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<document id="BDF9D8AC928D140D8C232C7E926F7B21" ID-CLB-Dataset="22253" ID-DOI="10.11646/zootaxa.4772.3.2" ID-GBIF-Dataset="983953d9-e7f1-48e8-b24d-b004ada2d986" ID-ISSN="1175-5326" ID-Zenodo-Dep="3819706" ID-ZooBank="403F169F-DD66-4A64-8887-571D55066E33" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589175668640" checkinUser="plazi" docAuthor="Anker, Arthur" docDate="2020" docId="038887EDFFEDFFDBFF081C5B02E2FC6F" docLanguage="en" docName="zootaxa.4772.3.2.pdf" docOrigin="Zootaxa 4772 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Triacanthoneus Anker 2010" docType="key" docVersion="8" lastPageNumber="467" masterDocId="FFB1FF95FFFDFFCAFF9F1A53077EFFA8" masterDocTitle="Taxonomic remarks on the alpheid shrimp genus Triacanthoneus Anker, 2010 with description of a second eastern Pacific species (Malacostraca: Decapoda)" masterLastPageNumber="468" masterPageNumber="450" pageNumber="466" updateTime="1698839100595" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="146D4FC7FC1E4D9602990B43BEC1D976">Taxonomic remarks on the alpheid shrimp genus Triacanthoneus Anker, 2010 with description of a second eastern Pacific species (Malacostraca: Decapoda)</mods:title>
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<mods:namePart id="DB2314782918225F3B8D334FEAA71DC8">Anker, Arthur</mods:namePart>
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<mods:title id="9F9E5741A67DDE0E8DEE4123E4C9E60C">Zootaxa</mods:title>
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<mods:date id="BED7ABBDAA51E0EE53B53E0ADA3413CB">2020</mods:date>
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<treatment id="038887EDFFEDFFDBFF081C5B02E2FC6F" ID-DOI="http://doi.org/10.5281/zenodo.3843775" ID-GBIF-Taxon="164035964" ID-Zenodo-Dep="3843775" LSID="urn:lsid:plazi:treatment:038887EDFFEDFFDBFF081C5B02E2FC6F" httpUri="http://treatment.plazi.org/id/038887EDFFEDFFDBFF081C5B02E2FC6F" lastPageId="17" lastPageNumber="467" pageId="16" pageNumber="466">
<subSubSection id="C33B6570FFEDFFDAFF081C5B033CF98A" box="[151,1090,1544,1571]" pageId="16" pageNumber="466" type="nomenclature">
<paragraph id="8B9E36FBFFEDFFDAFF081C5B033CF98A" blockId="16.[151,1090,1544,1571]" box="[151,1090,1544,1571]" pageId="16" pageNumber="466">
<heading id="D0D68197FFEDFFDAFF081C5B033CF98A" bold="true" box="[151,1090,1544,1571]" fontSize="11" level="1" pageId="16" pageNumber="466" reason="1">
<emphasis id="B955EAE9FFEDFFDAFF081C5B033CF98A" bold="true" box="[151,1090,1544,1571]" pageId="16" pageNumber="466">
Key to the presently known species assigned to
<taxonomicName id="4C214D78FFEDFFDAFD7A1C5B033CF98A" ID-CoL="85JMG" authority="Anker, 2010" authorityName="Anker" authorityYear="2010" box="[741,1090,1544,1571]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="466" phylum="Arthropoda" rank="genus">
<emphasis id="B955EAE9FFEDFFDAFD7A1C5B04DAF98B" bold="true" box="[741,932,1544,1571]" italics="true" pageId="16" pageNumber="466">Triacanthoneus</emphasis>
Anker, 2010
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33B6570FFEDFFDBFF081C1E02E2FC6F" lastPageId="17" lastPageNumber="467" pageId="16" pageNumber="466" type="key">
<keyStep id="30D52E5EFFEDFFDAFF081C1E02E2F912" pageId="16" pageNumber="466">
<paragraph id="8B9E36FBFFEDFFDAFF081C1E02E2F9D7" blockId="16.[151,1436,1613,1723]" pageId="16" pageNumber="466">
<keyLead id="30D095CEFFEDFFDAFF081C1E02E2F9D7" pageId="16" pageNumber="466">1A. Carapace with strong sharp dorsolateral teeth in hepatic or post-hepatic position; orbital teeth absent or present as broadly triangular lobes........................................................................................ 2</keyLead>
</paragraph>
<paragraph id="8B9E36FBFFEDFFDAFF081CD402E2F912" blockId="16.[151,1436,1613,1723]" pageId="16" pageNumber="466">
<keyLead id="30D095CEFFEDFFDAFF081CD402E2F912" pageId="16" pageNumber="466">1B. Carapace without strong sharp dorsolateral teeth in hepatic or post-hepatic position; orbital teeth present, acute, arising from or near post-orbital margin................................................................................ 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D52E5EFFEDFFDAFF081C8C02E2F82F" pageId="16" pageNumber="466">
<paragraph id="8B9E36FBFFEDFFDAFF081C8C02E2F887" blockId="16.[151,1437,1759,1928]" pageId="16" pageNumber="466">
<keyLead id="30D095CEFFEDFFDAFF081C8C02E2F887" pageId="16" pageNumber="466">
2A. Carapace with strong rostral + dorsal carina; eyestalk without anteromesial tubercle; cutting edges of major chela fingers with more than 20 minute teeth; third and fourth pereiopods with ischia unarmed [deep under coral rubble and in marine caves; depth range:
<quantity id="4CD99B1EFFEDFFDAFEBB1D490611F887" box="[292,367,1818,1840]" metricMagnitude="1" metricUnit="m" metricValue="1.15" metricValueMax="1.5" metricValueMin="0.8" pageId="16" pageNumber="466" unit="m" value="11.5" valueMax="15.0" valueMin="8.0">815 m</quantity>
;
<collectingCountry id="F336766BFFEDFFDAFEE61D4906C8F898" box="[377,438,1818,1840]" name="Belize" pageId="16" pageNumber="466">Belize</collectingCountry>
and
<collectingCountry id="F336766BFFEDFFDAFE7A1D4A0564F887" box="[485,538,1817,1839]" name="Cuba" pageId="16" pageNumber="466">Cuba</collectingCountry>
].............................................................
<taxonomicName id="4C214D78FFEDFFDAFA8E1D4802E2F887" ID-CoL="85ZFS" box="[1297,1436,1818,1840]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="466" phylum="Arthropoda" rank="species" species="chapelianus">
<emphasis id="B955EAE9FFEDFFDAFA8E1D4802E2F887" box="[1297,1436,1818,1840]" italics="true" pageId="16" pageNumber="466">T. chapelianus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9E36FBFFEDFFDAFF081D6402E2F82F" blockId="16.[151,1437,1759,1928]" pageId="16" pageNumber="466">
<keyLead id="30D095CEFFEDFFDAFF081D6402E2F82F" pageId="16" pageNumber="466">
2B. Carapace without rostral + dorsal carina; eyestalk with subtriangular anteromesial tubercle; cutting edges of major chela fingers with less than 10 shallow, bump-like teeth; third and fourth pereiopods with ischia armed with stout cuspidate setae [under coral rubble, in burrows among mangrove roots]; depth range:
<quantity id="4CD99B1EFFEDFFDAFC971D210427F82F" box="[776,857,1906,1928]" metricMagnitude="0" metricUnit="m" metricValue="1.25" metricValueMax="2.0" metricValueMin="0.5" pageId="16" pageNumber="466" unit="m" value="1.25" valueMax="2.0" valueMin="0.5">0.52 m</quantity>
; Caribbean coast of
<collectingCountry id="F336766BFFEDFFDAFB821D220312F82F" box="[1053,1132,1905,1927]" name="Panama" pageId="16" pageNumber="466">Panama</collectingCountry>
].....
<taxonomicName id="4C214D78FFEDFFDAFB5D1D200257F820" ID-CoL="874KX" authorityName="Anker" authorityYear="2020" baseAuthorityName="Anker" baseAuthorityYear="2010" box="[1218,1321,1907,1928]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="466" phylum="Arthropoda" rank="species" species="armatus" status="comb. nov.">
<emphasis id="B955EAE9FFEDFFDAFB5D1D200257F820" box="[1218,1321,1907,1928]" italics="true" pageId="16" pageNumber="466">T. armatus</emphasis>
</taxonomicName>
<emphasis id="B955EAE9FFEDFFDAFAB11D2102E2F82F" bold="true" box="[1326,1436,1906,1927]" pageId="16" pageNumber="466">
<taxonomicNameLabel id="A2665792FFEDFFDAFAB11D2102E2F82F" box="[1326,1436,1906,1927]" pageId="16" pageNumber="466" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D52E5EFFEDFFDBFF081DFE02E2FF65" lastPageId="17" lastPageNumber="467" pageId="16" pageNumber="466">
<paragraph id="8B9E36FBFFEDFFDAFF081DFE02E2F877" blockId="16.[151,1436,1965,2016]" pageId="16" pageNumber="466">
<keyLead id="30D095CEFFEDFFDAFF081DFE02E2F877" pageId="16" pageNumber="466">3A. Major chela with fingers much shorter than palm; antennular stylocerite with dorsal carina ending in acute or subacute tooth distally [East Pacific].................................................................................. 4</keyLead>
</paragraph>
<paragraph id="8B9E36FBFFECFFDBFF081AC902E2FF65" blockId="17.[151,1436,153,205]" pageId="17" pageNumber="467">
<keyLead id="30D095CEFFECFFDBFF081AC902E2FF65" pageId="17" pageNumber="467">3B. Major chela with fingers as long as to much longer than palm; antennular stylocerite with dorsal carina, if present, unarmed distally [West Atlantic]................................................................................ 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D52E5EFFECFFDBFF081AA102E2FE31" pageId="17" pageNumber="467">
<paragraph id="8B9E36FBFFECFFDBFF081AA102E3FEE9" blockId="17.[151,1437,241,410]" pageId="17" pageNumber="467">
<keyLead id="30D095CEFFECFFDBFF081AA102E3FEE9" pageId="17" pageNumber="467">
4A. Carapace with one robust seta in front of mid-dorsal tooth; rostrum horizontal or slightly descending, not ascending; telson narrow, about 3.3 times as long as maximal (proximal) width; minor cheliped carpus about 3.5 times as long as distal width [under large rocks on muddy sand; depth range:
<quantity id="4CD99B1EFFECFFDBFDF11B7F05D3FEEA" box="[622,685,300,322]" metricMagnitude="0" metricUnit="m" metricValue="1.0" metricValueMax="2.0" metricValueMin="0.0" pageId="17" pageNumber="467" unit="m" value="1.0" valueMax="2.0" valueMin="0.0">02 m</quantity>
;
<collectingCountry id="F336766BFFECFFDBFD281B7E047DFEEA" box="[695,771,301,322]" name="Panama" pageId="17" pageNumber="467">Panama</collectingCountry>
and
<collectingCountry id="F336766BFFECFFDBFCAD1B7804EDFEE9" box="[818,915,299,321]" name="Colombia" pageId="17" pageNumber="467">Colombia</collectingCountry>
]................................
<taxonomicName id="4C214D78FFECFFDBFAB11B7E02E3FEE9" ID-CoL="85ZFT" authorityName="Anker" authorityYear="2010" box="[1326,1437,300,322]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="467" phylum="Arthropoda" rank="species" species="pacificus">
<emphasis id="B955EAE9FFECFFDBFAB11B7E02E3FEE9" box="[1326,1437,300,322]" italics="true" pageId="17" pageNumber="467">T. pacificus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9E36FBFFECFFDBFF081B1902E2FE31" blockId="17.[151,1437,241,410]" pageId="17" pageNumber="467">
<keyLead id="30D095CEFFECFFDBFF081B1902E2FE31" pageId="17" pageNumber="467">
4B. Carapace with three robust setae in front of mid-dorsal tooth; rostrum ascending; telson moderately broad, about 2.6 times as long as maximal (proximal) width; minor cheliped carpus about 4.5 times as long as distal width [under large rocks on coarse sand; depth:
<quantity id="4CD99B1EFFECFFDBFEC21BD706D6FE32" box="[349,424,388,410]" metricMagnitude="0" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="17" pageNumber="467" unit="m" value="9.0" valueMax="10.0" valueMin="8.0">810 m</quantity>
;
<collectingCountry id="F336766BFFECFFDBFE2D1BD0057FFE31" box="[434,513,387,409]" name="Panama" pageId="17" pageNumber="467">Panama</collectingCountry>
].............................................................
<taxonomicName id="4C214D78FFECFFDBFB6E1BD60237FE31" ID-CoL="874KG" authority="Anker, 2020" authorityName="Anker" authorityYear="2020" box="[1265,1353,388,410]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="467" phylum="Arthropoda" rank="species" species="blanca" status="sp. nov.">
<emphasis id="B955EAE9FFECFFDBFB6E1BD60237FE31" box="[1265,1353,388,410]" italics="true" pageId="17" pageNumber="467">T. blanca</emphasis>
</taxonomicName>
<emphasis id="B955EAE9FFECFFDBFACF1BD702E2FE31" bold="true" box="[1360,1436,388,409]" pageId="17" pageNumber="467">
<taxonomicNameLabel id="A2665792FFECFFDBFACF1BD702E2FE31" box="[1360,1436,388,409]" pageId="17" pageNumber="467" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D52E5EFFECFFDBFF081BEC02E2FD0A" pageId="17" pageNumber="467">
<paragraph id="8B9E36FBFFECFFDBFF081BEC02E3FD84" blockId="17.[151,1437,447,674]" pageId="17" pageNumber="467">
<keyLead id="30D095CEFFECFFDBFF081BEC02E3FD84" pageId="17" pageNumber="467">
5A. Rostrum with ventral tooth situated at rostral mid-length; major chela fingers extremely elongate, much longer than palm, distally strongly crossing, cutting edges with very conspicuous slender raptorial teeth intercalated between smaller teeth; pleural region with acute process above coxa of fifth pereiopod; epiopod of second maxilliped with podobranch; large-sized species (cl 8.0
<quantity id="4CD99B1EFFECFFDBFE9218440629FD84" box="[269,343,535,556]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" pageId="17" pageNumber="467" unit="mm" value="9.1">9.1 mm</quantity>
) [anchialine caves;
<collectingCountry id="F336766BFFECFFDBFD891845051CFD84" box="[534,610,534,556]" name="Mexico" pageId="17" pageNumber="467">Mexico</collectingCountry>
].......................................................
<taxonomicName id="4C214D78FFECFFDBFA91184402E3FD84" ID-CoL="85ZFQ" authorityName="Alvarez, Iliffe, Gonzalez &amp; Villalobos" authorityYear="2012" box="[1294,1437,535,556]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="467" phylum="Arthropoda" rank="species" species="akumalensis">
<emphasis id="B955EAE9FFECFFDBFA91184402E3FD84" box="[1294,1437,535,556]" italics="true" pageId="17" pageNumber="467">T. akumalensis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9E36FBFFECFFDBFF08186702E2FD0A" blockId="17.[151,1437,447,674]" pageId="17" pageNumber="467">
<keyLead id="30D095CEFFECFFDBFF08186702E2FD0A" pageId="17" pageNumber="467">
5B. Rostrum with ventral tooth in subdistal position; major chela fingers subequal to palm to slightly longer, moderately elongate, distally not strongly crossing, cutting edges evenly serrated with small teeth, without raptorial teeth; pleural region without acute process above coxa of fifth pereiopod; epipod of second maxilliped without podobranch; small-sized species (cl under
<quantity id="4CD99B1EFFECFFDBFF7D18DF0653FD0A" box="[226,301,652,674]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="17" pageNumber="467" unit="mm" value="4.5">4.5 mm</quantity>
) [coral reefs and adjacent rubble flats, sometimes close to mangroves; depth range:
<quantity id="4CD99B1EFFECFFDBFBCC18DF03CEFD0A" box="[1107,1200,652,674]" metricMagnitude="0" metricUnit="m" metricValue="7.75" metricValueMax="15.0" metricValueMin="0.5" pageId="17" pageNumber="467" unit="m" value="7.75" valueMax="15.0" valueMin="0.5">0.515 m</quantity>
; Caribbean Sea].... 6
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D52E5EFFECFFDBFF08189402E2FC6F" pageId="17" pageNumber="467">
<paragraph id="8B9E36FBFFECFFDBFF08189402E3FC9C" blockId="17.[151,1437,711,967]" pageId="17" pageNumber="467">
<keyLead id="30D095CEFFECFFDBFF08189402E3FC9C" pageId="17" pageNumber="467">
6A. Dorsal surface of telson with two pairs of small cuspidate setae, both pairs situated in posterior third of telson; eyestalk with cornea not particularly reduced, anteromesial surface with very small blunt tubercle; major and minor cheliped ischia with two spiniform setae; orbital teeth present in form of low bumps or subtriangular lobes [coral reefs and adjacent rubble flats; depth range:
<quantity id="4CD99B1EFFECFFDBFEBB194C06FFFC9C" box="[292,385,799,821]" metricMagnitude="0" metricUnit="m" metricValue="7.75" metricValueMax="15.0" metricValueMin="0.5" pageId="17" pageNumber="467" unit="m" value="7.75" valueMax="15.0" valueMin="0.5">0.515 m</quantity>
; southern Gulf of Mexico and
<collectingCountry id="F336766BFFECFFDBFD37194D05A3FC9C" box="[680,733,798,820]" name="Cuba" pageId="17" pageNumber="467">Cuba</collectingCountry>
]...............................................
<taxonomicName id="4C214D78FFECFFDBFAB9194C02E3FC9C" ID-CoL="85ZFR" box="[1318,1437,799,820]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="467" phylum="Arthropoda" rank="species" species="alacranes">
<emphasis id="B955EAE9FFECFFDBFAB9194C02E3FC9C" box="[1318,1437,799,820]" italics="true" pageId="17" pageNumber="467">T. alacranes</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9E36FBFFECFFDBFF08196F02E2FC6F" blockId="17.[151,1437,711,967]" pageId="17" pageNumber="467">
<keyLead id="30D095CEFFECFFDBFF08196F02E2FC6F" pageId="17" pageNumber="467">
6B. Dorsal surface of telson with two pairs (rarely with three pairs) of fairly large and robust cuspidate setae, first pair situated at mid-length of telson; eyestalk with cornea somewhat reduced, anteromesial surface with conspicuous conical tubercle; major and minor cheliped ischia with three spiniform setae; orbital teeth absent [sand flats with abundant coral rubble close to sea grass and mangroves; depth range:
<quantity id="4CD99B1EFFECFFDBFDAA19C705F8FC02" box="[565,646,916,938]" metricMagnitude="0" metricUnit="m" metricValue="1.25" metricValueMax="2.0" metricValueMin="0.5" pageId="17" pageNumber="467" unit="m" value="1.25" valueMax="2.0" valueMin="0.5">0.52 m</quantity>
;
<collectingCountry id="F336766BFFECFFDBFD0F19C605A0FC02" box="[656,734,917,938]" name="Panama" pageId="17" pageNumber="467">Panama</collectingCountry>
, possibly also southern Gulf of Mexico, as
<taxonomicName id="4C214D78FFECFFDBFBE019C603A2FC02" ID-CoL="85ZFV" authorityName="in Anker" authorityYear="2010" box="[1151,1244,916,938]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="467" phylum="Arthropoda" rank="species" species="toro">
<emphasis id="B955EAE9FFECFFDBFBE019C603ECFC02" box="[1151,1170,917,938]" italics="true" pageId="17" pageNumber="467">T.</emphasis>
cf.
<emphasis id="B955EAE9FFECFFDBFB2A19C603A2FC02" box="[1205,1244,917,938]" italics="true" pageId="17" pageNumber="467">toro</emphasis>
</taxonomicName>
, see above].....................................................................................................
<taxonomicName id="4C214D78FFECFFDBFAC119E102E2FC6F" ID-CoL="85ZFV" authorityName="in Anker" authorityYear="2010" box="[1374,1436,946,967]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="467" phylum="Arthropoda" rank="species" species="toro">
<emphasis id="B955EAE9FFECFFDBFAC119E102E2FC6F" box="[1374,1436,946,967]" italics="true" pageId="17" pageNumber="467">T. toro</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>