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<mods:title id="52068905A3F88254F2AAED3ABBCB0151">“ Larger ” Benthic Foraminifera Of The Cenomanian. A Review Of The Identity And The Stratigraphic And Palaeogeographic Distribution Of Non-Fusiform Planispiral (Or Near-Planispiral) Forms</mods:title>
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<treatment id="03E587B6FFEEA228FCB6F8F6A187C4BA" ID-DOI="http://doi.org/10.5281/zenodo.10975543" ID-GBIF-Taxon="225698474" ID-Zenodo-Dep="10975543" LSID="urn:lsid:plazi:treatment:03E587B6FFEEA228FCB6F8F6A187C4BA" httpUri="http://treatment.plazi.org/id/03E587B6FFEEA228FCB6F8F6A187C4BA" lastPageId="59" lastPageNumber="98" pageId="55" pageNumber="94">
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<paragraph id="8BF336A0FFEEA224FCB6F8F6A1FAC012" blockId="55.[816,1436,1824,1881]" pageId="55" pageNumber="94">
<heading id="D0BB81CCFFEEA224FCB6F8F6A1FAC012" centered="true" fontSize="36" level="2" pageId="55" pageNumber="94" reason="4">
<emphasis id="B938EAB2FFEEA224FCB6F8F6A1FAC012" bold="true" pageId="55" pageNumber="94">
<taxonomicName id="4C4C4D23FFEEA224FCB6F8F6A1BAC073" ID-CoL="7ZWMK" authorityName="Calvez" authorityYear="1988" box="[816,1271,1824,1848]" class="Tubothalamea" family="Miliolidae" genus="Pseudonummoloculina" isUncertain="true" kingdom="Chromista" order="Miliolida" pageId="55" pageNumber="94" phylum="Foraminifera" rank="species" species="heimi">
<emphasis id="B938EAB2FFEEA224FCB6F8F6A170C073" bold="true" box="[816,1085,1824,1848]" italics="true" pageId="55" pageNumber="94">Pseudonummoloculina?</emphasis>
ex. grp.
<emphasis id="B938EAB2FFEEA224FB3FF8F6A1BAC073" bold="true" box="[1209,1271,1824,1848]" italics="true" pageId="55" pageNumber="94">heimi</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFDD4B51FFEEA224FA8BF8F6A0DAC073" author="Bonet, F." box="[1293,1431,1824,1848]" pageId="55" pageNumber="94" pagination="389 - 488" refId="ref74941" refString="Bonet, F., 1956. Zonification microfaunistica de las Calizas Cretacicas del este de Mexico. Boletin Asociation Mexicana de Geologos Petroleros: 389 - 488." type="book chapter" year="1956">Bonet 1956</bibRefCitation>
, emmend.
<bibRefCitation id="EFDD4B51FFEEA224FC26F897A1FDC012" author="Conkin, J. E. &amp; Conkin, B. M." box="[928,1200,1857,1881]" pageId="55" pageNumber="94" pagination="149 - 158" refId="ref77378" refString="Conkin, J. E. &amp; Conkin, B. M., 1958. Revision of the genus Nummoloculina and emendation of Nummoloculina heimi Bonet. Micropaleontology, 4 (2): 149 - 158." type="journal article" year="1958">Conkin &amp; Conkin, 1958</bibRefCitation>
)
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C356652BFFEEA224FCB6F855A62DC0B5" pageId="55" pageNumber="94" type="reference_group">
<paragraph id="8BF336A0FFEEA224FCB6F855A1FBC0D0" blockId="55.[816,1206,1923,1947]" box="[816,1206,1923,1947]" pageId="55" pageNumber="94">Reference Illustration &amp; Description</paragraph>
<paragraph id="8BF336A0FFEEA224FCB6F813A62DC0B5" blockId="55.[816,1437,1989,2046]" pageId="55" pageNumber="94">
<bibRefCitation id="EFDD4B51FFEEA224FCB6F813A101C096" author="Conkin, J. E. &amp; Conkin, B. M." box="[816,1100,1989,2013]" pageId="55" pageNumber="94" pagination="149 - 158" refId="ref77378" refString="Conkin, J. E. &amp; Conkin, B. M., 1958. Revision of the genus Nummoloculina and emendation of Nummoloculina heimi Bonet. Micropaleontology, 4 (2): 149 - 158." type="journal article" year="1958">Conkin &amp; Conkin (1958)</bibRefCitation>
, pl. 1, text figs. 1-25, p. 152- 156.
</paragraph>
</subSubSection>
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<paragraph id="8BF336A0FFE1A22BFEF1FCBAA1F1C4CA" blockId="56.[375,1212,875,897]" box="[375,1212,875,897]" pageId="56" pageNumber="95">
<emphasis id="B938EAB2FFE1A22BFEF1FCBAA4F0C4CA" bold="true" box="[375,445,876,897]" pageId="56" pageNumber="95">Fig. 42</emphasis>
Cenomanian paleogeographic distribution of
<taxonomicName id="4C4C4D23FFE1A22BFCF0FCBDA1F1C4CA" authorityName="Calvez" authorityYear="1988" box="[886,1212,875,897]" class="Tubothalamea" family="Miliolidae" genus="Pseudonummoloculina" kingdom="Chromista" order="Miliolida" pageId="56" pageNumber="95" phylum="Foraminifera" rank="species" species="aurigerica">Pseudonummoloculina aurigerica</taxonomicName>
</paragraph>
</caption>
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<paragraph id="8BF336A0FFE1A22BFF1CF8B9A118C08B" blockId="56.[154,1417,1903,1984]" pageId="56" pageNumber="95">
<emphasis id="B938EAB2FFE1A22BFF1CF8B9A5ADC0CF" bold="true" box="[154,224,1903,1924]" pageId="56" pageNumber="95">Fig. 43</emphasis>
Representative illustrations of
<taxonomicName id="4C4C4D23FFE1A22BFD8DF8B9A7AAC0CE" authorityName="Calvez" authorityYear="1988" box="[523,743,1903,1925]" class="Tubothalamea" family="Miliolidae" genus="Pseudonummoloculina" kingdom="Chromista" order="Miliolida" pageId="56" pageNumber="95" phylum="Foraminifera" rank="genus">
<emphasis id="B938EAB2FFE1A22BFD8DF8B9A7AAC0CE" box="[523,743,1903,1925]" italics="true" pageId="56" pageNumber="95">Pseudonummoloculina</emphasis>
</taxonomicName>
ex gr.
<taxonomicName id="4C4C4D23FFE1A22BFCACF8B9A612C0CE" authority="(Bonet)" authorityName="Bonet" authorityYear="1956" baseAuthorityName="Bonet" baseAuthorityYear="1956" box="[810,863,1903,1925]" class="Tubothalamea" family="Hauerinidae" genus="Nummoloculina" kingdom="Chromista" order="Miliolida" pageId="56" pageNumber="95" phylum="Foraminifera" rank="species" species="heimi">
<emphasis id="B938EAB2FFE1A22BFCACF8B9A612C0CE" box="[810,863,1903,1925]" italics="true" pageId="56" pageNumber="95">heimi</emphasis>
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:
<emphasis id="B938EAB2FFE1A22BFCEAF8B9A635C0CF" bold="true" box="[876,888,1903,1924]" pageId="56" pageNumber="95">a</emphasis>
Equatorial section,
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, pl. 1, fig. 1, Mexico);
<emphasis id="B938EAB2FFE1A22BFE96F85BA450C0E9" bold="true" box="[272,285,1933,1954]" pageId="56" pageNumber="95">b</emphasis>
Axial section (schematic),
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, text fig. 13, Texas);
<emphasis id="B938EAB2FFE1A22BFC58F85BA6A4C0E9" bold="true" box="[990,1001,1933,1954]" pageId="56" pageNumber="95">c</emphasis>
Axial section,
<bibRefCitation id="EFDD4B51FFE1A22BFBFCF85AA028C0E9" author="Conkin, J. E. &amp; Conkin, B. M." box="[1146,1381,1932,1954]" pageId="56" pageNumber="95" pagination="149 - 158" refId="ref77378" refString="Conkin, J. E. &amp; Conkin, B. M., 1958. Revision of the genus Nummoloculina and emendation of Nummoloculina heimi Bonet. Micropaleontology, 4 (2): 149 - 158." type="journal article" year="1958">Conkin &amp; Conkin (1958</bibRefCitation>
, pl. 1, fig. 6, Mexico);
<emphasis id="B938EAB2FFE1A22BFEC9F87DA411C08B" bold="true" box="[335,348,1963,1984]" pageId="56" pageNumber="95">d</emphasis>
Equatorial section (schematic),
<bibRefCitation id="EFDD4B51FFE1A22BFD14F87CA633C08B" author="Conkin, J. E. &amp; Conkin, B. M." box="[658,894,1962,1984]" pageId="56" pageNumber="95" pagination="149 - 158" refId="ref77378" refString="Conkin, J. E. &amp; Conkin, B. M., 1958. Revision of the genus Nummoloculina and emendation of Nummoloculina heimi Bonet. Micropaleontology, 4 (2): 149 - 158." type="journal article" year="1958">Conkin &amp; Conkin (1958</bibRefCitation>
, pl. 1, fig. 3, Mexico).
</paragraph>
</caption>
<subSubSection id="C356652BFFE0A229FF11FF48A4CBC637" lastPageId="58" lastPageNumber="97" pageId="57" pageNumber="96" type="description">
<paragraph id="8BF336A0FFE0A22AFF11FF48A7CBC54A" blockId="57.[151,772,158,2068]" pageId="57" pageNumber="96">
In addition to the correct generic classification of nummoloculinids in general, a separate issue relates to the status of the species
<taxonomicName id="4C4C4D23FFE0A22AFE0DFF36A48BC7B3" authority="(Bonet)" authorityName="Bonet" authorityYear="1956" baseAuthorityName="Bonet" baseAuthorityYear="1956" box="[395,454,224,248]" class="Tubothalamea" family="Hauerinidae" genus="Nummoloculina" kingdom="Chromista" order="Miliolida" pageId="57" pageNumber="96" phylum="Foraminifera" rank="species" species="heimi">
<emphasis id="B938EAB2FFE0A22AFE0DFF36A48BC7B3" box="[395,454,224,248]" italics="true" pageId="57" pageNumber="96">heimi</emphasis>
</taxonomicName>
.
<taxonomicName id="4C4C4D23FFE0A22AFE5CFF36A781C7B3" box="[474,716,224,248]" class="Tubothalamea" family="Hauerinidae" genus="Nummoloculina" kingdom="Chromista" order="Miliolida" pageId="57" pageNumber="96" phylum="Foraminifera" rank="species" species="heimi">
<emphasis id="B938EAB2FFE0A22AFE5CFF36A781C7B3" box="[474,716,224,248]" italics="true" pageId="57" pageNumber="96">Nummoloculina heimi</emphasis>
</taxonomicName>
was first described from the Albian-Cenomanian of
<collectingRegion id="4988F842FFE0A22AFD34FED7A5FFC671" country="Mexico" name="Distrito Federal" pageId="57" pageNumber="96">Mexico by</collectingRegion>
<bibRefCitation id="EFDD4B51FFE0A22AFF38FEF5A418C671" author="Bonet, F." box="[190,341,290,314]" pageId="57" pageNumber="96" pagination="389 - 488" refId="ref74941" refString="Bonet, F., 1956. Zonification microfaunistica de las Calizas Cretacicas del este de Mexico. Boletin Asociation Mexicana de Geologos Petroleros: 389 - 488." type="book chapter" year="1956">Bonet (1956)</bibRefCitation>
. Bonets descriptions and illustrations were according to
<bibRefCitation id="EFDD4B51FFE0A22AFEFDFE95A7DCC610" author="Conkin, J. E. &amp; Conkin, B. M." box="[379,657,323,347]" pageId="57" pageNumber="96" pagination="149 - 158" refId="ref77378" refString="Conkin, J. E. &amp; Conkin, B. M., 1958. Revision of the genus Nummoloculina and emendation of Nummoloculina heimi Bonet. Micropaleontology, 4 (2): 149 - 158." type="journal article" year="1958">Conkin &amp; Conkin (1958)</bibRefCitation>
unsatisfactory, and they emended the definition based on their own material which was also from
<collectingCountry id="F35B7630FFE0A22AFD9DFE53A720C6D6" box="[539,621,389,413]" name="Mexico" pageId="57" pageNumber="96">Mexico</collectingCountry>
and from the southern
<collectingCountry id="F35B7630FFE0A22AFF7DFE71A75DC6F4" box="[251,528,423,447]" name="United States of America" pageId="57" pageNumber="96">United States of America</collectingCountry>
. Some of this material came from the same lithostratigraphic unit in
<collectingCountry id="F35B7630FFE0A22AFD17FE1EA7AEC6AB" box="[657,739,456,480]" name="Mexico" pageId="57" pageNumber="96">Mexico</collectingCountry>
as Bonets, but not from the same (
<typeStatus id="54F78802FFE0A22AFE69FE3FA76CC54A" box="[495,545,489,513]" pageId="57" pageNumber="96">type</typeStatus>
) locality.
</paragraph>
<paragraph id="8BF336A0FFE0A22AFF11FDDCA4F4C5CE" blockId="57.[151,772,158,2068]" pageId="57" pageNumber="96">
<bibRefCitation id="EFDD4B51FFE0A22AFF11FDDCA4F9C569" author="Conkin, J. E. &amp; Conkin, B. M." box="[151,436,522,546]" pageId="57" pageNumber="96" pagination="149 - 158" refId="ref77378" refString="Conkin, J. E. &amp; Conkin, B. M., 1958. Revision of the genus Nummoloculina and emendation of Nummoloculina heimi Bonet. Micropaleontology, 4 (2): 149 - 158." type="journal article" year="1958">Conkin &amp; Conkin (1958)</bibRefCitation>
recognised two macrospheric morphotypes, essentially: (i) with a quinqueloculine nepionic stage followed by a planispiral coil and (ii) planispirally coiled throughout.
</paragraph>
<paragraph id="8BF336A0FFE0A22AFF11FD58A7D7C426" blockId="57.[151,772,158,2068]" pageId="57" pageNumber="96">
Subsequent studies by
<bibRefCitation id="EFDD4B51FFE0A22AFE0BFD58A732C5ED" author="Hottinger, L. &amp; Drobne, K. &amp; Caus, E." box="[397,639,654,678]" pageId="57" pageNumber="96" pagination="99 - 134" refId="ref84439" refString="Hottinger, L., Drobne, K. &amp; Caus, E., 1989. Late Cretaceous, larger complex Miliolids (foraminifera) endemic in the Pyrenean faunal province. Facies, 21: 99 - 134." type="journal article" year="1989">Hottinger et al. (1989)</bibRefCitation>
from
<collectingCountry id="F35B7630FFE0A22AFD44FD58A5FDC58C" name="Mexico" pageId="57" pageNumber="96">Mexico</collectingCountry>
resulted in them attributing forms to
<taxonomicName id="4C4C4D23FFE0A22AFDC4FD79A730C58C" authority="(Bonet)" authorityName="Bonet" authorityYear="1956" baseAuthorityName="Bonet" baseAuthorityYear="1956" box="[578,637,687,711]" class="Tubothalamea" family="Hauerinidae" genus="Nummoloculina" kingdom="Chromista" order="Miliolida" pageId="57" pageNumber="96" phylum="Foraminifera" rank="species" species="heimi">
<emphasis id="B938EAB2FFE0A22AFDC4FD79A730C58C" box="[578,637,687,711]" italics="true" pageId="57" pageNumber="96">heimi</emphasis>
</taxonomicName>
(“as revised by
<bibRefCitation id="EFDD4B51FFE0A22AFF3BFD07A482C5A2" author="Conkin, J. E. &amp; Conkin, B. M." box="[189,463,721,745]" pageId="57" pageNumber="96" pagination="149 - 158" refId="ref77378" refString="Conkin, J. E. &amp; Conkin, B. M., 1958. Revision of the genus Nummoloculina and emendation of Nummoloculina heimi Bonet. Micropaleontology, 4 (2): 149 - 158." type="journal article" year="1958">Conkin &amp; Conkin, 1958</bibRefCitation>
”) characterised by an early
<emphasis id="B938EAB2FFE0A22AFF11FD24A452C441" box="[151,287,754,778]" italics="true" pageId="57" pageNumber="96">streptospiral</emphasis>
(not quinqueloculine) stage followed by whorls which gradually stabilised their coiling axis without becoming completely stable; part of their so-called “stem miliolids” (
<bibRefCitation id="EFDD4B51FFE0A22AFED2FC83A771C426" author="Hottinger, L. &amp; Drobne, K. &amp; Caus, E." box="[340,572,853,877]" pageId="57" pageNumber="96" pagination="99 - 134" refId="ref84439" refString="Hottinger, L., Drobne, K. &amp; Caus, E., 1989. Late Cretaceous, larger complex Miliolids (foraminifera) endemic in the Pyrenean faunal province. Facies, 21: 99 - 134." type="journal article" year="1989">Hottinger et al., 1989</bibRefCitation>
; p. 103).
</paragraph>
<paragraph id="8BF336A0FFE0A22AFF11FCA0A45CC005" blockId="57.[151,772,158,2068]" pageId="57" pageNumber="96">
A separate but related issue also arises over the nature of the aperture. The presence of folds or notches/crenulations in the aperture is fundamental (among other things) for a generic assignment to
<taxonomicName id="4C4C4D23FFE0A22AFDDBFC0FA5A3C359" authorityName="Calvez" authorityYear="1988" class="Tubothalamea" family="Miliolidae" genus="Pseudonummoloculina" kingdom="Chromista" order="Miliolida" pageId="57" pageNumber="96" phylum="Foraminifera" rank="genus">
<emphasis id="B938EAB2FFE0A22AFDDBFC0FA5A3C359" italics="true" pageId="57" pageNumber="96">Pseudonummoloculina</emphasis>
</taxonomicName>
. Neither
<bibRefCitation id="EFDD4B51FFE0A22AFEE7FC2DA4BBC359" author="Bonet, F." box="[353,502,1018,1042]" pageId="57" pageNumber="96" pagination="389 - 488" refId="ref74941" refString="Bonet, F., 1956. Zonification microfaunistica de las Calizas Cretacicas del este de Mexico. Boletin Asociation Mexicana de Geologos Petroleros: 389 - 488." type="book chapter" year="1956">Bonet (1956)</bibRefCitation>
nor
<bibRefCitation id="EFDD4B51FFE0A22AFDB3FC2CA593C378" author="Conkin, J. E. &amp; Conkin, B. M." pageId="57" pageNumber="96" pagination="149 - 158" refId="ref77378" refString="Conkin, J. E. &amp; Conkin, B. M., 1958. Revision of the genus Nummoloculina and emendation of Nummoloculina heimi Bonet. Micropaleontology, 4 (2): 149 - 158." type="journal article" year="1958">Conkin &amp; Conkin (1958)</bibRefCitation>
observed such features in their North American material.
<bibRefCitation id="EFDD4B51FFE0A22AFF79FBEAA4BAC31F" author="Hottinger, L. &amp; Drobne, K. &amp; Caus, E." box="[255,503,1084,1108]" pageId="57" pageNumber="96" pagination="99 - 134" refId="ref84439" refString="Hottinger, L., Drobne, K. &amp; Caus, E., 1989. Late Cretaceous, larger complex Miliolids (foraminifera) endemic in the Pyrenean faunal province. Facies, 21: 99 - 134." type="journal article" year="1989">Hottinger et al. (1989)</bibRefCitation>
only observed a row of notches in one illustrated specimen [see
<figureCitation id="13772A25FFE0A22AFDE5FB8BA7F9C33E" box="[611,692,1117,1141]" captionStart="Fig" captionStartId="57.[816,849,1685,1706]" captionTargetBox="[830,1436,1187,1647]" captionTargetId="figure-1201@57.[816,1436,1187,1647]" captionTargetPageId="57" captionText="Fig. 44 Pseudonummoloculina heimi (Bonet) left after De Castro (1987: fig. 3-2) and right after Hottinger et al. (1989: pl. 22, fig. 6-part). Both specimens purported to show notched apertures. Neither specimen can, however, be attributed to heimi based on the criteria established by Piuz &amp; Vicedo (2020, see text). Those authors place both specimens in tentative synonymy with their new species Nummoloculinodonta akhdarensis." figureDoi="http://doi.org/10.5281/zenodo.10834277" httpUri="https://zenodo.org/record/10834277/files/figure.png" pageId="57" pageNumber="96">Fig. 44</figureCitation>
(right) herein] of a near-completely planispiral form but not observed at all in specimens with initial streptospiral coiling.
<bibRefCitation id="EFDD4B51FFE0A22AFF41FB17A48AC392" author="Piuz, A. &amp; Vicedo, V." box="[199,455,1217,1241]" pageId="57" pageNumber="96" refId="ref93491" refString="Piuz, A. &amp; Vicedo, V., 2020. New Cenomanian &quot; nummoloculinas &quot; of the Natih Formation of Oman. Cretaceous Research, 107, https: // doi. org / 10.1016 / j. cretres. 2019.104224" type="book" year="2020">Piuz and Vicedo (2020)</bibRefCitation>
tentatively suggested assigning this specimen to their new nummoloculinid species
<taxonomicName id="4C4C4D23FFE0A22AFF11FAD5A75FC250" authorityName="Piuz &amp; Vicedo" authorityYear="2020" box="[151,530,1283,1307]" class="Tubothalamea" family="Hauerinidae" genus="Nummoloculinodonta" higherTaxonomySource="CoL,GBIF" kingdom="Chromista" order="Miliolida" pageId="57" pageNumber="96" phylum="Foraminifera" rank="species" species="akhdarensis">
<emphasis id="B938EAB2FFE0A22AFF11FAD5A75FC250" box="[151,530,1283,1307]" italics="true" pageId="57" pageNumber="96">Nummoloculinodonta akhdarensis</emphasis>
</taxonomicName>
thus removing any suggestion of notched/crenulated apertures as a characteristic of
<taxonomicName id="4C4C4D23FFE0A22AFF6FFA93A469C216" authority="(Bonet)" authorityName="Bonet" authorityYear="1956" baseAuthorityName="Bonet" baseAuthorityYear="1956" box="[233,292,1349,1373]" class="Tubothalamea" family="Hauerinidae" genus="Nummoloculina" kingdom="Chromista" order="Miliolida" pageId="57" pageNumber="96" phylum="Foraminifera" rank="species" species="heimi">
<emphasis id="B938EAB2FFE0A22AFF6FFA93A469C216" box="[233,292,1349,1373]" italics="true" pageId="57" pageNumber="96">heimi</emphasis>
</taxonomicName>
forms. However,
<bibRefCitation id="EFDD4B51FFE0A22AFE6EFA93A7FDC216" author="Solak, C. &amp; Tasli, K. &amp; Koc, H." box="[488,688,1349,1373]" pageId="57" pageNumber="96" pagination="673 - 693" refId="ref100030" refString="Solak, C., Tasli, K. &amp; Koc, H., 2021. Benthic foraminifera from the Albian shallow-marine limestones in the Geyik Dagi area (Central Taurides), southern Turkey. Journal of Paleontology, 95 (4): 673 - 693." type="journal article" year="2021">Solak et al. (2021)</bibRefCitation>
contradict this by stating “…
<emphasis id="B938EAB2FFE0A22AFE0DFAB0A64EC235" box="[395,771,1382,1406]" italics="true" pageId="57" pageNumber="96">the widespread Cretaceous species</emphasis>
<taxonomicName id="4C4C4D23FFE0A22AFF11FA51A75AC2D4" authority="Bonet, 1956" authorityName="Bonet" authorityYear="1956" box="[151,535,1415,1439]" class="Tubothalamea" family="Hauerinidae" genus="Nummoloculina" kingdom="Chromista" order="Miliolida" pageId="57" pageNumber="96" phylum="Foraminifera" rank="species" species="heimi">
Nummoloculina heimi
<bibRefCitation id="EFDD4B51FFE0A22AFE14FA5EA75AC2D4" author="Bonet, F." box="[402,535,1415,1439]" pageId="57" pageNumber="96" pagination="389 - 488" refId="ref74941" refString="Bonet, F., 1956. Zonification microfaunistica de las Calizas Cretacicas del este de Mexico. Boletin Asociation Mexicana de Geologos Petroleros: 389 - 488." type="book chapter" year="1956">
<emphasis id="B938EAB2FFE0A22AFE14FA5EA75AC2D4" box="[402,535,1415,1439]" italics="true" pageId="57" pageNumber="96">Bonet, 1956</emphasis>
</bibRefCitation>
</taxonomicName>
<emphasis id="B938EAB2FFE0A22AFD91FA51A466C2A9" italics="true" pageId="57" pageNumber="96">
, with a clear notched aperture (
<bibRefCitation id="EFDD4B51FFE0A22AFE80FA7FA48DC28A" author="De Castro, P." box="[262,448,1449,1473]" pageId="57" pageNumber="96" pagination="109 - 124" refId="ref78786" refString="De Castro, P., 1987. On some foraminifera and algae in Apennine Upper Cretaceous and Paleocene: Memorie della Societa Geologica Italiana, 40: 109 - 124." type="journal article" year="1987">De Castro, 1987</bibRefCitation>
;
<bibRefCitation id="EFDD4B51FFE0A22AFE55FA7CA78AC28A" author="Hottinger, L. &amp; Drobne, K. &amp; Caus, E." box="[467,711,1449,1473]" pageId="57" pageNumber="96" pagination="99 - 134" refId="ref84439" refString="Hottinger, L., Drobne, K. &amp; Caus, E., 1989. Late Cretaceous, larger complex Miliolids (foraminifera) endemic in the Pyrenean faunal province. Facies, 21: 99 - 134." type="journal article" year="1989">Hottinger et al., 1989</bibRefCitation>
) was transferred to
</emphasis>
<taxonomicName id="4C4C4D23FFE0A22AFEB4FA1CA760C2A9" authorityName="Calvez" authorityYear="1988" box="[306,557,1482,1506]" class="Tubothalamea" family="Miliolidae" genus="Pseudonummoloculina" kingdom="Chromista" order="Miliolida" pageId="57" pageNumber="96" phylum="Foraminifera" rank="genus">Pseudonummoloculina</taxonomicName>
”. It should be noted that
<bibRefCitation id="EFDD4B51FFE0A22AFF4CFA3DA4D0C148" author="Solak, C. &amp; Tasli, K. &amp; Koc, H." box="[202,413,1515,1539]" pageId="57" pageNumber="96" pagination="673 - 693" refId="ref100030" refString="Solak, C., Tasli, K. &amp; Koc, H., 2021. Benthic foraminifera from the Albian shallow-marine limestones in the Geyik Dagi area (Central Taurides), southern Turkey. Journal of Paleontology, 95 (4): 673 - 693." type="journal article" year="2021">Solak et al. (2021)</bibRefCitation>
did not themselves observe the notched aperture in their own material either. De Castros material from the Cenomanian of Apennine
<collectingCountry id="F35B7630FFE0A22AFD03F9FBA7FAC10E" box="[645,695,1581,1605]" name="Italy" pageId="57" pageNumber="96">Italy</collectingCountry>
shows
<specimenCount id="9D4AFD29FFE0A22AFF11F999A461C12D" box="[151,300,1614,1638]" count="1" pageId="57" pageNumber="96" type="generic">one specimen</specimenCount>
(out of 4 examples) which shows an aperture with “wavy margins”.
<bibRefCitation id="EFDD4B51FFE0A22AFE3DF9B9A7FDC1CC" author="Hottinger, L. &amp; Drobne, K. &amp; Caus, E." box="[443,688,1647,1671]" pageId="57" pageNumber="96" pagination="99 - 134" refId="ref84439" refString="Hottinger, L., Drobne, K. &amp; Caus, E., 1989. Late Cretaceous, larger complex Miliolids (foraminifera) endemic in the Pyrenean faunal province. Facies, 21: 99 - 134." type="journal article" year="1989">Hottinger et al. (1989)</bibRefCitation>
s material from the Cenomanian of Mexico shows a single specimen (out of several tens of examples) with “notches in distal apertural margin”. This particular specimen (a subaxial section, bottom right corner of pl. 22, fig. 6 see
<figureCitation id="13772A25FFE0A22AFF11F8C3A45BC066" box="[151,278,1813,1837]" captionStart="Fig" captionStartId="57.[816,849,1685,1706]" captionTargetBox="[830,1436,1187,1647]" captionTargetId="figure-1201@57.[816,1436,1187,1647]" captionTargetPageId="57" captionText="Fig. 44 Pseudonummoloculina heimi (Bonet) left after De Castro (1987: fig. 3-2) and right after Hottinger et al. (1989: pl. 22, fig. 6-part). Both specimens purported to show notched apertures. Neither specimen can, however, be attributed to heimi based on the criteria established by Piuz &amp; Vicedo (2020, see text). Those authors place both specimens in tentative synonymy with their new species Nummoloculinodonta akhdarensis." figureDoi="http://doi.org/10.5281/zenodo.10834277" httpUri="https://zenodo.org/record/10834277/files/figure.png" pageId="57" pageNumber="96">Figure 44 b</figureCitation>
) is very similar to
<figureCitation id="13772A25FFE0A22AFE6AF8C3A70BC066" box="[492,582,1813,1837]" captionStart="Fig" captionStartId="63.[196,229,1868,1889]" captionTargetBox="[209,1452,980,1848]" captionTargetId="figure-72@63.[196,1463,967,1860]" captionTargetPageId="63" captionText="Fig. 48 Representative illustrations of Pseudonummoloculina? cf. irregularis: a Subequatorial section, Chiocchini et al. (2012, pl. 1, fig. 3, Italy); b Subaxial section, Chiocchini et al. (2012, pl. 120,fig. 2, Italy); c Subequatorial section, Chiocchini et al. (2012, pl. 120, fig. 4, Italy); d Pseudonummoloculina sp. sensu Solak et al. (2020, fig. 14V, Turkey)." figureDoi="http://doi.org/10.5281/zenodo.10834285" httpUri="https://zenodo.org/record/10834285/files/figure.png" pageId="57" pageNumber="96">Fig. 48b</figureCitation>
herein for
<emphasis id="B938EAB2FFE0A22AFD45F8C0A79EC066" box="[707,723,1814,1837]" italics="true" pageId="57" pageNumber="96">P</emphasis>
?
<taxonomicName id="4C4C4D23FFE0A22AFD6EF8C3A447C005" authorityName="sensu Chiocchini et al." authorityYear="2012" baseAuthorityName="Decrouez &amp; Radoicic" baseAuthorityYear="1977" class="Tubothalamea" family="Hauerinidae" genus="Nummoloculina" isUncertain="true" kingdom="Chromista" order="Miliolida" pageId="57" pageNumber="96" phylum="Foraminifera" rank="species" species="irregularis">
cf.
<emphasis id="B938EAB2FFE0A22AFF11F8E0A447C005" box="[151,266,1846,1870]" italics="true" pageId="57" pageNumber="96">irregularis</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="DF336628FFE0A22AFCB6F943A0CCC016" ID-DOI="http://doi.org/10.5281/zenodo.10834277" ID-Zenodo-Dep="10834277" httpUri="https://zenodo.org/record/10834277/files/figure.png" pageId="57" pageNumber="96" startId="57.[816,849,1685,1706]" targetBox="[830,1436,1187,1647]" targetPageId="57" targetType="figure">
<paragraph id="8BF336A0FFE0A22AFCB6F943A0CCC016" blockId="57.[816,1437,1685,1886]" pageId="57" pageNumber="96">
<emphasis id="B938EAB2FFE0A22AFCB6F943A631C1E1" bold="true" box="[816,892,1685,1706]" pageId="57" pageNumber="96">Fig. 44</emphasis>
<taxonomicName id="4C4C4D23FFE0A22AFC0EF943A1B7C1E0" authority="(Bonet, 1956)" box="[904,1274,1685,1707]" class="Tubothalamea" family="Miliolidae" genus="Pseudonummoloculina" kingdom="Chromista" order="Miliolida" pageId="57" pageNumber="96" phylum="Foraminifera" rank="species" species="heimi">
<emphasis id="B938EAB2FFE0A22AFC0EF943A1E9C1E0" box="[904,1188,1685,1707]" italics="true" pageId="57" pageNumber="96">Pseudonummoloculina heimi</emphasis>
(Bonet)
</taxonomicName>
left after
<bibRefCitation id="EFDD4B51FFE0A22AFA07F943A6FDC183" author="De Castro, P." pageId="57" pageNumber="96" pagination="109 - 124" refId="ref78786" refString="De Castro, P., 1987. On some foraminifera and algae in Apennine Upper Cretaceous and Paleocene: Memorie della Societa Geologica Italiana, 40: 109 - 124." type="journal article" year="1987">De Castro (1987</bibRefCitation>
: fig. 3-2) and right after
<bibRefCitation id="EFDD4B51FFE0A22AFB2EF964A036C183" author="Hottinger, L. &amp; Drobne, K. &amp; Caus, E." box="[1192,1403,1714,1736]" pageId="57" pageNumber="96" pagination="99 - 134" refId="ref84439" refString="Hottinger, L., Drobne, K. &amp; Caus, E., 1989. Late Cretaceous, larger complex Miliolids (foraminifera) endemic in the Pyrenean faunal province. Facies, 21: 99 - 134." type="journal article" year="1989">Hottinger et al. (1989</bibRefCitation>
: pl. 22, fig. 6-part). Both specimens purported to show notched apertures. Neither specimen can, however, be attributed to
<taxonomicName id="4C4C4D23FFE0A22AFAE1F938A0D1C04F" authority="(Bonet)" authorityName="Bonet" authorityYear="1956" baseAuthorityName="Bonet" baseAuthorityYear="1956" box="[1383,1436,1774,1796]" class="Tubothalamea" family="Miliolidae" genus="Pseudonummoloculina" kingdom="Chromista" order="Miliolida" pageId="57" pageNumber="96" phylum="Foraminifera" rank="species" species="heimi">
<emphasis id="B938EAB2FFE0A22AFAE1F938A0D1C04F" box="[1383,1436,1774,1796]" italics="true" pageId="57" pageNumber="96">heimi</emphasis>
</taxonomicName>
based on the criteria established by
<bibRefCitation id="EFDD4B51FFE0A22AFB1FF8DAA03CC069" author="Piuz, A. &amp; Vicedo, V." box="[1177,1393,1804,1826]" pageId="57" pageNumber="96" refId="ref93491" refString="Piuz, A. &amp; Vicedo, V., 2020. New Cenomanian &quot; nummoloculinas &quot; of the Natih Formation of Oman. Cretaceous Research, 107, https: // doi. org / 10.1016 / j. cretres. 2019.104224" type="book" year="2020">Piuz &amp; Vicedo (2020</bibRefCitation>
, see text). Those authors place both specimens in tentative synonymy with their new species
<taxonomicName id="4C4C4D23FFE0A22AFBB7F89EA036C015" authorityName="Piuz &amp; Vicedo" authorityYear="2020" box="[1073,1403,1864,1886]" class="Tubothalamea" family="Hauerinidae" genus="Nummoloculinodonta" higherTaxonomySource="CoL,GBIF" kingdom="Chromista" order="Miliolida" pageId="57" pageNumber="96" phylum="Foraminifera" rank="species" species="akhdarensis">
<emphasis id="B938EAB2FFE0A22AFBB7F89EA036C015" box="[1073,1403,1864,1886]" italics="true" pageId="57" pageNumber="96">Nummoloculinodonta akhdarensis</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BF336A0FFE0A22AFF11F881A076C652" blockId="57.[151,772,158,2068]" lastBlockId="57.[816,1436,158,480]" pageId="57" pageNumber="96">
<bibRefCitation id="EFDD4B51FFE0A22AFF11F881A4CBC024" author="Piuz, A. &amp; Vicedo, V." box="[151,390,1879,1903]" pageId="57" pageNumber="96" refId="ref93491" refString="Piuz, A. &amp; Vicedo, V., 2020. New Cenomanian &quot; nummoloculinas &quot; of the Natih Formation of Oman. Cretaceous Research, 107, https: // doi. org / 10.1016 / j. cretres. 2019.104224" type="book" year="2020">Piuz &amp; Vicedo (2020)</bibRefCitation>
discussed the implications of these observations (and others) and concluded that “…the different morphotypes mentioned above as
<taxonomicName id="4C4C4D23FFE0A22AFDE0F84CA789C0FA" baseAuthorityName="Bonet" baseAuthorityYear="1956" box="[614,708,1945,1969]" class="Tubothalamea" family="Miliolidae" genus="Pseudonummoloculina" kingdom="Chromista" order="Miliolida" pageId="57" pageNumber="96" phylum="Foraminifera" rank="species" species="heimi">
<emphasis id="B938EAB2FFE0A22AFDE0F84CA789C0FA" box="[614,708,1945,1969]" italics="true" pageId="57" pageNumber="96">P. heimi</emphasis>
</taxonomicName>
[are] likely separate taxa”. However, both they and we agree that many forms displaying this variety of morphological characteristics have been attributed to
<taxonomicName id="4C4C4D23FFE0A22AFDBCF82AA738CF5F" authority="(Bonet)" authorityName="Bonet" authorityYear="1956" baseAuthorityName="Bonet" baseAuthorityYear="1956" box="[570,629,2044,2068]" class="Tubothalamea" family="Miliolidae" genus="Pseudonummoloculina" kingdom="Chromista" order="Miliolida" pageId="57" pageNumber="96" phylum="Foraminifera" rank="species" species="heimi">
<emphasis id="B938EAB2FFE0A22AFDBCF82AA738CF5F" box="[570,629,2044,2068]" italics="true" pageId="57" pageNumber="96">heimi</emphasis>
</taxonomicName>
in the literature and that a comprehensive revision is required. The solution would require the adequate stratigraphic separation of similar morphotypes to determine if relationships were evolutionary or of intra-specific variability.
</paragraph>
<paragraph id="8BF336A0FFE0A22AFCB6FEF5A13CC694" blockId="57.[816,1436,158,480]" pageId="57" pageNumber="96">
We agree in part with
<bibRefCitation id="EFDD4B51FFE0A22AFB99FEF4A053C671" author="Piuz, A. &amp; Vicedo, V." box="[1055,1310,290,314]" pageId="57" pageNumber="96" refId="ref93491" refString="Piuz, A. &amp; Vicedo, V., 2020. New Cenomanian &quot; nummoloculinas &quot; of the Natih Formation of Oman. Cretaceous Research, 107, https: // doi. org / 10.1016 / j. cretres. 2019.104224" type="book" year="2020">Piuz &amp; Vicedos (2020)</bibRefCitation>
proposal to restrict
<emphasis id="B938EAB2FFE0A22AFC02FE95A11CC610" box="[900,1105,323,347]" italics="true" pageId="57" pageNumber="96">
<taxonomicName id="4C4C4D23FFE0A22AFC02FE95A6F2C610" authority="(Bonet)" authorityName="Bonet" authorityYear="1956" baseAuthorityName="Bonet" baseAuthorityYear="1956" box="[900,959,323,347]" class="Tubothalamea" family="Miliolidae" genus="Pseudonummoloculina" kingdom="Chromista" order="Miliolida" pageId="57" pageNumber="96" phylum="Foraminifera" rank="species" sensu="stricto" species="heimi">heimi</taxonomicName>
<taxonomicNameLabel id="A20B57C9FFE0A22AFC4EFE92A11CC610" box="[968,1105,324,347]" pageId="57" pageNumber="96" sensu="stricto">sensu stricto</taxonomicNameLabel>
</emphasis>
to the morphotypes described by
<bibRefCitation id="EFDD4B51FFE0A22AFCD0FEB3A6ABC637" author="Bonet, F." box="[854,998,356,380]" pageId="57" pageNumber="96" pagination="389 - 488" refId="ref74941" refString="Bonet, F., 1956. Zonification microfaunistica de las Calizas Cretacicas del este de Mexico. Boletin Asociation Mexicana de Geologos Petroleros: 389 - 488." type="book chapter" year="1956">Bonet (1956)</bibRefCitation>
and
<bibRefCitation id="EFDD4B51FFE0A22AFBA7FEB2A074C637" author="Conkin, J. E. &amp; Conkin, B. M." box="[1057,1337,356,380]" pageId="57" pageNumber="96" pagination="149 - 158" refId="ref77378" refString="Conkin, J. E. &amp; Conkin, B. M., 1958. Revision of the genus Nummoloculina and emendation of Nummoloculina heimi Bonet. Micropaleontology, 4 (2): 149 - 158." type="journal article" year="1958">Conkin &amp; Conkin (1958)</bibRefCitation>
essentially the North American specimens although recognising that these themselves may comprise several separate taxa. Their characteristics are:
</paragraph>
<paragraph id="8BF336A0FFE0A22AFCE7FE3DA14FC56F" blockId="57.[865,1436,491,754]" pageId="57" pageNumber="96">
<emphasis id="B938EAB2FFE0A22AFCE7FE3DA620C548" bold="true" box="[865,877,491,515]" pageId="57" pageNumber="96"></emphasis>
compressed axially and lacks umbilical axial thickening
</paragraph>
<paragraph id="8BF336A0FFE0A22AFCE7FDF9A12EC523" blockId="57.[865,1436,491,754]" pageId="57" pageNumber="96">
<emphasis id="B938EAB2FFE0A22AFCE7FDF9A620C50C" bold="true" box="[865,877,559,583]" pageId="57" pageNumber="96"></emphasis>
small quinqueloculine nepionic stage (max 4 whorls/8 chambers)
</paragraph>
<paragraph id="8BF336A0FFE0A22AFCE7FDA5A06FC5C0" blockId="57.[865,1436,491,754]" box="[865,1314,627,651]" pageId="57" pageNumber="96">
<emphasis id="B938EAB2FFE0A22AFCE7FDA5A620C5C0" bold="true" box="[865,877,627,651]" pageId="57" pageNumber="96"></emphasis>
numerous planispiral whorls (up to 7)
</paragraph>
<paragraph id="8BF336A0FFE0A22AFCE7FD40A01BC5E5" blockId="57.[865,1436,491,754]" box="[865,1366,662,686]" pageId="57" pageNumber="96">
<emphasis id="B938EAB2FFE0A22AFCE7FD40A620C5E5" bold="true" box="[865,877,662,686]" pageId="57" pageNumber="96"></emphasis>
numerous chambers per whorl (6 up to 16)
</paragraph>
<paragraph id="8BF336A0FFE0A22AFCE7FD6FA1D9C5B9" blockId="57.[865,1436,491,754]" pageId="57" pageNumber="96">
<emphasis id="B938EAB2FFE0A22AFCE7FD6FA620C59A" bold="true" box="[865,877,697,721]" pageId="57" pageNumber="96"></emphasis>
aperture with “stocky tooth” but no notch/ridge/crenulations
</paragraph>
<paragraph id="8BF336A0FFE0A22AFCB6FD2DA0CDC334" blockId="57.[816,1437,763,1152]" pageId="57" pageNumber="96">
Our observations suggest that, in the majority and possibly all cases, forms attributed to
<taxonomicName id="4C4C4D23FFE0A22AFB70FCCAA07CC47F" authority="(Bonet)" authorityName="Bonet" authorityYear="1956" baseAuthorityName="Bonet" baseAuthorityYear="1956" box="[1270,1329,796,820]" class="Tubothalamea" family="Miliolidae" genus="Pseudonummoloculina" kingdom="Chromista" order="Miliolida" pageId="57" pageNumber="96" phylum="Foraminifera" rank="species" species="heimi">
<emphasis id="B938EAB2FFE0A22AFB70FCCAA07CC47F" box="[1270,1329,796,820]" italics="true" pageId="57" pageNumber="96">heimi</emphasis>
</taxonomicName>
in Europe/Africa/Middle East do not appear to conform wholly to these criteria and that further taxonomic revision is required to determine if they are as we suspect - additional, separate taxa. Some may be attributable with further research to
<taxonomicName id="4C4C4D23FFE0A22AFC58FC14A000C491" authorityName="Piuz &amp; Vicedo" authorityYear="2020" box="[990,1357,962,986]" class="Tubothalamea" family="Hauerinidae" genus="Nummoloculinodonta" higherTaxonomySource="CoL,GBIF" kingdom="Chromista" order="Miliolida" pageId="57" pageNumber="96" phylum="Foraminifera" rank="species" species="akhdarensis">
<emphasis id="B938EAB2FFE0A22AFC58FC14A000C491" box="[990,1357,962,986]" italics="true" pageId="57" pageNumber="96">Nummoloculinodonta akhdarensis</emphasis>
</taxonomicName>
, others to
<taxonomicName id="4C4C4D23FFE0A22AFCC9FC32A1AFC4B0" authority="Chiocchini et al." authorityName="Chiocchini" box="[847,1250,995,1019]" class="Tubothalamea" family="Miliolidae" genus="Pseudonummoloculina" kingdom="Chromista" order="Miliolida" pageId="57" pageNumber="96" phylum="Foraminifera" rank="subSpecies" species="regularis" subSpecies="sensu">
<emphasis id="B938EAB2FFE0A22AFCC9FC32A612C4B0" box="[847,863,996,1019]" italics="true" pageId="57" pageNumber="96">P</emphasis>
.?
<emphasis id="B938EAB2FFE0A22AFCFDFC35A16FC4B0" box="[891,1058,995,1019]" italics="true" pageId="57" pageNumber="96">regularis sensu</emphasis>
Chiocchini et al.
</taxonomicName>
, or a completely new species. Such work is beyond the scope of this article, and we have therefore, reluctantly, placed
<taxonomicName id="4C4C4D23FFE0A22AFAC4FBF3A030C376" authority="(Bonet)" authorityName="Bonet" authorityYear="1956" baseAuthorityName="Bonet" baseAuthorityYear="1956" box="[1346,1405,1061,1085]" class="Tubothalamea" family="Miliolidae" genus="Pseudonummoloculina" kingdom="Chromista" order="Miliolida" pageId="57" pageNumber="96" phylum="Foraminifera" rank="species" species="heimi">
<emphasis id="B938EAB2FFE0A22AFAC4FBF3A030C376" box="[1346,1405,1061,1085]" italics="true" pageId="57" pageNumber="96">heimi</emphasis>
</taxonomicName>
in open nomenclature (“ex gr.”) and assigned it to an unsatisfactory genus, as a problem to be solved in the future.
</paragraph>
<paragraph id="8BF336A0FFE0A22AFCB6F870A621CF6B" blockId="57.[816,1437,1957,2080]" pageId="57" pageNumber="96">
<taxonomicName id="4C4C4D23FFE0A22AFCB6F870A6D5C0F6" baseAuthorityName="Bonet" baseAuthorityYear="1956" box="[816,920,1957,1981]" class="Tubothalamea" family="Miliolidae" genus="Pseudonummoloculina" kingdom="Chromista" order="Miliolida" pageId="57" pageNumber="96" phylum="Foraminifera" rank="species" species="heimi">
<emphasis id="B938EAB2FFE0A22AFCB6F870A6D5C0F6" box="[816,920,1957,1981]" italics="true" pageId="57" pageNumber="96">P.? heimi</emphasis>
</taxonomicName>
is more axially compressed than
<taxonomicName id="4C4C4D23FFE0A22AFA80F870A0D8C0F6" authorityName="Calvez" authorityYear="1988" box="[1286,1429,1958,1981]" class="Tubothalamea" family="Miliolidae" genus="Pseudonummoloculina" kingdom="Chromista" order="Miliolida" pageId="57" pageNumber="96" phylum="Foraminifera" rank="species" species="aurigerica">
<emphasis id="B938EAB2FFE0A22AFA80F870A0D8C0F6" box="[1286,1429,1958,1981]" italics="true" pageId="57" pageNumber="96">P. aurigerica</emphasis>
</taxonomicName>
, has more numerous planispiral whorls and chambers per whorl and does not appear to have notches or crenulations.
</paragraph>
<paragraph id="8BF336A0FFE3A229FF11FF48A4CBC637" blockId="58.[151,772,158,380]" pageId="58" pageNumber="97">
Despite having fewer chambers in each post-embryonic whorl (3-5 cf. 6-16)
<taxonomicName id="4C4C4D23FFE3A229FEF1FF16A64EC79C" authority="Chiocchini et al." authorityName="Chiocchini" box="[375,771,191,215]" class="Tubothalamea" family="Miliolidae" genus="Pseudonummoloculina" kingdom="Chromista" order="Miliolida" pageId="58" pageNumber="97" phylum="Foraminifera" rank="subSpecies" species="regularis" subSpecies="sensu">
<emphasis id="B938EAB2FFE3A229FEF1FF16A705C79C" box="[375,584,191,215]" italics="true" pageId="58" pageNumber="97">P.? regularis sensu</emphasis>
Chiocchini et al.
</taxonomicName>
can also appear similar in some random section orientations with a similar number of post embryonic planispiral coils (up to 7), although unlike
<emphasis id="B938EAB2FFE3A229FD84FEF5A76AC671" box="[514,551,291,314]" italics="true" pageId="58" pageNumber="97">P.?</emphasis>
ex. grp.
<taxonomicName id="4C4C4D23FFE3A229FD14FEF4A780C671" authority="(Bonet)" authorityName="Bonet" authorityYear="1956" baseAuthorityName="Bonet" baseAuthorityYear="1956" box="[658,717,290,314]" class="Tubothalamea" family="Miliolidae" genus="Pseudonummoloculina" kingdom="Chromista" order="Miliolida" pageId="58" pageNumber="97" phylum="Foraminifera" rank="species" species="heimi">
<emphasis id="B938EAB2FFE3A229FD14FEF4A780C671" box="[658,717,290,314]" italics="true" pageId="58" pageNumber="97">heimi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4C4D23FFE3A229FD59FEF5A749C610" authority="sensu Chiocchini et al." authorityName="sensu Chiocchini et al." authorityYear="2012" baseAuthorityName="Philippson" baseAuthorityYear="1887" class="Tubothalamea" family="Miliolidae" genus="Pseudonummoloculina" kingdom="Chromista" order="Miliolida" pageId="58" pageNumber="97" phylum="Foraminifera" rank="species" species="regularis">
<emphasis id="B938EAB2FFE3A229FD59FEF5A5B4C610" italics="true" pageId="58" pageNumber="97">P.? regularis</emphasis>
sensu Chiocchini et al.
</taxonomicName>
is planispiral virtually throughout its growth.
</paragraph>
</subSubSection>
<subSubSection id="C356652BFFE3A229FF11FE71A5B2C569" pageId="58" pageNumber="97" type="distribution">
<paragraph id="8BF336A0FFE3A229FF11FE71A4E5C6F4" blockId="58.[151,424,423,447]" box="[151,424,423,447]" pageId="58" pageNumber="97">Stratigraphic Distribution</paragraph>
<paragraph id="8BF336A0FFE3A229FF11FE3FA5B2C569" blockId="58.[151,772,489,2035]" pageId="58" pageNumber="97">(late Aptian?) Albian Cenomanian (?Turonian and younger).</paragraph>
</subSubSection>
<subSubSection id="C356652BFFE3A228FF11FDFDA6A7C583" lastPageId="59" lastPageNumber="98" pageId="58" pageNumber="97" type="description">
<paragraph id="8BF336A0FFE3A229FF11FDFDA7E4C4C5" blockId="58.[151,772,489,2035]" pageId="58" pageNumber="97">
Notwithstanding the comments made above, an evaluation of published records suggests
<emphasis id="B938EAB2FFE3A229FDA6FD9BA708C52F" box="[544,581,589,612]" italics="true" pageId="58" pageNumber="97">P.?</emphasis>
ex grp.
<taxonomicName id="4C4C4D23FFE3A229FD2DFD9AA7ABC52F" authority="(Bonet)" authorityName="Bonet" authorityYear="1956" baseAuthorityName="Bonet" baseAuthorityYear="1956" box="[683,742,588,612]" class="Tubothalamea" family="Miliolidae" genus="Pseudonummoloculina" kingdom="Chromista" order="Miliolida" pageId="58" pageNumber="97" phylum="Foraminifera" rank="species" species="heimi">
<emphasis id="B938EAB2FFE3A229FD2DFD9AA7ABC52F" box="[683,742,588,612]" italics="true" pageId="58" pageNumber="97">heimi</emphasis>
</taxonomicName>
is widely distributed with the majority from Albian-Cenomanian strata. Many records are accompanied by illustration but some of these illustrations do not necessarily confirm identity. In addition, the generic assignment (
<taxonomicName id="4C4C4D23FFE3A229FF64FD24A49BC441" authorityName="Calvez" authorityYear="1988" box="[226,470,754,778]" class="Tubothalamea" family="Miliolidae" genus="Pseudonummoloculina" kingdom="Chromista" order="Miliolida" pageId="58" pageNumber="97" phylum="Foraminifera" rank="genus">
<emphasis id="B938EAB2FFE3A229FF64FD24A49BC441" box="[226,470,754,778]" italics="true" pageId="58" pageNumber="97">Pseudonummoloculina</emphasis>
</taxonomicName>
or
<taxonomicName id="4C4C4D23FFE3A229FD83FD24A7E2C441" authorityName="emend. Conkin &amp; Conkin" authorityYear="1958" box="[517,687,754,778]" class="Tubothalamea" family="Hauerinidae" genus="Nummoloculina" kingdom="Chromista" order="Miliolida" pageId="58" pageNumber="97" phylum="Foraminifera" rank="genus">
<emphasis id="B938EAB2FFE3A229FD83FD24A7E2C441" box="[517,687,754,778]" italics="true" pageId="58" pageNumber="97">Nummoloculina</emphasis>
</taxonomicName>
) varies between authors. The upper age limit of this species is difficult to pinpoint although an extension into the Turonian appears possible. Records above this level need further evaluation (e.g.,
<bibRefCitation id="EFDD4B51FFE3A229FE1FFCA0A7D6C4C5" author="Tsaila-Monopolis, S." box="[409,667,886,910]" pageId="58" pageNumber="97" pagination="1 - 106" refId="ref101018" refString="Tsaila-Monopolis, S., 1977. Micropaleontological and stratigraphical study of the Tripolitza (Gavrovo) zone in the Peloponnesus. National Institute for Research and Development in Geology, Geophysics, Geochemistry and Remote Sensing, 20 (1): 1 - 106. [in Greek]" type="journal article" year="1977">Tsaila-Monopolis, 1977</bibRefCitation>
).
</paragraph>
<paragraph id="8BF336A0FFE3A229FF11FC41A402C33E" blockId="58.[151,772,489,2035]" pageId="58" pageNumber="97">
<taxonomicName id="4C4C4D23FFE3A229FF11FC41A4ADC4E4" ID-CoL="8ZF4K" baseAuthorityName="Bonet" baseAuthorityYear="1956" box="[151,480,919,943]" class="Tubothalamea" family="Miliolidae" genus="Pseudonummoloculina" kingdom="Chromista" order="Miliolida" pageId="58" pageNumber="97" phylum="Foraminifera" rank="species" species="heimi">
<emphasis id="B938EAB2FFE3A229FF11FC41A4ADC4E4" box="[151,480,919,943]" italics="true" pageId="58" pageNumber="97">Pseudonummoloculina? heimi</emphasis>
</taxonomicName>
was first described from the Albian Cenomanian El Abra formation of
<collectingCountry id="F35B7630FFE3A229FD34FC6EA64EC49B" box="[690,771,952,976]" name="Mexico" pageId="58" pageNumber="97">Mexico</collectingCountry>
(
<bibRefCitation id="EFDD4B51FFE3A229FF19FC0FA467C4BA" author="Bonet, F." box="[159,298,985,1009]" pageId="58" pageNumber="97" pagination="389 - 488" refId="ref74941" refString="Bonet, F., 1956. Zonification microfaunistica de las Calizas Cretacicas del este de Mexico. Boletin Asociation Mexicana de Geologos Petroleros: 389 - 488." type="book chapter" year="1956">Bonet, 1956</bibRefCitation>
).
<bibRefCitation id="EFDD4B51FFE3A229FEC7FC0FA720C4BA" author="Conkin, J. E. &amp; Conkin, B. M." box="[321,621,985,1009]" pageId="58" pageNumber="97" pagination="149 - 158" refId="ref77378" refString="Conkin, J. E. &amp; Conkin, B. M., 1958. Revision of the genus Nummoloculina and emendation of Nummoloculina heimi Bonet. Micropaleontology, 4 (2): 149 - 158." type="journal article" year="1958">Conkin &amp; Conkins (1958)</bibRefCitation>
material was from
<collectingCountry id="F35B7630FFE3A229FF51FC2CA465C359" box="[215,296,1018,1042]" name="Mexico" pageId="58" pageNumber="97">Mexico</collectingCountry>
(El Abra Formation), and
<collectingRegion id="4988F842FFE3A229FDD9FC2DA7D2C359" box="[607,671,1019,1042]" country="United States of America" name="Texas" pageId="58" pageNumber="97">Texas</collectingRegion>
(Devils River, Edwards and Glen Rose Formations) and
<collectingRegion id="4988F842FFE3A229FD30FBCDA64EC378" box="[694,771,1051,1075]" country="United States of America" name="Florida" pageId="58" pageNumber="97">Florida</collectingRegion>
(Fredericksburg Formation) of the
<collectingCountry id="F35B7630FFE3A229FD95FBEAA705C31F" box="[531,584,1084,1108]" name="United States of America" pageId="58" pageNumber="97">USA</collectingCountry>
(see also
<bibRefCitation id="EFDD4B51FFE3A229FD3FFBEAA40EC33E" author="Applin, P. L. &amp; Applin, E. R." pageId="58" pageNumber="97" pagination="1 - 84" refId="ref71877" refString="Applin, P. L. &amp; Applin, E. R., 1965. The Comanche Series and associated rocks in the subsurface in central and south Florida. U. S. Geological Survey Professional Paper 447, pp. 1 - 84." type="journal article" year="1965">Applin &amp; Applin, 1965</bibRefCitation>
).
</paragraph>
<paragraph id="8BF336A0FFE3A229FF11FBA9A752C182" blockId="58.[151,772,489,2035]" pageId="58" pageNumber="97">
Other confirmed illustrated records of this species from
<collectingCountry id="F35B7630FFE3A229FF11FB76A5A4C3F3" box="[151,233,1184,1208]" name="Mexico" pageId="58" pageNumber="97">Mexico</collectingCountry>
include
<bibRefCitation id="EFDD4B51FFE3A229FEDAFB76A7DDC3F3" author="Rosales-Dominguez, M." box="[348,656,1184,1208]" pageId="58" pageNumber="97" refId="ref95422" refString="Rosales-Dominguez, M. del C., 1989. Micropaleontology and paleogeography of the Upper Mural Limestone of southeastern Arizona and northern Sonora. Unpublished MSc thesis, University of Arizona, 61 pp." type="book" year="1989">Rosales-Dominguez (1989)</bibRefCitation>
;
<bibRefCitation id="EFDD4B51FFE3A229FD21FB76A4E8C392" author="Rosales-Dominguez, M. &amp; Bermudez-Santana, J. C. &amp; Aguilar-Pina, M." pageId="58" pageNumber="97" pagination="697 - 712" refId="ref95499" refString="Rosales-Dominguez, M. del C., Bermudez-Santana, J. C. &amp; Aguilar-Pina, M., 1997. Mid and Upper Cretaceous foraminiferal assemblages from the Sierra de Chiapas, southeastern Mexico. Cretaceous Research, 18: 697 - 712." type="journal article" year="1997">Rosales-Dominguez et al. (1997)</bibRefCitation>
and
<bibRefCitation id="EFDD4B51FFE3A229FE59FB17A7FBC392" author="Omana, L. &amp; Lopez-Doncel, R. &amp; Torres-Hernandez, J. R." box="[479,694,1217,1241]" pageId="58" pageNumber="97" pagination="1 - 94" refId="ref91696" refString="Omana, L., Lopez-Doncel, R., Torres-Hernandez, J. R., 2013, Late Cretaceous (Cenomanian-Santonian) foraminiferal biostratigraphy from W Valles-San Luis Platform, Mexico: 9 th Symposium of Cretaceous System, Ankara, Turkey, pp. 1 - 94." type="book chapter" year="2013">Omaña et al. (2013</bibRefCitation>
,
<bibRefCitation id="EFDD4B51FFE3A229FD42FB17A7B7C392" author="Omana, L. &amp; Lopez-Doncel, R. &amp; Torres, J. R. &amp; Alencaster, G. &amp; Lopez-Caballero, I." box="[708,762,1217,1241]" pageId="58" pageNumber="97" pagination="691 - 725" refId="ref91622" refString="Omana, L., Lopez-Doncel, R., Torres, J. R., Alencaster, G. &amp; Lopez-Caballero, I. 2019. Mid - late Cenomanian larger benthic foraminifers from the El Abra Formation W Valles-San Luis Potosi Platform, central - eastern Mexico: Taxonomy, biostratigraphy and paleoenvironmental implications. Boletin de la Sociedad Geologica Mexicana, 71 (3): 691 - 725." type="journal article" year="2019">2019</bibRefCitation>
) respectively from the Albian early Cenomanian Sierra Madre Formation and the Albian Cenomanian El Abra Formation. Scott &amp; Gonzalez-Leon (1991) recorded the species from the middle Albian Espinazo del Diablo and Nogal formations of the Lampazos region. Unillustrated records include
<bibRefCitation id="EFDD4B51FFE3A229FECEFA51A713C2D4" author="Ontiveros-Tarango, G." box="[328,606,1415,1439]" pageId="58" pageNumber="97" pagination="189 - 234" refId="ref92161" refString="Ontiveros-Tarango, G., 1973. Estudio estratigrafico de la porcin noroccidental de la Cuenca Morelos Guerrero. Boletin de la Associacion Mexicana de Geologos Petroleros, 25: 189 - 234." type="journal article" year="1973">Ontiveros-Tarango (1973</bibRefCitation>
; Cenomanian);
<bibRefCitation id="EFDD4B51FFE3A229FF11FA7FA4B8C28A" author="Hernandez-Romano, U. &amp; Aguilera-Franco, N. &amp; Medrano, M. &amp; Barcelo-Duarte, J." box="[151,501,1449,1473]" pageId="58" pageNumber="97" pagination="661 - 686" refId="ref83861" refString="Hernandez-Romano, U., Aguilera-Franco, N., Martinez- Medrano, M. &amp; Barcelo-Duarte, J., 1997. Guerrero-Morelos Platform drowning at the Cenomanian - Turonian boundary, Huitziltepec area, Guerrero State, southern Mexico. Cretaceous Research, 18: 661 - 686." type="journal article" year="1997">Hernández-Romano et al. (1997</bibRefCitation>
; late Cenomanian);
<bibRefCitation id="EFDD4B51FFE3A229FD57FA7FA45AC2A9" author="Cros, P. &amp; Michaud, F. &amp; Fourcade, E. &amp; Fleury, J. - J." pageId="58" pageNumber="97" pagination="311 - 332" refId="ref78074" refString="Cros, P., Michaud, F., Fourcade, E. &amp; Fleury, J. - J., 1998. Sedimentological evolution of the Cretaceous carbonate platform of Chiapas (Mexico). Journal of South American Earth Sciences, 11 (4): 311 - 332." type="journal article" year="1998">Cros et al. (1998</bibRefCitation>
; Cenomanian);
<bibRefCitation id="EFDD4B51FFE3A229FE43FA1CA64EC2A9" author="Aguilera-Franco, N. &amp; Hernandez-Romano, U. &amp; Allison, P. A." box="[453,771,1482,1506]" pageId="58" pageNumber="97" pagination="237 - 255" refId="ref70732" refString="Aguilera-Franco, N., Hernandez-Romano, U. &amp; Allison, P. A., 2001. Biostratigraphy and environmental changes across the Cenomanian-Turonian boundary, southern Mexico. Journal of South American Earth Sciences, 14: 237 - 255." type="journal article" year="2001">Aguilera-Franco et al. (2001)</bibRefCitation>
and
<bibRefCitation id="EFDD4B51FFE3A229FF48FA3DA49BC148" author="Aguilera-Franco, N." box="[206,470,1515,1539]" pageId="58" pageNumber="97" pagination="202 - 222" refId="ref70692" refString="Aguilera-Franco, N., 2003. Cenomanian - Coniacian zonation (foraminifers and calcareous algae) in the Guerrero-Morelos basin, southern Mexico. Revista Mexicana de Ciencias Geologicas, 20 (3): 202 - 222." type="journal article" year="2003">Aguilera-Franco (2003)</bibRefCitation>
(from the middle late Cenomanian); and
<bibRefCitation id="EFDD4B51FFE3A229FEF1F9DAA7B2C16F" author="Aguilera-Franco, N. &amp; Allison, P." box="[375,767,1548,1572]" pageId="58" pageNumber="97" pagination="25 - 50" refId="ref70580" refString="Aguilera-Franco, N. &amp; Allison, P., 2004. Events of the Cenomanian-Turonian succession, southern Mexico. Journal of Iberian Geology, 31: 25 - 50." type="journal article" year="2004">Aguilera-Franco &amp; Allison (2004</bibRefCitation>
; undated) from the
<collectingRegion id="4988F842FFE3A229FEEFF9FBA48FC10E" box="[361,450,1581,1605]" country="Mexico" name="Morelos" pageId="58" pageNumber="97">Morelos</collectingRegion>
Formation. However, the illustration in
<bibRefCitation id="EFDD4B51FFE3A229FEA7F998A76CC12D" author="Aguilera-Franco, N." box="[289,545,1614,1638]" pageId="58" pageNumber="97" refId="ref70663" refString="Aguilera-Franco, N., 2000. High Resolution Stratigraphy and Palaeoecology of the Cenomanian-Turonian Succession, Southern Mexico. Unpublished PhD thesis, Imperial College, London." type="book" year="2000">Aguilera-Franco (2000)</bibRefCitation>
is not identifiable at species level but appears incompatible with
<emphasis id="B938EAB2FFE3A229FD07F9A6A7EBC1CC" box="[641,678,1648,1671]" italics="true" pageId="58" pageNumber="97">P.?</emphasis>
ex. grp.
<taxonomicName id="4C4C4D23FFE3A229FF11F946A59FC1E3" ID-CoL="8ZF4K" authority="(Bonet)" authorityName="Bonet" authorityYear="1956" baseAuthorityName="Bonet" baseAuthorityYear="1956" box="[151,210,1680,1704]" class="Tubothalamea" family="Miliolidae" genus="Pseudonummoloculina" kingdom="Chromista" order="Miliolida" pageId="58" pageNumber="97" phylum="Foraminifera" rank="species" species="heimi">
<emphasis id="B938EAB2FFE3A229FF11F946A59FC1E3" box="[151,210,1680,1704]" italics="true" pageId="58" pageNumber="97">heimi</emphasis>
</taxonomicName>
as described herein. Lucas et al. (2015) provided an illustrated record from New
<collectingRegion id="4988F842FFE3A229FE41F967A751C182" box="[455,540,1713,1737]" country="Mexico" name="Mexico" pageId="58" pageNumber="97">Mexico</collectingRegion>
.
</paragraph>
<paragraph id="8BF336A0FFE3A229FF11F904A063C79C" blockId="58.[151,772,489,2035]" lastBlockId="58.[816,1437,158,2068]" pageId="58" pageNumber="97">
<bibRefCitation id="EFDD4B51FFE3A229FF11F904A41CC1A1" author="Ashworth, E. T." box="[151,337,1746,1770]" pageId="58" pageNumber="97" refId="ref72702" refString="Ashworth, E. T., 1974. Foraminifera from the Cretaceous of the Central Cordillera of Guatemala. Unpublished PhD thesis, Ohio State University." type="book" year="1974">Ashworth (1974)</bibRefCitation>
;
<bibRefCitation id="EFDD4B51FFE3A229FEE7F904A71CC1A1" author="Caceres Flores, A. F." box="[353,593,1746,1770]" pageId="58" pageNumber="97" refId="ref75990" refString="Caceres Flores, A. F., 2016. Estudio geologico - geotecnico para el diseno del soporte de los adit del Tunel 2 - Proyecto hidroeletrico renace II, San pedro Carcha, Alta Verapaz - Guatemala. Unpublished PhD thesis, Universidad Nacional de San Augustin." type="book" year="2016">
Caceres
<collectingCountry id="F35B7630FFE3A229FE38F904A74CC1A1" box="[446,513,1746,1770]" name="Indonesia" pageId="58" pageNumber="97">Flores</collectingCountry>
(2016)
</bibRefCitation>
and
<bibRefCitation id="EFDD4B51FFE3A229FD0CF904A46FC047" author="Radmacher, W. &amp; Vasquez, O. J. &amp; Tzalam, M. &amp; Jolomna, M. &amp; Molineros, A. &amp; Eldrett, J. S." pageId="58" pageNumber="97" refId="ref93753" refString="Radmacher, W., Vasquez, O. J., Tzalam, M., Jolomna, M., Molineros, A. &amp; Eldrett, J. S., 2021. What happened to the organic matter from the Upper Cretaceous succession in Guatemala, Central America? Marine and Petroleum Geology, 133, https: // doi. org / 10.1016 / j. marpetgeo. 2021.105246" type="book" year="2021">Radmacher et al. (2021)</bibRefCitation>
have plausible illustrated records from the Albian Cenomanian Coban Formation of
<collectingCountry id="F35B7630FFE3A229FD09F8C3A64FC066" box="[655,770,1813,1837]" name="Guatemala" pageId="58" pageNumber="97">Guatemala</collectingCountry>
(see also
<bibRefCitation id="EFDD4B51FFE3A229FF7AF8E0A4AFC005" author="Fourcade, E. &amp; Piccioni, L. &amp; Escriba, J. &amp; Rosselo, E." box="[252,482,1846,1870]" pageId="58" pageNumber="97" pagination="793 - 811" refId="ref81404" refString="Fourcade, E., Piccioni, L., Escriba, J. &amp; Rosselo, E., 1999. Cretaceous stratigraphy and palaeoenvironments of the southern Peten Basin, Guatemala: Cretaceous Research, 20: 793 - 811." type="journal article" year="1999">Fourcade et al., 1999</bibRefCitation>
, unillustrated, but Albian). However, the record by
<bibRefCitation id="EFDD4B51FFE3A229FE1AF881A717C024" author="Moeschler, B. G. C." box="[412,602,1879,1903]" pageId="58" pageNumber="97" refId="ref90248" refString="Moeschler, B. G. C., 2009. Evaluacionde la amenaza por inundacion en la parte alst de la Cuenca del Rio Cahabon. Unpublished PhD thesis. Universidad de San Carlos de Guatemala." type="book" year="2009">Moeschler (2009)</bibRefCitation>
is probably not this species (it may be
<taxonomicName id="4C4C4D23FFE3A229FE54F8AEA7DAC0DB" box="[466,663,1912,1936]" class="Globothalamea" family="Rzehakinidae" genus="Spiroloculina" kingdom="Chromista" order="Lituolida" pageId="58" pageNumber="97" phylum="Foraminifera" rank="species" species="undetermined">
<emphasis id="B938EAB2FFE3A229FE54F8AEA72EC0DB" box="[466,611,1912,1936]" italics="true" pageId="58" pageNumber="97">Spiroloculina</emphasis>
sp.
</taxonomicName>
).
<bibRefCitation id="EFDD4B51FFE3A229FD3DF8AEA712C0FA" author="Ayala Castanares, A. &amp; Furrazola-Bermudez, G." pageId="58" pageNumber="97" pagination="1 - 10" refId="ref72852" refString="Ayala Castanares, A. &amp; Furrazola-Bermudez, G., 1962. Nummoloculina heimi Bonet en el Cretacico Inferior de Cuba. Institute of Geology, National Autonomous University of Mexico, Paleontology, 12: 1 - 10." type="journal article" year="1962">Ayala-Castañares &amp; Furrazola-Bermúdez (1962)</bibRefCitation>
provide excellent illustrations of this species from the Albian Cenomanian of
<collectingCountry id="F35B7630FFE3A229FE89F80DA407C0B8" box="[271,330,2011,2035]" name="Cuba" pageId="58" pageNumber="97">Cuba</collectingCountry>
, whilst Diaz Otero et al. (2001) records but does not illustrate this species.
<bibRefCitation id="EFDD4B51FFE3A229FB3AFF48A0D1C7FD" author="Rogers, R. D. &amp; Mann, P. &amp; Scott, R. W. &amp; Patino, L." box="[1212,1436,158,182]" pageId="58" pageNumber="97" pagination="89 - 128" refId="ref95369" refString="Rogers, R. D., Mann, P., Scott, R. W. &amp; Patino, L., 2007. Cretaceous intra-arc rifting, sedimentation, and basin inversion in east-central Honduras. Geological Society of America, Special Paper, 428: 89 - 128." type="journal article" year="2007">Rogers et al. (2007)</bibRefCitation>
report the species from the Albian of
<collectingCountry id="F35B7630FFE3A229FB46FF69A066C79C" box="[1216,1323,191,215]" name="Honduras" pageId="58" pageNumber="97">Honduras</collectingCountry>
.
</paragraph>
<paragraph id="8BF336A0FFE3A229FCB6FF37A6A7C6D5" blockId="58.[816,1437,158,2068]" pageId="58" pageNumber="97">
In Western Europe records from
<collectingCountry id="F35B7630FFE3A229FB33FF36A042C7B3" box="[1205,1295,224,248]" name="Portugal" pageId="58" pageNumber="97">Portugal</collectingCountry>
by
<bibRefCitation id="EFDD4B51FFE3A229FAC1FF36A635C652" author="Berthou, P-Y." pageId="58" pageNumber="97" pagination="1 - 168" refId="ref74017" refString="Berthou, P-Y., 1973. Le Cenomanien de l'Estremadure Portugaise. Servicos Geologicos de Portugal, 23: 1 - 168." type="journal article" year="1973">Berthou (1973)</bibRefCitation>
and
<bibRefCitation id="EFDD4B51FFE3A229FC2BFED7A11BC652" author="Andrade, I. B." box="[941,1110,257,281]" pageId="58" pageNumber="97" refId="ref71641" refString="Andrade, I. B., 2018. Estudo paleoecologico e biostratigrafico do Cenomaniano da regiao de Lisboa (Lousa-Salemas) baseado em ostracodos e microfacies. Unpublished PhD thesis, University of Lisbon, 147 pp." type="book" year="2018">Andrade (2018)</bibRefCitation>
are illustrated but the illustration cannot be confirmed as
<emphasis id="B938EAB2FFE3A229FBE8FEF5A1DEC671" box="[1134,1171,291,314]" italics="true" pageId="58" pageNumber="97">P.?</emphasis>
ex. grp.
<taxonomicName id="4C4C4D23FFE3A229FB7CFEF4A078C671" authority="(Bonet)" authorityName="Bonet" authorityYear="1956" baseAuthorityName="Bonet" baseAuthorityYear="1956" box="[1274,1333,290,314]" class="Tubothalamea" family="Miliolidae" genus="Pseudonummoloculina" kingdom="Chromista" order="Miliolida" pageId="58" pageNumber="97" phylum="Foraminifera" rank="species" species="heimi">
<emphasis id="B938EAB2FFE3A229FB7CFEF4A078C671" box="[1274,1333,290,314]" italics="true" pageId="58" pageNumber="97">heimi</emphasis>
</taxonomicName>
. Records by
<bibRefCitation id="EFDD4B51FFE3A229FCD8FE95A1E3C610" author="Berthou, P-Y. &amp; Lauverjat, J." box="[862,1198,323,347]" pageId="58" pageNumber="97" pagination="121 - 144" refId="ref73919" refString="Berthou, P-Y. &amp; Lauverjat, J., 1979. Essai de synthese paleogeographique et paleobiostratigraphique du bassin occidental Portugais au cours du Cretace Superieur. Ciencias da Terra (UNL), Lisboa, 5: 121 - 144." type="journal article" year="1979">Berthou &amp; Lauverjat (1979)</bibRefCitation>
and Crosaz-Galletti (1979) are unillustrated. Most assign a middle late Cenomanian age.
</paragraph>
<paragraph id="8BF336A0FFE3A229FCB6FE71A1F4C359" blockId="58.[816,1437,158,2068]" pageId="58" pageNumber="97">
Records from
<collectingCountry id="F35B7630FFE3A229FC4DFE71A6B1C6F4" box="[971,1020,423,447]" name="Italy" pageId="58" pageNumber="97">Italy</collectingCountry>
are numerous (
<bibRefCitation id="EFDD4B51FFE3A229FB22FE71A016C6F4" author="De Castro, P." box="[1188,1371,423,447]" pageId="58" pageNumber="97" pagination="317 - 372" refId="ref78519" refString="De Castro, P., 1965. Su alcune Soritidae (Foraminiferida) del Cretacico della Campania. Societa dei Naturalisti in Napoli, 74: 317 - 372." type="journal article" year="1965">De Castro (1965</bibRefCitation>
;
<bibRefCitation id="EFDD4B51FFE3A229FAE1FE71A0D0C6F4" author="De Castro, P." box="[1383,1437,423,447]" pageId="58" pageNumber="97" pagination="109 - 124" refId="ref78786" refString="De Castro, P., 1987. On some foraminifera and algae in Apennine Upper Cretaceous and Paleocene: Memorie della Societa Geologica Italiana, 40: 109 - 124." type="journal article" year="1987">1987</bibRefCitation>
see comments above regarding synonymy with
<taxonomicName id="4C4C4D23FFE3A229FA02FE1FA6FCC54A" authorityName="Piuz &amp; Vicedo" authorityYear="2020" class="Tubothalamea" family="Hauerinidae" genus="Nummoloculinodonta" higherTaxonomySource="CoL,GBIF" kingdom="Chromista" order="Miliolida" pageId="58" pageNumber="97" phylum="Foraminifera" rank="species" species="akhdarensis">
<emphasis id="B938EAB2FFE3A229FA02FE1FA6FCC54A" italics="true" pageId="58" pageNumber="97">N. akhdarensis</emphasis>
</taxonomicName>
); Borghi &amp; Pignatti (2006);
<bibRefCitation id="EFDD4B51FFE3A229FB7AFE3FA636C569" author="Consorti, L. &amp; Caus, E. &amp; Frijia, G. &amp; Yazdi-Moghadam, M." pageId="58" pageNumber="97" pagination="378 - 389" refId="ref77762" refString="Consorti, L., Caus, E., Frijia, G. &amp; Yazdi-Moghadam, M., 2015. Praetaberina a new genus (type species Taberina bingistani Henson, 1948): a stratigraphic marker for the Late Cenomanian. Journal of Foraminiferal Research, 45 (4): 378 - 389." type="journal article" year="2015">Consorti et al. (2015)</bibRefCitation>
;
<bibRefCitation id="EFDD4B51FFE3A229FC0CFDDCA10EC569" author="Crescenti, U." box="[906,1091,522,546]" pageId="58" pageNumber="97" pagination="15 - 40" refId="ref78047" refString="Crescenti, U., 1969. Biostratigrafia delle facies Mesozoiche dell'Appennino Centrale: correlazioni. Geol. Rom., VIII: 15 - 40." type="journal article" year="1969">Crescenti (1969)</bibRefCitation>
; Di Stefano &amp; Ruberti (2000);
<bibRefCitation id="EFDD4B51FFE3A229FCB6FDFDA106C508" author="Spalluto, L. &amp; Caffau, M." box="[816,1099,555,579]" pageId="58" pageNumber="97" pagination="335 - 352" refId="ref100174" refString="Spalluto, L. &amp; Caffau, M., 2010. Stratigraphy of the mid-Cretaceous shallow-water limestones of the Apulia Carbonate Platform (Murge, Apulia, southern Italy). Italian Journal of Geosciences, 129 (3): 335 - 352." type="journal article" year="2010">Spalluto &amp; Caffau (2010)</bibRefCitation>
and
<bibRefCitation id="EFDD4B51FFE3A229FB03FDFDA07AC508" author="Spalluto, L." box="[1157,1335,555,579]" pageId="58" pageNumber="97" refId="ref100220" refString="Spalluto, L., 2011. Facies evolution and sequence chronostratigraphy of a &quot; mid &quot; - Cretaceous shallow-water carbonate succession of the Apulia Carbonate Platform from the northern Murge area (Apulia, southern Italy). Facies, DOI 10.1007 / s 10347 - 011 - 0266 - 0" type="book" year="2011">Spalluto (2011))</bibRefCitation>
but only the record of
<bibRefCitation id="EFDD4B51FFE3A229FC4FFD9AA1A9C52F" author="Spalluto, L. &amp; Caffau, M." box="[969,1252,588,612]" pageId="58" pageNumber="97" pagination="335 - 352" refId="ref100174" refString="Spalluto, L. &amp; Caffau, M., 2010. Stratigraphy of the mid-Cretaceous shallow-water limestones of the Apulia Carbonate Platform (Murge, Apulia, southern Italy). Italian Journal of Geosciences, 129 (3): 335 - 352." type="journal article" year="2010">Spalluto &amp; Caffau (2010)</bibRefCitation>
is confirmed by illustration. Ages assigned are from early late Cenomanian, but
<bibRefCitation id="EFDD4B51FFE3A229FC13FD58A10AC5ED" author="Crescenti, U." box="[917,1095,654,678]" pageId="58" pageNumber="97" pagination="15 - 40" refId="ref78047" refString="Crescenti, U., 1969. Biostratigrafia delle facies Mesozoiche dell'Appennino Centrale: correlazioni. Geol. Rom., VIII: 15 - 40." type="journal article" year="1969">Crescenti (1969)</bibRefCitation>
indicates this species ranges up to the “Senonian” (see also
<bibRefCitation id="EFDD4B51FFE3A229FBD9FD79A02DC58C" author="Tsaila-Monopolis, S." box="[1119,1376,687,711]" pageId="58" pageNumber="97" pagination="1 - 106" refId="ref101018" refString="Tsaila-Monopolis, S., 1977. Micropaleontological and stratigraphical study of the Tripolitza (Gavrovo) zone in the Peloponnesus. National Institute for Research and Development in Geology, Geophysics, Geochemistry and Remote Sensing, 20 (1): 1 - 106. [in Greek]" type="journal article" year="1977">Tsaila-Monopolis, 1977</bibRefCitation>
from
<collectingCountry id="F35B7630FFE3A229FCB6FD07A630C5A2" box="[816,893,721,745]" name="Greece" pageId="58" pageNumber="97">Greece</collectingCountry>
).
<bibRefCitation id="EFDD4B51FFE3A229FC1FFD07A1C8C5A2" author="Parente, M. &amp; Frijia, G. &amp; Di Lucia, M." box="[921,1157,721,745]" pageId="58" pageNumber="97" pagination="255 - 259" refId="ref92641" refString="Parente, M., Frijia, G. &amp; Di Lucia, M., 2010. Comment on &quot; Sea- level control on facies architecture in the Cenomanian - Coniacian Apulian margin (Western Tethys): A record of glacio-eustatic fluctuations during the Cretaceous greenhouse? &quot; by S. Galeotti, G. Rusciadelli, M. Sprovieri, L. Lanci, A. Gaudio &amp; S. Pekar. Palaeogeography, Palaeoclimatology, Palaeoecology, 293: 255 - 259." type="journal article" year="2010">Parente et al. (2010)</bibRefCitation>
provide an unillustrated record of “
<taxonomicName id="4C4C4D23FFE3A229FC2BFD24A18EC441" authorityName="Bonet" authorityYear="1956" box="[941,1219,754,778]" class="Tubothalamea" family="Hauerinidae" genus="Nummoloculina" isUncertain="true" kingdom="Chromista" order="Miliolida" pageId="58" pageNumber="97" phylum="Foraminifera" rank="species" species="heimi">
<emphasis id="B938EAB2FFE3A229FC2BFD24A11AC441" box="[941,1111,754,778]" italics="true" pageId="58" pageNumber="97">Nummoloculina</emphasis>
cf.
<emphasis id="B938EAB2FFE3A229FB0EFD24A18EC441" box="[1160,1219,754,778]" italics="true" pageId="58" pageNumber="97">heimi</emphasis>
</taxonomicName>
” from the late Turonian.
<bibRefCitation id="EFDD4B51FFE3A229FC0FFCC5A044C460" author="Chiocchini, M. &amp; Mancinelli, A." box="[905,1289,787,811]" pageId="58" pageNumber="97" pagination="109 - 152" refId="ref76867" refString="Chiocchini, M. &amp; Mancinelli, A., 1977. Microbiostratigrafia del Mesozoico in facies di piattaforma carbonatica dei Monti Aurunci (Lazio meridionale). Studi Geol. Camerti, 3: 109 - 152." type="journal article" year="1977">Chiocchini and Mancinelli (1977)</bibRefCitation>
mention the species as having biozonal value for the Turonian of the Apennines, but perhaps like other records, this may be because of loose use of the species concept of
<taxonomicName id="4C4C4D23FFE3A229FABDFCA0A03BC4C5" authority="(Bonet)" authorityName="Bonet" authorityYear="1956" baseAuthorityName="Bonet" baseAuthorityYear="1956" box="[1339,1398,886,910]" class="Tubothalamea" family="Hauerinidae" genus="Nummoloculina" kingdom="Chromista" order="Miliolida" pageId="58" pageNumber="97" phylum="Foraminifera" rank="species" species="heimi">
<emphasis id="B938EAB2FFE3A229FABDFCA0A03BC4C5" box="[1339,1398,886,910]" italics="true" pageId="58" pageNumber="97">heimi</emphasis>
</taxonomicName>
. In subsequent papers (e.g.,
<bibRefCitation id="EFDD4B51FFE3A229FBBFFC41A062C4E4" author="Chiocchini, M. &amp; Chiocchini, R. A. &amp; Didaskalou, P. &amp; Potetti, M." box="[1081,1327,919,943]" pageId="58" pageNumber="97" pagination="5 - 170" refId="ref76906" refString="Chiocchini, M., Chiocchini, R. A., Didaskalou, P. &amp; Potetti, M., 2008. Upper Triassic, Jurassic and Cretaceous microbiostratigraphy of the carbonatic platform facies in the central-southern Latium and Abruzzi: final revision. Memorie Descrittive della Carta Geologica d'Italia, 84: 5 - 170." type="journal article" year="2008">Chiocchini et al., 2008</bibRefCitation>
), the zonal index is called “
<taxonomicName id="4C4C4D23FFE3A229FB82FC6EA00BC49B" box="[1028,1350,952,976]" class="Tubothalamea" family="Hauerinidae" genus="Nummoloculina" isUncertain="true" kingdom="Chromista" order="Miliolida" pageId="58" pageNumber="97" phylum="Foraminifera" rank="species" species="irregulari">
<emphasis id="B938EAB2FFE3A229FB82FC6EA1E3C49B" box="[1028,1198,952,976]" italics="true" pageId="58" pageNumber="97">Nummoloculina</emphasis>
cf.
<emphasis id="B938EAB2FFE3A229FB5BFC6EA00BC49B" box="[1245,1350,952,976]" italics="true" pageId="58" pageNumber="97">irregulari</emphasis>
</taxonomicName>
s”. This may explain the illustrated Turonian record of “
<taxonomicName id="4C4C4D23FFE3A229FAC7FC0CA6A1C359" authorityName="Bonet" authorityYear="1956" class="Tubothalamea" family="Hauerinidae" genus="Nummoloculina" isUncertain="true" kingdom="Chromista" order="Miliolida" pageId="58" pageNumber="97" phylum="Foraminifera" rank="species" species="heimi">
<emphasis id="B938EAB2FFE3A229FAC7FC0CA6C5C359" italics="true" pageId="58" pageNumber="97">Nummoloculina</emphasis>
cf.
<emphasis id="B938EAB2FFE3A229FC37FC2CA6A1C359" box="[945,1004,1018,1042]" italics="true" pageId="58" pageNumber="97">heimi</emphasis>
</taxonomicName>
” by
<bibRefCitation id="EFDD4B51FFE3A229FBA6FC2CA1F8C359" author="Foglia, C." box="[1056,1205,1018,1042]" pageId="58" pageNumber="97" pagination="39 - 55" refId="ref81219" refString="Foglia, C., 1992. Correlazioni microbiostratigrafiche tra Alcune successioni Cretaceo-Paleogeniche del Massiccio del Pollina (confine Calabro-Lucano). Studi Geologici Camerti, XII: 39 - 55." type="journal article" year="1992">Foglia (1992)</bibRefCitation>
.
</paragraph>
<paragraph id="8BF336A0FFE3A229FCB6FBCDA0DDC047" blockId="58.[816,1437,158,2068]" pageId="58" pageNumber="97">
Records from
<collectingCountry id="F35B7630FFE3A229FC4BFBCDA167C378" box="[973,1066,1051,1075]" name="Slovenia" pageId="58" pageNumber="97">Slovenia</collectingCountry>
and the Balkans (mainly
<collectingCountry id="F35B7630FFE3A229FAC3FBCDA0D8C378" box="[1349,1429,1051,1075]" name="Croatia" pageId="58" pageNumber="97">Croatia</collectingCountry>
) are also numerous (e.g.,
<bibRefCitation id="EFDD4B51FFE3A229FBC6FBEAA1AAC31F" author="Radoicic, R." box="[1088,1255,1084,1108]" pageId="58" pageNumber="97" pagination="195 - 199" refId="ref93884" refString="Radoicic, R., 1965. Pianella turgida n. sp. from the Cenomanian of the Outer Dinarides. Geoloski Vjesnik, 18: 195 - 199." type="journal article" year="1965">Radoičić, 1965</bibRefCitation>
, illustrated from Cenomanian strata).
<bibRefCitation id="EFDD4B51FFE3A229FB9DFB8BA1BCC33E" author="Koch, R. &amp; Buser, E. S. &amp; Bucur, L. &amp; Bucur, I. I." box="[1051,1265,1117,1141]" pageId="58" pageNumber="97" pagination="1195 - 1215" refId="ref86835" refString="Koch, R., Buser, E. S., Bucur, L. &amp; Bucur, I. I., 1998. Biostratigraphy and facies development of Mid- to Late Cretaceous strata from the Nanos mountain (Western Slovenia, High Karst). Zentralblatt fur Geologie und Palaontologie, 1 (11 - 12): 1195 - 1215." type="journal article" year="1998">Koch et al. (1998)</bibRefCitation>
from
<collectingCountry id="F35B7630FFE3A229FAB9FB8BA0D1C33E" box="[1343,1436,1117,1141]" name="Slovenia" pageId="58" pageNumber="97">Slovenia</collectingCountry>
illustrates a form which is possibly
<emphasis id="B938EAB2FFE3A229FB48FBA9A1BEC3DD" box="[1230,1267,1151,1174]" italics="true" pageId="58" pageNumber="97">P.?</emphasis>
ex. grp.
<taxonomicName id="4C4C4D23FFE3A229FAE4FBA9A0D0C3DC" authority="(Bonet)" authorityName="Bonet" authorityYear="1956" baseAuthorityName="Bonet" baseAuthorityYear="1956" box="[1378,1437,1151,1175]" class="Tubothalamea" family="Hauerinidae" genus="Nummoloculina" kingdom="Chromista" order="Miliolida" pageId="58" pageNumber="97" phylum="Foraminifera" rank="species" species="heimi">
<emphasis id="B938EAB2FFE3A229FAE4FBA9A0D0C3DC" box="[1378,1437,1151,1175]" italics="true" pageId="58" pageNumber="97">heimi</emphasis>
</taxonomicName>
from the Cenomanian Turonian and
<bibRefCitation id="EFDD4B51FFE3A229FB62FB76A0D1C3F3" author="Jez, J. &amp; Otonicar, B. &amp; Fucek, L. &amp; Ogorelec, B." box="[1252,1436,1184,1208]" pageId="58" pageNumber="97" pagination="447 - 468" refId="ref85395" refString="Jez, J., Otonicar, B., Fucek, L. &amp; Ogorelec, B., 2011. Late Cretaceous sedimentary evolution of a northern sector of the Adriatic Carbonate Platform (Matarsko Podolje, SW Slovenia). Facies, 57: 447 - 468." type="journal article" year="2011">Jez et al. (2011)</bibRefCitation>
records unillustrated material from the late Cenomanian. Croatian records are more numerous with illustrated records:
<bibRefCitation id="EFDD4B51FFE3A229FCF0FAD5A1D7C250" author="Husinec A. &amp; Sokac, B." box="[886,1178,1283,1307]" pageId="58" pageNumber="97" pagination="418 - 441" refId="ref84611" refString="Husinec A. &amp; Sokac, B., 2006. Early Cretaceous benthic associations (foraminifera and calcareous algae) of a shallow tropical-water platform environment (Mljet Island, southern Croatia). Cretaceous Research, 27: 418 - 441." type="journal article" year="2006">Husinec &amp; Sokač (2006)</bibRefCitation>
(illustrated as “
<taxonomicName id="4C4C4D23FFE3A229FAE6FAD2A157C277" class="Tubothalamea" family="Miliolidae" genus="Pseudonummoloculina" kingdom="Chromista" order="Miliolida" pageId="58" pageNumber="97" phylum="Foraminifera" rank="species" species="undetermined">
<emphasis id="B938EAB2FFE3A229FAE6FAD2A6BCC277" italics="true" pageId="58" pageNumber="97">Pseudonummoloculina</emphasis>
sp.
</taxonomicName>
” but mentioned as
<taxonomicName id="4C4C4D23FFE3A229FB71FAF2A69DC216" authority="(Bonet, 1956)" class="Tubothalamea" family="Miliolidae" genus="Pseudonummoloculina" kingdom="Chromista" order="Miliolida" pageId="58" pageNumber="97" phylum="Foraminifera" rank="species" species="heimi">
<emphasis id="B938EAB2FFE3A229FB71FAF2A69DC216" italics="true" pageId="58" pageNumber="97">Pseudonummoloculina heimi</emphasis>
</taxonomicName>
in the text and range charts - Albian),
<bibRefCitation id="EFDD4B51FFE3A229FCB6FAB0A156C235" author="Tesovic, B. C. &amp; Glumac, B. &amp; Buckovic, D." box="[816,1051,1382,1406]" pageId="58" pageNumber="97" refId="ref100739" refString="Tesovic, B. C., Glumac, B. &amp; Buckovic, D., 2011. Integrated biostratigraphy and carbon isotope stratigraphy of the Lower Cretaceous (Barremian to Albian) Adriatic-Dinaridic carbonate platform deposits in Istria, Croatia. Cretaceous Research, doi: 10.1016 / j. cretres. 2010.12.011" type="book" year="2011">Tešović et al. (2011)</bibRefCitation>
from the early late Albian, and
<bibRefCitation id="EFDD4B51FFE3A229FCB6FA51A16DC2D4" author="Velic, I. &amp; Sokac, B." box="[816,1056,1415,1439]" pageId="58" pageNumber="97" pagination="185 - 190" refId="ref101388" refString="Velic, I. &amp; Sokac, B., 1979. Biostratigrafska istrazivanja donje krede Vanjskih Dinarida (II) Gornji alb otoka Korcule Geoloski Vjesnik, 31: 185 - 190." type="journal article" year="1979">Velić &amp; Sokač (1979)</bibRefCitation>
undated. The illustrated records of
<bibRefCitation id="EFDD4B51FFE3A229FCB6FA7FA69CC28A" author="Ritossa, A." box="[816,977,1449,1473]" pageId="58" pageNumber="97" refId="ref95103" refString="Ritossa, A., 2018. Donjokredne emerzijske pojave u kamenolomu Selina kraj Svetog Lovreca. Unpublished MSc thesis, University of Zagreb." type="book" year="2018">Ritossa (2018)</bibRefCitation>
, also undated, and
<bibRefCitation id="EFDD4B51FFE3A229FB26FA7FA02BC28A" author="Brcic, V. &amp; Glumac, B. &amp; Brlek, M. &amp; Fucek, L. &amp; Sparica Miko, M." box="[1184,1382,1449,1473]" pageId="58" pageNumber="97" refId="ref75508" refString="Brcic, V., Glumac, B., Brlek, M., Fucek, L. &amp; Sparica Miko, M., 2021. Cenomanian - Turonian oceanic anoxic event (OAE 2) imprint on the northwestern part of the Adriatic Carbonate Platform and a coeval intra-platform basin (Istria and Premuda Island, Croatia)." type="journal volume" year="2021">Brčić et al. (2021)</bibRefCitation>
(late Cenomanian) cannot be verified as this species. Additional records unconfirmed by illustration include
<bibRefCitation id="EFDD4B51FFE3A229FACFFA3DA6EEC16F" author="Brcic, V. &amp; Glumac, B. &amp; Fucek, L. &amp; Grizelj, A. &amp; Horvat, M. &amp; Posilovic, H. &amp; Misur, I." pageId="58" pageNumber="97" pagination="1 - 20" refId="ref75432" refString="Brcic, V., Glumac, B., Fucek, L., Grizelj, A., Horvat, M., Posilovic, H. &amp; Misur, I., 2017. The Cenomanian - Turonian boundary in the northwestern part of the Adriatic Carbonate Platform (Cicarija Mtn., Istria, Croatia): characteristics and implications. Facies, 63 (17): 1 - 20." type="journal article" year="2017">Brčić et al. (2017)</bibRefCitation>
;
<bibRefCitation id="EFDD4B51FFE3A229FC30F9DAA1EBC16F" author="Husinec, A. &amp; Velic, I. &amp; Fucek, L. &amp; Vlahovic, I. &amp; Maticec, D. &amp; Ostric, N. &amp; Korbar, T." box="[950,1190,1548,1572]" pageId="58" pageNumber="97" pagination="155 - 171" refId="ref84655" refString="Husinec, A., Velic, I., Fucek, L., Vlahovic, I., Maticec, D., Ostric, N. &amp; Korbar, T., 2000. Mid Cretaceous orbitolinid (Foraminiferida) record from the islands of Cres and Losinj (Croatia) and its regional stratigraphic correlation. Cretaceous Research, 21: 155 - 171." type="journal article" year="2000">Husinec et al. (2000</bibRefCitation>
,
<bibRefCitation id="EFDD4B51FFE3A229FB3EF9DAA1BFC16F" author="Husinec, A. &amp; Velic, I. &amp; Sokac, B." box="[1208,1266,1548,1572]" pageId="58" pageNumber="97" pagination="153 - 170" refId="ref84724" refString="Husinec, A., Velic, I. &amp; Sokac, B., 2009. Diversity patterns in Mid-Cretaceous benthic foraminifers and Dasycladacean algae of the southern part of the Mesozoic Adriatic Platform, Croatia. Geologic Problem Solving with Microfossils, SEPM Special Publication, 93: 153 - 170." type="journal article" year="2009">2009</bibRefCitation>
);
<bibRefCitation id="EFDD4B51FFE3A229FA8BF9DAA636C10E" author="Tisljar, J. &amp; Vlahovic, I. &amp; Velic &amp; Maticec, D. &amp; Robson, Z. &amp; Robson, J." pageId="58" pageNumber="97" pagination="137 - 152" refId="ref100843" refString="Tisljar, J., Vlahovic, I., Velic, Maticec, D., Robson, Z. &amp; Robson, J., 1998. Carbonate facies evolution from the late Albian to middle Cenomanian in southern Istria (Croatia): Influence of synsedimentary tectonics and extensive organic carbonate production. Facies, 38: 137 - 152." type="journal article" year="1998">Tišljar et al. (1998)</bibRefCitation>
;
<bibRefCitation id="EFDD4B51FFE3A229FC08F9FBA150C10E" author="Velic, I." box="[910,1053,1581,1605]" pageId="58" pageNumber="97" pagination="1 - 113" refId="ref101499" refString="Velic, I., 2007. Stratigraphy and Palaeobiogeography of Mesozoic Benthic Foraminifera of the Karst Dinarides (SE Europe). Geologia Croatica, 60 (1): 1 - 113." type="journal article" year="2007">Velić (2007)</bibRefCitation>
;
<bibRefCitation id="EFDD4B51FFE3A229FBB3F9FBA02EC10E" author="Korbar, T. &amp; Husinec, A." box="[1077,1379,1581,1605]" pageId="58" pageNumber="97" pagination="173 - 185" refId="ref87013" refString="Korbar, T. &amp; Husinec, A., 2003. Biostratigraphy of Turonian to (?) Coniacian Platform Carbonates: A Case Study from the Island of Cres (Northern Adriatic, Croatia). Geologia Croatica, 56 (2): 173 - 185." type="journal article" year="2003">Korbar &amp; Husinec (2003)</bibRefCitation>
, and
<bibRefCitation id="EFDD4B51FFE3A229FCB6F998A10AC12D" author="Velic, I. &amp; Vlahovic, I." box="[816,1095,1614,1638]" pageId="58" pageNumber="97" pagination="25 - 43" refId="ref101422" refString="Velic, I. &amp; Vlahovic, I., 1994. Foraminiferal assemblages in the Cenomanian of the Buzet-Savudrija area (northwestern Istria, Croatia). Geologia Croatica, 47 (1): 25 - 43." type="journal article" year="1994">Velić &amp; Vlahović (1994)</bibRefCitation>
. Assigned ages range from the early Albian to the early Campanian (e.g.,
<bibRefCitation id="EFDD4B51FFE3A229FA8BF9B9A0DDC1CC" author="Velic, I." box="[1293,1424,1647,1671]" pageId="58" pageNumber="97" pagination="1 - 113" refId="ref101499" refString="Velic, I., 2007. Stratigraphy and Palaeobiogeography of Mesozoic Benthic Foraminifera of the Karst Dinarides (SE Europe). Geologia Croatica, 60 (1): 1 - 113." type="journal article" year="2007">Velić, 2007</bibRefCitation>
). A record from the late Turonian Coniacian of
<collectingCountry id="F35B7630FFE3A229FAC8F946A0D1C1E3" box="[1358,1436,1680,1704]" name="Croatia" pageId="58" pageNumber="97">Croatia</collectingCountry>
(
<bibRefCitation id="EFDD4B51FFE3A229FCBEF967A6B0C182" author="Gusic, I. &amp; Jelaska, V. &amp; Velic, I." box="[824,1021,1713,1737]" pageId="58" pageNumber="97" pagination="447 - 456" refId="ref82704" refString="Gusic, I., Jelaska, V. &amp; Velic, I., 1988. Foraminiferal assemblages, facies, and environments in the Upper Cretaceous of the Island of Brac, Yugoslavia. Benthos' 86. Symposium international sur les Foraminiferes benthiques. Revue de Paleobiologie: 447 - 456." type="book chapter" year="1988">Gušić et al., 1988</bibRefCitation>
) is a nummoloculinid, but difficult to assign to a species. A single record from
<collectingCountry id="F35B7630FFE3A229FA9CF905A0D1C1A0" box="[1306,1436,1747,1771]" name="Serbia" pageId="58" pageNumber="97">Montenegro</collectingCountry>
(
<bibRefCitation id="EFDD4B51FFE3A229FCBFF922A69AC047" author="Bozovic, D. M." box="[825,983,1780,1804]" pageId="58" pageNumber="97" refId="ref75249" refString="Bozovic, D. M., 2016. Minerageny and potentiality of carbonate raw materials of mining area of Bjelopavlici (Montenegro). Unpublished PhD thesis, University of Belgrade." type="book" year="2016">Božović, 2016</bibRefCitation>
) is unillustrated and has no assigned age.
</paragraph>
<paragraph id="8BF336A0FFE3A228FCB6F8C3A7ADC670" blockId="58.[816,1437,158,2068]" lastBlockId="59.[151,772,158,2068]" lastPageId="59" lastPageNumber="98" pageId="58" pageNumber="97">
Most records from
<collectingCountry id="F35B7630FFE3A229FB85F8C3A103C066" box="[1027,1102,1813,1837]" name="Greece" pageId="58" pageNumber="97">Greece</collectingCountry>
(e.g.,
<bibRefCitation id="EFDD4B51FFE3A229FB13F8C3A06FC066" author="Fleury, J. - J." box="[1173,1314,1813,1837]" pageId="58" pageNumber="97" pagination="181 - 194" refId="ref81053" refString="Fleury, J. - J., 1971. Le Cenomanien a foraminiferes benthoniques du Massif du Varassova (zone de Gavrovo, Akarnanie, Grece continentale). Revue de Micropaleontologie, 14 (3): 181 - 194." type="journal article" year="1971">Fleury, 1971</bibRefCitation>
;
<bibRefCitation id="EFDD4B51FFE3A229FAA9F8C3A625C005" author="Decrouez, D." pageId="58" pageNumber="97" pagination="521 - 523" refId="ref79218" refString="Decrouez, D., 1976. Position stratigraphique de Pseudolituonella? guerneti nov. sp. en Eubee (Grece). Geobios, 9 (4): 521 - 523." type="journal article" year="1976">Decrouez, 1976</bibRefCitation>
, 1978; early late Cenomanian) are unillustrated, but that of
<bibRefCitation id="EFDD4B51FFE3A229FC20F881A1C9C024" author="Charvet, J. &amp; Decrouez, D. &amp; Polsak, A." box="[934,1156,1879,1903]" pageId="58" pageNumber="97" pagination="247 - 258" refId="ref76438" refString="Charvet, J., Decrouez, D. &amp; Polsak, A., 1976. Le Cretace du Foniakos (Argolide, Grece): examen paleontologique, repercussions stratigraphiques, paleogeographiques et tectoniques. Archives des Sciences, 29 (3): 247 - 258." type="journal article" year="1976">Charvet et al. (1976)</bibRefCitation>
is, but might be
<taxonomicName id="4C4C4D23FFE3A229FABFF88EA132C0DB" authority="sensu Chiocchini et al." authorityName="sensu Chiocchini et al." authorityYear="2012" baseAuthorityName="Philippson" baseAuthorityYear="1887" class="Tubothalamea" family="Miliolidae" genus="Pseudonummoloculina" kingdom="Chromista" order="Miliolida" pageId="58" pageNumber="97" phylum="Foraminifera" rank="species" species="regularis">
<emphasis id="B938EAB2FFE3A229FABFF88EA01CC024" box="[1337,1361,1880,1903]" italics="true" pageId="58" pageNumber="97">P.</emphasis>
?
<emphasis id="B938EAB2FFE3A229FAE5F88EA62CC0DB" italics="true" pageId="58" pageNumber="97">regularis</emphasis>
sensu Chiocchini et al.
</taxonomicName>
or
<taxonomicName id="4C4C4D23FFE3A229FB33F8AEA6FCC0FA" authorityName="Piuz &amp; Vicedo" authorityYear="2020" class="Tubothalamea" family="Hauerinidae" genus="Nummoloculinodonta" higherTaxonomySource="CoL,GBIF" kingdom="Chromista" order="Miliolida" pageId="58" pageNumber="97" phylum="Foraminifera" rank="species" species="akhdarensis">
<emphasis id="B938EAB2FFE3A229FB33F8AEA6FCC0FA" italics="true" pageId="58" pageNumber="97">Nummoloculinodonta akhdarensis</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFDD4B51FFE3A229FC46F84FA180C0FA" author="Tsaila-Monopolis, S." box="[960,1229,1945,1969]" pageId="58" pageNumber="97" pagination="1 - 106" refId="ref101018" refString="Tsaila-Monopolis, S., 1977. Micropaleontological and stratigraphical study of the Tripolitza (Gavrovo) zone in the Peloponnesus. National Institute for Research and Development in Geology, Geophysics, Geochemistry and Remote Sensing, 20 (1): 1 - 106. [in Greek]" type="journal article" year="1977">Tsaila-Monopolis (1977)</bibRefCitation>
illustrated the species from the “Cenomanian - Turonian” and “Senonian” of
<collectingCountry id="F35B7630FFE3A229FCD4F80DA6ECC0B8" box="[850,929,2011,2035]" name="Greece" pageId="58" pageNumber="97">Greece</collectingCountry>
. Some illustrations are more compatible with
<taxonomicName id="4C4C4D23FFE3A229FCB6F82BA064CF5F" authority="Chiocchini et al. (2012)" authorityName="Chiocchini" authorityYear="2012" box="[816,1321,2044,2068]" class="Tubothalamea" family="Miliolidae" genus="Pseudonummoloculina" kingdom="Chromista" order="Miliolida" pageId="58" pageNumber="97" phylum="Foraminifera" rank="subSpecies" species="regularis" subSpecies="sensu">
<emphasis id="B938EAB2FFE3A229FCB6F82BA605CF5F" box="[816,840,2045,2068]" italics="true" pageId="58" pageNumber="97">P.</emphasis>
?
<emphasis id="B938EAB2FFE3A229FCE6F82AA147CF5F" box="[864,1034,2044,2068]" italics="true" pageId="58" pageNumber="97">regularis sensu</emphasis>
<bibRefCitation id="EFDD4B51FFE3A229FB91F82AA064CF5F" author="Chiocchini, M. &amp; Pampaloni, M. L. &amp; Pichezzi, R. M." box="[1047,1321,2044,2068]" pageId="58" pageNumber="97" refId="ref77109" refString="Chiocchini, M., Pampaloni, M. L., &amp; Pichezzi, R. M., 2012. Microfacies e microfossili delle successioni carbonatiche mesozoiche del Lazio e dell &quot; Abruzzo (Italia centrale) - Cretacico. Memorie per servire alla descrizione della Carta Geologica d'Italia. ISPRA, Servizio Geologico d'Italia, Dipartimento Difesa del Suolo, Rome." type="book" year="2012">Chiocchini et al. (2012)</bibRefCitation>
</taxonomicName>
. Another record from
<collectingCountry id="F35B7630FFE2A228FEA0FF48A43CC7FD" box="[294,369,158,182]" name="Greece" pageId="59" pageNumber="98">Greece</collectingCountry>
(Zambetakis-Lekkas, 2006) is unillustrated and is said to range from the late Cenomanian to the Maastrichtian, although that seems unlikely.
<bibRefCitation id="EFDD4B51FFE2A228FD4FFF37A488C652" author="Zambetakis-Lekkas, A. &amp; Pomoni-Papaioannou, F. &amp; Alexopoulos, A." pageId="59" pageNumber="98" pagination="7 - 18" refId="ref103036" refString="Zambetakis-Lekkas, A., Pomoni-Papaioannou, F. &amp; Alexopoulos, A., 2006. New stratigraphic and palaeogeographic data from the Mesozoic strata of the Tripolitza platform in Central Crete. Evidence of subaerial exposures during Albian-Early Cenomanian. Hellenic Journal of Geosciences, 42: 7 - 18." type="journal article" year="2006">Zambetakis-Lekkas et al. (2006)</bibRefCitation>
also records the species (unillustrated) from
<collectingRegion id="4988F842FFE2A228FECAFEF4A4CBC671" box="[332,390,290,314]" country="Greece" name="Kriti" pageId="59" pageNumber="98">Crete</collectingRegion>
and assigns the same age range.
</paragraph>
<paragraph id="8BF336A0FFE2A228FF11FE95A733C461" blockId="59.[151,772,158,2068]" pageId="59" pageNumber="98">
North Africa records include from
<collectingCountry id="F35B7630FFE2A228FDA3FE95A738C610" box="[549,629,323,347]" name="Tunisia" pageId="59" pageNumber="98">Tunisia</collectingCountry>
(
<bibRefCitation id="EFDD4B51FFE2A228FD0EFE95A5B8C637" author="Bismuth, H. &amp; Bonnefous, J. &amp; Dufaure, P." pageId="59" pageNumber="98" refId="ref74651" refString="Bismuth, H., Bonnefous, J. &amp; Dufaure, P., 1967. Mesozoic microfacies of Tunisia. Petroloration Exploration Society of Libya, Guide-book to the Geology and History of Tunisia: 159 - 214, Tripoli." type="book" year="1967">Bismuth et al., 1967</bibRefCitation>
;
<bibRefCitation id="EFDD4B51FFE2A228FE80FEB2A4F2C637" author="Saidi, F. &amp; Ben Ismail, M. H. &amp; Negra, M. E. H. &amp; M'Rabet, A." box="[262,447,356,380]" pageId="59" pageNumber="98" pagination="17 - 41" refId="ref95842" refString="Saidi, F., Ben Ismail, M. H., Negra, M. E. H. &amp; M'Rabet, A., 1995. La plate forme carbonatee cenomanienne de Tunisie du Centre-Ouest. Facies, corteges argileux, paleoenvironnements et stratigraphie sequentielle. Geologie Mediterraneenne, 22 (1): 17 - 41." type="journal article" year="1995">Saïdi et al., 1995</bibRefCitation>
and
<bibRefCitation id="EFDD4B51FFE2A228FE71FEB2A7F7C637" author="Touir, J. &amp; Mechi, C. &amp; Ali, H. H." box="[503,698,356,380]" pageId="59" pageNumber="98" pagination="527 - 541" refId="ref100936" refString="Touir, J., Mechi, C. &amp; Ali, H. H., 2017. Changes in carbonate sedimentation and faunal assemblages in the Tunisian carbonate platform around the Cenomanian-Turonian boundary. Journal of African Earth Sciences, 129: 527 - 541." type="journal article" year="2017">Touir et al., 2017</bibRefCitation>
). Only the former record is confirmed by illustration and may include
<taxonomicName id="4C4C4D23FFE2A228FF76FE7EA79BC6F4" authority="Chiocchini et al. (2012)" authorityName="Chiocchini" authorityYear="2012" box="[240,726,423,447]" class="Tubothalamea" family="Miliolidae" genus="Pseudonummoloculina" kingdom="Chromista" order="Miliolida" pageId="59" pageNumber="98" phylum="Foraminifera" rank="subSpecies" species="regularis" subSpecies="sensu">
<emphasis id="B938EAB2FFE2A228FF76FE7EA44DC6F4" box="[240,256,424,447]" italics="true" pageId="59" pageNumber="98">P</emphasis>
.?
<emphasis id="B938EAB2FFE2A228FE9BFE71A489C6F4" box="[285,452,423,447]" italics="true" pageId="59" pageNumber="98">regularis sensu</emphasis>
<bibRefCitation id="EFDD4B51FFE2A228FE48FE71A79BC6F4" author="Chiocchini, M. &amp; Pampaloni, M. L. &amp; Pichezzi, R. M." box="[462,726,423,447]" pageId="59" pageNumber="98" refId="ref77109" refString="Chiocchini, M., Pampaloni, M. L., &amp; Pichezzi, R. M., 2012. Microfacies e microfossili delle successioni carbonatiche mesozoiche del Lazio e dell &quot; Abruzzo (Italia centrale) - Cretacico. Memorie per servire alla descrizione della Carta Geologica d'Italia. ISPRA, Servizio Geologico d'Italia, Dipartimento Difesa del Suolo, Rome." type="book" year="2012">Chiocchini et al. (2012)</bibRefCitation>
</taxonomicName>
but all authors assign a Cenomanian age. By contrast,
<bibRefCitation id="EFDD4B51FFE2A228FD30FE1EA452C54A" author="Luning, S. &amp; Grafe, K. - U. &amp; Bosence, D. &amp; Luciani, V. &amp; Craig, J." pageId="59" pageNumber="98" pagination="721 - 731" refId="ref88114" refString="Luning, S., Grafe, K. - U., Bosence, D., Luciani, V. &amp; Craig, J., 2000. Discovery of marine Late Cretaceous carbonates and evaporites in the Kufra Basin (Libya) redefines the southern limit of the Late Cretaceous transgression. Cretaceous Research, 21: 721 - 731." type="journal article" year="2000">Lüning et al. (2000)</bibRefCitation>
report the species from the
<collectingRegion id="4988F842FFE2A228FDD8FE3FA7D1C54A" box="[606,668,489,513]" country="Libya" name="Al Kufrah" pageId="59" pageNumber="98">Kufra</collectingRegion>
Basin of
<collectingCountry id="F35B7630FFE2A228FF11FDDCA598C569" box="[151,213,522,546]" name="Libya" pageId="59" pageNumber="98">Libya</collectingCountry>
and assign a Campanian? Maastrichtian age, although they provide no illustration but note that “
<emphasis id="B938EAB2FFE2A228FD28FDFDA59FC441" italics="true" pageId="59" pageNumber="98">
the present specimen is similar to those described in
<bibRefCitation id="EFDD4B51FFE2A228FD1DFD9BA45BC5CE" author="Hottinger, L. &amp; Drobne, K. &amp; Caus, E." pageId="59" pageNumber="98" pagination="99 - 134" refId="ref84439" refString="Hottinger, L., Drobne, K. &amp; Caus, E., 1989. Late Cretaceous, larger complex Miliolids (foraminifera) endemic in the Pyrenean faunal province. Facies, 21: 99 - 134." type="journal article" year="1989">Hottinger et al. (1989</bibRefCitation>
, pl. 22, fig. 6) and by
<bibRefCitation id="EFDD4B51FFE2A228FD83FDBBA7D2C5CE" author="Calvez, H." box="[517,671,621,645]" pageId="59" pageNumber="98" pagination="391 - 399" refId="ref76196" refString="Calvez, H., 1988. Pseudonummoloculina aurigerica n. gen. n. sp. et Dobrogelina? angulata n. sp., new foraminifera from Albian limestones of the French and Spanish Pyrenees. Revue de Paleobiologie, 2 (Benthos ' 86): 391 - 399." type="journal article" year="1988">Calvez (1988)</bibRefCitation>
; the only difference is the smaller size compared to the material described by Calvez. The specimen figured by
<bibRefCitation id="EFDD4B51FFE2A228FD15FD66A595C5A2" author="De Castro, P." pageId="59" pageNumber="98" pagination="109 - 124" refId="ref78786" refString="De Castro, P., 1987. On some foraminifera and algae in Apennine Upper Cretaceous and Paleocene: Memorie della Societa Geologica Italiana, 40: 109 - 124." type="journal article" year="1987">De Castro (1987</bibRefCitation>
, fig. 3) also shows all features of the present material.”
</emphasis>
In this respect it is worth noting that
<bibRefCitation id="EFDD4B51FFE2A228FDECFD24A64EC441" author="Calvez, H." box="[618,771,754,778]" pageId="59" pageNumber="98" pagination="391 - 399" refId="ref76196" refString="Calvez, H., 1988. Pseudonummoloculina aurigerica n. gen. n. sp. et Dobrogelina? angulata n. sp., new foraminifera from Albian limestones of the French and Spanish Pyrenees. Revue de Paleobiologie, 2 (Benthos ' 86): 391 - 399." type="journal article" year="1988">Calvez (1988)</bibRefCitation>
did not illustrate
<taxonomicName id="4C4C4D23FFE2A228FECBFCC5A4FEC460" baseAuthorityName="Bonet" baseAuthorityYear="1956" box="[333,435,787,811]" class="Tubothalamea" family="Miliolidae" genus="Pseudonummoloculina" kingdom="Chromista" order="Miliolida" pageId="59" pageNumber="98" phylum="Foraminifera" rank="species" species="heimi">
<emphasis id="B938EAB2FFE2A228FECBFCC5A4FEC460" box="[333,435,787,811]" italics="true" pageId="59" pageNumber="98">P.? heimi</emphasis>
</taxonomicName>
, but
<taxonomicName id="4C4C4D23FFE2A228FE6FFCC2A73AC461" authorityName="Calvez" authorityYear="1988" box="[489,631,787,811]" class="Tubothalamea" family="Miliolidae" genus="Pseudonummoloculina" kingdom="Chromista" order="Miliolida" pageId="59" pageNumber="98" phylum="Foraminifera" rank="species" species="aurigerica">
<emphasis id="B938EAB2FFE2A228FE6FFCC2A73AC461" box="[489,631,787,811]" italics="true" pageId="59" pageNumber="98">P. aurigerica</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BF336A0FFE2A228FF11FCE3A7FFC3DC" blockId="59.[151,772,158,2068]" pageId="59" pageNumber="98">
Numerous illustrated records occur from
<collectingCountry id="F35B7630FFE2A228FDD7FCE2A7D3C407" box="[593,670,820,844]" name="Turkey" pageId="59" pageNumber="98">Turkey</collectingCountry>
of which those of
<bibRefCitation id="EFDD4B51FFE2A228FF7FFC83A4FBC426" author="Sari, B. &amp; Tasli, K. &amp; Ozer, S." box="[249,438,853,877]" pageId="59" pageNumber="98" pagination="393 - 425" refId="ref96523" refString="Sari, B., Tasli, K. &amp; Ozer, S., 2009. Benthonic Foraminiferal Biostratigraphy of the Upper Cretaceous (Middle Cenomanian-Coniacian) Sequences of the Bey Daglari Carbonate Platform, Western Taurides, Turkey. Turkish Journal of Earth Sciences, 18: 393 - 425." type="journal article" year="2009">Sari et al. (2009)</bibRefCitation>
and Solak et al. (2020, 2021) are confirmed by illustration. The records of
<bibRefCitation id="EFDD4B51FFE2A228FD1AFCA0A471C4E4" author="Ozkan, R. &amp; Altiner, D." pageId="59" pageNumber="98" pagination="153 - 178" refId="ref92491" refString="Ozkan, R. &amp; Altiner, D., 2019. The Cretaceous Mardin Group carbonates in southeast Turkey: lithostratigraphy, foraminiferal biostratigraphy, microfacies and sequence stratigraphic evolution. Cretaceous Research, 98: 153 - 178." type="journal article" year="2019">Ozkan &amp; Altiner (2019)</bibRefCitation>
, Solak (2021) and
<bibRefCitation id="EFDD4B51FFE2A228FDA0FC41A7B1C4E4" author="Solak, C. &amp; Tasli, K. &amp; Koc, H." box="[550,764,919,943]" pageId="59" pageNumber="98" pagination="43 - 63" refId="ref99916" refString="Solak, C., Tasli, K. &amp; Koc, H., 2017. Biostratigraphy and facies analysis of the Upper Cretaceous-Danian? platform carbonate succession in the Kuyucak area, western Central Taurides, S Turkey. Cretaceous Research, 79: 43 - 63." type="journal article" year="2017">Solak et al. (2017</bibRefCitation>
,
<bibRefCitation id="EFDD4B51FFE2A228FF11FC6EA583C49B" author="Solak, C. &amp; Tasli, K. &amp; Ozer, S &amp; Koc, H." box="[151,206,952,976]" pageId="59" pageNumber="98" pagination="67 - 83" refId="ref100126" refString="Solak, C., Tasli, K., Ozer, S &amp; Koc, H., 2019. The Madenli (Central Taurides) Upper Cretaceous platform carbonate succession: benthic foraminiferal biostratigraphy and platform evolution. Geobios, 52: 67 - 83." type="journal article" year="2019">2019</bibRefCitation>
) are also illustrated but the specimens are only
<emphasis id="B938EAB2FFE2A228FD53FC6FA584C4BA" italics="true" pageId="59" pageNumber="98">possibly</emphasis>
of this species. Most records are assigned a middle late Cenomanian age although
<bibRefCitation id="EFDD4B51FFE2A228FE75FC2CA788C359" author="Solak, C. &amp; Tasli, K. &amp; Koc, H." box="[499,709,1018,1042]" pageId="59" pageNumber="98" pagination="673 - 693" refId="ref100030" refString="Solak, C., Tasli, K. &amp; Koc, H., 2021. Benthic foraminifera from the Albian shallow-marine limestones in the Geyik Dagi area (Central Taurides), southern Turkey. Journal of Paleontology, 95 (4): 673 - 693." type="journal article" year="2021">Solak et al. (2021)</bibRefCitation>
indicates an age as old as late Albian and
<bibRefCitation id="EFDD4B51FFE2A228FDA8FBCDA7AFC378" author="Sari, B. &amp; Tasli, K. &amp; Ozer, S." box="[558,738,1051,1075]" pageId="59" pageNumber="98" pagination="393 - 425" refId="ref96523" refString="Sari, B., Tasli, K. &amp; Ozer, S., 2009. Benthonic Foraminiferal Biostratigraphy of the Upper Cretaceous (Middle Cenomanian-Coniacian) Sequences of the Bey Daglari Carbonate Platform, Western Taurides, Turkey. Turkish Journal of Earth Sciences, 18: 393 - 425." type="journal article" year="2009">Sari et al. (2009)</bibRefCitation>
an age as young as Coniacian. The record of
<bibRefCitation id="EFDD4B51FFE2A228FDF7FBEAA455C33E" author="Ozkan, R. &amp; Altiner, D." pageId="59" pageNumber="98" pagination="153 - 178" refId="ref92491" refString="Ozkan, R. &amp; Altiner, D., 2019. The Cretaceous Mardin Group carbonates in southeast Turkey: lithostratigraphy, foraminiferal biostratigraphy, microfacies and sequence stratigraphic evolution. Cretaceous Research, 98: 153 - 178." type="journal article" year="2019">Ozkan &amp; Altiner (2019)</bibRefCitation>
is thought to be from the early Cenomanian.
<bibRefCitation id="EFDD4B51FFE2A228FF11FBA9A41CC3DC" author="Sinanoglu, D." box="[151,337,1151,1175]" pageId="59" pageNumber="98" pagination="267 - 293" refId="ref99477" refString="Sinanoglu, D., 2021. Systematic and biostratigraphic evaluation of the Late Cretaceous benthic foraminifera assemblages of southeastern Batman. Bulletin of Mineral Research and Exploration, 165: 267 - 293." type="journal article" year="2021">Sinanoglu (2021)</bibRefCitation>
provides an unillustrated record.
</paragraph>
<paragraph id="8BF336A0FFE2A228FF11FB76A59AC16F" blockId="59.[151,772,158,2068]" pageId="59" pageNumber="98">
In the eastern Mediterranean area
<emphasis id="B938EAB2FFE2A228FD97FB76A778C3FC" box="[529,565,1184,1207]" italics="true" pageId="59" pageNumber="98">P.?</emphasis>
ex. grp.
<taxonomicName id="4C4C4D23FFE2A228FD1DFB76A79BC3F3" authority="(Bonet)" authorityName="Bonet" authorityYear="1956" baseAuthorityName="Bonet" baseAuthorityYear="1956" box="[667,726,1184,1208]" class="Tubothalamea" family="Miliolidae" genus="Pseudonummoloculina" kingdom="Chromista" order="Miliolida" pageId="59" pageNumber="98" phylum="Foraminifera" rank="species" species="heimi">
<emphasis id="B938EAB2FFE2A228FD1DFB76A79BC3F3" box="[667,726,1184,1208]" italics="true" pageId="59" pageNumber="98">heimi</emphasis>
</taxonomicName>
has been reported from
<collectingCountry id="F35B7630FFE2A228FEF5FB17A4E3C392" box="[371,430,1217,1241]" name="Syria" pageId="59" pageNumber="98">Syria</collectingCountry>
,
<collectingCountry id="F35B7630FFE2A228FE3AFB17A754C392" box="[444,537,1217,1241]" name="Lebanon" pageId="59" pageNumber="98">Lebanon</collectingCountry>
, and
<collectingCountry id="F35B7630FFE2A228FDDCFB17A7D8C392" box="[602,661,1217,1241]" name="Israel" pageId="59" pageNumber="98">Israel</collectingCountry>
, although confirmed by illustration only from the first two (i.e.,
<bibRefCitation id="EFDD4B51FFE2A228FF11FAD5A4D3C250" author="Ghanem, H. &amp; Kuss, J." box="[151,414,1283,1307]" pageId="59" pageNumber="98" pagination="85 - 132" refId="ref82188" refString="Ghanem, H. &amp; Kuss, J., 2013. Stratigraphic control of the Aptian - Early Turonian sequences of the Levant Platform, Coastal Range, northwest Syria. GeoArabia, 18 (4): 85 - 132." type="journal article" year="2013">Ghanem &amp; Kuss, 2013</bibRefCitation>
, and
<bibRefCitation id="EFDD4B51FFE2A228FE66FAD5A7E1C250" author="Saint-Marc, P." box="[480,684,1283,1307]" pageId="59" pageNumber="98" pagination="1 - 342" refId="ref96009" refString="Saint-Marc, P., 1974 a. Etude stratigraphique et micropaleontologique de l'Albien, du Cenomanien et du Turonien du Liban. Notes et Memoires sur le Moyen- Orient, XIII, Museum National d'Histoire Naturelle, Paris, pp. 1 - 342." type="book chapter" year="1974">
Saint-Marc, 1974
<bibRefCitation id="EFDD4B51FFE2A228FD26FAD2A7E1C250" author="Arnaud, A. &amp; Berthout &amp; Brunt, L. &amp; Cherchi, A. &amp; Chiocchini, M. &amp; De Castro, P." box="[672,684,1284,1307]" pageId="59" pageNumber="98" pagination="383 - 393" refId="ref72126" refString="Arnaud, A., Berthout, Brunt, L., Cherchi, A., Chiocchini, M., De Castro, P. &amp; 9 others, 1981. Tableau de repartition stratigraphique des grands foraminiferes caracteristiques du Cretace Moyen de la region Mediterraneenne. Cretaceous Research, 2: 383 - 393." type="journal article" year="1981">a</bibRefCitation>
</bibRefCitation>
, 1981). Note the illustrated records of
<bibRefCitation id="EFDD4B51FFE2A228FE6AFAF2A795C277" author="Ghanem, H. &amp; Mouty, M. &amp; Kuss, J." box="[492,728,1316,1340]" pageId="59" pageNumber="98" pagination="155 - 184" refId="ref82231" refString="Ghanem, H., Mouty, M. &amp; Kuss, J., 2012. Biostratigraphy and carbon-isotope stratigraphy of the uppermost Aptian to Upper Cenomanian strata of the South Palmyrides, Syria. GeoArabia, 17 (2): 155 - 184." type="journal article" year="2012">Ghanem et al. (2012)</bibRefCitation>
are insufficient to confirm their identity. Unconfirmed Israeli records are from
<bibRefCitation id="EFDD4B51FFE2A228FED7FAB0A4B7C235" author="Hamaoui, M." box="[337,506,1382,1406]" pageId="59" pageNumber="98" pagination="1 - 27" refId="ref83150" refString="Hamaoui, M., 1965. Biostratigraphy of the Cenomanian type Hazera Formation. Geological Survey of Israel, stratigraphic section no. 2 b, pp. 1 - 27." type="journal article" year="1965">Hamaoui (1965</bibRefCitation>
,
<bibRefCitation id="EFDD4B51FFE2A228FD81FAB0A70CC235" author="Hamaoui, M." box="[519,577,1382,1406]" pageId="59" pageNumber="98" refId="ref83183" refString="Hamaoui, M., 1966. Microfossils from Cenomanian sections in the Neguev. Geological Survey of Israel, no. Pal / 3 / 66, 12 p." type="book" year="1966">1966</bibRefCitation>
). All of these records are from throughout the Cenomanian, with Turonian records (e.g.,
<bibRefCitation id="EFDD4B51FFE2A228FEB5FA7FA4B8C28A" author="Saint-Marc, P." box="[307,501,1449,1473]" pageId="59" pageNumber="98" pagination="85 - 94" refId="ref95980" refString="Saint-Marc, P., 1970. Sur quelques foraminiferes Cenomaniens et Turoniens du Liban. Revue de Micropaleontologie, 13 (2): 85 - 94." type="journal article" year="1970">Saint-Marc, 1970</bibRefCitation>
) being revisable as late Cenomanian (e.g.,
<bibRefCitation id="EFDD4B51FFE2A228FEE4FA1CA76DC2A9" author="Saint-Marc, P." box="[354,544,1482,1506]" pageId="59" pageNumber="98" pagination="111 - 118" refId="ref96132" refString="Saint-Marc, P., 1978. Biostratigraphy of Albian, Cenomanian and Turonian of Lebanon. Annales Mines et Geologie, Tunis, 28: 111 - 118." type="journal article" year="1978">Saint-Marc, 1978</bibRefCitation>
).
<bibRefCitation id="EFDD4B51FFE2A228FDB5FA1CA64FC2A9" author="Mouty, M. &amp; Al-Maleh, A. K. &amp; Laban, H. A." box="[563,770,1482,1506]" pageId="59" pageNumber="98" pagination="429 - 443" refId="ref90890" refString="Mouty, M., Al-Maleh, A. K. &amp; Laban, H. A., 2003. Le Cretace moyen de la chaine des Palmyrides (Syrie centrale). Geodiversitas, 25 (3): 429 - 443." type="journal article" year="2003">Mouty et al. (2003)</bibRefCitation>
have reported the species from the late Cenomanian of
<collectingCountry id="F35B7630FFE2A228FF11F9DAA59EC16F" box="[151,211,1548,1572]" name="Syria" pageId="59" pageNumber="98">Syria</collectingCountry>
.
</paragraph>
<paragraph id="8BF336A0FFE2A228FF11F9FBA0C7C6D6" blockId="59.[151,772,158,2068]" lastBlockId="59.[816,1437,158,712]" pageId="59" pageNumber="98">
Records from the Sarvak Formation of
<collectingCountry id="F35B7630FFE2A228FDBAF9F8A72AC10E" box="[572,615,1582,1605]" name="Iran" pageId="59" pageNumber="98">Iran</collectingCountry>
and the Mishrif Formation in
<collectingCountry id="F35B7630FFE2A228FED2F999A432C12D" box="[340,383,1615,1638]" name="Iraq" pageId="59" pageNumber="98">Iraq</collectingCountry>
are numerous, especially from the former, but the quality of confirmatory illustrations is variable. Records with good or plausible illustration in the sense of
<emphasis id="B938EAB2FFE2A228FEA3F964A478C182" box="[293,309,1714,1737]" italics="true" pageId="59" pageNumber="98">P</emphasis>
.? ex. grp.
<taxonomicName id="4C4C4D23FFE2A228FE37F967A4A1C182" authority="(Bonet)" authorityName="Bonet" authorityYear="1956" baseAuthorityName="Bonet" baseAuthorityYear="1956" box="[433,492,1713,1737]" class="Tubothalamea" family="Miliolidae" genus="Pseudonummoloculina" kingdom="Chromista" order="Miliolida" pageId="59" pageNumber="98" phylum="Foraminifera" rank="species" species="heimi">
<emphasis id="B938EAB2FFE2A228FE37F967A4A1C182" box="[433,492,1713,1737]" italics="true" pageId="59" pageNumber="98">heimi</emphasis>
</taxonomicName>
include
<bibRefCitation id="EFDD4B51FFE2A228FDC8F967A5ADC1A1" author="Esfandyari, M. &amp; Mohseni, H. &amp; Heidari, M." pageId="59" pageNumber="98" refId="ref80463" refString="Esfandyari, M., Mohseni, H. &amp; Heidari, M., 2023. Facies analysis, depositional sequences and platform evolution of the Sarvak Formation (late Albian-Turonian) in the Zagros Basin, West of Iran. Journal of African Earth Sciences, doi: https: // doi. org / 10.1016 / j. jafrearsci. 2022.104811" type="book" year="2023">Esfandyari et al. (2023)</bibRefCitation>
,
<bibRefCitation id="EFDD4B51FFE2A228FF76F904A701C1A1" author="Rahimpour-Bonab, H. &amp; Mehrabi, H. &amp; Navidtalab, A. &amp; Omidvar, M. &amp; Enayati-Bidgoli, A. H. &amp; Sonei, R. &amp; Sajjadi, F. &amp; Amiri-Bakhtyar, H. &amp; Arzani, N. &amp; Mazidi, E." box="[240,588,1746,1770]" pageId="59" pageNumber="98" pagination="335 - 362" refId="ref94321" refString="Rahimpour-Bonab, H., Mehrabi, H., Navidtalab, A., Omidvar, M., Enayati-Bidgoli, A. H., Sonei, R., Sajjadi, F., Amiri-Bakhtyar, H., Arzani, N. &amp; Izadi- Mazidi, E., 2013. Palaeo-exposure surfaces in Cenomanian-Santonian carbonate reservoirs in the Dezful Embayment, SW Iran. Journal of Petroleum Geology, 36 (4): 335 - 362." type="journal article" year="2013">Rahimpour-Bonab et al. (2013)</bibRefCitation>
and Mohajer et al., (2021b). Records with questionable or no illustration include
<bibRefCitation id="EFDD4B51FFE2A228FF6BF8C3A4F3C066" author="Assadi, A. &amp; Honarmand, J. &amp; Moallemi, S. - A. &amp; Abdollahie-Fard, I." box="[237,446,1813,1837]" pageId="59" pageNumber="98" pagination="26" refId="ref72730" refString="Assadi, A., Honarmand, J., Moallemi, S. - A. &amp; Abdollahie-Fard, I., 2016. Depositional environments and sequence stratigraphy of the Sarvak Formation in an oil field in the Abadan Plain, SW Iran. Facies, 62: 26, doi 10.1007 / s 10347 - 016 - 0477 - 5" type="journal article" year="2016">Assadi et al. (2016)</bibRefCitation>
(probably
<taxonomicName id="4C4C4D23FFE2A228FDB3F8C0A406C005" authority="Chiocchini et al." authorityName="Chiocchini" class="Tubothalamea" family="Miliolidae" genus="Pseudonummoloculina" kingdom="Chromista" order="Miliolida" pageId="59" pageNumber="98" phylum="Foraminifera" rank="subSpecies" species="regularis" subSpecies="sensu">
<emphasis id="B938EAB2FFE2A228FDB3F8C0A708C066" box="[565,581,1814,1837]" italics="true" pageId="59" pageNumber="98">P</emphasis>
.?
<emphasis id="B938EAB2FFE2A228FDD9F8C3A64FC066" box="[607,770,1813,1837]" italics="true" pageId="59" pageNumber="98">regularis sensu</emphasis>
Chiocchini et al.
</taxonomicName>
);
<bibRefCitation id="EFDD4B51FFE2A228FEE5F8E0A72CC005" author="Daneshian, J. &amp; Roshanpour, E. &amp; Asadi Mehmandosti, E." box="[355,609,1846,1870]" pageId="59" pageNumber="98" refId="ref78381" refString="Daneshian, J., Roshanpour, E. &amp; Asadi Mehmandosti, E., 2016. Foraminiferal Biostratigraphy on the passing boundary of the Sarvak- Ilam formations in Chenareh Anticline, Northwest of Andimeshk. The 34 th National &amp; 2 th International Geosciences Congress, Geological survey of Iran, Tehran, Iran." type="proceedings paper" year="2016">Daneshian et al. (2016)</bibRefCitation>
;
<bibRefCitation id="EFDD4B51FFE2A228FDF7F8E0A44CC024" author="Kiarostami, K. &amp; Baghbani, D. &amp; Aleali, S. M. &amp; Aghanabati, S. A. &amp; Parandavar, M." pageId="59" pageNumber="98" pagination="155 - 164" refId="ref86516" refString="Kiarostami, K., Baghbani, D., Aleali, S. M., Aghanabati, S. A. &amp; Parandavar, M., 2019. Investigation of lithostratigraphic and biostratigraphic of the Sarvak Formation at type section. Geosciences, 29 (113): 155 - 164." type="journal article" year="2019">Kiarostami et al. (2019)</bibRefCitation>
;
<bibRefCitation id="EFDD4B51FFE2A228FE97F881A762C024" author="Saeedi Razavi, B. &amp; Askari, F. &amp; Kamali, M. R. &amp; Kazemzadeh, E." box="[273,559,1879,1903]" pageId="59" pageNumber="98" pagination="91 - 108" refId="ref95576" refString="Saeedi Razavi, B., Askari, F., Kamali, M. R. &amp; Kazemzadeh, E., 2019. Biostratigraphy, microfacies, depositional environment and sequence stratigraphy of the Sarvak Formation in one of the oilfields, southwest Iran. Sedimentary facies, 12 (1): 91 - 108 [in Farsi, English abstract]." type="journal article" year="2019">Saeedi Razavi et al. (2019</bibRefCitation>
,
<bibRefCitation id="EFDD4B51FFE2A228FDB9F881A739C024" author="Saeedi Razavi, B. &amp; Rikhtehgarzadeh, M. &amp; Senemari, S." box="[575,628,1879,1903]" pageId="59" pageNumber="98" pagination="241 - 254" refId="ref95642" refString="Saeedi Razavi, B., Rikhtehgarzadeh, M. &amp; Senemari, S., 2021. Biostratigraphy correlation of the Sarvak and Ilam Formations in middle restricted Dezful embayment, Southwest of Iran. Scientific Quarterly Journal of Geosciences, 30 (118): 241 - 254, [in Arabic, English abstract] DOI: 10.22071 / GSJ. 2020.214861.1739" type="journal article" year="2021">2021</bibRefCitation>
);
<bibRefCitation id="EFDD4B51FFE2A228FD0BF881A45BC0DB" author="Mohajer, M. D. &amp; Afghah, M. &amp; Dehghanian, M. &amp; Zakariaii, S. J. S." pageId="59" pageNumber="98" pagination="37 - 40" refId="ref90470" refString="Mohajer, M. D., Afghah, M., Dehghanian, M., &amp; Zakariaii, S. J. S., 2022 b. Biozonation, microfacies analysis and depositional environment of the Cenomanian sediments (Sarvak Formation) in South Zagros Basin (SW Iran). Carbonates and Evaporites, 37 - 40, https: // doi. org / 10.1007 / s 13146 - 022 - 00786 - 1" type="book chapter" year="2022">Mohajer et al. (2022b)</bibRefCitation>
;
<bibRefCitation id="EFDD4B51FFE2A228FEB6F8AEA4BEC0DB" author="Omidvar, M. &amp; Mehrabi, H. &amp; Sajjadi, F." box="[304,499,1912,1936]" pageId="59" pageNumber="98" pagination="158 - 177" refId="ref92031" refString="Omidvar, M., Mehrabi, H. &amp; Sajjadi, F., 2014 a. Depositional environment and biostratigraphy of the upper Sarvak Formation in Ahwaz Oilfield (Well No. 63). Sedimentary Facies, 7 (2): 158 - 177." type="journal article" year="2014">Omidvar (2014a)</bibRefCitation>
;
<bibRefCitation id="EFDD4B51FFE2A228FD8AF8AEA595C0FA" author="Rikhtegarzadeh, M. &amp; Vaziri, S. H. &amp; Aleali, M. &amp; Amiri Bakhtiar, H. &amp; Jahani, D." pageId="59" pageNumber="98" pagination="194 - 208" refId="ref94890" refString="Rikhtegarzadeh, M., Vaziri, S. H., Aleali, M., Amiri Bakhtiar, H. &amp; Jahani, D., 2016. Microbiostratigraphy, microfacies and depositional environment of the Sarvak Formation in Bi Bi Hakimeh Oil Field (well no. 29), southwest Iran. International Journal of Geography and Geology, 5 (10): 194 - 208." type="journal article" year="2016">Rikhtegarzadeh et al. (2016</bibRefCitation>
,
<bibRefCitation id="EFDD4B51FFE2A228FF6AF84FA46EC0FA" author="Rikhtegarzadeh, M. &amp; Vaziri, S. H. &amp; Aleali, M. &amp; Amiri Bakhtiar, H. &amp; Jahani, D." box="[236,291,1945,1969]" pageId="59" pageNumber="98" pagination="413 - 428" refId="ref94962" refString="Rikhtegarzadeh, M., Vaziri, S. H., Aleali, M., Amiri Bakhtiar, H. &amp; Jahani, D., 2017. Microbiostratigraphy, Microfacies and Depositional Environment of the Sarvak and Ilam Formations in the Gachsaran Oilfield, southwest Iran. Micropaleontology, 63 (6): 413 - 428." type="journal article" year="2017">2017</bibRefCitation>
) and
<bibRefCitation id="EFDD4B51FFE2A228FEF4F84FA7F5C0FA" author="Al-Salihi, A. &amp; Ibrahim, Y. K." box="[370,696,1945,1969]" pageId="59" pageNumber="98" pagination="196 - 207" refId="ref71294" refString="Al-Salihi, A. &amp; Ibrahim, Y. K., 2023. Larger Foraminifera of Middle and Late Cenomanian (Biostrome Association), Southern Iraq. Iraqi Geological Journal, 56 (1 A): 196 - 207." type="journal article" year="2023">Al-Salihi &amp; Ibrahim (2023)</bibRefCitation>
. Even though considered of biozonal value by the authors, the illustrations of
<emphasis id="B938EAB2FFE2A228FECEF80AA420C0B8" box="[328,365,2012,2035]" italics="true" pageId="59" pageNumber="98">P.?</emphasis>
ex. grp.
<taxonomicName id="4C4C4D23FFE2A228FE64F80DA750C0B8" authority="(Bonet)" authorityName="Bonet" authorityYear="1956" baseAuthorityName="Bonet" baseAuthorityYear="1956" box="[482,541,2011,2035]" class="Tubothalamea" family="Miliolidae" genus="Pseudonummoloculina" kingdom="Chromista" order="Miliolida" pageId="59" pageNumber="98" phylum="Foraminifera" rank="species" species="heimi">
<emphasis id="B938EAB2FFE2A228FE64F80DA750C0B8" box="[482,541,2011,2035]" italics="true" pageId="59" pageNumber="98">heimi</emphasis>
</taxonomicName>
(as
<emphasis id="B938EAB2FFE2A228FDDFF80DA596CF5F" italics="true" pageId="59" pageNumber="98">
<taxonomicName id="4C4C4D23FFE2A228FDDFF80DA59ECF5F" authorityName="Bonet" authorityYear="1956" class="Tubothalamea" family="Hauerinidae" genus="Nummoloculina" kingdom="Chromista" order="Miliolida" pageId="59" pageNumber="98" phylum="Foraminifera" rank="species" species="heimi">Nummoloculina heimi</taxonomicName>
)
</emphasis>
by Afghah et al. (2014) and
<bibRefCitation id="EFDD4B51FFE2A228FDBBF82AA63AC7FD" author="Afghah, M. &amp; Fadaei, H. R." box="[573,887,158,2068]" pageId="59" pageNumber="98" refId="ref70404" refString="Afghah, M. &amp; Fadaei, H. R., 2014. Biostratigraphy of Cenomanian succession in Zagros area (south west of Iran). Geosciences Journal, DOI 10.1007 / s 12303 - 014 - 0045 - 3" type="book" year="2014">Afghah &amp; Fadaei (2014)</bibRefCitation>
cannot be confirmed as being of this species. The illustration by
<bibRefCitation id="EFDD4B51FFE2A228FC53FF69A1A6C79C" author="Afghah, M. &amp; Dookh, R." box="[981,1259,191,215]" pageId="59" pageNumber="98" pagination="5 - 19" refId="ref70352" refString="Afghah, M. &amp; Dookh, R., 2014. Microbiostratigraphy of the Sarvak Formation in the east and north-east of Shiraz (Kuh-e-Siah and Kuh-e-Pichakan), SW of Iranian Journal of Science I. A. U. (JSIAU), 24: 5 - 19." type="journal article" year="2014">Afghah &amp; Dookh (2014)</bibRefCitation>
is of an alveolinid. These records are predominantly assigned a Cenomanian age, although
<bibRefCitation id="EFDD4B51FFE2A228FBB3FED7A0D1C652" author="Rahimpour-Bonab, H. &amp; Mehrabi, H. &amp; Navidtalab, A. &amp; Omidvar, M. &amp; Enayati-Bidgoli, A. H. &amp; Sonei, R. &amp; Sajjadi, F. &amp; Amiri-Bakhtyar, H. &amp; Arzani, N. &amp; Mazidi, E." box="[1077,1436,257,281]" pageId="59" pageNumber="98" pagination="335 - 362" refId="ref94321" refString="Rahimpour-Bonab, H., Mehrabi, H., Navidtalab, A., Omidvar, M., Enayati-Bidgoli, A. H., Sonei, R., Sajjadi, F., Amiri-Bakhtyar, H., Arzani, N. &amp; Izadi- Mazidi, E., 2013. Palaeo-exposure surfaces in Cenomanian-Santonian carbonate reservoirs in the Dezful Embayment, SW Iran. Journal of Petroleum Geology, 36 (4): 335 - 362." type="journal article" year="2013">Rahimpour-Bonab et al. (2013)</bibRefCitation>
provide a combined total range of late Albian Turonian. An illustrated record by
<bibRefCitation id="EFDD4B51FFE2A228FBD6FE95A063C610" author="Gollesstaneh, A." box="[1104,1326,323,347]" pageId="59" pageNumber="98" refId="ref82472" refString="Gollesstaneh, A. 1965. A micropalaeontological study of the Upper lurassic and Lower Cretaceous of Southern Iran. Unpublished PhD thesis, University of London." type="book" year="1965">Gollesstaneh (1965)</bibRefCitation>
from the “late Aptian early Albian” of the Iranian Zagros, represents one of the oldest records of this species
<emphasis id="B938EAB2FFE2A228FA91FE50A0CFC6D6" box="[1303,1410,389,413]" italics="true" pageId="59" pageNumber="98">sensu lato</emphasis>
.
</paragraph>
<paragraph id="8BF336A0FFE2A228FCB6FE71A14CC508" blockId="59.[816,1437,158,712]" pageId="59" pageNumber="98">
Records from
<collectingCountry id="F35B7630FFE2A228FC4DFE71A6BBC6F5" box="[971,1014,423,446]" name="Iraq" pageId="59" pageNumber="98">Iraq</collectingCountry>
are fewer and include illustrated forms by
<bibRefCitation id="EFDD4B51FFE2A228FCD5FE1EA187C6AB" author="Al-Dulaimy, R. T. &amp; Al-Sheikhly, S. S." box="[851,1226,456,480]" pageId="59" pageNumber="98" pagination="1 - 14" refId="ref70996" refString="Al-Dulaimy, R. T. &amp; Al-Sheikhly, S. S., 2013. Biostratigraphy of the Mishrif Formation from Well Amarah- 1 southeastern Iraq. Iraqi Bulletin of Geology and Mining, 9 (2): 1 - 14." type="journal article" year="2013">Al-Dulaimy &amp; Al-Sheikhly (2013)</bibRefCitation>
which are possibly
<emphasis id="B938EAB2FFE2A228FCB6FE3CA618C54A" box="[816,853,490,513]" italics="true" pageId="59" pageNumber="98">P.?</emphasis>
ex. grp.
<taxonomicName id="4C4C4D23FFE2A228FC4CFE3FA148C54A" authority="(Bonet)" authorityName="Bonet" authorityYear="1956" baseAuthorityName="Bonet" baseAuthorityYear="1956" box="[970,1029,489,513]" class="Tubothalamea" family="Hauerinidae" genus="Nummoloculina" kingdom="Chromista" order="Miliolida" pageId="59" pageNumber="98" phylum="Foraminifera" rank="species" species="heimi">
<emphasis id="B938EAB2FFE2A228FC4CFE3FA148C54A" box="[970,1029,489,513]" italics="true" pageId="59" pageNumber="98">heimi</emphasis>
</taxonomicName>
. Illustrations by
<bibRefCitation id="EFDD4B51FFE2A228FB54FE3FA63AC569" author="Al-Dulaimy, S. I. M. &amp; Ibrahim, Y. K. &amp; Abdallah, F. T." pageId="59" pageNumber="98" pagination="79 - 89" refId="ref71040" refString="Al-Dulaimy, S. I. M., Ibrahim, Y. K. &amp; Abdallah, F. T., 2022. Biozonation (benthic foraminifera) of Mishrif Formation at Majnoon and Zubair oil fields, southern Iraq. Bulletin of the Geological Society of Malaysia, 73: 79 - 89." type="journal article" year="2022">Al-Dulaimy et al. (2022)</bibRefCitation>
from the late Cenomanian Mishrif Formation may well be alveolinids.
</paragraph>
<paragraph id="8BF336A0FFE2A228FCB6FD9AA6A7C583" blockId="59.[816,1437,158,712]" pageId="59" pageNumber="98">
Additional unillustrated records occur from the Natih Formation of
<collectingCountry id="F35B7630FFE2A228FC4DFDBBA141C5CE" box="[971,1036,621,645]" name="Oman" pageId="59" pageNumber="98">Oman</collectingCountry>
(
<bibRefCitation id="EFDD4B51FFE2A228FB98FDBBA01BC5CE" author="Kennedy, W. J. &amp; Simmons, M. D." box="[1054,1366,621,645]" pageId="59" pageNumber="98" pagination="127 - 154" refId="ref86200" refString="Kennedy, W. J. &amp; Simmons, M. D., 1991. Mid-Cretaceous ammonites and associated microfossils from the Central Oman Mountains. Newsletters on Stratigraphy, 25 (3): 127 - 154." type="journal article" year="1991">Kennedy &amp; Simmons, 1991</bibRefCitation>
;
<bibRefCitation id="EFDD4B51FFE2A228FAEFFDBBA15CC5ED" author="Simmons, M. D. &amp; Hart, M. B." pageId="59" pageNumber="98" pagination="176 - 207" refId="ref98973" refString="Simmons, M. D. &amp; Hart, M. B., 1987. The biostratigraphy and microfacies of the Early to mid-Cretaceous carbonates of Wadi Mi'aidin, Central Oman Mountains. In: Hart, M. B. (ed), Micropalaeontology of Carbonate Environments, John Wiley &amp; Sons: 176 - 207." type="book chapter" year="1987">Simmons &amp; Hart, 1987</bibRefCitation>
) and are assigned a middle late Cenomanian age.
</paragraph>
</subSubSection>
<subSubSection id="C356652BFFE2A228FCB6FD24A05BC407" pageId="59" pageNumber="98" type="distribution">
<paragraph id="8BF336A0FFE2A228FCB6FD24A1BAC441" blockId="59.[816,1271,754,778]" box="[816,1271,754,778]" pageId="59" pageNumber="98">Cenomanian Paleogeographic Distribution</paragraph>
<paragraph id="8BF336A0FFE2A228FCB6FCE2A05BC407" blockId="59.[816,1437,820,1010]" box="[816,1302,820,844]" pageId="59" pageNumber="98">Pan-Neotethyan, America and the Caribbean.</paragraph>
</subSubSection>
<subSubSection id="C356652BFFE2A228FCB6FC83A187C4BA" pageId="59" pageNumber="98" type="description">
<paragraph id="8BF336A0FFE2A228FCB6FC83A187C4BA" blockId="59.[816,1437,820,1010]" pageId="59" pageNumber="98">
See references above. As a note, pre-Cenomanian records can be extended further since
<bibRefCitation id="EFDD4B51FFE2A228FBF4FCA0A69DC4E4" author="Arnaud-Vanneau, A. &amp; Premoli-Silva, I." pageId="59" pageNumber="98" pagination="199 - 219" refId="ref72480" refString="Arnaud-Vanneau, A. &amp; Premoli-Silva, I., 1995. Biostratigraphy and systematic description of benthic foraminifers from Mid-Cretaceous shallow-water carbonate platform sediments at sites 878 and 879 (Mit and Takuyo-daisan Guyots). Proceedings of the Ocean Drilling Program, Scientific Results, 144: 199 - 219." type="journal article" year="1995">Arnaud-Vanneau &amp; Premoli-Silva (1995)</bibRefCitation>
note that their “
<taxonomicName id="4C4C4D23FFE2A228FB0CFC41A013C4E4" box="[1162,1374,919,943]" class="Tubothalamea" family="Hauerinidae" genus="Nummoloculina" kingdom="Chromista" order="Miliolida" pageId="59" pageNumber="98" phylum="Foraminifera" rank="species" species="undetermined">
<emphasis id="B938EAB2FFE2A228FB0CFC41A079C4E4" box="[1162,1332,919,943]" italics="true" pageId="59" pageNumber="98">Nummoloculina</emphasis>
sp.
</taxonomicName>
” recorded from the late Albian (?) of MIT Guyot in the Pacific is very similar to
<emphasis id="B938EAB2FFE2A228FB83FC0CA167C4BA" box="[1029,1066,986,1009]" italics="true" pageId="59" pageNumber="98">P.?</emphasis>
ex. grp.
<emphasis id="B938EAB2FFE2A228FB01FC0FA187C4BA" box="[1159,1226,985,1009]" italics="true" pageId="59" pageNumber="98">
<taxonomicName id="4C4C4D23FFE2A228FB01FC0FA18BC4BA" authority="(Bonet)" authorityName="Bonet" authorityYear="1956" baseAuthorityName="Bonet" baseAuthorityYear="1956" box="[1159,1222,985,1009]" class="Tubothalamea" family="Hauerinidae" genus="Nummoloculina" kingdom="Chromista" order="Miliolida" pageId="59" pageNumber="98" phylum="Foraminifera" rank="species" species="heimi">heimi</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>