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<document id="E8AA1248F9F4A2D2F31F51D16B38061B" ID-DOI="10.5281/zenodo.198244" ID-GBIF-Dataset="2145a6ff-d31c-45f1-bea1-af8477a51da6" ID-ISSN="1175-5326" ID-Zenodo-Dep="198244" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460290874690" checkinUser="plazi" docAuthor="Gharali, Babak, Kamali, Karim, Evenhuis, Neal, Talebi, Ali Asghar &amp; Khalgani, Jafar" docDate="2010" docId="03B987B2FFEC5E3FFF50F4DEFE7555C4" docLanguage="en" docName="zt02627p019.pdf" docOrigin="Zootaxa 2627" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Empidideicus greatheadi Gharali &amp; Evenhuis, sp. nov." docType="treatment" docVersion="9" lastPageNumber="10" masterDocId="FF80FFCAFFEB5E36FFC7F449FFC05658" masterDocTitle="First record of the genus Empidideicus (Diptera: Bombylioidea: Mythicomyiidae) from Iran, with description of six new species" masterLastPageNumber="19" masterPageNumber="1" pageNumber="8" updateTime="1698251303231" updateUser="plazi">
<mods:mods id="46500E0C0A29800DE7B47844A2570615" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C074C8458199A3A8D20796BB8E3569FC">
<mods:title id="32600FA268392BFF7E9FE175FC1F5AC9">First record of the genus Empidideicus (Diptera: Bombylioidea: Mythicomyiidae) from Iran, with description of six new species</mods:title>
</mods:titleInfo>
<mods:name id="F37A305E42266E3824B3C3BDE501A85B" type="personal">
<mods:role id="BE01AE259D3A3779B70DCB1FC9DCE128">
<mods:roleTerm id="032273A0C5D1AB2DE8BBD59733387AF8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CF100D57AE119CE7BCB6DC7C3D052F22">Gharali, Babak</mods:namePart>
</mods:name>
<mods:name id="DC3B09DE4D59CB38D71AC575E533A3CD" type="personal">
<mods:role id="7DF509BF08BCEB252409D6CD927A69EE">
<mods:roleTerm id="FC37419528B5061D2D60E8F6B03ACE18">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D3443142FB854686A5B5330F4B5BF84B">Kamali, Karim</mods:namePart>
</mods:name>
<mods:name id="0B5C1B4606D8F152446A9CBD53945D54" type="personal">
<mods:role id="85C3578596A5F5380BBFD3B0838E9273">
<mods:roleTerm id="ADCEF51C0A702A707723FF91F9C22A56">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="75B83F0C7D581817DD3AD34E3AFE60A1">Evenhuis, Neal</mods:namePart>
</mods:name>
<mods:name id="E86D9FCEC899D69B865FE6F1AEFEFF89" type="personal">
<mods:role id="E29510256B6E70D99585B973A6881947">
<mods:roleTerm id="B685B5C49680F7BFE6CA763FA1E1E558">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="97EB968E744556A774249E5BDDA34FE2">Talebi, Ali Asghar</mods:namePart>
</mods:name>
<mods:name id="AD504D8ACAFFD8BBA5E9016023408FAE" type="personal">
<mods:role id="8EC37EE540403A14322C06BB79B66579">
<mods:roleTerm id="CA55607CD926116AD32D23E79A427DAC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C0360CDDCDAC597B2A46808DFC23CBB2">Khalgani, Jafar</mods:namePart>
</mods:name>
<mods:typeOfResource id="C98E7EBB68C2E3919A2435C3E1A39C70">text</mods:typeOfResource>
<mods:relatedItem id="266E03317A1B86BDC03C49DC100B8B9E" type="host">
<mods:titleInfo id="A06FA967D1CDF1B57AB08670C3317E0F">
<mods:title id="FAB66C1D9FDB90D04C5A07AD867975AA">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="F11883331E02984D552BEA86DDF0FBD3">
<mods:date id="D6158EB2027435EEF9BB50B5C7A729AC">2010</mods:date>
<mods:detail id="645C21EA1B8C3D4D2BBF8BE37679B704" type="volume">
<mods:number id="99B1F83F4F6AC0A920F322FE0BA36D29">2627</mods:number>
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<mods:extent id="C0408F5D222306C92B99B47DB30458B9" unit="page">
<mods:start id="46719C61FC19A38E0638D977C6527160">1</mods:start>
<mods:end id="161A84B63DEDA54D3723AD79314669CD">19</mods:end>
</mods:extent>
</mods:part>
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<mods:classification id="9D238BCFB2A52B6585776B52C81EA068">journal article</mods:classification>
<mods:identifier id="22FAB7049C0224A33688A34360F1F77F" type="DOI">10.5281/zenodo.198244</mods:identifier>
<mods:identifier id="577481246824665AF00AEB4C47A0B1F8" type="GBIF-Dataset">2145a6ff-d31c-45f1-bea1-af8477a51da6</mods:identifier>
<mods:identifier id="40EC578F1AECC757A8518E78136D4FB7" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="D0C082F398C06C0D0382AC78FC8CCB59" type="Zenodo-Dep">198244</mods:identifier>
</mods:mods>
<treatment id="03B987B2FFEC5E3FFF50F4DEFE7555C4" ID-DOI="http://doi.org/10.5281/zenodo.6209827" ID-GBIF-Taxon="119398623" ID-Zenodo-Dep="6209827" LSID="urn:lsid:plazi:treatment:03B987B2FFEC5E3FFF50F4DEFE7555C4" httpUri="http://treatment.plazi.org/id/03B987B2FFEC5E3FFF50F4DEFE7555C4" lastPageId="9" lastPageNumber="10" pageId="7" pageNumber="8">
<subSubSection id="C30A652FFFEC5E31FF50F4DEFEEA54CC" pageId="7" pageNumber="8" type="nomenclature">
<paragraph id="8BAF36A4FFEC5E31FF50F4DEFCF356E9" blockId="7.[151,819,151,212]" box="[151,819,151,177]" pageId="7" pageNumber="8">
<heading id="D0E781C8FFEC5E31FF50F4DEFCF356E9" bold="true" box="[151,819,151,177]" fontSize="11" level="1" pageId="7" pageNumber="8" reason="1">
<emphasis id="B964EAB6FFEC5E31FF50F4DEFCF356E9" bold="true" box="[151,819,151,177]" pageId="7" pageNumber="8">
<taxonomicName id="4C104D27FFEC5E31FF50F4DEFD0856E9" authority="Gharali &amp; Evenhuis" authorityName="Gharali &amp; Evenhuis" box="[151,712,151,177]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="greatheadi" status="sp. nov.">
<emphasis id="B964EAB6FFEC5E31FF50F4DEFE0456E9" bold="true" box="[151,452,151,177]" italics="true" pageId="7" pageNumber="8">Empidideicus greatheadi</emphasis>
Gharali &amp; Evenhuis
</taxonomicName>
,
<taxonomicNameLabel id="A25757CDFFEC5E31FD12F4DEFCF356E9" box="[725,819,151,177]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF36A4FFEC5E31FF50F4F3FEBB568C" blockId="7.[151,819,151,212]" box="[151,379,186,212]" pageId="7" pageNumber="8">
(
<figureCitation id="132B2A21FFEC5E31FF58F4F3FF2F568C" box="[159,239,186,212]" captionStart="FIGURE 3" captionStartId="8.[151,255,1968,1992]" captionTargetBox="[165,1418,198,1920]" captionTargetId="figure@8.[165,1422,170,1946]" captionTargetPageId="8" captionText="FIGURE 3. Empidideicus greatheadi: a—antennae lateral view, b—wing, c—phallic complex dorsal view, d—phallic complex lateral view, e—epandrium dorsal view, f—gonocoxa dorsal view, g—female genitalia." httpUri="https://zenodo.org/record/198247/files/figure.png" pageId="7" pageNumber="8">Figs. 3</figureCitation>
a–g, 7c, 8d)
</paragraph>
<paragraph id="8BAF36A4FFEC5E31FF50F54CFEEA54CC" blockId="7.[151,1438,261,1979]" pageId="7" pageNumber="8">
<emphasis id="B964EAB6FFEC5E31FF50F54CFE665747" bold="true" box="[151,422,261,287]" pageId="7" pageNumber="8">Specimens examined:</emphasis>
<typeStatus id="54AB8806FFEC5E31FE69F54CFDDB5747" box="[430,539,261,287]" pageId="7" pageNumber="8" type="holotype">Holotype</typeStatus>
female (dried by HMDS) and
<specimenCount id="9D16FD2DFFEC5E31FC4EF54CFC2A5747" box="[905,1002,261,287]" pageId="7" pageNumber="8" type="male">10 male</specimenCount>
,
<specimenCount id="9D16FD2DFFEC5E31FC30F54CFBAD5747" box="[1015,1133,261,287]" pageId="7" pageNumber="8" type="female">10 female</specimenCount>
<typeStatus id="54AB8806FFEC5E31FBB2F54CFB265747" box="[1141,1254,261,287]" pageId="7" pageNumber="8" type="paratype">paratypes</typeStatus>
(
<specimenCount id="9D16FD2DFFEC5E31FB30F54CFA935747" box="[1271,1363,261,287]" pageId="7" pageNumber="8" type="male">5 males</specimenCount>
and
<specimenCount id="9D16FD2DFFEC5E31FA4AF54CFF32571C" pageId="7" pageNumber="8" type="female">5 females</specimenCount>
dried by HMDS, remainder preserved in alcohol),
<collectingCountry id="F3077634FFEC5E31FC8AF563FC5A571C" box="[845,922,298,324]" name="Iran" pageId="7" pageNumber="8">IRAN</collectingCountry>
: Abazar, Abazar village, Ghazvin province,
<geoCoordinate id="EE245063FFEC5E31FF50F519FF395732" box="[151,249,336,362]" direction="north" orientation="latitude" pageId="7" pageNumber="8" precision="925" value="36.283333">N36°17'</geoCoordinate>
<geoCoordinate id="EE245063FFEC5E31FF39F519FEA35732" box="[254,355,336,362]" direction="east" orientation="longitude" pageId="7" pageNumber="8" precision="925" value="50.166668">E50° 10'</geoCoordinate>
,
<quantity id="4CE89B41FFEC5E31FEBFF519FE105732" box="[376,464,336,362]" metricMagnitude="3" metricUnit="m" metricValue="1.463" pageId="7" pageNumber="8" unit="m" value="1463.0">1463 m</quantity>
asl,
<date id="FFAE1064FFEC5E31FDC1F519FD645732" box="[518,676,336,362]" pageId="7" pageNumber="8" value="2009-06-15">15 June 2009</date>
, Pan trapping, leg. B. Gharali (all in TMUC);
<specimenCount id="9D16FD2DFFEC5E31FB7AF519FADD5732" box="[1213,1309,336,362]" pageId="7" pageNumber="8" type="male">15 male</specimenCount>
,
<specimenCount id="9D16FD2DFFEC5E31FAEFF519FA5C5732" box="[1320,1436,336,362]" pageId="7" pageNumber="8" type="female">25 female</specimenCount>
<typeStatus id="54AB8806FFEC5E31FF50F53CFECA57D7" box="[151,266,373,399]" pageId="7" pageNumber="8" type="paratype">paratypes</typeStatus>
(preserved in alcohol),
<collectingCountry id="F3077634FFEC5E31FDECF53CFDB957D7" box="[555,633,373,399]" name="Iran" pageId="7" pageNumber="8">IRAN</collectingCountry>
: Barajin,
<quantity id="4CE89B41FFEC5E31FD2AF53CFCEB57D7" box="[749,811,373,399]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="8" unit="km" value="8.0">8 km</quantity>
North of Ghazvin City, Ghazvin province,
<geoCoordinate id="EE245063FFEC5E31FAFEF53CFF0157EC" direction="north" orientation="latitude" pageId="7" pageNumber="8" precision="15" value="36.34722">N36°20' 50&quot;</geoCoordinate>
<geoCoordinate id="EE245063FFEC5E31FF06F5D3FE8C57EC" box="[193,332,410,436]" direction="east" orientation="longitude" pageId="7" pageNumber="8" precision="15" value="350.07086">E350°4'15&quot;</geoCoordinate>
,
<quantity id="4CE89B41FFEC5E31FE9EF5D3FE7457EC" box="[345,436,410,436]" metricMagnitude="3" metricUnit="m" metricValue="1.512" pageId="7" pageNumber="8" unit="m" value="1512.0">1512 m</quantity>
asl.,
<date id="FFAE1064FFEC5E31FE33F5D3FD5557EC" box="[500,661,410,436]" pageId="7" pageNumber="8" value="2009-06-23">23 June 2009</date>
, leg. B. Gharali (BPBM);
<specimenCount id="9D16FD2DFFEC5E31FC1BF5D3FBFE57EC" box="[988,1086,410,436]" pageId="7" pageNumber="8" type="male">11 male</specimenCount>
,
<specimenCount id="9D16FD2DFFEC5E31FB8DF5D3FB0157EC" box="[1098,1217,410,436]" pageId="7" pageNumber="8" type="female">10 female</specimenCount>
<typeStatus id="54AB8806FFEC5E31FB0EF5D3FAFE57EC" box="[1225,1342,410,436]" pageId="7" pageNumber="8" type="paratype">paratypes</typeStatus>
,
<collectingCountry id="F3077634FFEC5E31FA8CF5D3FA5857EC" box="[1355,1432,410,436]" name="Iran" pageId="7" pageNumber="8">IRAN</collectingCountry>
: Nodahak rangeland, Nodahak village, Ghazvin province,
<geoCoordinate id="EE245063FFEC5E31FC8AF589FC715782" box="[845,945,448,474]" direction="north" orientation="latitude" pageId="7" pageNumber="8" precision="925" value="35.95">N35°57'</geoCoordinate>
<geoCoordinate id="EE245063FFEC5E31FC7EF589FBE45782" box="[953,1060,448,474]" direction="east" orientation="longitude" pageId="7" pageNumber="8" precision="925" value="49.583332">E49° 35'</geoCoordinate>
,
<quantity id="4CE89B41FFEC5E31FBF6F589FB4B5782" box="[1073,1163,448,474]" metricMagnitude="3" metricUnit="m" metricValue="1.328" pageId="7" pageNumber="8" unit="m" value="1328.0">1328 m</quantity>
asl.,
<date id="FFAE1064FFEC5E31FB0AF589FAA35782" box="[1229,1379,448,474]" pageId="7" pageNumber="8" value="2009-06-03">3 June 2009</date>
, Pan trapping, leg. B. Gharali (BPBM);
<specimenCount id="9D16FD2DFFEC5E31FDFBF5ACFD7757A7" box="[572,695,485,511]" pageId="7" pageNumber="8" type="female">12 female</specimenCount>
,
<specimenCount id="9D16FD2DFFEC5E31FD04F5ACFCD457A7" box="[707,788,485,511]" pageId="7" pageNumber="8" type="male">8 male</specimenCount>
<typeStatus id="54AB8806FFEC5E31FCDBF5ACFC5257A7" box="[796,914,485,511]" pageId="7" pageNumber="8" type="paratype">paratypes</typeStatus>
, same locality data as
<typeStatus id="54AB8806FFEC5E31FB65F5ACFACB57A7" box="[1186,1291,485,511]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
(ZMHB);
<specimenCount id="9D16FD2DFFEC5E31FA4AF5ACFF2A547C" pageId="7" pageNumber="8" type="female">5 female</specimenCount>
,
<specimenCount id="9D16FD2DFFEC5E31FF32F643FE85547C" box="[245,325,522,548]" pageId="7" pageNumber="8" type="male">8 male</specimenCount>
<typeStatus id="54AB8806FFEC5E31FE8AF643FE00547C" box="[333,448,522,548]" pageId="7" pageNumber="8" type="paratype">paratypes</typeStatus>
, same locality data as
<typeStatus id="54AB8806FFEC5E31FD0FF643FCF2547C" box="[712,818,522,548]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
, personal collection of Jorge Almeida (
<collectingCountry id="F3077634FFEC5E31FAC7F643FAAA547C" box="[1280,1386,522,548]" name="Portugal" pageId="7" pageNumber="8">Portugal</collectingCountry>
);
<specimenCount id="9D16FD2DFFEC5E31FAB8F643FF155412" pageId="7" pageNumber="8" type="male">10 male</specimenCount>
,
<specimenCount id="9D16FD2DFFEC5E31FF25F679FE9C5412" box="[226,348,560,586]" pageId="7" pageNumber="8" type="female">10 female</specimenCount>
<typeStatus id="54AB8806FFEC5E31FEA2F679FE1A5412" box="[357,474,560,586]" pageId="7" pageNumber="8" type="paratype">paratypes</typeStatus>
(preserved in alcohol),
<collectingCountry id="F3077634FFEC5E31FCC6F679FC905412" box="[769,848,560,586]" name="Iran" pageId="7" pageNumber="8">IRAN</collectingCountry>
: Nodahak rangeland, Nodahak village, Ghazvin province,
<geoCoordinate id="EE245063FFEC5E31FEC8F61CFEBA5437" box="[271,378,597,623]" direction="north" orientation="latitude" pageId="7" pageNumber="8" precision="925" value="35.95">N35° 57'</geoCoordinate>
<geoCoordinate id="EE245063FFEC5E31FE45F61CFE245437" box="[386,484,597,623]" direction="east" orientation="longitude" pageId="7" pageNumber="8" precision="925" value="49.583332">E49°35'</geoCoordinate>
,
<quantity id="4CE89B41FFEC5E31FE37F61CFD8B5437" box="[496,587,597,623]" metricMagnitude="3" metricUnit="m" metricValue="1.328" pageId="7" pageNumber="8" unit="m" value="1328.0">1328 m</quantity>
asl.,
<date id="FFAE1064FFEC5E31FD4BF61CFCE05437" box="[652,800,597,623]" pageId="7" pageNumber="8" value="2009-06-03">3 June 2009</date>
, Pan trapping, leg. B. Gharali (personal collection of first author).
</paragraph>
</subSubSection>
<subSubSection id="C30A652FFFEC5E31FF01F6D6FEC35252" pageId="7" pageNumber="8" type="diagnosis">
<paragraph id="8BAF36A4FFEC5E31FF01F6D6FEC35252" blockId="7.[151,1438,261,1979]" pageId="7" pageNumber="8">
<emphasis id="B964EAB6FFEC5E31FF01F6D6FE8654E1" bold="true" box="[198,326,671,697]" pageId="7" pageNumber="8">Diagnosis.</emphasis>
<taxonomicName id="4C104D27FFEC5E31FE8AF6E9FDB254E1" box="[333,626,672,697]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="greatheadi">
<emphasis id="B964EAB6FFEC5E31FE8AF6E9FDB254E1" box="[333,626,672,697]" italics="true" pageId="7" pageNumber="8">Empidideicus greatheadi</emphasis>
</taxonomicName>
is distinguished from congeners in
<collectingCountry id="F3077634FFEC5E31FBD3F6E9FB8354E2" box="[1044,1091,672,698]" name="Iran" pageId="7" pageNumber="8">Iran</collectingCountry>
by the black mesonotum and completely yellow scutellum. Using the key to species of the
<collectingCountry id="F3077634FFEC5E31FCACF68CFC675487" box="[875,935,709,735]" name="United Arab Emirates" pageId="7" pageNumber="8">UAE</collectingCountry>
(
<bibRefCitation id="EF814B55FFEC5E31FC70F68CFBB35487" author="Evenhuis" box="[951,1139,709,735]" pageId="7" pageNumber="8" refString="Evenhuis, N. L. (2009) Order Diptera, family Mythicomyiidae. In: van Harten, A. (Ed.), Arthropod fauna of the UAE, Vol. 2. Dar Al Ummah, Adu Dhabi, pp. 714 - 740." type="book" year="2009">Evenhuis, 2009</bibRefCitation>
), our new species runs to
<taxonomicName id="4C104D27FFEC5E31FF50F6A3FE00555C" authority="Evenhuis" authorityName="Evenhuis" box="[151,448,746,772]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="anahorticus">
<emphasis id="B964EAB6FFEC5E31FF50F6A3FE86555B" box="[151,326,746,771]" italics="true" pageId="7" pageNumber="8">E. anahorticus</emphasis>
Evenhuis
</taxonomicName>
; but
<taxonomicName id="4C104D27FFEC5E31FE3CF6A3FD5C555B" box="[507,668,746,771]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="greatheadi">
<emphasis id="B964EAB6FFEC5E31FE3CF6A3FD5C555B" box="[507,668,746,771]" italics="true" pageId="7" pageNumber="8">E. greatheadi</emphasis>
</taxonomicName>
is easily separated from this Arabian species by the color of the scutellum, which is yellow in our species and reddish brown in the Arabian one. According to Engel’s (1933) key, our new species runs to
<taxonomicName id="4C104D27FFEC5E31FE3FF77CFC4E5517" authority="Thalhammer, 1911" authorityName="Thalhammer" authorityYear="1911" box="[504,910,821,847]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="hungaricus">
<emphasis id="B964EAB6FFEC5E31FE3FF77CFD625516" box="[504,674,821,846]" italics="true" pageId="7" pageNumber="8">E. hungaricus</emphasis>
Thalhammer, 1911
</taxonomicName>
but it is easily distinguished from it by the completely black occiput (occiput yellow laterally in
<taxonomicName id="4C104D27FFEC5E31FCE2F713FC10552B" box="[805,976,858,883]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="hungaricus">
<emphasis id="B964EAB6FFEC5E31FCE2F713FC10552B" box="[805,976,858,883]" italics="true" pageId="7" pageNumber="8">E. hungaricus</emphasis>
</taxonomicName>
).
<taxonomicName id="4C104D27FFEC5E31FC2CF713FA5C552C" authority="Zaitzev, 1971" authorityName="Zaitzev" authorityYear="1971" box="[1003,1436,858,884]" class="Insecta" family="Mythicomyiidae" genus="Epidideicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="asiaticus">
<emphasis id="B964EAB6FFEC5E31FC2CF713FB2E552B" box="[1003,1262,858,883]" italics="true" pageId="7" pageNumber="8">Epidideicus asiaticus</emphasis>
<bibRefCitation id="EF814B55FFEC5E31FB30F713FA5C552C" author="Zaitzev" box="[1271,1436,858,884]" pageId="7" pageNumber="8" refString="Zaitzev, V. F. (1971) New bombyliid species (Diptera, Bombyliidae) from Soviet central Asia and Mongolia. Entomologicheskoe Obozrenie, 50, 509 - 510." type="journal article" year="1971">Zaitzev, 1971</bibRefCitation>
</taxonomicName>
occurring in Middle Asia is similar to
<taxonomicName id="4C104D27FFEC5E31FDA6F7C9FCC255C1" box="[609,770,896,921]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="greatheadi">
<emphasis id="B964EAB6FFEC5E31FDA6F7C9FCC255C1" box="[609,770,896,921]" italics="true" pageId="7" pageNumber="8">E. greatheadi</emphasis>
</taxonomicName>
but is separated from Iranian species by yellow pedicel (dark brown in
<taxonomicName id="4C104D27FFEC5E31FE8AF7ECFE2C55E6" box="[333,492,933,958]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="greatheadi">
<emphasis id="B964EAB6FFEC5E31FE8AF7ECFE2C55E6" box="[333,492,933,958]" italics="true" pageId="7" pageNumber="8">E. greatheadi</emphasis>
</taxonomicName>
) and brown mark in the middle of scutellum (scutellum completely yellow in
<taxonomicName id="4C104D27FFEC5E31FA44F7ECFED855BB" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="greatheadi">
<emphasis id="B964EAB6FFEC5E31FA44F7ECFED855BB" italics="true" pageId="7" pageNumber="8">E. greatheadi</emphasis>
</taxonomicName>
).
<bibRefCitation id="EF814B55FFEC5E31FEF5F783FE2455BC" author="Zaitzev" box="[306,484,970,996]" pageId="7" pageNumber="8" refString="Zaitzev, V. F. (1971) New bombyliid species (Diptera, Bombyliidae) from Soviet central Asia and Mongolia. Entomologicheskoe Obozrenie, 50, 509 - 510." type="journal article" year="1971">Zaitzev (1971)</bibRefCitation>
did not illustrate the genitalia of his species so we are not able to compare the genitalia.
</paragraph>
</subSubSection>
<subSubSection id="C30A652FFFEC5E3FFF01F05CFAE854E4" lastPageId="9" lastPageNumber="10" pageId="7" pageNumber="8" type="description">
<paragraph id="8BAF36A4FFEC5E31FF01F05CFBF95277" blockId="7.[151,1438,261,1979]" box="[198,1081,1045,1071]" pageId="7" pageNumber="8">
<emphasis id="B964EAB6FFEC5E31FF01F05CFE9F5277" bold="true" box="[198,351,1045,1071]" pageId="7" pageNumber="8">Description.</emphasis>
Female. Length
<quantity id="4CE89B41FFEC5E31FDE2F05CFD4E5277" box="[549,654,1045,1071]" metricMagnitude="-3" metricUnit="m" metricValue="2.12" pageId="7" pageNumber="8" unit="mm" value="2.12">2.12 mm</quantity>
(Female) –
<quantity id="4CE89B41FFEC5E31FCDBF05CFC445277" box="[796,900,1045,1071]" metricMagnitude="-3" metricUnit="m" metricValue="1.63" pageId="7" pageNumber="8" unit="mm" value="1.63">1.63 mm</quantity>
(Male) (n=20).
</paragraph>
<paragraph id="8BAF36A4FFEC5E31FF02F073FD0D5302" blockId="7.[151,1438,261,1979]" pageId="7" pageNumber="8">
Head (
<figureCitation id="132B2A21FFEC5E31FED2F073FE9F520C" box="[277,351,1082,1108]" captionStart="FIGURE 7" captionStartId="15.[151,256,1963,1987]" captionTargetBox="[175,1410,194,1941]" captionTargetId="figure@15.[175,1410,194,1942]" captionTargetPageId="15" captionText="FIGURE 7. Lateral view of Empidideicus adults (females): a— E. amicus, b— E. ebellicus, c— E. greatheadi, d— E. matricarius, e— E. legulicoxa, f— E. persicus." httpUri="https://zenodo.org/record/198250/files/figure.png" pageId="7" pageNumber="8">Fig. 7</figureCitation>
c). Longer than high (1.3 times); eyes dichoptic, separated at vertex by 1.8 times distance between lateral ocelli; occiput and postgena black, with minute yellowish hairs; frons slightly depressed medially, yellow with semi-ellipsoid brown mark medially; face, oral margin, and mentum yellow; antennae (
<figureCitation id="132B2A21FFEC5E31FF58F0E3FF24529C" box="[159,228,1194,1220]" captionStart="FIGURE 3" captionStartId="8.[151,255,1968,1992]" captionTargetBox="[165,1418,198,1920]" captionTargetId="figure@8.[165,1422,170,1946]" captionTargetPageId="8" captionText="FIGURE 3. Empidideicus greatheadi: a—antennae lateral view, b—wing, c—phallic complex dorsal view, d—phallic complex lateral view, e—epandrium dorsal view, f—gonocoxa dorsal view, g—female genitalia." httpUri="https://zenodo.org/record/198247/files/figure.png" pageId="7" pageNumber="8">Fig. 3</figureCitation>
a) dark brown except scape yellow, set in deep pocket; scape short, subtrapezoidal; pedicel subconical, wider than long; first flagellomere oval, length 1.5 times greatest width; second flagellomere 0.6 times length of first flagellomere with minute, bullet-shaped, transparent apical style; clypeus yellow with brown area basally; about as long as labrum; proboscis brown; length 1.25 times head length; labrum sclerotised, stiff, pointed apically, length subequal to head length.
</paragraph>
<paragraph id="8BAF36A4FFEC5E31FF02F12CFC4E504C" blockId="7.[151,1438,261,1979]" pageId="7" pageNumber="8">
Thorax (
<figureCitation id="132B2A21FFEC5E31FEEEF12CFEAE5327" box="[297,366,1381,1407]" captionStart="FIGURE 8" captionStartId="16.[151,256,1295,1319]" captionTargetBox="[153,1436,189,1267]" captionTargetId="figure@16.[153,1438,189,1268]" captionTargetPageId="16" captionText="FIGURE 8. Dorsal view of Empidideicus mesonotum (females): a, b— E. amicus, c— E. ebellicus, d— E. greatheadi, e— E. matricarius, f— E. persicus." httpUri="https://zenodo.org/record/198251/files/figure.png" pageId="7" pageNumber="8">Fig. 8</figureCitation>
d). Slightly wider than head; mesonotum subshiny, yellow with 3 black longitudinal stripes, middle stripe extends from anterior margin to 4/5 length of mesonotum; lateral stripes extend from humeral callus to postalar callus; scutellum yellow; pleura yellow with black color on following: anepisternum as a crescent mark anteroventrally, lateral and ventral margins of anepimeron; lower three-fourths of katepisternum and meron; halter stem and knob yellowish white.
</paragraph>
<paragraph id="8BAF36A4FFEC5E31FF02F269FE925007" blockId="7.[151,1438,261,1979]" pageId="7" pageNumber="8">Legs. Yellow except basal three-fourths of femora faintly brown; tarsal segments except metatarsi brownish black.</paragraph>
<paragraph id="8BAF36A4FFEC5E31FF02F223FD835113" blockId="7.[151,1438,261,1979]" pageId="7" pageNumber="8">
Wings (
<figureCitation id="132B2A21FFEC5E31FEE3F223FEAC50DC" box="[292,364,1642,1668]" captionStart="FIGURE 3" captionStartId="8.[151,255,1968,1992]" captionTargetBox="[165,1418,198,1920]" captionTargetId="figure@8.[165,1422,170,1946]" captionTargetPageId="8" captionText="FIGURE 3. Empidideicus greatheadi: a—antennae lateral view, b—wing, c—phallic complex dorsal view, d—phallic complex lateral view, e—epandrium dorsal view, f—gonocoxa dorsal view, g—female genitalia." httpUri="https://zenodo.org/record/198247/files/figure.png" pageId="7" pageNumber="8">Fig. 3</figureCitation>
b). Hyaline; length (from base to end of R4+5) 2.7 times width (from end of R1 to CuA2); veins brown; costa, Sc, R1, R4+5 well sclerotised; veins M1, M2, M1+2 thin, less sclerotised; vein separating basal cells, base of Rs, and A1 evanescent; costa ends slightly beyond end of R4+5; M1 curved to wing margin; M2 slightly curved to wing margin; cell dm open; A1 straight to wing margin; anal cell open in wing margin by width subequal to r-m crossvein; fringe of hairs on posterior margin of wing shortest at wing tip becoming longer and more sparse toward base.
</paragraph>
<paragraph id="8BAF36A4FFEC5E31FF02F31FFCCB51E3" blockId="7.[151,1438,261,1979]" pageId="7" pageNumber="8">Abdomen. With minute sparse yellow hairs; tergites blackish brown with posterior margin widely yellow, yellow color extends much forward laterally and toward abdomen tip; sternites yellowish white except the last sternite with faintly brown areas basally and laterally.</paragraph>
<caption id="DF6F662CFFE35E3EFF50F3F9FB5C51BF" httpUri="https://zenodo.org/record/198247/files/figure.png" pageId="8" pageNumber="9" targetBox="[165,1418,198,1920]" targetPageId="8">
<paragraph id="8BAF36A4FFE35E3EFF50F3F9FB5C51BF" blockId="8.[151,1435,1968,2023]" pageId="8" pageNumber="9">
<emphasis id="B964EAB6FFE35E3EFF50F3F9FEDA5190" bold="true" box="[151,282,1968,1992]" pageId="8" pageNumber="9">FIGURE 3.</emphasis>
<taxonomicName id="4C104D27FFE35E3EFEE5F3F9FDF0519F" box="[290,560,1968,1991]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="greatheadi">
<emphasis id="B964EAB6FFE35E3EFEE5F3F9FDF0519F" box="[290,560,1968,1991]" italics="true" pageId="8" pageNumber="9">Empidideicus greatheadi</emphasis>
</taxonomicName>
: a—antennae lateral view, b—wing, c—phallic complex dorsal view, d—phallic complex lateral view, e—epandrium dorsal view, f—gonocoxa dorsal view, g—female genitalia.
</paragraph>
</caption>
<paragraph id="8BAF36A4FFE25E3FFF02F4D1FBE757CA" blockId="9.[151,1437,152,924]" pageId="9" pageNumber="10">
Genitalia (
<figureCitation id="132B2A21FFE25E3FFE81F4D1FE5056EA" box="[326,400,152,178]" captionStart="FIGURE 3" captionStartId="8.[151,255,1968,1992]" captionTargetBox="[165,1418,198,1920]" captionTargetId="figure@8.[165,1422,170,1946]" captionTargetPageId="8" captionText="FIGURE 3. Empidideicus greatheadi: a—antennae lateral view, b—wing, c—phallic complex dorsal view, d—phallic complex lateral view, e—epandrium dorsal view, f—gonocoxa dorsal view, g—female genitalia." httpUri="https://zenodo.org/record/198247/files/figure.png" pageId="9" pageNumber="10">
Fig.
<quantity id="4CE89B41FFE25E3FFE47F4D1FE5056EA" box="[384,400,152,178]" metricMagnitude="-3" metricUnit="kg" metricValue="3.0" pageId="9" pageNumber="10" unit="g" value="3.0">3</quantity>
</figureCitation>
g). Spermathecal reservoir subglobular, slightly wider than long, with shallow apical cylindrical invagination, width of invagination 0.4 times diameter of reservoir, its depth one-third reservoir height with floor puffed up; apical spermathecal duct membranous about 3.5 times reservoir height, basally slightly sclerotised and brownish; sperm pump striated, gradually narrowing toward base, with well sclerotised apical valve, 0.8 times apical duct in length; basal spermathecal duct membranous, subequal to reservoir height; common spermathecal duct absent; furca U-shaped; lateral arms broadening as triangular sclerotised plate apically, subapically with clavate processes oriented mesally.
</paragraph>
<paragraph id="8BAF36A4FFE25E3FFF02F5D4FA5C57EF" blockId="9.[151,1437,152,924]" box="[197,1436,413,439]" pageId="9" pageNumber="10">Male. Similar to female except darker and smaller; mesonotum with faint brown mark in prescutellar area.</paragraph>
<paragraph id="8BAF36A4FFE25E3FFF01F58BFAE854E4" blockId="9.[151,1437,152,924]" pageId="9" pageNumber="10">
Male genitalia (
<figureCitation id="132B2A21FFE25E3FFE47F58BFE105784" box="[384,464,450,476]" captionStart="FIGURE 3" captionStartId="8.[151,255,1968,1992]" captionTargetBox="[165,1418,198,1920]" captionTargetId="figure@8.[165,1422,170,1946]" captionTargetPageId="8" captionText="FIGURE 3. Empidideicus greatheadi: a—antennae lateral view, b—wing, c—phallic complex dorsal view, d—phallic complex lateral view, e—epandrium dorsal view, f—gonocoxa dorsal view, g—female genitalia." httpUri="https://zenodo.org/record/198247/files/figure.png" pageId="9" pageNumber="10">Figs. 3</figureCitation>
c–f). Epandrium yellow, dark brown basally, ellipsoid in dorsal view, with slightly pointed tip; Gonocoxa fused, well sclerotised marginally except anterior margin, with two triangular inwardly oriented long processes, gonostylus L-shaped, basal part subquadrate, apical part narrowly pointed apically; gonocoxal apodeme extends to tip of aedeagal apodeme; basal aedeagal apodeme with two paired rami basally; lateral apodeme one-third length of median apodeme; aedeagal bulb large, rounded with two small processes; lateral processes shorter than lateral apodemes; epiphallus consists of less sclerotised, apically rounded plate dorsally and serrate plate apicomarginally with a finger-shaped process apicomesally.
</paragraph>
</subSubSection>
<subSubSection id="C30A652FFFE25E3FFF02F68EFBF5555F" pageId="9" pageNumber="10" type="etymology">
<paragraph id="8BAF36A4FFE25E3FFF02F68EFBF5555F" blockId="9.[151,1437,152,924]" pageId="9" pageNumber="10">
<emphasis id="B964EAB6FFE25E3FFF02F68EFE9354B9" bold="true" box="[197,339,711,737]" pageId="9" pageNumber="10">Etymology.</emphasis>
The new species is named in the honor of the famous bee fly taxonomist, David Greathead, who passed away just a day after sending some of his papers to the first author.
</paragraph>
</subSubSection>
<subSubSection id="C30A652FFFE25E3FFF02F75BFE7555C4" pageId="9" pageNumber="10" type="distribution">
<paragraph id="8BAF36A4FFE25E3FFF02F75BFB455574" blockId="9.[151,1437,152,924]" box="[197,1157,786,812]" pageId="9" pageNumber="10">
<emphasis id="B964EAB6FFE25E3FFF02F75BFEA55574" bold="true" box="[197,357,786,812]" pageId="9" pageNumber="10">Distribution.</emphasis>
This species is currently known only from
<collectingCountry id="F3077634FFE25E3FFCA5F75BFC515574" box="[866,913,786,812]" name="Iran" pageId="9" pageNumber="10">Iran</collectingCountry>
(Ghazvin province).
</paragraph>
<paragraph id="8BAF36A4FFE25E3FFF01F77EFE7555C4" blockId="9.[151,1437,152,924]" pageId="9" pageNumber="10">
<emphasis id="B964EAB6FFE25E3FFF01F77EFE925509" bold="true" box="[198,338,823,849]" pageId="9" pageNumber="10">Variability.</emphasis>
Lateral black stripes on mesonotum sometimes joined to median stripe in anterior section or entire length so mesonotum seems completely black on disc with two large interhumeral marks and prescutellar area yellow.
</paragraph>
</subSubSection>
</treatment>
</document>
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