279 lines
38 KiB
XML
279 lines
38 KiB
XML
<document id="67F0DE3B9045F47BCE1469DB8731610D" ID-CLB-Dataset="21248" ID-DOI="10.1080/00222933.2016.1198838" ID-GBIF-Dataset="13edff6d-715f-48b3-8c10-9f3547dcf061" ID-ISSN="1464-5262" ID-Zenodo-Dep="3993161" ID-ZooBank="76FF4B91-5624-44A3-B121-2B269CB75C6F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1597931016685" checkinUser="carolina" docAuthor="Ariyama, Hiroyuki" docDate="2016" docId="038887F8FFF2BB50E2BCFB7BFB0A44BD" docLanguage="en" docName="JNATHIST.50.35-36.2277-2297.pdf.imf" docOrigin="Journal of Natural History 50" docStyle="DocumentStyle:3C762F9783B768A90048B82ACD135A62.3:JNatHist.2017-.journal_article.0cover" docStyleId="3C762F9783B768A90048B82ACD135A62" docStyleName="JNatHist.2017-.journal_article.0cover" docStyleVersion="3" docTitle="Paraniphargus Tattersall 1925" docType="treatment" docVersion="8" lastPageNumber="2289" masterDocId="FFB1FF80FFF8BB5CE313FF9AFF8E4477" masterDocTitle="Two new species of eyeless amphipods from a coastal area in Japan (Crustacea: Amphipoda: Hadziidae, Melitidae), with reinstatement of the genus Paraniphargus Tattersall, 1925" masterLastPageNumber="2297" masterPageNumber="2277" pageNumber="2287" updateTime="1698844888288" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="B81BA8B131E49BE2AEBB833866F16F94">Two new species of eyeless amphipods from a coastal area in Japan (Crustacea: Amphipoda: Hadziidae, Melitidae), with reinstatement of the genus Paraniphargus Tattersall, 1925</mods:title>
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<mods:namePart id="D126385396F8F1D976C8EFA5CD580EB3">Ariyama, Hiroyuki</mods:namePart>
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<mods:title id="D3F337D4BF1E44EAE0700E5D6F7B845E">Journal of Natural History</mods:title>
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<mods:date id="7F566A977A681C6EDDDAE42AF858A0E3">2016</mods:date>
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<treatment id="038887F8FFF2BB50E2BCFB7BFB0A44BD" ID-DOI="http://doi.org/10.5281/zenodo.4328448" ID-GBIF-Taxon="167284534" ID-Zenodo-Dep="4328448" LSID="urn:lsid:plazi:treatment:038887F8FFF2BB50E2BCFB7BFB0A44BD" httpUri="http://treatment.plazi.org/id/038887F8FFF2BB50E2BCFB7BFB0A44BD" lastPageId="12" lastPageNumber="2289" pageId="10" pageNumber="2287">
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<subSubSection id="C33B6565FFF2BB56E2BCFB7BFCFD408D" box="[431,883,1248,1274]" pageId="10" pageNumber="2287" type="nomenclature">
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<paragraph id="8B9E36EEFFF2BB56E2BCFB7BFCFD408D" blockId="10.[431,883,1214,1274]" box="[431,883,1248,1274]" pageId="10" pageNumber="2287">
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<heading id="D0D68182FFF2BB56E2BCFB7BFCFD408D" box="[431,883,1248,1274]" centered="true" fontSize="11" level="2" pageId="10" pageNumber="2287" reason="2">
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Genus
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<taxonomicName id="4C214D6DFFF2BB56E110FB7AFCFD408D" ID-CoL="7PS3K" authority="Tattersall, 1925" authorityName="Tattersall" authorityYear="1925" box="[515,883,1248,1274]" class="Malacostraca" family="Melitidae" genus="Paraniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="10" pageNumber="2287" phylum="Arthropoda" rank="genus">
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<emphasis id="B955EAFCFFF2BB56E110FB7AFD36408D" bold="true" box="[515,696,1248,1274]" italics="true" pageId="10" pageNumber="2287">Paraniphargus</emphasis>
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<bibRefCitation id="EFB04B1FFFF2BB56E1D2FB7BFCFD408D" author="Tattersall WM" box="[705,883,1249,1274]" pageId="10" pageNumber="2287" pagination="241 - 247" refId="ref9338" refString="Tattersall WM. 1925. Freshwater Amphipoda from the Andaman Isles. Rec Ind Mus. 27: 241 - 247." type="journal article" year="1925">Tattersall, 1925</bibRefCitation>
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||
</taxonomicName>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C33B6565FFF2BB56E3B3FABEFCEC4128" pageId="10" pageNumber="2287" type="reference_group">
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||
<paragraph id="8B9E36EEFFF2BB56E3B3FABEFCEC4128" blockId="10.[160,1106,1316,1375]" pageId="10" pageNumber="2287">
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<treatmentCitationGroup id="AB3111C0FFF2BB56E3B3FABEFCEC4128" pageId="10" pageNumber="2287">
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<treatmentCitation id="0A8010FFFFF2BB56E3B3FABEFDA2414B" author="Tattersall WM" box="[160,556,1316,1340]" page="241" pageId="10" pageNumber="2287" year="1925">
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<taxonomicName id="4C214D6DFFF2BB56E3B3FABEFDA2414B" ID-CoL="7PS3K" authority="Tattersall, 1925: 241" authorityName="Tattersall" authorityPageNumber="241" authorityYear="1925" box="[160,556,1316,1340]" class="Malacostraca" family="Melitidae" genus="Paraniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="10" pageNumber="2287" phylum="Arthropoda" rank="genus">
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<emphasis id="B955EAFCFFF2BB56E3B3FABEFEB1414B" box="[160,319,1316,1340]" italics="true" pageId="10" pageNumber="2287">Paraniphargus</emphasis>
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||
<bibRefCitation id="EFB04B1FFFF2BB56E25BFABEFDA2414B" author="Tattersall WM" box="[328,556,1316,1340]" pageId="10" pageNumber="2287" pagination="241 - 247" refId="ref9338" refString="Tattersall WM. 1925. Freshwater Amphipoda from the Andaman Isles. Rec Ind Mus. 27: 241 - 247." type="journal article" year="1925">Tattersall, 1925: 241</bibRefCitation>
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||
</taxonomicName>
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||
</treatmentCitation>
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||
;
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||
<treatmentCitation id="0A8010FFFFF2BB56E12BFABEFC2D414B" author="Barnard JL & Barnard CM" box="[568,931,1316,1340]" page="675" pageId="10" pageNumber="2287" year="1983">
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||
<bibRefCitation id="EFB04B1FFFF2BB56E12BFABEFC2D414B" author="Barnard JL & Barnard CM" box="[568,931,1316,1340]" pageId="10" pageNumber="2287" refId="ref8323" refString="Barnard JL, Barnard CM. 1983. Freshwater Amphipoda of the World, I. Evolutionary patterns and II. Handbook of bibliography. Mt. Vernon, VA: Hayfield Associates." type="book" year="1983">Barnard and Barnard, 1983: 675</bibRefCitation>
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</treatmentCitation>
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, fig. 5, map 54.
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<emphasis id="B955EAFCFFF2BB56E3B3FADDFF6D4128" box="[160,227,1351,1375]" italics="true" pageId="10" pageNumber="2287">Melita</emphasis>
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:
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<treatmentCitation id="0A8010FFFFF2BB56E3E1FADDFE7D4128" author="Sawicki TR & Holsinger JR & Iliffe TM" box="[242,499,1351,1375]" page="66" pageId="10" pageNumber="2287" year="2005">
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<bibRefCitation id="EFB04B1FFFF2BB56E3E1FADDFE7D4128" author="Sawicki TR & Holsinger JR & Iliffe TM" box="[242,499,1351,1375]" pageId="10" pageNumber="2287" pagination="49 - 74" refId="ref9106" refString="Sawicki TR, Holsinger JR, Iliffe TM. 2005. New species of amphipod crustaceans in the genera Tegano and Melita (Hadzoidea: Melitidae) from subterranean groundwaters in Guam, Palau, and the Philippines. J Crust Biol. 25: 49 - 74." type="journal article" year="2005">Sawicki et al., 2005: 66</bibRefCitation>
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</treatmentCitation>
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||
(in part) [not
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||
<treatmentCitation id="0A8010FFFFF2BB56E185FADDFCD74128" author="Leach WE" box="[662,857,1351,1375]" page="403" pageId="10" pageNumber="2287" year="1814">
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<bibRefCitation id="EFB04B1FFFF2BB56E185FADDFCD74128" author="Leach WE" box="[662,857,1351,1375]" pageId="10" pageNumber="2287" pagination="383 - 437" refId="ref8786" refString="Leach WE. 1814. Crustaceology. In: Brewster D, editor. The Edinburgh encyclopaedia, Volume 7. Edinburgh: William Blackwood; p. 383 - 437." type="book chapter" year="1814">Leach, 1814: 403</bibRefCitation>
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</treatmentCitation>
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].
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C33B6565FFF2BB57E3B3FA14FEB54631" lastPageId="11" lastPageNumber="2288" pageId="10" pageNumber="2287" type="type_taxon">
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||
<paragraph id="8B9E36EEFFF2BB56E3B3FA14FEBA41D0" blockId="10.[160,797,1422,1482]" box="[160,308,1422,1447]" pageId="10" pageNumber="2287">
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||
<heading id="D0D68182FFF2BB56E3B3FA14FEBA41D0" box="[160,308,1422,1447]" fontSize="11" level="3" pageId="10" pageNumber="2287" reason="5">
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||
<emphasis id="B955EAFCFFF2BB56E3B3FA14FEBA41D0" bold="true" box="[160,308,1422,1447]" inLineHeading="true" italics="true" pageId="10" pageNumber="2287" reason="3">
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||
<typeStatus id="549A884CFFF2BB56E3B3FA14FF5641D0" box="[160,216,1422,1447]" pageId="10" pageNumber="2287">Type</typeStatus>
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species
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</emphasis>
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</heading>
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||
</paragraph>
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||
<paragraph id="8B9E36EEFFF2BB56E3B3FA28FC9341BD" blockId="10.[160,797,1422,1482]" box="[160,797,1458,1482]" pageId="10" pageNumber="2287">
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||
<taxonomicName id="4C214D6DFFF2BB56E3B3FA28FDF841BD" ID-CoL="4D668" authority="Tattersall, 1925" authorityName="Tattersall" authorityYear="1925" box="[160,630,1458,1482]" class="Malacostraca" family="Melitidae" genus="Paraniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="10" pageNumber="2287" phylum="Arthropoda" rank="species" species="annandalei">
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||
<emphasis id="B955EAFCFFF2BB56E3B3FA28FE4D41BD" box="[160,451,1458,1482]" italics="true" pageId="10" pageNumber="2287">Paraniphargus annandalei</emphasis>
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||
<bibRefCitation id="EFB04B1FFFF2BB56E2D8FA28FDF841BD" author="Tattersall WM" box="[459,630,1458,1482]" pageId="10" pageNumber="2287" pagination="241 - 247" refId="ref9338" refString="Tattersall WM. 1925. Freshwater Amphipoda from the Andaman Isles. Rec Ind Mus. 27: 241 - 247." type="journal article" year="1925">Tattersall, 1925</bibRefCitation>
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||
</taxonomicName>
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by monotypy.
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||
</paragraph>
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||
<paragraph id="8B9E36EEFFF2BB56E3B3FA63FE094265" blockId="10.[160,1157,1529,1727]" box="[160,391,1529,1554]" pageId="10" pageNumber="2287">
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||
<heading id="D0D68182FFF2BB56E3B3FA63FE094265" box="[160,391,1529,1554]" fontSize="11" level="3" pageId="10" pageNumber="2287" reason="5">
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||
<emphasis id="B955EAFCFFF2BB56E3B3FA63FE094265" bold="true" box="[160,391,1529,1554]" inLineHeading="true" italics="true" pageId="10" pageNumber="2287" reason="3">Emended diagnosis</emphasis>
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</heading>
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||
</paragraph>
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||
<paragraph id="8B9E36EEFFF2BB57E3B3F987FEB54631" blockId="10.[160,1157,1529,1727]" lastBlockId="11.[160,1157,142,582]" lastPageId="11" lastPageNumber="2288" pageId="10" pageNumber="2287">
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||
Head, rostrum weak, eyes absent. Antennae slender; antenna l peduncular article 1 longer than article 3, accessory flagellum 2- or 3-articulated; antenna 2 shorter than 1, flagellum short. Mandibular palp slender, with 3 articles; article 1 short, articles 2
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<emphasis id="B955EAFCFFF2BB56E3BDF91EFF3542EB" box="[174,187,1668,1692]" italics="true" pageId="10" pageNumber="2287">–</emphasis>
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3 long, article 3 linear, weakly setose. Lower lip with developed inner lobes. Maxilla 1, inner plate ovate, with 3
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<emphasis id="B955EAFCFFF2BB56E150F93DFDDE42C8" box="[579,592,1703,1727]" italics="true" pageId="10" pageNumber="2287">–</emphasis>
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4 apical setae, not setose medially; outer plate with 6
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<emphasis id="B955EAFCFFF3BB57E3F9FF14FF7944D1" box="[234,247,142,166]" italics="true" pageId="11" pageNumber="2288">–</emphasis>
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9 robust setae distally; palp 2-articulated, with apical robust setae. Maxilla 2, inner plate without facial or medial setae. Coxae relatively long, coxa 4 lobate. Gnathopod 1 small, subchelate; carpus weakly elongate, unlobed; propodus shorter than carpus, trapezoidal, palm transverse, dactylus fitting palm; no sexual dimorphism. Gnathopod 2 enlarged, subchelate; carpus of medium length, weakly lobate; propodus large, roundish rectangular, palm oblique; dactylus long, curved posteriorly; no sexual dimorphism. Pereopods 3
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<emphasis id="B955EAFCFFF3BB57E1A6FEC4FD4C4501" box="[693,706,350,374]" italics="true" pageId="11" pageNumber="2288">–</emphasis>
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4 slender; bases of pereopods 5
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<emphasis id="B955EAFCFFF3BB57E77BFEC4FBFB4501" box="[1128,1141,350,374]" italics="true" pageId="11" pageNumber="2288">–</emphasis>
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7 relatively expanded, slightly lobate posterodistally, posterior margins serrate. Uropods biramous; uropods 1
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<emphasis id="B955EAFCFFF3BB57E10BFE39FDAB45CC" box="[536,549,419,443]" italics="true" pageId="11" pageNumber="2288">–</emphasis>
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2 rami extending subequally, with robust setae marginally; uropod 3 elongate; peduncle short; parviramous, outer ramus long, 1- articulated, with marginal and terminal setae, inner ramus shorter than 20% length of outer. Telson of ordinary length, fully cleft, lobes tapering, each with 2 apical robust setae.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33B6565FFF3BB50E3B3FD12FB0A44BD" lastPageId="12" lastPageNumber="2289" pageId="11" pageNumber="2288" type="discussion">
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||
<paragraph id="8B9E36EEFFF3BB57E3B3FD12FE8846D6" blockId="11.[160,1156,648,1713]" box="[160,262,648,673]" pageId="11" pageNumber="2288">
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||
<heading id="D0D68182FFF3BB57E3B3FD12FE8846D6" box="[160,262,648,673]" fontSize="11" level="3" pageId="11" pageNumber="2288" reason="5">
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||
<emphasis id="B955EAFCFFF3BB57E3B3FD12FE8846D6" bold="true" box="[160,262,648,673]" inLineHeading="true" italics="true" pageId="11" pageNumber="2288" reason="3">Remarks</emphasis>
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||
</heading>
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||
</paragraph>
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||
<paragraph id="8B9E36EEFFF3BB57E3B3FD36FC5947C1" blockId="11.[160,1156,648,1713]" pageId="11" pageNumber="2288">
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||
Four species have been described in the genus
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<emphasis id="B955EAFCFFF3BB57E033FD36FC4246B3" box="[800,972,684,708]" italics="true" pageId="11" pageNumber="2288">
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||
<taxonomicName id="4C214D6DFFF3BB57E033FD36FC4B46B3" authorityName="Tattersall" authorityYear="1925" box="[800,965,684,708]" class="Malacostraca" family="Melitidae" genus="Paraniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="11" pageNumber="2288" phylum="Arthropoda" rank="genus">Paraniphargus</taxonomicName>
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:
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</emphasis>
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<taxonomicName id="4C214D6DFFF3BB57E0CDFD36FED94691" authority="Tattersall, 1925" authorityName="Tattersall" authorityYear="1925" class="Malacostraca" family="Melitidae" genus="Paraniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="11" pageNumber="2288" phylum="Arthropoda" rank="species" species="annandalei">
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<emphasis id="B955EAFCFFF3BB57E0CDFD36FB0A46B3" box="[990,1156,684,708]" italics="true" pageId="11" pageNumber="2288">P. annandalei</emphasis>
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||
<bibRefCitation id="EFB04B1FFFF3BB57E3B3FD54FED94691" author="Tattersall WM" box="[160,343,718,742]" pageId="11" pageNumber="2288" pagination="241 - 247" refId="ref9338" refString="Tattersall WM. 1925. Freshwater Amphipoda from the Andaman Isles. Rec Ind Mus. 27: 241 - 247." type="journal article" year="1925">Tattersall, 1925</bibRefCitation>
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||
</taxonomicName>
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from a freshwater stream in the Andaman Islands;
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<taxonomicName id="4C214D6DFFF3BB57E0CDFD54FE53477E" authority="Schellenberg, 1931" authorityName="Schellenberg" authorityYear="1931" class="Malacostraca" family="Melitidae" genus="Paraniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="11" pageNumber="2288" phylum="Arthropoda" rank="species" species="ruttneri">
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||
<emphasis id="B955EAFCFFF3BB57E0CDFD54FF7A477E" italics="true" pageId="11" pageNumber="2288">Paraniphargus ruttneri</emphasis>
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||
<bibRefCitation id="EFB04B1FFFF3BB57E3EEFD6BFE53477E" author="Schellenberg A" box="[253,477,753,777]" pageId="11" pageNumber="2288" pagination="493 - 511" refId="ref9155" refString="Schellenberg A. 1931. Amphipoden der Sunda-Expeditionen Thienemann und Rensch. Arch Hydrobiol. Suppl. 8: 493 - 511." type="journal article" year="1931">Schellenberg, 1931</bibRefCitation>
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||
</taxonomicName>
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from a freshwater lake in Java;
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<taxonomicName id="4C214D6DFFF3BB57E045FD6BFEB3475B" authority="Monod, 1970" authorityName="Monod" authorityYear="1970" class="Malacostraca" family="Melitidae" genus="Paraniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="11" pageNumber="2288" phylum="Arthropoda" rank="species" species="leleuporum">
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||
<emphasis id="B955EAFCFFF3BB57E045FD6BFB0A477E" box="[854,1156,753,777]" italics="true" pageId="11" pageNumber="2288">Paraniphargus leleuporum</emphasis>
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||
<bibRefCitation id="EFB04B1FFFF3BB57E3B3FC8EFEB3475B" author="Monod T" box="[160,317,788,812]" pageId="11" pageNumber="2288" pagination="11 - 53" refId="ref8922" refString="Monod T. 1970. Sur quelques crustaces Malacostraces des Iles Galapagos recoltes par N. et J. Leleup (1964 - 1965). Mission zoologique belge aux iles Galapagos et en Ecuador (N. et J. Leleup, 1964 - 1965). 2: 11 - 53." type="journal article" year="1970">Monod, 1970</bibRefCitation>
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</taxonomicName>
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from groundwater in the Galapagos Islands;
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<taxonomicName id="4C214D6DFFF3BB57E072FC8EFEFE4739" authority="Bamber, 2003" authorityName="Bamber" authorityYear="2003" class="Malacostraca" family="Melitidae" genus="Paraniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="11" pageNumber="2288" phylum="Arthropoda" rank="species" species="vermiamicus">
|
||
<emphasis id="B955EAFCFFF3BB57E072FC8EFF4B4739" italics="true" pageId="11" pageNumber="2288">Paraniphargus vermiamicus</emphasis>
|
||
<bibRefCitation id="EFB04B1FFFF3BB57E3DDFCACFEFE4739" author="Bamber RN" box="[206,368,822,846]" pageId="11" pageNumber="2288" pagination="195 - 207" refId="ref8154" refString="Bamber RN. 2003. New species of Amphipoda from Hong Kong shores. In: Morton B, editor. Perspectives on marine environmental change in Hong Kong and Southern China, 1997 - 2001. Hong Kong: Hong Kong University Press; p. 195 - 207." type="book chapter" year="2003">Bamber, 2003</bibRefCitation>
|
||
</taxonomicName>
|
||
from marine waters in
|
||
<collectingCountry id="F336767EFFF3BB57E199FCACFC9D4739" box="[650,787,822,846]" name="Hong Kong" pageId="11" pageNumber="2288">Hong Kong</collectingCountry>
|
||
.
|
||
<taxonomicName id="4C214D6DFFF3BB57E00CFCACFBC04739" authorityName="Monod" authorityYear="1970" box="[799,1102,822,846]" class="Malacostraca" family="Melitidae" genus="Paraniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="11" pageNumber="2288" phylum="Arthropoda" rank="species" species="leleuporum">
|
||
<emphasis id="B955EAFCFFF3BB57E00CFCACFBC04739" box="[799,1102,822,846]" italics="true" pageId="11" pageNumber="2288">Paraniphargus leleuporum</emphasis>
|
||
</taxonomicName>
|
||
was subsequently transferred to a new genus,
|
||
<taxonomicName id="4C214D6DFFF3BB57E1A7FCC3FC7E4706" authority="Barnard, 1976" authorityName="Barnard" authorityYear="1976" box="[692,1008,857,881]" class="Malacostraca" family="Melitidae" genus="Galapsiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="11" pageNumber="2288" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B955EAFCFFF3BB57E1A7FCC3FCB54706" box="[692,827,857,881]" italics="true" pageId="11" pageNumber="2288">Galapsiellus</emphasis>
|
||
<bibRefCitation id="EFB04B1FFFF3BB57E05BFCC3FC7E4706" author="Barnard JL" box="[840,1008,857,881]" pageId="11" pageNumber="2288" pagination="421 - 432" refId="ref8261" refString="Barnard JL. 1976. Affinities of Paraniphargus lelouparum Monod, a blind anchialine amphipod (Crustacea) from the Galapagos Islands. Proc Biol Soc Wash. 89: 421 - 432." type="journal article" year="1976">Barnard, 1976</bibRefCitation>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFB04B1FFFF3BB57E73BFCC3FF6447E3" author="Barnard JL" pageId="11" pageNumber="2288" pagination="421 - 432" refId="ref8261" refString="Barnard JL. 1976. Affinities of Paraniphargus lelouparum Monod, a blind anchialine amphipod (Crustacea) from the Galapagos Islands. Proc Biol Soc Wash. 89: 421 - 432." type="journal article" year="1976">Barnard (1976)</bibRefCitation>
|
||
, and
|
||
<taxonomicName id="4C214D6DFFF3BB57E221FCE6FE6F47E3" authorityName="Bamber" authorityYear="2003" box="[306,481,892,916]" class="Malacostraca" family="Melitidae" genus="Paraniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="11" pageNumber="2288" phylum="Arthropoda" rank="species" species="vermiamicus">
|
||
<emphasis id="B955EAFCFFF3BB57E221FCE6FE6F47E3" box="[306,481,892,916]" italics="true" pageId="11" pageNumber="2288">P. vermiamicus</emphasis>
|
||
</taxonomicName>
|
||
was transferred to
|
||
<taxonomicName id="4C214D6DFFF3BB57E1C6FCE6FB0A47E3" authority="Barnard and Karaman, 1982" authorityName="Barnard and Karaman" authorityYear="1982" box="[725,1156,892,916]" class="Malacostraca" family="Melitidae" genus="Tegano" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="11" pageNumber="2288" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B955EAFCFFF3BB57E1C6FCE6FCA747E3" box="[725,809,892,916]" italics="true" pageId="11" pageNumber="2288">Tegano</emphasis>
|
||
<bibRefCitation id="EFB04B1FFFF3BB57E027FCE6FB0A47E3" author="Barnard JL & Karaman GS" box="[820,1156,892,916]" pageId="11" pageNumber="2288" pagination="167 - 187" refId="ref8357" refString="Barnard JL, Karaman GS. 1982. Classificatory revisions in gammaridean Amphipoda (Crustacea), part 2. Proc Biol Soc Wash. 95: 167 - 187." type="journal article" year="1982">Barnard and Karaman, 1982</bibRefCitation>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFB04B1FFFF3BB57E3D4FC04FE7847C1" author="Horton T & Lowry J" box="[199,502,926,950]" pageId="11" pageNumber="2288" pagination="10" refId="ref8654" refString="Horton T, Lowry J. 2012. WoRMS taxon details: Tegano vermiamicus (Bamber, 2003) [Internet]. [cited 2016 Feb 10]. Available from: http: // www. marinespecies. org / aphia. php? p = taxdetails & id = 537259" type="journal article" year="2012">Horton and Lowry (2012)</bibRefCitation>
|
||
, leaving only two species in the genus.
|
||
</paragraph>
|
||
<paragraph id="8B9E36EEFFF3BB57E3D3FC5BFE5E42C6" blockId="11.[160,1156,648,1713]" pageId="11" pageNumber="2288">
|
||
Several studies have dealt with the difference between
|
||
<taxonomicName id="4C214D6DFFF3BB57E022FC5BFC5E47AE" authorityName="Tattersall" authorityYear="1925" box="[817,976,961,985]" class="Malacostraca" family="Melitidae" genus="Paraniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="11" pageNumber="2288" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B955EAFCFFF3BB57E022FC5BFC5E47AE" box="[817,976,961,985]" italics="true" pageId="11" pageNumber="2288">Paraniphargus</emphasis>
|
||
</taxonomicName>
|
||
and the related genus,
|
||
<emphasis id="B955EAFCFFF3BB57E3E2FC7EFEBA478B" box="[241,308,996,1020]" italics="true" pageId="11" pageNumber="2288">Melita</emphasis>
|
||
<bibRefCitation id="EFB04B1FFFF3BB57E228FC7EFE4E478B" author="Leach WE" box="[315,448,996,1020]" pageId="11" pageNumber="2288" pagination="383 - 437" refId="ref8786" refString="Leach WE. 1814. Crustaceology. In: Brewster D, editor. The Edinburgh encyclopaedia, Volume 7. Edinburgh: William Blackwood; p. 383 - 437." type="book chapter" year="1814">Leach, 1814</bibRefCitation>
|
||
.
|
||
<bibRefCitation id="EFB04B1FFFF3BB57E2DEFC7EFD22478B" author="Schellenberg A" box="[461,684,996,1020]" pageId="11" pageNumber="2288" pagination="493 - 511" refId="ref9155" refString="Schellenberg A. 1931. Amphipoden der Sunda-Expeditionen Thienemann und Rensch. Arch Hydrobiol. Suppl. 8: 493 - 511." type="journal article" year="1931">Schellenberg (1931)</bibRefCitation>
|
||
pointed out that they differ only in the 1- articulated outer ramus of the uropod 3, the lack of the anteroventral cusp on the head, the weak mandibular palp, and the naked medial margin of the inner lobes of both maxillae.
|
||
<bibRefCitation id="EFB04B1FFFF3BB57E219FBD6FDCE4013" author="Barnard JL & Barnard CM" box="[266,576,1100,1124]" pageId="11" pageNumber="2288" refId="ref8323" refString="Barnard JL, Barnard CM. 1983. Freshwater Amphipoda of the World, I. Evolutionary patterns and II. Handbook of bibliography. Mt. Vernon, VA: Hayfield Associates." type="book" year="1983">Barnard and Barnard (1983)</bibRefCitation>
|
||
stated that many species of
|
||
<emphasis id="B955EAFCFFF3BB57E094FBD6FC444013" box="[903,970,1100,1124]" italics="true" pageId="11" pageNumber="2288">Melita</emphasis>
|
||
have lost article 2 on the uropod 3 outer ramus, and that
|
||
<taxonomicName id="4C214D6DFFF3BB57E166FBF4FC9A40F1" authorityName="Tattersall" authorityYear="1925" box="[629,788,1134,1158]" class="Malacostraca" family="Melitidae" genus="Paraniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="11" pageNumber="2288" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B955EAFCFFF3BB57E166FBF4FC9A40F1" box="[629,788,1134,1158]" italics="true" pageId="11" pageNumber="2288">Paraniphargus</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<emphasis id="B955EAFCFFF3BB57E0B5FBF4FC6740F1" box="[934,1001,1134,1158]" italics="true" pageId="11" pageNumber="2288">Melita</emphasis>
|
||
in the loss of eyes and medial maxillary setae.
|
||
<bibRefCitation id="EFB04B1FFFF3BB57E10EFB0BFC9E40DE" author="Stock JH & Ilife TM" box="[541,784,1169,1193]" pageId="11" pageNumber="2288" pagination="245 - 269" refId="ref9258" refString="Stock JH, Ilife TM. 1995. Melitidae (Crustacea, Amphipoda) from anchialine limestone caves in New Caledonia. Cont Zool. 65: 245 - 269." type="journal article" year="1995">Stock and Ilife (1995)</bibRefCitation>
|
||
suggested that there were only slight differences between
|
||
<emphasis id="B955EAFCFFF3BB57E2C9FB29FD4540BC" box="[474,715,1203,1227]" italics="true" pageId="11" pageNumber="2288">
|
||
<taxonomicName id="4C214D6DFFF3BB57E2C9FB29FDF440BC" authorityName="Tattersall" authorityYear="1925" box="[474,634,1203,1227]" class="Malacostraca" family="Melitidae" genus="Paraniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="11" pageNumber="2288" phylum="Arthropoda" rank="genus">Paraniphargus</taxonomicName>
|
||
, Melita
|
||
</emphasis>
|
||
and a third genus
|
||
<taxonomicName id="4C214D6DFFF3BB57E0A3FB29FF594099" authority="Ruffo, 1985" authorityName="Ruffo" authorityYear="1985" class="Malacostraca" family="Melitidae" genus="Josephosella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="11" pageNumber="2288" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B955EAFCFFF3BB57E0A3FB29FBB640BC" box="[944,1080,1203,1227]" italics="true" pageId="11" pageNumber="2288">Josephosella</emphasis>
|
||
<bibRefCitation id="EFB04B1FFFF3BB57E751FB29FF594099" author="Ruffo S" pageId="11" pageNumber="2288" pagination="485 - 509" refId="ref9076" refString="Ruffo S. 1985. Nuovi Anfipodi mesopsammici delle Isole Andamane (Crust. Amphipoda). Boll Mus Civ Stor Nat Verona. 10: 485 - 509." type="journal article" year="1985">Ruffo, 1985</bibRefCitation>
|
||
</taxonomicName>
|
||
. More recently,
|
||
<bibRefCitation id="EFB04B1FFFF3BB57E299FB4CFDE94099" author="Sawicki TR & Holsinger JR & Iliffe TM" box="[394,615,1238,1262]" pageId="11" pageNumber="2288" pagination="49 - 74" refId="ref9106" refString="Sawicki TR, Holsinger JR, Iliffe TM. 2005. New species of amphipod crustaceans in the genera Tegano and Melita (Hadzoidea: Melitidae) from subterranean groundwaters in Guam, Palau, and the Philippines. J Crust Biol. 25: 49 - 74." type="journal article" year="2005">Sawicki et al. (2005)</bibRefCitation>
|
||
synonymized
|
||
<taxonomicName id="4C214D6DFFF3BB57E01BFB4CFC294099" authorityName="Tattersall" authorityYear="1925" box="[776,935,1238,1262]" class="Malacostraca" family="Melitidae" genus="Paraniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="11" pageNumber="2288" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B955EAFCFFF3BB57E01BFB4CFC294099" box="[776,935,1238,1262]" italics="true" pageId="11" pageNumber="2288">Paraniphargus</emphasis>
|
||
</taxonomicName>
|
||
with
|
||
<emphasis id="B955EAFCFFF3BB57E0F5FB4CFBA74099" box="[998,1065,1238,1262]" italics="true" pageId="11" pageNumber="2288">Melita</emphasis>
|
||
citing a lack of distinction in the setation of maxilla 2 and the number of articles in the uropod 3 outer ramus between these genera. The present study recognizes the sexually monomorphic gnathopod 2 as a distinctive generic character of
|
||
<taxonomicName id="4C214D6DFFF3BB57E058FAA4FC644121" authorityName="Tattersall" authorityYear="1925" box="[843,1002,1342,1366]" class="Malacostraca" family="Melitidae" genus="Paraniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="11" pageNumber="2288" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B955EAFCFFF3BB57E058FAA4FC644121" box="[843,1002,1342,1366]" italics="true" pageId="11" pageNumber="2288">Paraniphargus</emphasis>
|
||
</taxonomicName>
|
||
separating it from
|
||
<emphasis id="B955EAFCFFF3BB57E3CCFAFBFEAC410E" box="[223,290,1377,1401]" italics="true" pageId="11" pageNumber="2288">Melita</emphasis>
|
||
species that have a sexually dimorphic gnathopod 2 (
|
||
<bibRefCitation id="EFB04B1FFFF3BB57E086FAFBFF5941EC" author="Barnard JL & Barnard CM" pageId="11" pageNumber="2288" refId="ref8323" refString="Barnard JL, Barnard CM. 1983. Freshwater Amphipoda of the World, I. Evolutionary patterns and II. Handbook of bibliography. Mt. Vernon, VA: Hayfield Associates." type="book" year="1983">Barnard and Barnard 1983</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB04B1FFFF3BB57E3F7FA19FD8441EC" author="Jarrett NE & Bousfield EL" box="[228,522,1411,1435]" pageId="11" pageNumber="2288" pagination="3 - 74" refId="ref8707" refString="Jarrett NE, Bousfield EL. 1996. The amphipod superfamily Hadzioidea on the Pacific coast of North America: family Melitidae. Part I. The Melita group: systematics and distributional ecology. Amphipacifica. 2: 3 - 74." type="journal article" year="1996">Jarrett and Bousfield 1996</bibRefCitation>
|
||
). Although the original description of
|
||
<taxonomicName id="4C214D6DFFF3BB57E0A8FA19FBDF41EC" authorityName="Tattersall" authorityYear="1925" box="[955,1105,1411,1435]" class="Malacostraca" family="Melitidae" genus="Paraniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="11" pageNumber="2288" phylum="Arthropoda" rank="species" species="annandalei">
|
||
<emphasis id="B955EAFCFFF3BB57E0A8FA19FBDF41EC" box="[955,1105,1411,1435]" italics="true" pageId="11" pageNumber="2288">P. annandalei</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C214D6DFFF3BB57E3B3FA3CFE9E41C9" authorityName="Schellenberg" authorityYear="1931" box="[160,272,1446,1470]" class="Malacostraca" family="Melitidae" genus="Paraniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="11" pageNumber="2288" phylum="Arthropoda" rank="species" species="ruttneri">
|
||
<emphasis id="B955EAFCFFF3BB57E3B3FA3CFE9E41C9" box="[160,272,1446,1470]" italics="true" pageId="11" pageNumber="2288">P. ruttneri</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFB04B1FFFF3BB57E255FA3CFE7141C9" author="Tattersall WM" box="[326,511,1446,1470]" pageId="11" pageNumber="2288" pagination="241 - 247" refId="ref9338" refString="Tattersall WM. 1925. Freshwater Amphipoda from the Andaman Isles. Rec Ind Mus. 27: 241 - 247." type="journal article" year="1925">Tattersall (1925)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFB04B1FFFF3BB57E150FA3CFCA441C9" author="Schellenberg A" box="[579,810,1446,1470]" pageId="11" pageNumber="2288" pagination="493 - 511" refId="ref9155" refString="Schellenberg A. 1931. Amphipoden der Sunda-Expeditionen Thienemann und Rensch. Arch Hydrobiol. Suppl. 8: 493 - 511." type="journal article" year="1931">Schellenberg (1931)</bibRefCitation>
|
||
, respectively, do not include details of the presence or absence of sexually dimorphic characters,
|
||
<bibRefCitation id="EFB04B1FFFF3BB57E0CAFA53FB0A4196" author="Barnard JL" box="[985,1156,1481,1505]" pageId="11" pageNumber="2288" pagination="421 - 432" refId="ref8261" refString="Barnard JL. 1976. Affinities of Paraniphargus lelouparum Monod, a blind anchialine amphipod (Crustacea) from the Galapagos Islands. Proc Biol Soc Wash. 89: 421 - 432." type="journal article" year="1976">Barnard (1976)</bibRefCitation>
|
||
proposed the absence of sexual dimorphism in the gnathopods without stating the basis for this observation. The medial margin of the maxilla 2 inner plate is quite bare of setae in all species of
|
||
<taxonomicName id="4C214D6DFFF3BB57E28DF9ABFDB3423E" authorityName="Tattersall" authorityYear="1925" box="[414,573,1585,1609]" class="Malacostraca" family="Melitidae" genus="Paraniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="11" pageNumber="2288" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B955EAFCFFF3BB57E28DF9ABFDB3423E" box="[414,573,1585,1609]" italics="true" pageId="11" pageNumber="2288">Paraniphargus</emphasis>
|
||
</taxonomicName>
|
||
including
|
||
<taxonomicName id="4C214D6DFFF3BB57E1ABF9ABFC90423E" authorityName="Ariyama" authorityYear="2016" box="[696,798,1585,1609]" class="Malacostraca" family="Melitidae" genus="Paraniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="11" pageNumber="2288" phylum="Arthropoda" rank="species" species="shiosai" status="sp. nov.">
|
||
<emphasis id="B955EAFCFFF3BB57E1ABF9ABFC90423E" box="[696,798,1585,1609]" italics="true" pageId="11" pageNumber="2288">P. shiosai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2665787FFF3BB57E034F9ABFC0C423E" box="[807,898,1585,1609]" pageId="11" pageNumber="2288" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, whereas in the
|
||
<emphasis id="B955EAFCFFF3BB57E753F9ABFB0D423E" box="[1088,1155,1585,1609]" italics="true" pageId="11" pageNumber="2288">Melita</emphasis>
|
||
species, number of setae on the medial margin varies from very many to a few but never without setae. Based on the above characters
|
||
<taxonomicName id="4C214D6DFFF3BB57E1DEF9ECFCE242F9" authorityName="Tattersall" authorityYear="1925" box="[717,876,1654,1678]" class="Malacostraca" family="Melitidae" genus="Paraniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="11" pageNumber="2288" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B955EAFCFFF3BB57E1DEF9ECFCE242F9" box="[717,876,1654,1678]" italics="true" pageId="11" pageNumber="2288">Paraniphargus</emphasis>
|
||
</taxonomicName>
|
||
is here reinstated as a distinct genus from
|
||
<emphasis id="B955EAFCFFF3BB57E294F903FE4442C6" box="[391,458,1689,1713]" italics="true" pageId="11" pageNumber="2288">Melita</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9E36EEFFF4BB50E3B3FF14FECD44D0" blockId="12.[160,1156,142,202]" box="[160,323,142,167]" pageId="12" pageNumber="2289">
|
||
<heading id="D0D68182FFF4BB50E3B3FF14FECD44D0" box="[160,323,142,167]" fontSize="11" level="3" pageId="12" pageNumber="2289" reason="5">
|
||
<emphasis id="B955EAFCFFF4BB50E3B3FF14FECD44D0" bold="true" box="[160,323,142,167]" inLineHeading="true" italics="true" pageId="12" pageNumber="2289" reason="3">Included taxa</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B9E36EEFFF4BB50E3B3FF28FB0A44BD" blockId="12.[160,1156,142,202]" box="[160,1156,178,202]" pageId="12" pageNumber="2289">
|
||
Three species:
|
||
<taxonomicName id="4C214D6DFFF4BB50E22DFF28FDF644BD" authority="Tattersall, 1925" authorityName="Tattersall" authorityYear="1925" box="[318,632,178,202]" class="Malacostraca" family="Melitidae" genus="Paraniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="12" pageNumber="2289" phylum="Arthropoda" rank="species" species="annandalei">
|
||
<emphasis id="B955EAFCFFF4BB50E22DFF28FE4044BD" box="[318,462,178,202]" italics="true" pageId="12" pageNumber="2289">P. annandalei</emphasis>
|
||
<bibRefCitation id="EFB04B1FFFF4BB50E2C7FF28FDF644BD" author="Tattersall WM" box="[468,632,178,202]" pageId="12" pageNumber="2289" pagination="241 - 247" refId="ref9338" refString="Tattersall WM. 1925. Freshwater Amphipoda from the Andaman Isles. Rec Ind Mus. 27: 241 - 247." type="journal article" year="1925">Tattersall, 1925</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C214D6DFFF4BB50E190FF28FC3144BD" authority="Schellenberg, 1931" authorityName="Schellenberg" authorityYear="1931" box="[643,959,178,202]" class="Malacostraca" family="Melitidae" genus="Paraniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="12" pageNumber="2289" phylum="Arthropoda" rank="species" species="ruttneri">
|
||
<emphasis id="B955EAFCFFF4BB50E190FF28FD6744BD" box="[643,745,178,202]" italics="true" pageId="12" pageNumber="2289">P. ruttneri</emphasis>
|
||
<bibRefCitation id="EFB04B1FFFF4BB50E1FCFF28FC3144BD" author="Schellenberg A" box="[751,959,178,202]" pageId="12" pageNumber="2289" pagination="493 - 511" refId="ref9155" refString="Schellenberg A. 1931. Amphipoden der Sunda-Expeditionen Thienemann und Rensch. Arch Hydrobiol. Suppl. 8: 493 - 511." type="journal article" year="1931">Schellenberg, 1931</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C214D6DFFF4BB50E0D9FF28FBA544BD" authorityName="Ariyama" authorityYear="2016" box="[970,1067,178,202]" class="Malacostraca" family="Melitidae" genus="Paraniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="12" pageNumber="2289" phylum="Arthropoda" rank="species" species="shiosai" status="sp. nov.">
|
||
<emphasis id="B955EAFCFFF4BB50E0D9FF28FBA544BD" box="[970,1067,178,202]" italics="true" pageId="12" pageNumber="2289">P. shiosai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2665787FFF4BB50E723FF28FB0A44BD" box="[1072,1156,178,202]" pageId="12" pageNumber="2289" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |