231 lines
24 KiB
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231 lines
24 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.6303238" ID-GBIF-Dataset="564eaad8-6245-4f42-9a88-42b97bf30807" ID-GBIF-Taxon="193278222" ID-Zenodo-Dep="6303238" approvedBy="tatiana" checkinTime="1644003985773" checkinUser="tatiana" docAuthor="Boudinot, Brendon E., Borowiec, Marek L. & Prebus, Matthew M." docDate="2022" docId="03B987A9436FFF9CFCA4FA32FAAED8CA" docLanguage="en" docName="systEnt.47.113-151.pdf.imf" docOrigin="Systematic Entomology 47" docTitle="Sussudio Boudinot & Borowiec & Prebus 2022, gen.nov." docType="treatment" docVersion="5" lastPageId="30" lastPageNumber="143" masterDocId="FF80FFD14372FF82FFF5FF95FF99D837" masterDocTitle="Phylogeny, evolution, and classification of the ant genus Lasius, the tribe Lasiini and the subfamily Formicinae (Hymenoptera: Formicidae)" masterLastPageNumber="151" masterPageNumber="113" pageId="29" pageNumber="142" updateTime="1646046328535" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Phylogeny, evolution, and classification of the ant genus Lasius, the tribe Lasiini and the subfamily Formicinae (Hymenoptera: Formicidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Boudinot, Brendon E.</mods:namePart>
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<mods:affiliation>Department of Entomology and Nematology, University of California, Davis CA, U. S. A. & Friedrich-Schiller-Universität Jena, Institut für Spezielle Zoologie, Jena, Germany</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Borowiec, Marek L.</mods:namePart>
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<mods:affiliation>Department of Entomology and Nematology, University of California, Davis CA, U. S. A. & Department of Plant Pathology, Entomology and Nematology, University of Idaho, Moscow ID, U. S. A. & Institute for Bioinformatics and Evolutionary Studies, University of Idaho, Moscow ID, U. S. A.</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Prebus, Matthew M.</mods:namePart>
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<mods:affiliation>Department of Entomology and Nematology, University of California, Davis CA, U. S. A. & School of Life Sciences, Arizona State University, Tempe AZ, U. S. A.</mods:affiliation>
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<mods:nameIdentifier type="email">mprebus@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Systematic Entomology</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-01-10</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>47</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>113</mods:start>
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<mods:end>151</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1111/syen.12522</mods:identifier>
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<mods:identifier type="GBIF-Dataset">564eaad8-6245-4f42-9a88-42b97bf30807</mods:identifier>
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<mods:identifier type="Zenodo-Dep">5975346</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6303238" ID-GBIF-Taxon="193278222" ID-Zenodo-Dep="6303238" LSID="urn:lsid:plazi:treatment:03B987A9436FFF9CFCA4FA32FAAED8CA" httpUri="http://treatment.plazi.org/id/03B987A9436FFF9CFCA4FA32FAAED8CA" lastPageId="30" lastPageNumber="143" pageId="29" pageNumber="142">
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<subSubSection box="[849,1110,1447,1469]" pageId="29" pageNumber="142" type="nomenclature">
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<paragraph blockId="29.[830,1437,276,1618]" box="[849,1110,1447,1469]" pageId="29" pageNumber="142">
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<emphasis bold="true" box="[849,1110,1447,1469]" pageId="29" pageNumber="142">
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Genus †
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<taxonomicName authority="Boudinot & Borowiec & Prebus, 2022" authorityName="Boudinot & Borowiec & Prebus" authorityYear="2022" box="[932,1021,1447,1468]" class="Insecta" family="Formicidae" genus="Sussudio" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="142" phylum="Arthropoda" rank="genus" status="gen.nov.">
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<emphasis bold="true" box="[932,1021,1447,1468]" italics="true" pageId="29" pageNumber="142">Sussudio</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[1027,1110,1447,1469]" pageId="29" pageNumber="142" rank="genus">gen.nov.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="29" pageNumber="142" type="type_taxon">
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<paragraph blockId="29.[830,1437,276,1618]" pageId="29" pageNumber="142">
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<emphasis box="[849,1094,1477,1498]" italics="true" pageId="29" pageNumber="142">Type and included species</emphasis>
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. †
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<taxonomicName authority="Wheeler, 1915" authorityName="Wheeler" authorityYear="1915" box="[1117,1349,1477,1498]" class="Insecta" family="Formicidae" genus="Sussudio" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="142" phylum="Arthropoda" rank="species" species="boreus">
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<emphasis box="[1117,1203,1477,1498]" italics="true" pageId="29" pageNumber="142">P. boreus</emphasis>
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<bibRefCitation author="Wheeler, W. M." box="[1207,1349,1477,1498]" journalOrPublisher="Schriften der Physikalisch-Okonomischen Gesellschaft zu Konigsberg" pageId="29" pageNumber="142" pagination="1 - 142" part="55" refId="ref39440" refString="Wheeler, W. M. (1915 1914) The ants of the Baltic Amber. Schriften der Physikalisch-Okonomischen Gesellschaft zu Konigsberg, 55, 1 - 142." title="The ants of the Baltic Amber" type="journal article" year="1915">Wheeler, 1915</bibRefCitation>
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</taxonomicName>
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. Original designation.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="30" lastPageNumber="143" pageId="29" pageNumber="142" type="description">
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<paragraph blockId="29.[830,1437,276,1618]" pageId="29" pageNumber="142">
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<emphasis box="[849,996,1537,1558]" italics="true" pageId="29" pageNumber="142">ZooBank LSID</emphasis>
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. http://zoobank.org/urn:lsid:zoobank.org:act: BFF515E7-9966-443F-8A34-555A4B41906B
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</paragraph>
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<paragraph blockId="29.[830,1437,276,1618]" box="[849,1040,1597,1618]" pageId="29" pageNumber="142">
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<emphasis box="[849,1034,1597,1618]" italics="true" pageId="29" pageNumber="142">Definition (worker)</emphasis>
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.
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</paragraph>
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<paragraph blockId="29.[844,1437,1661,1863]" box="[844,1291,1661,1682]" pageId="29" pageNumber="142">
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1. With characters of the
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<taxonomicName baseAuthorityName="Lepeletier de Saint-Fargeau" box="[1090,1199,1661,1682]" class="Insecta" family="Formicidae" genus="Formicinae" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="150" phylum="Arthropoda" rank="subFamily" subFamily="Formicinae">Formicinae</taxonomicName>
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(Note 1).
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</paragraph>
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<paragraph blockId="29.[844,1437,1661,1863]" pageId="29" pageNumber="142">
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2.
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<emphasis box="[873,1094,1692,1713]" italics="true" pageId="29" pageNumber="142">Cranium size variable</emphasis>
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,
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<emphasis italics="true" pageId="29" pageNumber="142">width somewhat broader than to about twice that of mesosoma</emphasis>
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(Note 2).
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</paragraph>
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<paragraph blockId="29.[844,1437,1661,1863]" box="[844,1223,1752,1773]" pageId="29" pageNumber="142">3. Mandible triangular, with 7–8 teeth.</paragraph>
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<paragraph blockId="29.[844,1437,1661,1863]" pageId="29" pageNumber="142">4. Dorsal mandibular groove extending along lateral mandibular margin in dorsal view.</paragraph>
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<paragraph blockId="29.[844,1437,1661,1863]" box="[844,1437,1842,1863]" pageId="29" pageNumber="142">5. Palps reduced, not reaching midlength of postgenal bridge.</paragraph>
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<paragraph blockId="30.[153,758,172,581]" box="[165,616,172,193]" pageId="30" pageNumber="143">6. Frontal carinae raised above antennal toruli.</paragraph>
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<paragraph blockId="30.[153,758,172,581]" box="[165,670,202,223]" pageId="30" pageNumber="143">7. Antennal toruli abutting posterior clypeal margin.</paragraph>
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<paragraph blockId="30.[153,758,172,581]" box="[165,571,232,253]" pageId="30" pageNumber="143">8. Antennomere count unknown (Note 3).</paragraph>
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<paragraph blockId="30.[153,758,172,581]" box="[165,536,262,283]" pageId="30" pageNumber="143">9. Antennomere III longer than broad.</paragraph>
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<paragraph blockId="30.[153,758,172,581]" box="[153,706,291,312]" pageId="30" pageNumber="143">10. Compound eyes situated posterior to head midlength.</paragraph>
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<paragraph blockId="30.[153,758,172,581]" box="[153,741,321,342]" pageId="30" pageNumber="143">11. Compound eyes not enormous and long axes subparallel.</paragraph>
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<paragraph blockId="30.[153,758,172,581]" box="[153,519,351,372]" pageId="30" pageNumber="143">12. Ocelli minutely present or absent.</paragraph>
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<paragraph blockId="30.[153,758,172,581]" box="[153,711,381,402]" pageId="30" pageNumber="143">13. Promesonotum domed, raised well above propodeum.</paragraph>
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<paragraph blockId="30.[153,758,172,581]" box="[153,523,411,432]" pageId="30" pageNumber="143">14. Metanotal groove well-developed.</paragraph>
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<paragraph blockId="30.[153,758,172,581]" pageId="30" pageNumber="143">15. Petiolar foramen in profile view clearly raised and margined with conspicuous thickened carina or lip.</paragraph>
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<paragraph blockId="30.[153,758,172,581]" pageId="30" pageNumber="143">16. Petiolar node squamiform, tall; dorsoventral height equal to or possibly exceeding dorsum of propodeum.</paragraph>
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<paragraph blockId="30.[153,758,172,581]" box="[153,751,560,581]" pageId="30" pageNumber="143">17. Abdominal segment III without raised tergosternal fusion.</paragraph>
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</subSubSection>
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<subSubSection pageId="30" pageNumber="143" type="discussion">
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<paragraph blockId="30.[151,759,618,1863]" box="[151,350,618,640]" pageId="30" pageNumber="143">
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<heading bold="true" box="[151,350,618,640]" fontSize="9" level="1" pageId="30" pageNumber="143" reason="6">
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<emphasis bold="true" box="[151,343,618,640]" pageId="30" pageNumber="143">Notes on definition</emphasis>
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:
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</heading>
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</paragraph>
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<paragraph blockId="30.[151,759,618,1863]" pageId="30" pageNumber="143">
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<emphasis box="[151,229,649,670]" italics="true" pageId="30" pageNumber="143">Note 1.</emphasis>
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Several characters were not possible to evaluate. These include the following: Reduction of third tooth from mandibular apex, palp formula, the metapleural gland, ventral length of the petiolar foramen, relative separation of the meso- and metacoxae, presence or absence of setal double-row on ventral tibial surfaces, petiolar apodeme conformation, cross-sectional shape of the petiolar sternum, and presence or absence of the posthelcial sulcus.
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</paragraph>
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<paragraph blockId="30.[151,759,618,1863]" pageId="30" pageNumber="143">
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<emphasis box="[151,222,887,908]" italics="true" pageId="30" pageNumber="143">Note 2.</emphasis>
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The massive head of †
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<taxonomicName authorityName="Wheeler" authorityYear="1915" box="[448,536,887,908]" class="Insecta" family="Formicidae" genus="Sussudio" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="143" phylum="Arthropoda" rank="species" species="boreus">
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<emphasis box="[448,536,887,908]" italics="true" pageId="30" pageNumber="143">P. boreus</emphasis>
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</taxonomicName>
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led
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<bibRefCitation author="Wheeler, W. M." box="[580,732,887,908]" journalOrPublisher="Schriften der Physikalisch-Okonomischen Gesellschaft zu Konigsberg" pageId="30" pageNumber="143" pagination="1 - 142" part="55" refId="ref39440" refString="Wheeler, W. M. (1915 1914) The ants of the Baltic Amber. Schriften der Physikalisch-Okonomischen Gesellschaft zu Konigsberg, 55, 1 - 142." title="The ants of the Baltic Amber" type="journal article" year="1915">Wheeler (1915)</bibRefCitation>
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to erroneously assume that the specimens were majors of
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<taxonomicName authorityName="Emery" authorityYear="1887" class="Insecta" family="Formicidae" genus="Pseudolasius" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="143" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="30" pageNumber="143">Pseudolasius</emphasis>
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</taxonomicName>
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; Wheeler did note the posteriorly set eyes of †
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<taxonomicName authorityName="Wheeler" authorityYear="1915" class="Insecta" family="Formicidae" genus="Sussudio" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="143" phylum="Arthropoda" rank="species" species="boreus">
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<emphasis italics="true" pageId="30" pageNumber="143">P. boreus</emphasis>
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</taxonomicName>
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, one of the distinctions between it and
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<taxonomicName authorityName="Emery" authorityYear="1887" box="[625,751,977,998]" class="Insecta" family="Formicidae" genus="Pseudolasius" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="143" phylum="Arthropoda" rank="genus">
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<emphasis box="[625,751,977,998]" italics="true" pageId="30" pageNumber="143">Pseudolasius</emphasis>
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</taxonomicName>
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, but vacillated based on the insufficient knowledge of the latter genus at that time.
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</paragraph>
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<paragraph blockId="30.[151,759,618,1863]" pageId="30" pageNumber="143">
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<emphasis box="[151,226,1066,1087]" italics="true" pageId="30" pageNumber="143">Note 3.</emphasis>
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The examined specimens are missing their terminal antennomeres, and no previous descriptions of †
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<taxonomicName authorityName="Wheeler" authorityYear="1915" box="[667,757,1096,1117]" class="Insecta" family="Formicidae" genus="Sussudio" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="143" phylum="Arthropoda" rank="species" species="boreus">
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<emphasis box="[667,757,1096,1117]" italics="true" pageId="30" pageNumber="143">P. boreus</emphasis>
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</taxonomicName>
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include an antennomere count.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="30" pageNumber="143" type="etymology">
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<paragraph blockId="30.[151,759,618,1863]" pageId="30" pageNumber="143">
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<emphasis box="[170,272,1156,1177]" italics="true" pageId="30" pageNumber="143">Etymology</emphasis>
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. Asimplification of ‘pseudo-
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<taxonomicName authorityName="Emery" authorityYear="1887" box="[623,749,1156,1177]" class="Insecta" family="Formicidae" genus="Pseudolasius" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="143" phylum="Arthropoda" rank="genus">
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<emphasis box="[623,749,1156,1177]" italics="true" pageId="30" pageNumber="143">Pseudolasius</emphasis>
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</taxonomicName>
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’ inspired by the musician Phil Collins. Masculine.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="30" pageNumber="143" type="discussion">
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<paragraph blockId="30.[151,759,618,1863]" lastBlockId="30.[821,1427,172,253]" pageId="30" pageNumber="143">
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<emphasis box="[170,271,1216,1237]" italics="true" pageId="30" pageNumber="143">Comments</emphasis>
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. †
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<taxonomicName authorityName="Boudinot & Borowiec & Prebus" authorityYear="2022" box="[305,470,1216,1237]" class="Insecta" family="Formicidae" genus="Sussudio" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="143" phylum="Arthropoda" rank="species" species="boreus" status="comb.nov.">
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<emphasis box="[305,470,1216,1237]" italics="true" pageId="30" pageNumber="143">Sussudio boreus</emphasis>
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</taxonomicName>
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<emphasis bold="true" box="[486,588,1215,1237]" pageId="30" pageNumber="143">
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<taxonomicNameLabel box="[486,588,1215,1237]" pageId="30" pageNumber="143" rank="species">comb.nov.</taxonomicNameLabel>
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</emphasis>
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was previously placed in the
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<taxonomicName authorityName="Ashmead" authorityYear="1905" box="[278,344,1245,1266]" class="Insecta" family="Formicidae" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="143" phylum="Arthropoda" rank="tribe" tribe="Lasiini">Lasiini</taxonomicName>
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(
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<bibRefCitation author="Emery, C." box="[356,481,1245,1266]" journalOrPublisher="Subfam. Formicinae. Genera Insectorum" pageId="30" pageNumber="143" pagination="1 - 302" part="183" refId="ref33484" refString="Emery, C. (1925) Hymenoptera. Fam. Formicidae. Subfam. Formicinae. Genera Insectorum, 183, 1 - 302." title="Hymenoptera. Fam. Formicidae" type="journal article" year="1925">Emery, 1925</bibRefCitation>
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,
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<bibRefCitation author="Dlussky, G. M. & Fedoseeva, E. B." bookContentInfo="232 pp." box="[491,753,1245,1266]" editor="Ponomarenko, A. G." journalOrPublisher="Moskva: Nauka" pageId="30" pageNumber="143" pagination="70 - 144" refId="ref33007" refString="Dlussky, G. M. & Fedoseeva, E. B. (1988) Origin and early stages of evolution in ants [in Russian]. Pp. 70 - 144 in: Ponomarenko, A. G. (ed.) Cretaceous Biocenotic Crisis and Insect Evolution. [in Russian.]. Moskva: Nauka, 232 pp." title="Origin and early stages of evolution in ants" type="book chapter" volumeTitle="Cretaceous Biocenotic Crisis and Insect Evolution" year="1988">Dlussky & Fedoseeva 1988</bibRefCitation>
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||
,
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<bibRefCitation author="Bolton, B." bookContentInfo="p. 222" box="[151,281,1275,1296]" journalOrPublisher="Harvard University Press, Cambridge" pageId="30" pageNumber="143" refId="ref31447" refString="Bolton, B. (1994) Identification Guide to the Ant Genera of the World, p. 222. Harvard University Press, Cambridge." title="Identification Guide to the Ant Genera of the World" type="book" year="1994">Bolton, 1994</bibRefCitation>
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), and has been considered to be a member of the genus
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<taxonomicName authorityName="Emery" authorityYear="1887" box="[213,339,1305,1326]" class="Insecta" family="Formicidae" genus="Pseudolasius" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="143" phylum="Arthropoda" rank="genus">
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<emphasis box="[213,339,1305,1326]" italics="true" pageId="30" pageNumber="143">Pseudolasius</emphasis>
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</taxonomicName>
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||
(
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||
<bibRefCitation author="Wheeler, W. M." box="[351,495,1305,1326]" journalOrPublisher="Schriften der Physikalisch-Okonomischen Gesellschaft zu Konigsberg" pageId="30" pageNumber="143" pagination="1 - 142" part="55" refId="ref39440" refString="Wheeler, W. M. (1915 1914) The ants of the Baltic Amber. Schriften der Physikalisch-Okonomischen Gesellschaft zu Konigsberg, 55, 1 - 142." title="The ants of the Baltic Amber" type="journal article" year="1915">Wheeler, 1915</bibRefCitation>
|
||
;
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<bibRefCitation author="LaPolla, J. S. & Dlussky, G. M." box="[504,747,1305,1326]" journalOrPublisher="Proceedings of the Entomological Society of Washington" pageId="30" pageNumber="143" pagination="258 - 273" part="112" refId="ref35305" refString="LaPolla, J. S. & Dlussky, G. M. (2010) Review of fossil Prenolepis genus-group species (Hymenoptera: Formicidae). Proceedings of the Entomological Society of Washington, 112, 258 - 273." title="Review of fossil Prenolepis genus-group species (Hymenoptera: Formicidae)" type="journal article" year="2010">LaPolla & Dlussky, 2010</bibRefCitation>
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). Based on an examination of a syntype (GZG-BST04646) and a nontype (NHMW1984-31-211) worker, as well as the original description from
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<bibRefCitation author="Wheeler, W. M." box="[323,479,1394,1415]" journalOrPublisher="Schriften der Physikalisch-Okonomischen Gesellschaft zu Konigsberg" pageId="30" pageNumber="143" pagination="1 - 142" part="55" refId="ref39440" refString="Wheeler, W. M. (1915 1914) The ants of the Baltic Amber. Schriften der Physikalisch-Okonomischen Gesellschaft zu Konigsberg, 55, 1 - 142." title="The ants of the Baltic Amber" type="journal article" year="1915">Wheeler (1915)</bibRefCitation>
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, we reject both placements. Neither specimen can be mistaken due to the unique combination of characters they display, including setation, high petiolar nodes, massive crania, and mesosomal form. †
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<taxonomicName authorityName="Boudinot & Borowiec & Prebus" authorityYear="2022" box="[601,757,1484,1505]" class="Insecta" family="Formicidae" genus="Sussudio" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="143" phylum="Arthropoda" rank="species" species="boreus">
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<emphasis box="[601,757,1484,1505]" italics="true" pageId="30" pageNumber="143">Sussudio boreus</emphasis>
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</taxonomicName>
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does not display the modifications of the third abdominal segment characteristic of the
|
||
<taxonomicName box="[459,562,1544,1565]" class="Insecta" family="Formicidae" genus="Prenolepis" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="143" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[459,562,1544,1565]" italics="true" pageId="30" pageNumber="143">Prenolepis</emphasis>
|
||
</taxonomicName>
|
||
genus group, also rejecting its original placement in
|
||
<taxonomicName authorityName="Emery" authorityYear="1887" box="[481,607,1573,1594]" class="Insecta" family="Formicidae" genus="Pseudolasius" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="143" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[481,607,1573,1594]" italics="true" pageId="30" pageNumber="143">Pseudolasius</emphasis>
|
||
</taxonomicName>
|
||
(see diagnostic Note 2 above). †
|
||
<taxonomicName authorityName="Boudinot & Borowiec & Prebus" authorityYear="2022" box="[316,401,1603,1624]" class="Insecta" family="Formicidae" genus="Sussudio" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="143" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[316,401,1603,1624]" italics="true" pageId="30" pageNumber="143">Sussudio</emphasis>
|
||
</taxonomicName>
|
||
is excluded from the
|
||
<taxonomicName authorityName="Ashmead" authorityYear="1905" box="[617,683,1603,1624]" class="Insecta" family="Formicidae" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="143" phylum="Arthropoda" rank="tribe" tribe="Lasiini">Lasiini</taxonomicName>
|
||
overall by two additional characters: (i) The petiolar node, which is completely vertical and very tall dorsoventrally, is as tall as or possibly taller than the propodeum; and (ii) the raised and conspicuously lipped petiolar foramen. No lasiine, extant or extinct, has such a tall petiolar node, nor do any species display the petiolar foramen conformation observed in †
|
||
<taxonomicName authorityName="Boudinot & Borowiec & Prebus" authorityYear="2022" box="[627,712,1782,1803]" class="Insecta" family="Formicidae" genus="Sussudio" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="143" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[627,712,1782,1803]" italics="true" pageId="30" pageNumber="143">Sussudio</emphasis>
|
||
</taxonomicName>
|
||
. We cannot confidently place †
|
||
<taxonomicName authorityName="Boudinot & Borowiec & Prebus" authorityYear="2022" box="[412,497,1812,1833]" class="Insecta" family="Formicidae" genus="Sussudio" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="143" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[412,497,1812,1833]" italics="true" pageId="30" pageNumber="143">Sussudio</emphasis>
|
||
</taxonomicName>
|
||
in any extant tribe, therefore, we consider †
|
||
<emphasis box="[342,570,1842,1863]" italics="true" pageId="30" pageNumber="143">
|
||
<taxonomicName authority="Boudinot & Borowiec & Prebus, 2022" authorityName="Boudinot & Borowiec & Prebus" authorityYear="2022" box="[342,427,1842,1863]" class="Insecta" family="Formicidae" genus="Sussudio" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="143" phylum="Arthropoda" rank="genus" status="incertae sedis">Sussudio</taxonomicName>
|
||
<taxonomicNameLabel box="[435,570,1842,1863]" pageId="30" pageNumber="143">incertae sedis</taxonomicNameLabel>
|
||
</emphasis>
|
||
in the
|
||
<taxonomicName baseAuthorityName="Lepeletier de Saint-Fargeau" box="[643,755,1842,1863]" class="Insecta" family="Formicidae" genus="Formicinae" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="150" phylum="Arthropoda" rank="subFamily" subFamily="Formicinae">Formicinae</taxonomicName>
|
||
. As with other fossil taxa, which are of uncertain placement in the
|
||
<taxonomicName baseAuthorityName="Lepeletier de Saint-Fargeau" box="[858,969,202,223]" class="Insecta" family="Formicidae" genus="Formicinae" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="150" phylum="Arthropoda" rank="subFamily" subFamily="Formicinae">Formicinae</taxonomicName>
|
||
, micro-CT scanning could be used to evaluate the form of the helcium and third abdominal sternum.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |