="13" pageId="4" pageNumber="229">80.6</td>
<td box="[1277,1325,626,645]" gridcol="3" gridrow="13" pageId="4" pageNumber="229">96.3</td>
<td box="[1350,1399,626,645]" gridcol="4" gridrow="13" pageId="4" pageNumber="229">89.1</td>
<td box="[1422,1449,626,645]" gridcol="5" gridrow="13" pageId="4" pageNumber="229">5.5</td>
</tr>
<tr box="[818,1449,650,669]" gridrow="14" pageId="4" pageNumber="229">
<td box="[818,1041,650,669]" gridcol="0" gridrow="14" pageId="4" pageNumber="229">Head depth</td>
<td box="[1064,1180,650,669]" gridcol="1" gridrow="14" pageId="4" pageNumber="229">33.0</td>
<td box="[1205,1253,650,669]" gridcol="2" gridrow="14" pageId="4" pageNumber="229">33.0</td>
<td box="[1277,1325,650,669]" gridcol="3" gridrow="14" pageId="4" pageNumber="229">50.3</td>
<td box="[1350,1399,650,669]" gridcol="4" gridrow="14" pageId="4" pageNumber="229">40.2</td>
<td box="[1422,1449,650,669]" gridcol="5" gridrow="14" pageId="4" pageNumber="229">5.3</td>
</tr>
<tr box="[818,1449,675,694]" gridrow="15" pageId="4" pageNumber="229">
<td box="[818,1041,675,694]" gridcol="0" gridrow="15" pageId="4" pageNumber="229">Interorbital</td>
<td box="[1064,1180,675,694]" gridcol="1" gridrow="15" pageId="4" pageNumber="229">19.5</td>
<td box="[1205,1253,675,694]" gridcol="2" gridrow="15" pageId="4" pageNumber="229">17.8</td>
<td box="[1277,1325,675,694]" gridcol="3" gridrow="15" pageId="4" pageNumber="229">23.2</td>
<td box="[1350,1399,675,694]" gridcol="4" gridrow="15" pageId="4" pageNumber="229">20.6</td>
<td box="[1422,1449,675,694]" gridcol="5" gridrow="15" pageId="4" pageNumber="229">1.7</td>
</tr>
<tr box="[818,1449,699,719]" gridrow="16" pageId="4" pageNumber="229">
<td box="[818,1041,699,719]" gridcol="0" gridrow="16" pageId="4" pageNumber="229">Snout length</td>
<td box="[1064,1180,699,719]" gridcol="1" gridrow="16" pageId="4" pageNumber="229">31.9</td>
<td box="[1205,1253,699,719]" gridcol="2" gridrow="16" pageId="4" pageNumber="229">23.0</td>
<td box="[1277,1325,699,719]" gridcol="3" gridrow="16" pageId="4" pageNumber="229">37.2</td>
<td box="[1350,1399,699,719]" gridcol="4" gridrow="16" pageId="4" pageNumber="229">31.4</td>
<td box="[1422,1449,699,719]" gridcol="5" gridrow="16" pageId="4" pageNumber="229">4.6</td>
</tr>
<tr box="[818,1449,724,743]" gridrow="17" pageId="4" pageNumber="229">
<td box="[818,1041,724,743]" gridcol="0" gridrow="17" pageId="4" pageNumber="229">Nasal babel length</td>
<td box="[1064,1180,724,743]" gridcol="1" gridrow="17" pageId="4" pageNumber="229">30.8</td>
<td box="[1205,1253,724,743]" gridcol="2" gridrow="17" pageId="4" pageNumber="229">26.5</td>
<td box="[1277,1325,724,743]" gridcol="3" gridrow="17" pageId="4" pageNumber="229">39.8</td>
<td box="[1350,1399,724,743]" gridcol="4" gridrow="17" pageId="4" pageNumber="229">32.1</td>
<td box="[1422,1449,724,743]" gridcol="5" gridrow="17" pageId="4" pageNumber="229">4.6</td>
</tr>
<tr box="[818,1449,748,768]" gridrow="18" pageId="4" pageNumber="229">
<td box="[818,1041,748,768]" gridcol="0" gridrow="18" pageId="4" pageNumber="229">Maxillary babel length</td>
<td box="[1064,1180,748,768]" gridcol="1" gridrow="18" pageId="4" pageNumber="229">41.6</td>
<td box="[1205,1253,748,768]" gridcol="2" gridrow="18" pageId="4" pageNumber="229">37.0</td>
<td box="[1277,1325,748,768]" gridcol="3" gridrow="18" pageId="4" pageNumber="229">55.0</td>
<td box="[1350,1399,748,768]" gridcol="4" gridrow="18" pageId="4" pageNumber="229">45.9</td>
<td box="[1422,1449,748,768]" gridcol="5" gridrow="18" pageId="4" pageNumber="229">6.0</td>
</tr>
<tr box="[818,1449,773,792]" gridrow="19" pageId="4" pageNumber="229">
<td box="[818,1041,773,792]" gridcol="0" gridrow="19" pageId="4" pageNumber="229">Rictal barbel length</td>
<td box="[1064,1180,773,792]" gridcol="1" gridrow="19" pageId="4" pageNumber="229">34.1</td>
<td box="[1205,1253,773,792]" gridcol="2" gridrow="19" pageId="4" pageNumber="229">32.2</td>
<td box="[1277,1325,773,792]" gridcol="3" gridrow="19" pageId="4" pageNumber="229">53.9</td>
<td box="[1350,1399,773,792]" gridcol="4" gridrow="19" pageId="4" pageNumber="229">39.8</td>
<td box="[1422,1449,773,792]" gridcol="5" gridrow="19" pageId="4" pageNumber="229">7.4</td>
</tr>
<tr box="[818,1449,797,817]" gridrow="20" pageId="4" pageNumber="229">
<td box="[818,1041,797,817]" gridcol="0" gridrow="20" pageId="4" pageNumber="229">Mouth width</td>
<td box="[1064,1180,797,817]" gridcol="1" gridrow="20" pageId="4" pageNumber="229">31.9</td>
<td box="[1205,1253,797,817]" gridcol="2" gridrow="20" pageId="4" pageNumber="229">22.5</td>
<td box="[1277,1325,797,817]" gridcol="3" gridrow="20" pageId="4" pageNumber="229">38.8</td>
<td box="[1350,1399,797,817]" gridcol="4" gridrow="20" pageId="4" pageNumber="229">28.6</td>
<td box="[1422,1449,797,817]" gridcol="5" gridrow="20" pageId="4" pageNumber="229">4.9</td>
</tr>
<tr box="[818,1449,822,842]" gridrow="21" pageId="4" pageNumber="229">
<td box="[818,1041,822,842]" gridcol="0" gridrow="21" pageId="4" pageNumber="229">Eye diameter</td>
<td box="[1064,1180,822,842]" gridcol="1" gridrow="21" pageId="4" pageNumber="229">15.1</td>
<td box="[1205,1253,822,842]" gridcol="2" gridrow="21" pageId="4" pageNumber="229">13.6</td>
<td box="[1277,1325,822,842]" gridcol="3" gridrow="21" pageId="4" pageNumber="229">19.0</td>
<td box="[1350,1399,822,842]" gridcol="4" gridrow="21" pageId="4" pageNumber="229">15.9</td>
<td box="[1422,1449,822,842]" gridcol="5" gridrow="21" pageId="4" pageNumber="229">1.7</td>
</tr>
</table>
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4647125" ID-Zenodo-Dep="4647125" httpUri="https://zenodo.org/record/4647125/files/figure.png" pageId="4" pageNumber="229" startId="4.[136,175,798,822]" targetBox="[151,771,189,759]" targetPageId="4">
<paragraph blockId="4.[136,779,798,887]" pageId="4" pageNumber="229">
<emphasis bold="true" box="[136,205,798,822]" pageId="4" pageNumber="229">Fig. 4.</emphasis>
Opercle and Interopercle of
<taxonomicName box="[506,772,798,822]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="229" phylum="Chordata" rank="species" species="anhanga">
<emphasis box="[506,772,798,822]" italics="true" pageId="4" pageNumber="229">Trichomycterus anhanga</emphasis>
</taxonomicName>
, lateral view, left side, INPA 36117, paratype. OP = opercle; IO = interopercle. Scale bar = 0.1 mm.
</paragraph>
</caption>
<paragraph blockId="4.[136,779,976,1096]" pageId="4" pageNumber="229">protector of animals. Its presence can be detected by a whistle and thereafter, the animal that was being hunted disappears. Anhangá is thus regarded as a protector of forest life. A noun in apposition.</paragraph>
</subSubSection>
</treatment>
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