278 lines
26 KiB
XML
278 lines
26 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1030.62544" ID-GBIF-Dataset="91fec855-7fa9-4ad6-84d7-9b9d35e23e8b" ID-PMC="PMC8060247" ID-Pensoft-Pub="1313-2970-1030-1" ID-Pensoft-UUID="875199397EEE5F7898EA1DB25DA62D25" ID-PubMed="33958904" ModsDocID="1313-2970-1030-1" checkinTime="1618371561543" checkinUser="pensoft" docAuthor="Means, Jackson C., Hennen, Derek A. & Marek, Paul E." docDate="2021" docId="07444855538E52A68D6FA30699B51E02" docLanguage="en" docName="ZooKeys 1030: 1-180" docOrigin="ZooKeys 1030" docPubDate="2021-04-13" docSource="http://dx.doi.org/10.3897/zookeys.1030.62544" docTitle="Nannaria oblonga" docType="treatment" docVersion="5" id="875199397EEE5F7898EA1DB25DA62D25" lastPageNumber="1" masterDocId="875199397EEE5F7898EA1DB25DA62D25" masterDocTitle="A revision of the minor species group in the millipede genus Nannaria Chamberlin, 1918 (Diplopoda, Polydesmida, Xystodesmidae)" masterLastPageNumber="180" masterPageNumber="1" pageNumber="1" updateTime="1673961214978" updateUser="julia">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A revision of the minor species group in the millipede genus Nannaria Chamberlin, 1918 (Diplopoda, Polydesmida, Xystodesmidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Means, Jackson C.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7377-0696</mods:nameIdentifier>
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<mods:affiliation>Virginia Tech, Department of Entomology, Blacksburg, Virginia 24061, USA</mods:affiliation>
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<mods:nameIdentifier type="email">mjacks4@vt.edu</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Hennen, Derek A.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7005-1151</mods:nameIdentifier>
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<mods:affiliation>Virginia Tech, Department of Entomology, Blacksburg, Virginia 24061, USA</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Marek, Paul E.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-7048-2514</mods:nameIdentifier>
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<mods:affiliation>Virginia Tech, Department of Entomology, Blacksburg, Virginia 24061, USA</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-04-13</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1030</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>180</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1030.62544</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1030.62544</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1030-1</mods:identifier>
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<mods:identifier type="Pensoft-UUID">875199397EEE5F7898EA1DB25DA62D25</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="181004764" LSID="urn:lsid:plazi:treatment:07444855538E52A68D6FA30699B51E02" httpUri="http://treatment.plazi.org/id/07444855538E52A68D6FA30699B51E02" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName ID-CoL="45M4W" LSID="07444855-538E-52A6-8D6F-A30699B51E02" authority="(Koch, 1847)" baseAuthorityName="Koch" baseAuthorityYear="1847" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Nannaria oblonga" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="oblonga">Nannaria oblonga (Koch, 1847)</taxonomicName>
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<figureCitation captionStart="Figure 110" captionStartId="F110" captionText="Figure 110. Nannaria oblonga syntype ♂ (MFN) left gonopod A anterior view; red triangle indicates crochet-like acropodite tip B medial view C posterior view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1030.62544.figure110" httpUri="https://binary.pensoft.net/fig/531213" pageId="0" pageNumber="1">Fig. 110 Vernacular Name: "The Pennsylvania Twisted-Claw Millipede"</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="reference_group">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Koch" authorityYear="1847" class="Diplopoda" family="Xystodesmidae" genus="Fontaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fontaria oblonga" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="oblonga">Fontaria oblonga</taxonomicName>
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C.L. Koch, 1847: 142. C.L.
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<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.10639" author="Koch, CL" journalOrPublisher="Getreu Nach Der Natur Abgebildet Und Beschrieben" pageId="0" pageNumber="1" pagination="1 - 134" refId="B35" refString="Koch, CL, 1863. Die Myriapoden. Getreu Nach Der Natur Abgebildet Und Beschrieben 1: 1 - 134, DOI: https://doi.org/10.5962/bhl.title.10639" title="Die Myriapoden." url="https://doi.org/10.5962/bhl.title.10639" volume="1" year="1863">Koch 1863</bibRefCitation>
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: 73-74, fig. 64.
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<bibRefCitation author="Attems, CG" journalOrPublisher="Denkschriften der Kaiserlichen Akademieder Wissenschaften zu Wien, Mathematisch-Naturwissenschaftlische Classe" pageId="0" pageNumber="1" pagination="221 - 482" refId="B2" refString="Attems, CG, 1898. System der Polydesmiden, I. Theil. Denkschriften der Kaiserlichen Akademieder Wissenschaften zu Wien, Mathematisch-Naturwissenschaftlische Classe 67: 221 - 482" title="System der Polydesmiden, I. Theil." volume="67" year="1898">Attems 1898</bibRefCitation>
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: 263.
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<bibRefCitation DOI="https://doi.org/10.1515/9783111430645-003" author="Attems, CG" journalOrPublisher="Das Tierreich" pageId="0" pageNumber="1" pagination="1 - 487" refId="B3" refString="Attems, CG, 1938. Polydesmoidea II. Fam. Leptodesmidae, Platyrhachidae, Oxydesmidae, Gomphodesmidae. Das Tierreich 69: 1 - 487, DOI: https://doi.org/10.1515/9783111430645-003" title="Polydesmoidea II. Fam. Leptodesmidae, Platyrhachidae, Oxydesmidae, Gomphodesmidae." url="https://doi.org/10.1515/9783111430645-003" volume="69" year="1938">Attems 1938</bibRefCitation>
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: 167.
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<bibRefCitation DOI="https://doi.org/10.5479/si.03629236.212" author="Chamberlin, RV" journalOrPublisher="United States National Museum Bulletin" pageId="0" pageNumber="1" pagination="1 - 236" refId="B22" refString="Chamberlin, RV, Hoffman, RL, 1958. Checklist of the millipeds of North America. United States National Museum Bulletin 212: 1 - 236, DOI: https://doi.org/10.5479/si.03629236.212" title="Checklist of the millipeds of North America." url="https://doi.org/10.5479/si.03629236.212" volume="212" year="1958">Chamberlin and Hoffman 1958</bibRefCitation>
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: 55.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName baseAuthorityName="Koch" baseAuthorityYear="1847" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Nannaria oblonga" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="oblonga">Nannaria oblonga</taxonomicName>
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:
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<bibRefCitation author="Hoffman, RL" journalOrPublisher="Virginia Museum of Natural History Special Publication Number" pageId="0" pageNumber="1" pagination="1 - 584" refId="B33" refString="Hoffman, RL, 1999. Checklist of the millipeds of North and Middle America. Virginia Museum of Natural History Special Publication Number 8: 1 - 584" title="Checklist of the millipeds of North and Middle America." volume="8" year="1999">Hoffman 1999</bibRefCitation>
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: 367.
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<bibRefCitation author="Marek, PE" journalOrPublisher="Virginia Museum of Natural History Publications" pageId="0" pageNumber="1" pagination="1 - 117" refId="B46" refString="Marek, PE, Tanabe, T, Sierwald, P, 2014. A species catalog of the millipede family Xystodesmidae (Diplopoda: Polydesmida). Virginia Museum of Natural History Publications 17: 1 - 117" title="A species catalog of the millipede family Xystodesmidae (Diplopoda: Polydesmida)." volume="17" year="2014">Marek et al. 2014</bibRefCitation>
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: 37.
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<bibRefCitation DOI="https://doi.org/10.1093/isd/ixab003" author="Means, JC" journalOrPublisher="Insect Systematics and Diversity" pageId="0" pageNumber="1" pagination="1 - 26" refId="B52" refString="Means, JC, Hennen, DA, Tanabe, T, Marek, PE, 2021. Phylogenetic systematics of the millipede family Xystodesmidae. Insect Systematics and Diversity 5: 1 - 26, DOI: https://doi.org/10.1093/isd/ixab003" title="Phylogenetic systematics of the millipede family Xystodesmidae." url="https://doi.org/10.1093/isd/ixab003" volume="5" year="2021">Means et al. 2021</bibRefCitation>
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: S71.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<materialsCitation ID-GBIF-Occurrence="3071355349" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="United States" location="Pennsylvania" specimenCount="♂" stateProvince="Pennsylvania" typeStatus="Syntype">
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<emphasis bold="true" pageId="0" pageNumber="1">
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<typeStatus>Syntype</typeStatus>
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</emphasis>
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:
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<collectingCountry name="United States of America">United States</collectingCountry>
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-
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<emphasis bold="true" pageId="0" pageNumber="1">
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<collectingRegion country="United States of America" name="Pennsylvania">Pennsylvania</collectingRegion>
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</emphasis>
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•
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<specimenCount>♂</specimenCount>
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; from an unknown locality in
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<normalizedToken originalValue="“Pennsylvania”">"Pennsylvania"</normalizedToken>
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; MFN
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Adult males of
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<taxonomicName baseAuthorityName="Koch" baseAuthorityYear="1847" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria oblonga" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="oblonga">
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<emphasis italics="true" pageId="0" pageNumber="1">Nannaria oblonga</emphasis>
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</taxonomicName>
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are distinct from other
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<taxonomicName authorityName="Means & Hennen & Marek" authorityYear="2021" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Nannaria</emphasis>
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</taxonomicName>
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, the possibly sympatric
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<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. fowleri" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="fowleri">
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<emphasis italics="true" pageId="0" pageNumber="1">N. fowleri</emphasis>
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</taxonomicName>
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, and the nearby
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<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. shenandoa" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="shenandoa">
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<emphasis italics="true" pageId="0" pageNumber="1">N. shenandoa</emphasis>
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</taxonomicName>
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, based on the following combination of characters:
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Gonopods</emphasis>
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.
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</emphasis>
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Gonopodal acropodite gently curving medially before apex, not strongly curved as in
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<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. shenandoa" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="shenandoa">
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<emphasis italics="true" pageId="0" pageNumber="1">N. shenandoa</emphasis>
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</taxonomicName>
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. Distal zone short, simple, curving smoothly posteriorly, with a crochet-like appearance (Fig.
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<figureCitation captionStart="Figure 110" captionStartId="F110" captionText="Figure 110. Nannaria oblonga syntype ♂ (MFN) left gonopod A anterior view; red triangle indicates crochet-like acropodite tip B medial view C posterior view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1030.62544.figure110" httpUri="https://binary.pensoft.net/fig/531213" pageId="0" pageNumber="1">110A</figureCitation>
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, red triangle). Distal zone not directed medially as in
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<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. fowleri" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="fowleri">
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<emphasis italics="true" pageId="0" pageNumber="1">N. fowleri</emphasis>
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</taxonomicName>
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, or elongated, with flanges as in
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<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. shenandoa" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="shenandoa">
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<emphasis italics="true" pageId="0" pageNumber="1">N. shenandoa</emphasis>
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</taxonomicName>
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. Telopodite basal zone ca. 1/3 length of acropodite, not ca. 1/4 as in
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<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. shenandoa" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="shenandoa">
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<emphasis italics="true" pageId="0" pageNumber="1">N. shenandoa</emphasis>
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</taxonomicName>
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. Prefemur with prefemoral process thin, ca. 1/2 length of acropodite, curving medially to parallel acropodite near tip, not elongated, 1/3 length of acropodite and paralleling acropodite majority of length as in
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<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. fowleri" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="fowleri">
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<emphasis italics="true" pageId="0" pageNumber="1">N. fowleri</emphasis>
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</taxonomicName>
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, or curving laterally as in
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<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. shenandoa" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="shenandoa">
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<emphasis italics="true" pageId="0" pageNumber="1">N. shenandoa</emphasis>
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</taxonomicName>
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. Prefemoral spine lacking, not pronounced as tooth-like structure as in
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<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. fowleri" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="fowleri">
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<emphasis italics="true" pageId="0" pageNumber="1">N. fowleri</emphasis>
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</taxonomicName>
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.
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Color</emphasis>
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.
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</emphasis>
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<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.49866" author="Koch, CL" journalOrPublisher="Baendchen, Regensburg" pageId="0" pageNumber="1" refId="B34" refString="Koch, CL, 1847. System der Myriapoden mit den Verzeichnissen und Berichtigungen zu Deutschlands Crustaceen, Myriapoden und Arachniden, in: Kritische Revision der Insectenfaune Deutschlands, III. Baendchen, Regensburg, DOI: https://doi.org/10.5962/bhl.title.49866" title="System der Myriapoden mit den Verzeichnissen und Berichtigungen zu Deutschlands Crustaceen, Myriapoden und Arachniden, in: Kritische Revision der Insectenfaune Deutschlands, III." url="https://doi.org/10.5962/bhl.title.49866" year="1847">Koch (1847)</bibRefCitation>
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describes
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<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. oblonga" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="oblonga">
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<emphasis italics="true" pageId="0" pageNumber="1">N. oblonga</emphasis>
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</taxonomicName>
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as being dark brown with white paranota.
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</paragraph>
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<caption doi="10.3897/zookeys.1030.62544.figure110" httpUri="https://binary.pensoft.net/fig/531213" pageId="0" pageNumber="1" start="Figure 110" startId="F110">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 110.</emphasis>
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<taxonomicName baseAuthorityName="Koch" baseAuthorityYear="1847" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria oblonga" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="oblonga">
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<emphasis italics="true" pageId="0" pageNumber="1">Nannaria oblonga</emphasis>
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</taxonomicName>
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syntype ♂ (MFN) left gonopod
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<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
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anterior view; red triangle indicates crochet-like acropodite tip
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<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
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medial view
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<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
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posterior view. Scale bar: 0.5 mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Measurements.</paragraph>
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<paragraph pageId="0" pageNumber="1">The type is too fragmented for accurate measurement.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="variation">
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<paragraph pageId="0" pageNumber="1">Variation.</paragraph>
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<paragraph pageId="0" pageNumber="1">No known variation.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="distribution">
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<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Known only from the type locality (Pennsylvania, Fig.
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<figureCitation captionStart="Figure 127" captionStartId="F127" captionText="Figure 127. Distribution map of Nannaria minor species group collection localities in the northern Appalachians." figureDoi="10.3897/zookeys.1030.62544.figure127" httpUri="https://binary.pensoft.net/fig/531230" pageId="0" pageNumber="1">127</figureCitation>
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). Distribution area: N/A; status: MRE.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="ecology">
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<paragraph pageId="0" pageNumber="1">Ecology.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.49866" author="Koch, CL" journalOrPublisher="Baendchen, Regensburg" pageId="0" pageNumber="1" refId="B34" refString="Koch, CL, 1847. System der Myriapoden mit den Verzeichnissen und Berichtigungen zu Deutschlands Crustaceen, Myriapoden und Arachniden, in: Kritische Revision der Insectenfaune Deutschlands, III. Baendchen, Regensburg, DOI: https://doi.org/10.5962/bhl.title.49866" title="System der Myriapoden mit den Verzeichnissen und Berichtigungen zu Deutschlands Crustaceen, Myriapoden und Arachniden, in: Kritische Revision der Insectenfaune Deutschlands, III." url="https://doi.org/10.5962/bhl.title.49866" year="1847">Koch (1847)</bibRefCitation>
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does not mention any ecological notes in his description of the species.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="etymology">
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<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.49866" author="Koch, CL" journalOrPublisher="Baendchen, Regensburg" pageId="0" pageNumber="1" refId="B34" refString="Koch, CL, 1847. System der Myriapoden mit den Verzeichnissen und Berichtigungen zu Deutschlands Crustaceen, Myriapoden und Arachniden, in: Kritische Revision der Insectenfaune Deutschlands, III. Baendchen, Regensburg, DOI: https://doi.org/10.5962/bhl.title.49866" title="System der Myriapoden mit den Verzeichnissen und Berichtigungen zu Deutschlands Crustaceen, Myriapoden und Arachniden, in: Kritische Revision der Insectenfaune Deutschlands, III." url="https://doi.org/10.5962/bhl.title.49866" year="1847">Koch (1847)</bibRefCitation>
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does not give an etymology for the name
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<emphasis italics="true" pageId="0" pageNumber="1">oblonga</emphasis>
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in his 1863 publication, he goes into more detail about the syntype, and describes the species as
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<normalizedToken originalValue="“länglich,”">"laenglich,"</normalizedToken>
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meaning elongated or oblong in German. It is therefore reasonable to assume that the name is in reference to the general oblong shape of
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<taxonomicName authorityName="Means & Hennen & Marek" authorityYear="2021" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Nannaria</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="note on clade membership">
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<paragraph pageId="0" pageNumber="1">Note on clade membership.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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No specimens of
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<taxonomicName baseAuthorityName="Koch" baseAuthorityYear="1847" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria oblonga" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="oblonga">
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<emphasis italics="true" pageId="0" pageNumber="1">Nannaria oblonga</emphasis>
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</taxonomicName>
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were collected for this study, and while
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<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. oblonga" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="oblonga">
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<emphasis italics="true" pageId="0" pageNumber="1">N. oblonga</emphasis>
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</taxonomicName>
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shares some gonopodal characters with
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<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. tenuis" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="tenuis">
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<emphasis italics="true" pageId="0" pageNumber="1">N. tenuis</emphasis>
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</taxonomicName>
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sp. nov. (such as the caudally-directed acropodite tip and a lack of a prefemoral spine)
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<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. oblonga" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="oblonga">
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<emphasis italics="true" pageId="0" pageNumber="1">N. oblonga</emphasis>
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</taxonomicName>
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is geographically disjunct from the
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<taxonomicName authorityName="Means & Hennen & Marek" authorityYear="2021" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria ignis" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ignis">
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<emphasis italics="true" pageId="0" pageNumber="1">Nannaria ignis</emphasis>
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</taxonomicName>
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clade, to which
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<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. tenuis" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="tenuis">
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<emphasis italics="true" pageId="0" pageNumber="1">N. tenuis</emphasis>
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</taxonomicName>
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sp. nov. belongs. Due to a lack of molecular data and a paucity of shared morphological characters, clade membership is uncertain.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type locality">
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<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
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<paragraph pageId="0" pageNumber="1">United States, Pennsylvania; MFN.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="notes">
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<paragraph pageId="0" pageNumber="1">Notes.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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In the original publication,
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<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.49866" author="Koch, CL" journalOrPublisher="Baendchen, Regensburg" pageId="0" pageNumber="1" refId="B34" refString="Koch, CL, 1847. System der Myriapoden mit den Verzeichnissen und Berichtigungen zu Deutschlands Crustaceen, Myriapoden und Arachniden, in: Kritische Revision der Insectenfaune Deutschlands, III. Baendchen, Regensburg, DOI: https://doi.org/10.5962/bhl.title.49866" title="System der Myriapoden mit den Verzeichnissen und Berichtigungen zu Deutschlands Crustaceen, Myriapoden und Arachniden, in: Kritische Revision der Insectenfaune Deutschlands, III." url="https://doi.org/10.5962/bhl.title.49866" year="1847">Koch (1847</bibRefCitation>
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: 142) did not designate a type or mention the number of specimens he examined and later on,
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<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.10639" author="Koch, CL" journalOrPublisher="Getreu Nach Der Natur Abgebildet Und Beschrieben" pageId="0" pageNumber="1" pagination="1 - 134" refId="B35" refString="Koch, CL, 1863. Die Myriapoden. Getreu Nach Der Natur Abgebildet Und Beschrieben 1: 1 - 134, DOI: https://doi.org/10.5962/bhl.title.10639" title="Die Myriapoden." url="https://doi.org/10.5962/bhl.title.10639" volume="1" year="1863">Koch (1863)</bibRefCitation>
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described both a male and female specimen in greater detail. The specimen in the MFN collection was labeled
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<normalizedToken originalValue="“Holotypus.”">"Holotypus."</normalizedToken>
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Without knowing the number of specimens Koch examined for his original description of the species, we cannot be sure if it is a holotype by monotypy, and hence refrain from designating a lectotype.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |