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<document id="1F248F1BFA0164E9CDFDA2703F5D4BBF" ID-CLB-Dataset="7001" ID-DOI="10.11646/zootaxa.4965.3.1" ID-GBIF-Dataset="6cadad22-abf2-4abe-9165-156d5c63ece3" ID-ISSN="1175-5326" ID-Zenodo-Dep="4752531" ID-ZooBank="94FB89CF-2083-4FAC-AE60-B8CCF1D5FE8E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1619690781738" checkinUser="plazi" docAuthor="Bennett, Robb, Copley, Claudia &amp; Copley, Darren" docDate="2021" docId="038B87C31913BE3FFF3A6BF2FE5D9BAA" docLanguage="en" docName="zootaxa.4965.3.1.pdf" docOrigin="Zootaxa 4965 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cybaeus penedentatus Bennett 2009" docType="treatment" docVersion="7" lastPageNumber="417" masterDocId="FFB2FFBB1903BE2FFFAD6F15FFC49C5B" masterDocTitle="Cybaeus (Araneae: Cybaeidae): the consocius species group of the Californian clade" masterLastPageNumber="436" masterPageNumber="401" pageNumber="417" updateTime="1698937978772" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="3FA9684DA373AB6E1B7AA4B774A3010D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7CA9DCC806E0D4C4CDA187C019B48566">
<mods:title id="97693AD7E9D24D98C1DE9840B52F71A7">Cybaeus (Araneae: Cybaeidae): the consocius species group of the Californian clade</mods:title>
</mods:titleInfo>
<mods:name id="06649E5237CB889AE85AF52F1391E861" type="personal">
<mods:role id="BBB8F0DF1E1DC85B7862507A1E0DE21B">
<mods:roleTerm id="AF7B5F4A27DD8F6F0261B44FED9139B1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="004FF6736DAC822F8B35815E4A8AC1E4">Bennett, Robb</mods:namePart>
<mods:nameIdentifier id="8547A71714864039F28E57752C29F93E" type="ORCID">0000-0002-6587-7079</mods:nameIdentifier>
<mods:nameIdentifier id="4354D60ADFE4AC01B135A41EA72BA5CE" type="email">ennett@shaw.ca</mods:nameIdentifier>
</mods:name>
<mods:name id="77D80C880CD0CEF9EF74A50C01539E59" type="personal">
<mods:role id="D11852B504C409310EFEC6944C124B0A">
<mods:roleTerm id="FCA73C80B870837807066886FE22F3B5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EB04B7769BBFEB4E5C6D3E0280FBC645">Copley, Claudia</mods:namePart>
<mods:nameIdentifier id="0701363C4DB258461336AF3E8C6A9BDC" type="ORCID">0000-0002-8184-5819</mods:nameIdentifier>
<mods:affiliation id="26BAECBDEFF3DA98B19BDE4577C9E7C3">ccopley @ royalbcmuseum. bc. ca; https: // orcid. org / 0000 - 0002 - 8184 - 5819</mods:affiliation>
<mods:nameIdentifier id="2B216E4413339B03508732315D77E731" type="email">ccopley@royalbcmuseum.bc.ca</mods:nameIdentifier>
</mods:name>
<mods:name id="1A6F2509E6AFA44AC04296620AAF1D81" type="personal">
<mods:role id="0102C8F90A7666E2FB91EDA73ADA646D">
<mods:roleTerm id="0B7231AAFD4BC7A17EA2937C8AE55C54">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1BE39DD1A042127B9A7B27B3E64FD4A9">Copley, Darren</mods:namePart>
<mods:nameIdentifier id="93E9D0ACB35C7AAD88D04B32ECF1D915" type="ORCID">0000-0002-1944-4272</mods:nameIdentifier>
<mods:affiliation id="2B7D342D7FE2E9D7D76866A6F5AE6BE6">dcopley @ royalbcmuseum. bc. ca; https: // orcid. org / 0000 - 0002 - 1944 - 4272</mods:affiliation>
<mods:nameIdentifier id="74CA84E01E31BFE2BD810E53DF8E9ACC" type="email">dcopley@royalbcmuseum.bc.ca</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="1BF36AD5E62ECDC510D91857D7F98717">text</mods:typeOfResource>
<mods:relatedItem id="08FBC90CA2AE456FDB4405089A172EDB" type="host">
<mods:titleInfo id="49727B21116D91780502EB59707740B9">
<mods:title id="0EABAF441275075A2330B5536F960838">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="1C8008B8AB082F5B3D111B14F4E58CE4">
<mods:date id="BBEA3D5CA966ED5C460774F9735C52A0">2021</mods:date>
<mods:detail id="A40BEEFE48152E3A1D702FBEFBC1C5E6" type="pubDate">
<mods:number id="535F1786A871DD8520D067D3BC3342D6">2021-04-29</mods:number>
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<mods:detail id="02237D225DB3CFA0BFF2F338E79DD260" type="volume">
<mods:number id="AE474A158ABCFCA20DE2EB7B131F2CDE">4965</mods:number>
</mods:detail>
<mods:detail id="C06711CE463CE082FCFB9BD51C934DE8" type="issue">
<mods:number id="709D435D121F1569CF6191621ABAE944">3</mods:number>
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<mods:start id="DC754EBDA7F25F665831084A3C1A5265">401</mods:start>
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<mods:classification id="799521CACF331EAFF8629622E657E2B6">journal article</mods:classification>
<mods:identifier id="26FCF1777F5B7E314858393F81FEA8BF" type="CLB-Dataset">7001</mods:identifier>
<mods:identifier id="06354CA3A4F6D558B2050346ED9BBD16" type="DOI">10.11646/zootaxa.4965.3.1</mods:identifier>
<mods:identifier id="FC5BD1D30B926CDCF01F04F62916A679" type="GBIF-Dataset">6cadad22-abf2-4abe-9165-156d5c63ece3</mods:identifier>
<mods:identifier id="65FB9E5EAC38FFB1BB81182A16CBA823" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
<treatment id="038B87C31913BE3FFF3A6BF2FE5D9BAA" ID-DOI="http://doi.org/10.5281/zenodo.4752660" ID-GBIF-Taxon="181983460" ID-Zenodo-Dep="4752660" LSID="urn:lsid:plazi:treatment:038B87C31913BE3FFF3A6BF2FE5D9BAA" httpUri="http://treatment.plazi.org/id/038B87C31913BE3FFF3A6BF2FE5D9BAA" lastPageNumber="417" pageId="16" pageNumber="417">
<subSubSection id="C338655E1913BE3FFF3A6BF2FDCA9959" box="[151,526,1255,1282]" pageId="16" pageNumber="417" type="nomenclature">
<paragraph id="8B9D36D51913BE3FFF3A6BF2FDCA9959" blockId="16.[151,526,1255,1319]" box="[151,526,1255,1282]" pageId="16" pageNumber="417">
<heading id="D0D581B91913BE3FFF3A6BF2FDCA9959" bold="true" box="[151,526,1255,1282]" fontSize="11" level="1" pageId="16" pageNumber="417" reason="1">
<taxonomicName id="4C224D561913BE3FFF3A6BF2FDCA9959" ID-CoL="32RGT" authority="Bennett" authorityName="Bennett" authorityYear="2009" box="[151,526,1255,1282]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="penedentatus">
<emphasis id="B956EAC71913BE3FFF3A6BF2FDCA9959" bold="true" box="[151,526,1255,1282]" pageId="16" pageNumber="417">
<emphasis id="B956EAC71913BE3FFF3A6BF2FE619959" bold="true" box="[151,421,1255,1282]" italics="true" pageId="16" pageNumber="417">Cybaeus penedentatus</emphasis>
Bennett
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C338655E1913BE3FFF3A6A19FC4B9932" pageId="16" pageNumber="417" type="description">
<paragraph id="8B9D36D51913BE3FFF3A6A19FEAC997D" blockId="16.[151,526,1255,1319]" box="[151,360,1292,1319]" pageId="16" pageNumber="417">
<figureCitation id="13192A501913BE3FFF3A6A19FED3997C" box="[151,279,1292,1319]" captionStart-0="FIGURES 42–45" captionStart-1="FIGURES 46–51" captionStart-2="FIGURES 52–54" captionStartId-0="17.[151,264,1806,1830]" captionStartId-1="18.[151,264,1787,1811]" captionStartId-2="19.[151,264,1076,1100]" captionTargetBox-0="[151,1419,155,1775]" captionTargetBox-1="[151,1433,181,1759]" captionTargetBox-2="[176,1401,188,1050]" captionTargetId-0="figure-17@17.[151,1436,151,1783]" captionTargetId-1="figure-27@18.[151,1437,181,1763]" captionTargetId-2="figure-17@19.[176,1411,181,1052]" captionTargetPageId-0="17" captionTargetPageId-1="18" captionTargetPageId-2="19" captionText-0="FIGURES 42–45. Cybaeus penedentatus, male from Holy City, California, left palpus. 42–43 Patella and tibia (42 retrolateral, 43 dorsal). 44–45 Genital bulb (44 ventral, 45 retrolateral). PA—proximal arm of tegular apophysis. PTA—patellar apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrows indicate ventral keel of proximal arm of tegular apophysis." captionText-1="FIGURES 46–51. Cybaeus penedentatus, females from California, copulatory organ (46–48 from Ben Lomond, 49 from Pescadero Creek, 50–51 from Mount Madonna). 46 Epigynum, ventral. 47–51 Vulva (47, 49–50 ventral; 48, 51 dorsal). AT—atrium." captionText-2="FIGURES 52–54. Cybaeus penedentatus, female from Brookdale, California, copulatory organ. 52 Epigynum, ventral. 53–54 Vulva (53 ventral, 54 dorsal). AT—atrium." figureDoi-0="http://doi.org/10.5281/zenodo.4752582" figureDoi-1="http://doi.org/10.5281/zenodo.4752586" figureDoi-2="http://doi.org/10.5281/zenodo.4752592" httpUri-0="https://zenodo.org/record/4752582/files/figure.png" httpUri-1="https://zenodo.org/record/4752586/files/figure.png" httpUri-2="https://zenodo.org/record/4752592/files/figure.png" pageId="16" pageNumber="417">Figs 42–54</figureCitation>
,
<figureCitation id="13192A501913BE3FFE8F6A19FE84997C" box="[290,320,1292,1319]" captionStart="FIGURES 55–58" captionStartId="20.[151,264,1895,1919]" captionTargetBox="[177,1409,381,1872]" captionTargetId="figure-139@20.[177,1411,381,1872]" captionTargetPageId="20" captionText="FIGURES 55–58. Cybaeus spp. males from California, left palpus, proximal arm of tegular apophysis (55 C. opulentus spec. nov. from Riverton, 56 C. penedentatus from Holy City, 57 C. vulpinus from Columbia, 58 C. pan from Calpine) (55–56 ventral, 57–58 prolateral). Arrows indicate diagnostic characters (55 slightly twisted tip and prominent basal protrusion; 56 acuminate slightly twisted tip and longitudinal ventral keel; 57 smoothly curved, twisted tip; 58 triangular keel dorsally on tip)." figureDoi="http://doi.org/10.5281/zenodo.4752596" httpUri="https://zenodo.org/record/4752596/files/figure.png" pageId="16" pageNumber="417">56</figureCitation>
,
<figureCitation id="13192A501913BE3FFEE16A19FEAC997D" box="[332,360,1292,1318]" captionStart="FIGURE 84" captionStartId="28.[151,250,1943,1967]" captionTargetBox="[164,1421,188,1913]" captionTargetId="figure-27@28.[158,1429,181,1919]" captionTargetPageId="28" captionText="FIGURE 84. Distribution of Cybaeus consocius, C. hesper, C. hummeli spec. nov., C. penedentatus, and C. ubicki spec. nov. AZ—Arizona, CA—California, ID—Idaho, OR—Oregon, NV—Nevada, UT—Utah, WA—Washington. California county names capitalized." figureDoi="http://doi.org/10.5281/zenodo.4752622" httpUri="https://zenodo.org/record/4752622/files/figure.png" pageId="16" pageNumber="417">84</figureCitation>
</paragraph>
<paragraph id="8B9D36D51913BE3FFF3A6A44FC4B9932" blockId="16.[151,911,1360,1385]" box="[151,911,1360,1385]" pageId="16" pageNumber="417">
Bennett in
<bibRefCitation id="EFB34B241913BE3FFEAB6A45FE339933" author="Copley, C. R. &amp; Bennett, R. &amp; Perlman, S. J." box="[262,503,1360,1385]" pageId="16" pageNumber="417" pagination="367 - 401" refId="ref18027" refString="Copley, C. R., Bennett, R. &amp; Perlman, S. J. (2009) Systematics of Nearctic Cybaeus (Araneae: Cybaeidae). Invertebrate Systematics, 23, 367 - 401. https: // doi. org / 10.1071 / IS 09001" type="journal article" year="2009">
Copley
<emphasis id="B956EAC71913BE3FFEFB6A44FE4D9933" box="[342,393,1360,1384]" italics="true" pageId="16" pageNumber="417">et al.</emphasis>
2009: 386
</bibRefCitation>
, figs 67–78.
<bibRefCitation id="EFB34B241913BE3FFDD46A45FC4B9932" author="World Spider Catalog" box="[633,911,1360,1385]" pageId="16" pageNumber="417" refId="ref20275" refString="World Spider Catalog (2021) World Spider Catalog. Version 22.0. Natural History Museum, Bern. Available from: http: // wsc. nmbe. ch, (accessed 29 January 2021) https: // doi. org / 10.24436 / 2" type="book" year="2021">World Spider Catalog 2021</bibRefCitation>
</paragraph>
</subSubSection>
<subSubSection id="C338655E1913BE3FFF3A6A82FCAD9A1A" pageId="16" pageNumber="417" type="materials_examined">
<paragraph id="8B9D36D51913BE3FFF3A6A82FF2D99A2" blockId="16.[151,1437,1430,2033]" pageId="16" pageNumber="417">
<emphasis id="B956EAC71913BE3FFF3A6A82FE0799EB" bold="true" box="[151,451,1430,1457]" pageId="16" pageNumber="417">Type material examined.</emphasis>
<materialsCitation id="3B4A3C881913BE3FFE616A83FCC5998E" ID-GBIF-Occurrence="3110777326" collectingDate="1966-12-06" collectionCode="V, CAS" collectorName="D. Roth &amp; Ferguson" country="United States of America" county="Santa Clara County" location="Highway" municipality="Holy City" pageId="16" pageNumber="417" specimenCount="1" stateProvince="California" typeStatus="holotype">
<emphasis id="B956EAC71913BE3FFE616A83FCA299EA" bold="true" box="[460,870,1430,1457]" pageId="16" pageNumber="417">
<collectingCountry id="F33576451913BE3FFE616A83FDD999EB" box="[460,541,1430,1456]" name="United States of America" pageId="16" pageNumber="417">U.S.A.</collectingCountry>
:
<emphasis id="B956EAC71913BE3FFD816A83FD6099EA" bold="true" box="[556,676,1430,1457]" italics="true" pageId="16" pageNumber="417">
<collectingRegion id="49E6F8371913BE3FFD816A83FD6099EA" box="[556,676,1430,1457]" country="United States of America" name="California" pageId="16" pageNumber="417">California</collectingRegion>
</emphasis>
:
<typeStatus id="549988771913BE3FFD1B6A82FCE799EA" box="[694,803,1431,1457]" pageId="16" pageNumber="417" type="holotype">Holotype</typeStatus>
male
</emphasis>
.
<collectingCounty id="62FC4E591913BE3FFCDB6A82FB9399EA" box="[886,1111,1431,1457]" pageId="16" pageNumber="417">Santa Clara County</collectingCounty>
,
<quantity id="4CDA9B301913BE3FFBC96A82FB7C99EA" box="[1124,1208,1431,1457]" metricMagnitude="3" metricUnit="m" metricValue="3.218688" pageId="16" pageNumber="417" unit="mi" value="2.0">2 miles</quantity>
north of
<collectingMunicipality id="6BF9ACAF1913BE3FFA8B6A82FA5C99EA" box="[1318,1432,1431,1457]" pageId="16" pageNumber="417">Holy City</collectingMunicipality>
,
<location id="8EFD600E1913BE3FFF3A6AAEFF3A998E" LSID="urn:lsid:plazi:treatment:038B87C31913BE3FFF3A6BF2FE5D9BAA:8EFD600E1913BE3FFF3A6AAEFF3A998E" box="[151,254,1467,1493]" country="United States of America" county="Santa Clara County" municipality="Holy City" name="Highway" pageId="16" pageNumber="417" stateProvince="California">Highway</location>
17,
<date id="FF9C10151913BE3FFE836AAEFE66998E" box="[302,418,1467,1493]" pageId="16" pageNumber="417" value="1966-12-06">
<collectingDate id="EFD8E9FD1913BE3FFE836AAEFE66998E" box="[302,418,1467,1493]" pageId="16" pageNumber="417" value="1966-12-06">6.xii.1966</collectingDate>
</date>
,
<collectionCode id="ED33AE101913BE3FFE006AAEFE05998E" box="[429,449,1467,1493]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="16" pageNumber="417" type="Museum">V</collectionCode>
.
<collectorName id="26D753031913BE3FFE666AAEFDE7998E" box="[459,547,1467,1493]" pageId="16" pageNumber="417">D. Roth</collectorName>
&amp;
<collectorName id="26D753031913BE3FFDEA6AAEFD6B998E" box="[583,687,1467,1493]" pageId="16" pageNumber="417">Ferguson</collectorName>
(
<collectionCode id="ED33AE101913BE3FFD106AAEFD3D998E" box="[701,761,1467,1493]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="16" pageNumber="417" type="Museum">CAS</collectionCode>
)
</materialsCitation>
.
<emphasis id="B956EAC71913BE3FFCA16AAEFC49998E" bold="true" box="[780,909,1467,1493]" pageId="16" pageNumber="417">
<typeStatus id="549988771913BE3FFCA16AAEFC4C998E" box="[780,904,1467,1493]" pageId="16" pageNumber="417" type="paratype">Paratypes</typeStatus>
:
</emphasis>
Specimens and locality data as in
<bibRefCitation id="EFB34B241913BE3FFAA06AAEFF2199A2" author="Copley, C. R. &amp; Bennett, R. &amp; Perlman, S. J." pageId="16" pageNumber="417" pagination="367 - 401" refId="ref18027" refString="Copley, C. R., Bennett, R. &amp; Perlman, S. J. (2009) Systematics of Nearctic Cybaeus (Araneae: Cybaeidae). Invertebrate Systematics, 23, 367 - 401. https: // doi. org / 10.1071 / IS 09001" type="journal article" year="2009">
Copley
<emphasis id="B956EAC71913BE3FFAC86AAEFA59998E" box="[1381,1437,1467,1493]" italics="true" pageId="16" pageNumber="417">et al.</emphasis>
(2009)
</bibRefCitation>
.
</paragraph>
<paragraph id="8B9D36D51913BE3FFF6A6917FCAD9A1A" blockId="16.[151,1437,1430,2033]" pageId="16" pageNumber="417">
<emphasis id="B956EAC71913BE3FFF6A6917FE3E9A47" bold="true" box="[199,506,1538,1564]" pageId="16" pageNumber="417">Other material examined.</emphasis>
<materialsCitation id="3B4A3C881913BE3FFDAD6917FDF89A1A" ID-GBIF-Occurrence="3110777373" collectingDate="1985-02-24" collectingDateMax="1997-01-05" collectingDateMin="1985-02-24" collectionCode="CAS, V" collectorName="D. Ubick" country="United States of America" location="San Juan Grade Rd." pageId="16" pageNumber="417" specimenCount="4" specimenCount-female="4" stateProvince="California">
<emphasis id="B956EAC71913BE3FFDAD6917FD1A9A46" bold="true" box="[512,734,1538,1565]" pageId="16" pageNumber="417">
<collectingCountry id="F33576451913BE3FFDAD6917FD959A47" box="[512,593,1538,1564]" name="United States of America" pageId="16" pageNumber="417">U.S.A.</collectingCountry>
:
<emphasis id="B956EAC71913BE3FFDF06917FD119A46" bold="true" box="[605,725,1538,1565]" italics="true" pageId="16" pageNumber="417">
<collectingRegion id="49E6F8371913BE3FFDF06917FD119A46" box="[605,725,1538,1565]" country="United States of America" name="California" pageId="16" pageNumber="417">California</collectingRegion>
</emphasis>
:
</emphasis>
Monterey.
<specimenCount id="9D24FD5C1913BE3FFCF16916FC459A46" box="[860,897,1539,1565]" pageId="16" pageNumber="417" type="female">2♀</specimenCount>
,
<location id="8EFD600E1913BE3FFC206916FBAF9A46" LSID="urn:lsid:plazi:treatment:038B87C31913BE3FFF3A6BF2FE5D9BAA:8EFD600E1913BE3FFC206916FBAF9A46" box="[909,1131,1539,1565]" country="United States of America" name="San Juan Grade Rd." pageId="16" pageNumber="417" stateProvince="California">San Juan Grade Rd.</location>
,
<date id="FF9C10151913BE3FFBDB6917FB109A46" box="[1142,1236,1538,1565]" pageId="16" pageNumber="417" value="1997-01-05">
<collectingDate id="EFD8E9FD1913BE3FFBDB6917FB109A46" box="[1142,1236,1538,1565]" pageId="16" pageNumber="417" value="1997-01-05">5.i.1997</collectingDate>
</date>
,
<collectorName id="26D753031913BE3FFB726916FA819A46" box="[1247,1349,1539,1565]" pageId="16" pageNumber="417">D. Ubick</collectorName>
(
<collectionCode id="ED33AE101913BE3FFAF96916FA549A46" box="[1364,1424,1539,1565]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="16" pageNumber="417" type="Museum">CAS</collectionCode>
); Santa Cruz.
<specimenCount id="9D24FD5C1913BE3FFE8C6932FE819A1A" box="[289,325,1575,1601]" pageId="16" pageNumber="417" type="female">2♀</specimenCount>
, Felton,
<date id="FF9C10151913BE3FFE0A6933FDDF9A1A" box="[423,539,1574,1601]" pageId="16" pageNumber="417" value="1985-02-24">
<collectingDate id="EFD8E9FD1913BE3FFE0A6933FDDF9A1A" box="[423,539,1574,1601]" pageId="16" pageNumber="417" value="1985-02-24">24.ii.1985</collectingDate>
</date>
,
<collectionCode id="ED33AE101913BE3FFD8B6932FDF89A1A" box="[550,572,1575,1601]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="16" pageNumber="417" type="Museum">V</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B4A3C881913BE3FFDED6932FCA19A1A" ID-GBIF-Occurrence="3110777350" box="[576,869,1575,1601]" collectionCode="CAS" collectorName="D. Roth &amp; B. Roth" country="United States of America" pageId="16" pageNumber="417" specimenCount="1" stateProvince="California">
<collectorName id="26D753031913BE3FFDED6932FD529A1A" box="[576,662,1575,1601]" pageId="16" pageNumber="417">D. Roth</collectorName>
&amp;
<collectorName id="26D753031913BE3FFD176932FCD59A1A" box="[698,785,1575,1601]" pageId="16" pageNumber="417">B. Roth</collectorName>
(
<collectionCode id="ED33AE101913BE3FFC8C6932FC999A1A" box="[801,861,1575,1601]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="16" pageNumber="417" type="Museum">CAS</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C338655E1913BE3FFF6A695FFE3D9B3A" pageId="16" pageNumber="417" type="diagnosis">
<paragraph id="8B9D36D51913BE3FFF6A695FFE3D9B3A" blockId="16.[151,1437,1430,2033]" pageId="16" pageNumber="417">
<emphasis id="B956EAC71913BE3FFF6A695FFE869A3F" bold="true" box="[199,322,1610,1636]" pageId="16" pageNumber="417">Diagnosis.</emphasis>
Both sexes of
<taxonomicName id="4C224D561913BE3FFE5E695EFD6E9A3E" authorityName="Bennett" authorityYear="2009" box="[499,682,1611,1637]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="penedentatus">
<emphasis id="B956EAC71913BE3FFE5E695EFD6E9A3E" box="[499,682,1611,1637]" italics="true" pageId="16" pageNumber="417">C. penedentatus</emphasis>
</taxonomicName>
are distinctive and unlikely to be confused with any of the other species in the
<taxonomicName id="4C224D561913BE3FFE97697AFE6C9AD3" authorityName="Chamberlin &amp; Ivie" authorityYear="1932" box="[314,424,1647,1672]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="consocius">
<emphasis id="B956EAC71913BE3FFE97697AFE6C9AD3" box="[314,424,1647,1672]" italics="true" pageId="16" pageNumber="417">consocius</emphasis>
</taxonomicName>
group. The male is diagnosed by its elongate patellar apophysis (as long as the width of the patella) with very few peg setae (
<figureCitation id="13192A501913BE3FFD936986FD549AF6" box="[574,656,1683,1709]" captionStart="FIGURES 42–45" captionStartId="17.[151,264,1806,1830]" captionTargetBox="[151,1419,155,1775]" captionTargetId="figure-17@17.[151,1436,151,1783]" captionTargetPageId="17" captionText="FIGURES 42–45. Cybaeus penedentatus, male from Holy City, California, left palpus. 42–43 Patella and tibia (42 retrolateral, 43 dorsal). 44–45 Genital bulb (44 ventral, 45 retrolateral). PA—proximal arm of tegular apophysis. PTA—patellar apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrows indicate ventral keel of proximal arm of tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.4752582" httpUri="https://zenodo.org/record/4752582/files/figure.png" pageId="16" pageNumber="417">Fig. 43</figureCitation>
) as well as the posteriorly directed basal flange, longitudinal ventral keel, and acuminate slightly twisted tip of the proximal arm of the tegular apophysis (
<figureCitation id="13192A501913BE3FFBC469A2FB2A9A8A" box="[1129,1262,1718,1745]" captionStart="FIGURES 42–45" captionStartId="17.[151,264,1806,1830]" captionTargetBox="[151,1419,155,1775]" captionTargetId="figure-17@17.[151,1436,151,1783]" captionTargetPageId="17" captionText="FIGURES 42–45. Cybaeus penedentatus, male from Holy City, California, left palpus. 42–43 Patella and tibia (42 retrolateral, 43 dorsal). 44–45 Genital bulb (44 ventral, 45 retrolateral). PA—proximal arm of tegular apophysis. PTA—patellar apophysis, RTA—retrolateral tibial apophysis. Unlabelled arrows indicate ventral keel of proximal arm of tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.4752582" httpUri="https://zenodo.org/record/4752582/files/figure.png" pageId="16" pageNumber="417">Figs 44–45</figureCitation>
). The female is diagnosed primarily by the very small, transverse, medially located atrium (
<figureCitation id="13192A501913BE3FFC4C69CEFB729AAE" box="[993,1206,1754,1781]" captionStart="FIGURES 46–51" captionStartId="18.[151,264,1787,1811]" captionTargetBox="[151,1433,181,1759]" captionTargetId="figure-27@18.[151,1437,181,1763]" captionTargetPageId="18" captionText="FIGURES 46–51. Cybaeus penedentatus, females from California, copulatory organ (46–48 from Ben Lomond, 49 from Pescadero Creek, 50–51 from Mount Madonna). 46 Epigynum, ventral. 47–51 Vulva (47, 49–50 ventral; 48, 51 dorsal). AT—atrium." figureDoi="http://doi.org/10.5281/zenodo.4752586" httpUri="https://zenodo.org/record/4752586/files/figure.png" pageId="16" pageNumber="417">Figs 46–47, 49–50</figureCitation>
,
<figureCitation id="13192A501913BE3FFB6D69CFFB199AAE" box="[1216,1245,1754,1781]" captionStart="FIGURES 52–54" captionStartId="19.[151,264,1076,1100]" captionTargetBox="[176,1401,188,1050]" captionTargetId="figure-17@19.[176,1411,181,1052]" captionTargetPageId="19" captionText="FIGURES 52–54. Cybaeus penedentatus, female from Brookdale, California, copulatory organ. 52 Epigynum, ventral. 53–54 Vulva (53 ventral, 54 dorsal). AT—atrium." figureDoi="http://doi.org/10.5281/zenodo.4752592" httpUri="https://zenodo.org/record/4752592/files/figure.png" pageId="16" pageNumber="417">52</figureCitation>
) and secondarily by the membranous copulatory ducts extending contiguously or in close proximity from the atrium to the anterior margin of the vulva before diverging and heading laterally and posteriorly to merge with the spermathecal heads and stalks (
<figureCitation id="13192A501913BE3FFEB86852FE529B3A" box="[277,406,1862,1889]" captionStart="FIGURES 46–51" captionStartId="18.[151,264,1787,1811]" captionTargetBox="[151,1433,181,1759]" captionTargetId="figure-27@18.[151,1437,181,1763]" captionTargetPageId="18" captionText="FIGURES 46–51. Cybaeus penedentatus, females from California, copulatory organ (46–48 from Ben Lomond, 49 from Pescadero Creek, 50–51 from Mount Madonna). 46 Epigynum, ventral. 47–51 Vulva (47, 49–50 ventral; 48, 51 dorsal). AT—atrium." figureDoi="http://doi.org/10.5281/zenodo.4752586" httpUri="https://zenodo.org/record/4752586/files/figure.png" pageId="16" pageNumber="417">Figs 47–51</figureCitation>
,
<figureCitation id="13192A501913BE3FFE0F6853FE299B3A" box="[418,493,1862,1889]" captionStart="FIGURES 52–54" captionStartId="19.[151,264,1076,1100]" captionTargetBox="[176,1401,188,1050]" captionTargetId="figure-17@19.[176,1411,181,1052]" captionTargetPageId="19" captionText="FIGURES 52–54. Cybaeus penedentatus, female from Brookdale, California, copulatory organ. 52 Epigynum, ventral. 53–54 Vulva (53 ventral, 54 dorsal). AT—atrium." figureDoi="http://doi.org/10.5281/zenodo.4752592" httpUri="https://zenodo.org/record/4752592/files/figure.png" pageId="16" pageNumber="417">53–54</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C338655E1913BE3FFF6A687FFDBC9BDE" box="[199,632,1898,1925]" pageId="16" pageNumber="417" type="description">
<paragraph id="8B9D36D51913BE3FFF6A687FFDBC9BDE" blockId="16.[151,1437,1430,2033]" box="[199,632,1898,1925]" pageId="16" pageNumber="417">
<emphasis id="B956EAC71913BE3FFF6A687FFE9D9BDF" bold="true" box="[199,345,1898,1924]" pageId="16" pageNumber="417">Description.</emphasis>
See
<bibRefCitation id="EFB34B241913BE3FFE22687EFDB09BDE" author="Copley, C. R. &amp; Bennett, R. &amp; Perlman, S. J." box="[399,628,1899,1925]" pageId="16" pageNumber="417" pagination="367 - 401" refId="ref18027" refString="Copley, C. R., Bennett, R. &amp; Perlman, S. J. (2009) Systematics of Nearctic Cybaeus (Araneae: Cybaeidae). Invertebrate Systematics, 23, 367 - 401. https: // doi. org / 10.1071 / IS 09001" type="journal article" year="2009">
Copley
<emphasis id="B956EAC71913BE3FFE4A687EFDDB9BDE" box="[487,543,1899,1925]" italics="true" pageId="16" pageNumber="417">et al.</emphasis>
(2009)
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C338655E1913BE3FFF6A689BFE5D9BAA" pageId="16" pageNumber="417" type="distribution">
<paragraph id="8B9D36D51913BE3FFF6A689BFE5D9BAA" blockId="16.[151,1437,1430,2033]" pageId="16" pageNumber="417">
<emphasis id="B956EAC71913BE3FFF6A689BFEA59BF3" bold="true" box="[199,353,1934,1960]" pageId="16" pageNumber="417">Distribution.</emphasis>
Coastal central
<collectingRegion id="49E6F8371913BE3FFDBE689AFD419BF2" box="[531,645,1935,1961]" country="United States of America" name="California" pageId="16" pageNumber="417">California</collectingRegion>
west of San Francisco Bay south to Monterey Bay in San Mateo, Santa Clara and Santa Cruz Counties (
<figureCitation id="13192A501913BE3FFDA868A6FD9E9B95" box="[517,602,1971,1998]" captionStart="FIGURE 86" captionStartId="30.[151,250,1943,1967]" captionTargetBox="[164,1426,185,1915]" captionTargetId="figure-27@30.[156,1431,181,1919]" captionTargetPageId="30" captionText="FIGURE 86. Distributions of Cybaeus pan spec. nov., C. vulpinus, and C. opulentus spec. nov. AZ—Arizona, CA—California, ID—Idaho, OR—Oregon, NV—Nevada, UT—Utah, WA—Washington. California county names capitalized." figureDoi="http://doi.org/10.5281/zenodo.4752630" httpUri="https://zenodo.org/record/4752630/files/figure.png" pageId="16" pageNumber="417">Fig. 86</figureCitation>
). The
<typeStatus id="549988771913BE3FFD0F68A6FCC39B96" box="[674,775,1971,1997]" pageId="16" pageNumber="417" type="holotype">holotype</typeStatus>
, the single known male specimen of
<taxonomicName id="4C224D561913BE3FFB0168A6FAA69B96" authorityName="Bennett" authorityYear="2009" box="[1196,1378,1971,1997]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="penedentatus">
<emphasis id="B956EAC71913BE3FFB0168A6FAA69B96" box="[1196,1378,1971,1997]" italics="true" pageId="16" pageNumber="417">C. penedentatus</emphasis>
</taxonomicName>
, was collected in December.
</paragraph>
</subSubSection>
</treatment>
</document>
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