274 lines
31 KiB
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274 lines
31 KiB
XML
<document ID-CLB-Dataset="51924" ID-DOI="http://dx.doi.org/10.3897/zookeys.1142.90097" ID-GBIF-Dataset="c2e00414-9633-4890-ab72-cb2aa3e9d174" ID-PMC="PMC10208988" ID-Pensoft-Pub="1313-2970-1142-1" ID-Pensoft-UUID="1D4BDF04F72B59B9984E00D0B4FB338D" ID-PubMed="37250203" ID-ZooBank="A1129EE50F9941CFB73AE771B66E2486" ModsDocID="1313-2970-1142-1" checkinTime="1674655519517" checkinUser="pensoft" docAuthor="Tongkerd, Piyoros, Tumpeesuwan, Sakboworn, Inkhavilay, Khamla, Prasankok, Pongpun, Jeratthitikul, Ekgachai, Panha, Somsak & Sutcharit, Chirasak" docDate="2023" docId="D1130B9897CF5F868B096212955B437C" docLanguage="en" docName="ZooKeys 1142: 1-144" docOrigin="ZooKeys 1142" docPubDate="2023-01-24" docSource="http://dx.doi.org/10.3897/zookeys.1142.90097" docTitle="Rhiostoma platymorpha Tongkerd & Tumpeesuwan 2023, sp. nov." docType="treatment" docUuid="13B875AD-3E60-431E-B4B2-51F3FC5E03A0" docUuidSource="ZooBank" docVersion="5" id="1D4BDF04F72B59B9984E00D0B4FB338D" lastPageNumber="1" masterDocId="1D4BDF04F72B59B9984E00D0B4FB338D" masterDocTitle="Systematic revision of the snorkel snail genus Rhiostoma Benson, 1860 (Gastropoda, Caenogastropoda, Cyclophoridae) with descriptions of new species" masterLastPageNumber="144" masterPageNumber="1" pageNumber="1" updateTime="1732652259105" updateUser="ExternalLinkService">
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<mods:titleInfo id="403C71FF1BC02806087609965E6ADC51">
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<mods:title id="CC6FC2797E819A53D4154394DCE396E4">Systematic revision of the snorkel snail genus Rhiostoma Benson, 1860 (Gastropoda, Caenogastropoda, Cyclophoridae) with descriptions of new species</mods:title>
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<mods:name id="0E7E70522445DD187E2FC825AB554F6D" type="personal">
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<mods:roleTerm id="82491150AA710BC49806DEA6A9A29434">Author</mods:roleTerm>
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<mods:namePart id="E600F4CC3966559A8BD1B50D0FB355A3">Tongkerd, Piyoros</mods:namePart>
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<mods:affiliation id="384838B337543B3CD5E3947A8DD5EB4D">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:name id="985AAD49C91B78B2F85129982AA74DFF" type="personal">
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<mods:roleTerm id="F1E7268382AB977C7246C2DEE23D0B5C">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7521D670B15DC8ED46C407F1A03377F2">Tumpeesuwan, Sakboworn</mods:namePart>
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<mods:affiliation id="938028222CC94BF7A3020A02D81B713E">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand & Present address: Department of Biology, Faculty of Science, Mahasarakham University, Kantharawichai, Maha Sarakham 44150 Thailand</mods:affiliation>
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</mods:name>
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<mods:name id="55E5BACBDA000095AA46888F30375363" type="personal">
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<mods:role id="250323149BB2CE3FE3EAB59C243479E2">
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<mods:roleTerm id="9F5C86613CF8A5C56091D95C8824982B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E21C3453521322763C0BE1C53BC4191E">Inkhavilay, Khamla</mods:namePart>
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<mods:affiliation id="C0CCA3F2ACA1DA407D8CE8A3BA3B8F36">Department of Biology, Faculty of Natural Science, National University of Laos, P. O. Box 7322, Dongdok, Vientiane, Laos</mods:affiliation>
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</mods:name>
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<mods:name id="2D25B63E6825E2DE310CBAFACDBFD9A6" type="personal">
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<mods:roleTerm id="D517096AADFCF70DE866D4B6D9B595B5">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F0C1D4FDE0A6CAE6DBB1426AA51EA690">Prasankok, Pongpun</mods:namePart>
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<mods:affiliation id="FB8E7DFD3E05C94A55F7A4D68626EC1C">School of Biology, Institute of Science, Suranaree University of Technology, Nakhon Ratchasima 30000, Thailand</mods:affiliation>
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</mods:name>
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<mods:name id="619247D6D0F7FC7DDDE28B09018C0896" type="personal">
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<mods:namePart id="54649D7982AE59AFC8039949C779E719">Jeratthitikul, Ekgachai</mods:namePart>
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<mods:affiliation id="EEA226374F1C133EC9E57197831D374C">Animal Systematics and Molecular Ecology Laboratory, Department of Biology, Faculty of Science, Mahidol University, Bangkok 10400, Thailand</mods:affiliation>
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</mods:name>
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<mods:name id="827F85E00D6254139E03A26F91855F21" type="personal">
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<mods:namePart id="C1B6EFE99FE69AECD18A1454D54A9AE3">Panha, Somsak</mods:namePart>
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<mods:affiliation id="205ED10A922DB29D2C17068FF3D68E84">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:nameIdentifier id="3988B30C8173457F19073F750EFB3E7B" type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
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<mods:namePart id="122C873C3AFE4237291A9F13BBD29FD5">Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation id="F398816D1355A618B996F04EF8594274">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:nameIdentifier id="CF1DBDAE6D7EFBCFC6DBCBCFF8FAE1C5" type="email">jirasak4@yahoo.com</mods:nameIdentifier>
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<mods:typeOfResource id="221D406BA82B10F4542056C9024F6DDC">text</mods:typeOfResource>
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<mods:title id="A499804C37EF5F90ADA4114809136CD1">ZooKeys</mods:title>
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<mods:part id="C3320B402954539562E0A1B12D542057">
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<mods:date id="3EB99112FAED2772FDD8C6AB0E7546E0">2023</mods:date>
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<mods:number id="8CE0F2B5ADE2795AABFD4E0C69CF2887">2023-01-24</mods:number>
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<mods:detail id="32E6BF0DD2A39DECCC6932C8AB866B8F" type="volume">
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<mods:number id="98501D81B68DF5D1A780DA2FFE93C483">1142</mods:number>
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<mods:start id="095C38E6530E1D68209413EB8370F759">1</mods:start>
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<mods:end id="E5D3287A8DFB17CF61AD46814031D0E6">144</mods:end>
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<mods:url id="3F7EE78FB2F799541869A8B089C05D47">http://dx.doi.org/10.3897/zookeys.1142.90097</mods:url>
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<mods:classification id="591C494A5EDB4D766D3017DF59DA374D">journal article</mods:classification>
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<mods:identifier id="F6EBF3558CC8599A24520FF03145E85B" type="DOI">http://dx.doi.org/10.3897/zookeys.1142.90097</mods:identifier>
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<mods:identifier id="3C2554EBB219C71AC2F85C8EC2F5249F" type="Pensoft-Pub">1313-2970-1142-1</mods:identifier>
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<mods:identifier id="C93AE7C389EFC1358FA759A7BC338168" type="ZooBank">A1129EE50F9941CFB73AE771B66E2486</mods:identifier>
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<treatment id="D1130B9897CF5F868B096212955B437C" ID-GBIF-Taxon="209102256" LSID="urn:lsid:zoobank.org:act:13B875AD-3E60-431E-B4B2-51F3FC5E03A0" httpUri="http://treatment.plazi.org/id/D1130B9897CF5F868B096212955B437C" lastPageNumber="1" pageId="0" pageNumber="1" scope_class="Gastropoda" scope_family="Cyclophoridae" scope_subClass="Caenogastropoda" scope_subFamily="Cyclophorinae">
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<subSubSection id="8998B6BE5AB682DBE07260EE87434F06" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="4409DB847402D5A51F26BF1BF94DCB95" pageId="0" pageNumber="1">
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20.
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<taxonomicName id="552614505D0041F264C6C91BDA2DA7EE" ID-CoL="B3CK8" LSID="https://zoobank.org/13B875AD-3E60-431E-B4B2-51F3FC5E03A0" authority="Tongkerd & Tumpeesuwan" authorityName="Tongkerd & Tumpeesuwan" authorityYear="2023" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Rhiostoma platymorpha" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="platymorpha" status="sp. nov.">Rhiostoma platymorpha Tongkerd & Tumpeesuwan</taxonomicName>
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<taxonomicNameLabel id="526FD52BB83112AFE3B92E589F4BBA85" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="D6FED9076E3F64B2BB55C79F12904B72" pageId="0" pageNumber="1" type="description">
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<paragraph id="B8C1F66E31A0DD57D9E9941AFCFEE650" pageId="0" pageNumber="1">
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<figureCitation id="5B8392A5E871D5FE98ABFE0A3BCB5D21" captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Distribution map of Rhiostoma asiphon - species group." figureDoi="10.3897/zookeys.1142.90097.figure34" httpUri="https://binary.pensoft.net/fig/799892" pageId="0" pageNumber="1">Figs 34</figureCitation>
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<figureCitation id="702B7823B5E860FAD03159F98C6D4BB9" captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Shell of Rhiostoma platymorpha sp. nov. A holotype CUMZ 4498 / 1 from Tham Muang On, Mae On, Chiang Mai B paratype CUMZ 4763 from type locality C, D paratypes CUMZ 4498 / 2 from type locality E, F specimens CUMZ 4783 from Tham Bua Tong, Mae Tang, Chiang Mai (with F albinistic shell form)." figureDoi="10.3897/zookeys.1142.90097.figure41" httpUri="https://binary.pensoft.net/fig/799899" pageId="0" pageNumber="1">, 41</figureCitation>
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<figureCitation id="3DD25EFE189182C8B4E06DFE3C9414D8" captionStart="Figure 42" captionStartId="F42" captionText="Figure 42. Radula morphology A Rhiostoma asiphon CUMZ 4826 from Angthong Islands, Surat Thani B Rhiostoma strubelli specimen CUMZ 10200 from Vieng Phuka, Luang Namtha, Laos C Rhiostoma breviocollar sp. nov. paratypes CUMZ 4491 from type locality D Rhiostoma anceyi sp. nov. paratype CUMZ 4494 / 2 from type locality E Rhiostoma furfurosum sp. nov. paratype CUMZ 4705 / 2 from type locality F Rhiostoma platymorpha sp. nov. paratype CUMZ 4763 from type locality." figureDoi="10.3897/zookeys.1142.90097.figure42" httpUri="https://binary.pensoft.net/fig/799900" pageId="0" pageNumber="1">, 42F</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="537AE77231994B4BB3D59377CD573F12" pageId="0" pageNumber="1" type="reference_group">
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<paragraph id="940E64F35608F823004654EFA77FB21E" pageId="0" pageNumber="1">
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<taxonomicName id="6D34EA61D72A73CF82CEBC7F09700161" authority="Tongkerd & Tumpeesuwan & Inkhavilay & Prasankok & Jeratthitikul & Panha & Sutcharit, 2023" authorityName="Tongkerd & Tumpeesuwan & Inkhavilay & Prasankok & Jeratthitikul & Panha & Sutcharit" authorityYear="2023" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Rhiostoma" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus" status="sp. nov.">Rhiostoma</taxonomicName>
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sp. 1-
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<bibRefCitation id="B3F2B90845C22022F52F873E2B8A9CB9" author="Tumpeesuwan, S" journalOrPublisher="Master Thesis, Department of Biology, Faculty of Science, Chulalongkorn University" pageId="0" pageNumber="1" refId="B175" refString="Tumpeesuwan, S, 2001. Taxonomy of snorkel snails genus Rhiostoma Benson, 1860 in Thailand. Master Thesis, Department of Biology, Faculty of Science, Chulalongkorn University" title="Taxonomy of snorkel snails genus Rhiostoma Benson, 1860 in Thailand." year="2001">Tumpeesuwan 2001</bibRefCitation>
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: 59-61, figs 4.19-4.21 (in part).
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</paragraph>
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</subSubSection>
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<subSubSection id="E65C54E7B1D7EF1436AAD7E38E692AE3" pageId="0" pageNumber="1" type="type material">
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<paragraph id="6F916F9953BEA8BAA88D135A4F0695C1" pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph id="5E46CE2725D1E6582B0B7E2A6EE3ADC7" pageId="0" pageNumber="1">
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<emphasis id="60C17BA527E0FA4030EC78ADB53F6758" bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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CUMZ 4498/1 (cW 19.0 mm, cH 8.4 mm, dL 7.2 mm; Fig.
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<figureCitation id="3A46243736BD5FDA7186C6231F726260" captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Shell of Rhiostoma platymorpha sp. nov. A holotype CUMZ 4498 / 1 from Tham Muang On, Mae On, Chiang Mai B paratype CUMZ 4763 from type locality C, D paratypes CUMZ 4498 / 2 from type locality E, F specimens CUMZ 4783 from Tham Bua Tong, Mae Tang, Chiang Mai (with F albinistic shell form)." figureDoi="10.3897/zookeys.1142.90097.figure41" httpUri="https://binary.pensoft.net/fig/799899" pageId="0" pageNumber="1">41A</figureCitation>
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).
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<emphasis id="D897B242E55B38ECA9785F6CD8082C71" bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
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CUMZ 3976 (22 shells), CUMZ 4349 (1 shell), CUMZ 4436 (4 adults + 1 juvenile), CUMZ 4498/2 (6 shells; Fig.
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<figureCitation id="EF808FB8DF029849A5A7FAF546A0E216" captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Shell of Rhiostoma platymorpha sp. nov. A holotype CUMZ 4498 / 1 from Tham Muang On, Mae On, Chiang Mai B paratype CUMZ 4763 from type locality C, D paratypes CUMZ 4498 / 2 from type locality E, F specimens CUMZ 4783 from Tham Bua Tong, Mae Tang, Chiang Mai (with F albinistic shell form)." figureDoi="10.3897/zookeys.1142.90097.figure41" httpUri="https://binary.pensoft.net/fig/799899" pageId="0" pageNumber="1">41C, D</figureCitation>
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), CUMZ 4731 (1 adult + 1 juvenile), CUMZ 4763 (4 shells; Figs
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<figureCitation id="068ABACF92A490E78865F5EEE72C2AD4" captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Shell of Rhiostoma platymorpha sp. nov. A holotype CUMZ 4498 / 1 from Tham Muang On, Mae On, Chiang Mai B paratype CUMZ 4763 from type locality C, D paratypes CUMZ 4498 / 2 from type locality E, F specimens CUMZ 4783 from Tham Bua Tong, Mae Tang, Chiang Mai (with F albinistic shell form)." figureDoi="10.3897/zookeys.1142.90097.figure41" httpUri="https://binary.pensoft.net/fig/799899" pageId="0" pageNumber="1">41B</figureCitation>
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,
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<figureCitation id="8267B15189BED4E8E447B6DA546D092E" captionStart="Figure 42" captionStartId="F42" captionText="Figure 42. Radula morphology A Rhiostoma asiphon CUMZ 4826 from Angthong Islands, Surat Thani B Rhiostoma strubelli specimen CUMZ 10200 from Vieng Phuka, Luang Namtha, Laos C Rhiostoma breviocollar sp. nov. paratypes CUMZ 4491 from type locality D Rhiostoma anceyi sp. nov. paratype CUMZ 4494 / 2 from type locality E Rhiostoma furfurosum sp. nov. paratype CUMZ 4705 / 2 from type locality F Rhiostoma platymorpha sp. nov. paratype CUMZ 4763 from type locality." figureDoi="10.3897/zookeys.1142.90097.figure42" httpUri="https://binary.pensoft.net/fig/799900" pageId="0" pageNumber="1">42F</figureCitation>
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), CUMZ 4801 (48 shells), CUMZ 4809 (74 shells), CUMZ 4892 (10 adults + 1 juveniles), CUMZ 10196 (3 shells), CUMZ 10198 (12 adults + 2 juveniles), CUMZ 10199 (1 shell), NHMUK 20220445 (5 shells), and SMF 368680 (5 shells). All paratypes are from the type locality.
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</paragraph>
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<caption id="AFFAEDF5C43FB2E3DDB524EA7E279FAE" doi="10.3897/zookeys.1142.90097.figure41" httpUri="https://binary.pensoft.net/fig/799899" pageId="0" pageNumber="1" start="Figure 41" startId="F41">
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<paragraph id="6728BABB66AC893892CF5BF9E2FF056A" pageId="0" pageNumber="1">
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<emphasis id="49CA96F5CC28529FF6902BC6A2CF70D4" bold="true" pageId="0" pageNumber="1">Figure 41.</emphasis>
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Shell of
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<taxonomicName id="C55DE851A1B802A8F10A62C79FFABA88" authorityName="Tongkerd & Tumpeesuwan" authorityYear="2023" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma platymorpha" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="platymorpha">
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<emphasis id="581EC7810C1B3988630EC747F1B19952" italics="true" pageId="0" pageNumber="1">Rhiostoma platymorpha</emphasis>
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</taxonomicName>
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sp. nov.
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<emphasis id="62887315C238BD1AA4703597006DCC9F" bold="true" pageId="0" pageNumber="1">A</emphasis>
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holotype CUMZ 4498/1 from Tham Muang On, Mae On, Chiang Mai
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<emphasis id="772228E0BF19B35D14161818683D8209" bold="true" pageId="0" pageNumber="1">B</emphasis>
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paratype CUMZ 4763 from type locality
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<emphasis id="3F6D9791FF019143A1F98A2E49FBA01E" bold="true" pageId="0" pageNumber="1">C, D</emphasis>
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paratypes CUMZ 4498/2 from type locality
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<emphasis id="7B137758FDDA1598BFCFCF6CC392460B" bold="true" pageId="0" pageNumber="1">E, F</emphasis>
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specimens CUMZ 4783 from Tham Bua Tong, Mae Tang, Chiang Mai (with
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<emphasis id="8D123A88D39591A8D67E968488659293" bold="true" pageId="0" pageNumber="1">F</emphasis>
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albinistic shell form).
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</paragraph>
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</caption>
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<caption id="4B0F9CA58B085787ACA7119A791DCC84" doi="10.3897/zookeys.1142.90097.figure42" httpUri="https://binary.pensoft.net/fig/799900" pageId="0" pageNumber="1" start="Figure 42" startId="F42">
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<paragraph id="12C83EC526F44360914B5000D650E0E0" pageId="0" pageNumber="1">
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<emphasis id="B5190F6A69E778262A7C3F66561AB991" bold="true" pageId="0" pageNumber="1">Figure 42.</emphasis>
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Radula morphology
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<emphasis id="C50D479E9F7966A16817614EBDBBACB1" bold="true" pageId="0" pageNumber="1">A</emphasis>
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<taxonomicName id="ACBDBB33117115E6ACE1667180B7959C" authorityName="Moellendorff" authorityYear="1893" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma asiphon" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="asiphon">
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|
<emphasis id="8AB8BCBCAF64E41334122098CD88F6A3" italics="true" pageId="0" pageNumber="1">Rhiostoma asiphon</emphasis>
|
|
</taxonomicName>
|
|
CUMZ 4826 from Angthong Islands, Surat Thani
|
|
<emphasis id="1BFF5F3583CD7A17128436F144811C10" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
|
<taxonomicName id="2EB02BAF6D125C927522375E496F6825" authorityName="Moellendorff" authorityYear="1899" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma strubelli" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="strubelli">
|
|
<emphasis id="B9EC41B446C5D575C90686D00866E7A8" italics="true" pageId="0" pageNumber="1">Rhiostoma strubelli</emphasis>
|
|
</taxonomicName>
|
|
specimen CUMZ 10200 from Vieng Phuka, Luang Namtha, Laos
|
|
<emphasis id="DF7AC9901CE127B9311734109580938A" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
|
<taxonomicName id="ABB449111B678A41240820CBC9E1FD6B" authorityName="Tongkerd & Tumpeesuwan" authorityYear="2023" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma breviocollar" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="breviocollar">
|
|
<emphasis id="DABBEFB7A67FDB07D19F5928A5FD0237" italics="true" pageId="0" pageNumber="1">Rhiostoma breviocollar</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. paratypes CUMZ 4491 from type locality
|
|
<emphasis id="645CD7F27B67E6A1C08F7931A7EE2E17" bold="true" pageId="0" pageNumber="1">D</emphasis>
|
|
<taxonomicName id="364A10154FAA87703E983C6EA2622873" authorityName="Tongkerd & Inkhavilay" authorityYear="2023" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma anceyi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="anceyi">
|
|
<emphasis id="C2959CFF926C795FF11835D9732EF7B9" italics="true" pageId="0" pageNumber="1">Rhiostoma anceyi</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. paratype CUMZ 4494/2 from type locality
|
|
<emphasis id="E1F18B7DBFA4373D4A981C95B3D9EA1D" bold="true" pageId="0" pageNumber="1">E</emphasis>
|
|
<taxonomicName id="F739CEE886A80DC73B0CE2D3EE4C76C7" authorityName="Tongkerd & Panha" authorityYear="2023" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma furfurosum" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="furfurosum">
|
|
<emphasis id="F793B27807A9F53108C4AE4821610002" italics="true" pageId="0" pageNumber="1">Rhiostoma furfurosum</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. paratype CUMZ 4705/2 from type locality
|
|
<emphasis id="B4874E7F29FFB947C5AB9C1FC5C03734" bold="true" pageId="0" pageNumber="1">F</emphasis>
|
|
<taxonomicName id="214AFF818E65F95B1E482A44B09C33C1" authorityName="Tongkerd & Tumpeesuwan" authorityYear="2023" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma platymorpha" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="platymorpha">
|
|
<emphasis id="54CAFEE5B9506E5907972EA6BE6C38B5" italics="true" pageId="0" pageNumber="1">Rhiostoma platymorpha</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. paratype CUMZ 4763 from type locality.
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection id="89D003462822B964929CE1815FF5EDB5" pageId="0" pageNumber="1" type="type locality">
|
|
<paragraph id="BD3B38F6D7E142FA3759BDEF08A48C82" pageId="0" pageNumber="1">Type locality.</paragraph>
|
|
<paragraph id="20AE2DA008C787C043337B4EB13BFB16" pageId="0" pageNumber="1">
|
|
Tham Muang On, Mae On District, Chiang Mai Province, Thailand (
|
|
<geoCoordinate id="93C677A017FF506CC79E7146153F4A26" degrees="18" direction="north" minutes="47" orientation="latitude" precision="1" seconds="10.6" value="18.786278">18°47'10.6"N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="2384FE7AE430CC772758A1B37B873629" degrees="99" direction="east" minutes="14" orientation="longitude" precision="1" seconds="17.1" value="99.23808">99°14'17.1"E</geoCoordinate>
|
|
). Limestone hill with dry evergreen forest. The locality is surrounded by a timber plantation and farms for annual crops.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="009186A8A8FCB0729B7EB7E30437E4EB" pageId="0" pageNumber="1" type="materials_examined">
|
|
<paragraph id="51B5D80D0BCFAD4B421D6F803FA672FC" pageId="0" pageNumber="1">Other material examined.</paragraph>
|
|
<paragraph id="B140401D8BC16BE4FEE8913BE3FF837B" pageId="0" pageNumber="1">
|
|
<emphasis id="BB356071B4F97869F890E22FE71F6205" bold="true" pageId="0" pageNumber="1">
|
|
<collectingCountry id="D7D8DCD7E559708842B2FAB955ADEE91" name="Thailand">Thailand</collectingCountry>
|
|
</emphasis>
|
|
: Tham Boa Tong, Mae Tang,
|
|
<collectingRegion id="9C6ED27C305450119CC7B0BBC922191F" country="Thailand" name="Chiang Mai">Chiang Mai</collectingRegion>
|
|
: CUMZ 4437, 4783 (Fig.
|
|
<figureCitation id="5F0807CF3F1200A34674581BEC1E019F" captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Shell of Rhiostoma platymorpha sp. nov. A holotype CUMZ 4498 / 1 from Tham Muang On, Mae On, Chiang Mai B paratype CUMZ 4763 from type locality C, D paratypes CUMZ 4498 / 2 from type locality E, F specimens CUMZ 4783 from Tham Bua Tong, Mae Tang, Chiang Mai (with F albinistic shell form)." figureDoi="10.3897/zookeys.1142.90097.figure41" httpUri="https://binary.pensoft.net/fig/799899" pageId="0" pageNumber="1">41E, F</figureCitation>
|
|
), 10197.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="24A03BE04B552FF87E6FB7D9C80BCADD" pageId="0" pageNumber="1" type="diagnosis">
|
|
<paragraph id="BC5F40DB50D5889C5E971426D79334AC" pageId="0" pageNumber="1">Diagnosis.</paragraph>
|
|
<paragraph id="1091054B8B831EA2AB569A95F9AF6D00" pageId="0" pageNumber="1">Shell small, thin, and flattened; detached whorl shorter than apertural width. Shell colour with brownish zigzag patterns on both sides. Breathing device with incomplete tube shape (rarely short tubular) and attached to preceding whorl.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="D4AD413B9380C4D3446A598797DEF605" pageId="0" pageNumber="1" type="differential diagnosis">
|
|
<paragraph id="DD3569765A4C8E7A65C04F8000A0637C" pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
|
|
<paragraph id="613750A42B978486034C21367D06536F" pageId="0" pageNumber="1">
|
|
This new species can be distinguished from
|
|
<taxonomicName id="713065C31ED864C993E070C8B7ABAB09" lsidName="R. strubelli" pageId="0" pageNumber="1" rank="species" species="strubelli">
|
|
<emphasis id="A4A9F16577D40FBB312D492DF6B65474" italics="true" pageId="0" pageNumber="1">R. strubelli</emphasis>
|
|
</taxonomicName>
|
|
by having a flattened shell, short detached whorl, and brownish zigzag colour pattern, usually on both the dorsal and ventral sides of the shell. In contrast,
|
|
<taxonomicName id="0443B23483B8515C5ACF40FD5D655519" lsidName="R. strubelli" pageId="0" pageNumber="1" rank="species" species="strubelli">
|
|
<emphasis id="34E56B31708E587BD59516443825CB73" italics="true" pageId="0" pageNumber="1">R. strubelli</emphasis>
|
|
</taxonomicName>
|
|
has a sub-discoidal to discoidal shell shape, and brownish zigzag pattern only on the dorsal side.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="F565390EC76F45E6045842DBB68622B4" pageId="0" pageNumber="1" type="description">
|
|
<paragraph id="0654D3E49A5A1D9E598B5201EF4EB5EE" pageId="0" pageNumber="1">Description.</paragraph>
|
|
<paragraph id="F52575C2180C3AA324A27C06B7C3D8C6" pageId="0" pageNumber="1">
|
|
<emphasis id="1D23D981C0A600F1003E0F10A3EED5F6" bold="true" pageId="0" pageNumber="1">
|
|
<emphasis id="D4822BA4EDA8A821032ABE9F80729015" italics="true" pageId="0" pageNumber="1">Shell</emphasis>
|
|
.
|
|
</emphasis>
|
|
Shell small, cW 17.2-20.2 mm, cH 8.8-11.3 mm, thin, and flattened to sub-discoidal shape; detached-whorl length 2.0-6.0 mm. Apex acute; spire nearly flattened to slightly elevated. Whorls 4 to 5, convex, increasing regularly; suture wide and shallow; last whorl rounded and slender. Shell surface with irregular growth lines. Periostracum corneous and transparent. Shell colour with dark to brownish zigzag pattern, and ventral side with paler pattern; narrow dark brown spiral band on periphery. Detached whorl shorter than apertural width and descending. Peristome circular and double; lip slightly thickened and not expanded. Breathing device with incomplete tube shape and its tip usually attached to preceding whorl; outer lip forming a long and nearly closed tube; inner lip with deep incision. Umbilicus widely opened and deep. Operculum calcareous, low cup-shaped, and multispiral (Fig.
|
|
<figureCitation id="0A56A44FAD27750C76A3ABA05BBA0B56" captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Shell of Rhiostoma platymorpha sp. nov. A holotype CUMZ 4498 / 1 from Tham Muang On, Mae On, Chiang Mai B paratype CUMZ 4763 from type locality C, D paratypes CUMZ 4498 / 2 from type locality E, F specimens CUMZ 4783 from Tham Bua Tong, Mae Tang, Chiang Mai (with F albinistic shell form)." figureDoi="10.3897/zookeys.1142.90097.figure41" httpUri="https://binary.pensoft.net/fig/799899" pageId="0" pageNumber="1">41</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="6B50A8E83DF534FF38A20374AE7B2E2D" pageId="0" pageNumber="1">
|
|
<emphasis id="D234233C0E9CCCF49D34A753908307DA" bold="true" pageId="0" pageNumber="1">
|
|
<emphasis id="A751DD3AC930D262244E42E2D303032E" italics="true" pageId="0" pageNumber="1">Radula</emphasis>
|
|
.
|
|
</emphasis>
|
|
Teeth arrangement and morphology are similar to those of
|
|
<taxonomicName id="015C939A79C04148173579F628FD0D31" lsidName="R. asiphon" pageId="0" pageNumber="1" rank="species" species="asiphon">
|
|
<emphasis id="387FB9B266486EAEE3C595D60DC32AFF" italics="true" pageId="0" pageNumber="1">R. asiphon</emphasis>
|
|
</taxonomicName>
|
|
. Central tooth with large central cusp and two lateral cusps on each side. Lateral teeth composed of four cusps; central cusp large with pointed tip; two inner cusps with pointed tips; one outer cusp very small. Marginal teeth each composed of three pointed cusps (Fig.
|
|
<figureCitation id="520B4CA5EE38BD495B9FFE0422F84B41" captionStart="Figure 42" captionStartId="F42" captionText="Figure 42. Radula morphology A Rhiostoma asiphon CUMZ 4826 from Angthong Islands, Surat Thani B Rhiostoma strubelli specimen CUMZ 10200 from Vieng Phuka, Luang Namtha, Laos C Rhiostoma breviocollar sp. nov. paratypes CUMZ 4491 from type locality D Rhiostoma anceyi sp. nov. paratype CUMZ 4494 / 2 from type locality E Rhiostoma furfurosum sp. nov. paratype CUMZ 4705 / 2 from type locality F Rhiostoma platymorpha sp. nov. paratype CUMZ 4763 from type locality." figureDoi="10.3897/zookeys.1142.90097.figure42" httpUri="https://binary.pensoft.net/fig/799900" pageId="0" pageNumber="1">42F</figureCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="729AC84CC0B26B8BB8103BE00B2A75EC" pageId="0" pageNumber="1" type="etymology">
|
|
<paragraph id="28F455862653B0AB500B8D7381FDBE45" pageId="0" pageNumber="1">Etymology.</paragraph>
|
|
<paragraph id="680AC5CF76BE8584D42B7F57E8A42F9C" pageId="0" pageNumber="1">
|
|
The subspecific name
|
|
<emphasis id="BDFBF2B3BB583C85F8BF012E6212F8E6" italics="true" pageId="0" pageNumber="1">platymorpha</emphasis>
|
|
comes from two Latin words
|
|
<emphasis id="F3BD51521481458F176E207CACDE0661" italics="true" pageId="0" pageNumber="1">platy</emphasis>
|
|
, which means flat, and
|
|
<emphasis id="DC1B3313EAF7503CD505072590409B14" italics="true" pageId="0" pageNumber="1">morpha</emphasis>
|
|
meaning form. It refers to the prominent flattened shell shape of this species.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="D058C1B04A66994CABDFD33CB86A977E" pageId="0" pageNumber="1" type="distribution">
|
|
<paragraph id="1EDFADF4927E5B1A257B1B3172A6B9F6" pageId="0" pageNumber="1">Distribution.</paragraph>
|
|
<paragraph id="FED759C14426AC3B802D968695E56C8D" pageId="0" pageNumber="1">This new species is known from the type locality, limestone hills in Chiang Mai Province.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="7193FF33108D92C077FA709208C5A377" pageId="0" pageNumber="1" type="remarks">
|
|
<paragraph id="D924F80B7841B5BCEC213FFCC226E645" pageId="0" pageNumber="1">Remarks.</paragraph>
|
|
<paragraph id="F18FD1AA388BE1F724CD7B02D5D3DD82" pageId="0" pageNumber="1">
|
|
The specimens from Tham Buatong, Chaing Mai (Fig.
|
|
<figureCitation id="48624B417DB08A018EF185A71AEDA571" captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Shell of Rhiostoma platymorpha sp. nov. A holotype CUMZ 4498 / 1 from Tham Muang On, Mae On, Chiang Mai B paratype CUMZ 4763 from type locality C, D paratypes CUMZ 4498 / 2 from type locality E, F specimens CUMZ 4783 from Tham Bua Tong, Mae Tang, Chiang Mai (with F albinistic shell form)." figureDoi="10.3897/zookeys.1142.90097.figure41" httpUri="https://binary.pensoft.net/fig/799899" pageId="0" pageNumber="1">41E, F</figureCitation>
|
|
), tend to differ from the type specimen in having sub-discoidal shells. However, the brownish zigzag colour pattern appears on the dorsal and ventral shell surface, and an incomplete tube-shaped breathing device suggests they are closely related to this species. That being noted, a few specimens with monochrome whitish shells also occur in this population; this is probably an albinistic shell (Fig.
|
|
<figureCitation id="44B2B50550E08504DE5D245E3E07B5FD" captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Shell of Rhiostoma platymorpha sp. nov. A holotype CUMZ 4498 / 1 from Tham Muang On, Mae On, Chiang Mai B paratype CUMZ 4763 from type locality C, D paratypes CUMZ 4498 / 2 from type locality E, F specimens CUMZ 4783 from Tham Bua Tong, Mae Tang, Chiang Mai (with F albinistic shell form)." figureDoi="10.3897/zookeys.1142.90097.figure41" httpUri="https://binary.pensoft.net/fig/799899" pageId="0" pageNumber="1">41F</figureCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |