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<mods:title id="D6ADA2C25859919A59304A10CCCBF7BB">Scorpions of Sri Lanka (Scorpiones Buthidae, Chaerilidae, Scorpionidae) with description of four new species of the genera Charmus Karsch, 1879 and Reddyanus Vachon, 1972, stat n</mods:title>
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<taxonomicName id="FFD84D4AA876FFF1FEECFE5D9CD7FE1F" ID-CoL="T9Z5" authority="Pocock, 1894" authorityName="Pocock" authorityYear="1894" box="[300,693,394,418]" class="Arachnida" family="Chaerilidae" genus="Chaerilus" kingdom="Animalia" order="Scorpiones" pageId="97" pageNumber="96" phylum="Arthropoda" rank="species" species="ceylonensis">
<emphasis id="0AACEADBA876FFF1FEECFE5D9C7FFE1F" bold="true" box="[300,541,394,418]" italics="true" pageId="97" pageNumber="96">Chaerilus ceylonensis</emphasis>
<bibRefCitation id="5C494B38A876FFF1FDE4FE5C9CD7FE1F" author="Pocock" box="[548,693,395,418]" firstAuthor="Pocock" journalOrPublisher="Annals and Magazine of Natural History" pageId="97" pageNumber="96" pagination="72 - 84" part="6" refId="ref43722" refString="POCOCK, R. I. 1894. A small contribution to our knowledge of the scorpions of India. Annals and Magazine of Natural History, 6 (13): 72 - 84." title="A small contribution to our knowledge of the scorpions of India" type="journal article" year="1894">Pocock, 1894</bibRefCitation>
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(
<figureCitation id="A0E32A4CA876FFF1FEB4FE7D9FAEFE7C" box="[372,460,426,449]" captionStart="Figures 1215" captionStartId="6.[192,278,1885,1909]" captionTargetBox="[191,1440,167,1856]" captionTargetId="figure-11@6.[192,1439,168,1855]" captionTargetPageId="6" captionText="Figures 1215: Maps showing the known geographical distribution of the Sri Lankan species." figureDoi="http://doi.org/10.5281/zenodo.7124476" httpUri="https://zenodo.org/record/7124476/files/figure.png" pageId="97" pageNumber="96">Figs. 13</figureCitation>
,
<figureCitation id="A0E32A4CA876FFF1FE19FE7D9C66FE7C" box="[473,516,426,449]" captionStart="Figures 193200" captionStartId="48.[192,278,1070,1094]" captionTargetBox="[213,1446,171,1037]" captionTargetId="figure-11@48.[205,1452,168,1041]" captionTargetPageId="48" captionText="Figures 193200: Distal segments of leg IV, retroventral view of Sri Lankan Buthidae and Chaerilidae genera. Buthoscorpio sarasinorum, male from locality 15CF (193), Charmus laneus, male from locality 15CO (194), Hottentotta tamulus, male from locality 15CK (195), Isometrus maculatus, female from locality 15CI (196), Isometrus thwaitesi, female from locality 15CO (197), Lychas srilankensis, male from locality 15CJ (198), Reddyanus basilicus, male from locality 15CS (199), and Chaerilus ceylonensis, male from locality 15CD (200)." figureDoi="http://doi.org/10.5281/zenodo.7124608" httpUri="https://zenodo.org/record/7124608/files/figure.png" pageId="97" pageNumber="96">200</figureCitation>
,
<figureCitation id="A0E32A4CA876FFF1FDCEFE7D9C0EFE7C" box="[526,620,426,449]" captionStart-0="Figures 403429" captionStart-1="Figures 430435" captionStart-2="Figures 436439" captionStart-3="Figures 440445" captionStart-4="Figures 446453" captionStart-5="Figures 454457" captionStartId-0="85.[192,278,744,768]" captionStartId-1="87.[192,278,1684,1708]" captionStartId-2="88.[192,278,1829,1853]" captionStartId-3="89.[192,278,1198,1222]" captionStartId-4="90.[192,278,1664,1688]" captionStartId-5="91.[192,278,1853,1877]" captionTargetBox-1="[212,1427,152,1653]" captionTargetBox-2="[254,1381,199,1801]" captionTargetBox-3="[201,1437,172,1165]" captionTargetBox-4="[200,1434,229,1621]" captionTargetBox-5="[213,1417,187,1825]" captionTargetId-0="figure-11@84.[172,1464,185,1878]" captionTargetId-1="figure-6@87.[192,1439,144,1654]" captionTargetId-2="figure-84@88.[255,1380,200,1800]" captionTargetId-3="figure-706@89.[192,1439,168,1169]" captionTargetId-4="figure-244@90.[192,1439,227,1632]" captionTargetId-5="figure-46@91.[214,1416,188,1824]" captionTargetPageId-0="84" captionTargetPageId-1="87" captionTargetPageId-2="88" captionTargetPageId-3="89" captionTargetPageId-4="90" captionTargetPageId-5="91" captionText-0="Figures 403429: Telson, lateral view of Sri Lankan Buthidae and Chaerilidae species. Figures 403404. Isometrus maculatus, male from locality 15CP (403) and female from locality 15CI (404). Figures 405406. I. thwaitesi, male from locality 15CH (405) and female from locality 15CO (406). Figures 407408. Lychas srilankensis, male (407) and female (408) from locality 15CJ. Figures 409410. Reddyanus basilicus, male (409) and female (410) from locality 15CR. Figures 411412. R. besucheti, male holotype (411) and female from locality 15CG (412). Figures 413414. R. ceylonensis sp. n., male holotype (413) and female paratype (414) from locality 15CI. Figures 415416. R. jayarathnei sp. n., male (415) and female (416) paratypes. Figures 417418. R. loebli, male (417) and female (418) from locality 15CG. Figures 419420. R. ranawanai sp. n., male holotype (419) and female paratype (420). Figures 421422: Buthoscorpio sarasinorum, male from locality 15CF (421) and female from locality 15CG (422). Figures 423424. Charmus laneus, male (423) and female (424) from locality 15CO. Figures 425426. Charmus saradieli sp. n. male paratype (425) (MHNG) and female holotype (426). Figurs 427. Hottentotta tamulus, male from locality 15CK (427). Figures 428429. Chaerilus ceylonensis, male (428) and female (429) from locality 15CD." captionText-1="Figures 430435: Chaerilus ceylonensis from locality 15CD. Figures 430431. Male in dorsal (430) and ventral (431) views. Figures 432435. Female, dorsal (432) and ventral (433) views, and chelicera ventral (434) and dorsal (435) views." captionText-2="Figures 436439: Chaerilus ceylonensis from locality 15CD. Figures 436, 438. Male, chelicerae, carapace and tergites III (436), and sternopectinal region with sternite III (438). Figures 437, 439. Female, chelicerae, carapace and tergites III (437), and sternopectinal region (439)." captionText-3="Figures 440445: Chaerilus ceylonensis from locality 15CD. Figures 440442. Male, metasoma and telson, lateral (440), ventral (441), and dorsal (442) views. Figures 443445. Female, metasoma and telson, lateral (443), ventral (444), and dorsal (445) views." captionText-4="Figures 446453: Chaerilus ceylonensis from locality 15CD. Figures 446449. Male. Pedipalp chela, dorsal (446), external (447) and ventral (448) views. Movable finger (449). Figures 450453. Female. Pedipalp chela, dorsal (450), external (451) and ventral (452) views. Movable finger (453)." captionText-5="Figures 454457: Chaerilus ceylonensis from locality 15CD, male (454), female (455), female with juveniles after first ecdysis (456), and female with newborns (457)." figureDoi-0="http://doi.org/10.5281/zenodo.7124696" figureDoi-1="http://doi.org/10.5281/zenodo.7124698" figureDoi-2="http://doi.org/10.5281/zenodo.7124702" figureDoi-3="http://doi.org/10.5281/zenodo.7124704" figureDoi-4="http://doi.org/10.5281/zenodo.7124708" figureDoi-5="http://doi.org/10.5281/zenodo.7124710" httpUri-0="https://zenodo.org/record/7124696/files/figure.png" httpUri-1="https://zenodo.org/record/7124698/files/figure.png" httpUri-2="https://zenodo.org/record/7124702/files/figure.png" httpUri-3="https://zenodo.org/record/7124704/files/figure.png" httpUri-4="https://zenodo.org/record/7124708/files/figure.png" httpUri-5="https://zenodo.org/record/7124710/files/figure.png" pageId="97" pageNumber="96">428457</figureCitation>
)
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<emphasis id="0AACEADBA876FFF1FF00FE3A9FD1FDB9" box="[192,435,493,516]" italics="true" pageId="97" pageNumber="96">Chaerilus ceylonensis</emphasis>
<bibRefCitation id="5C494B38A876FFF1FE01FE3A9CD1FDB9" author="Pocock" box="[449,691,493,516]" firstAuthor="Pocock" journalOrPublisher="Annals and Magazine of Natural History" page="83 - 84" pageId="97" pageNumber="96" pagination="72 - 84" part="6" refId="ref43722" refString="POCOCK, R. I. 1894. A small contribution to our knowledge of the scorpions of India. Annals and Magazine of Natural History, 6 (13): 72 - 84." title="A small contribution to our knowledge of the scorpions of India" type="journal article" year="1894">Pocock, 1894: 8384</bibRefCitation>
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;
<treatmentCitation id="B97910D8A876FFF1FD08FE3A9F3EFD9E" author="VACHON, M." page="102" pageId="97" pageNumber="96" year="1982">
<bibRefCitation id="5C494B38A876FFF1FD08FE3A9F3EFD9E" author="Vachon" firstAuthor="Vachon" journalOrPublisher="Revue suisse de Zoologie" page="102" pageId="97" pageNumber="96" pagination="77 - 114" part="89" refId="ref45190" refString="VACHON, M. 1982. Les scorpions de Sri Lanka (Recherches sur les scorpions appartenant ou deposes au Museum d`Histoire naturelle de Geneve III.). Revue suisse de Zoologie, 89: 77 - 114." title="Les scorpions de Sri Lanka (Recherches sur les scorpions appartenant ou deposes au Museum d`Histoire naturelle de Geneve III.)" type="journal article" year="1982">Vachon, 1982: 102</bibRefCitation>
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, figs. 9495;
<treatmentCitation id="B97910D8A876FFF1FE2FFDDB9F2FFDFF" author="TIKADER B. K. &amp; D. B. BASTAWADE" page="326" pageId="97" pageNumber="96" year="1983">
<bibRefCitation id="5C494B38A876FFF1FE2FFDDB9F2FFDFF" author="Tikader &amp; Bastawade" bookContentInfo="668 pp." firstAuthor="Tikader" journalOrPublisher="Zoological Survey of India, Calcutta" page="326 - 332" pageId="97" pageNumber="96" refId="ref44789" refString="TIKADER B. K. &amp; D. B. BASTAWADE. 1983. The Fauna of India: Scorpions. Scorpionida, Arachnida. Vol III. Zoological Survey of India, Calcutta, 668 pp." title="The Fauna of India: Scorpions. Scorpionida, Arachnida. Vol III" type="book" year="1983">Tikader &amp; Bastawade, 1983: 326332</bibRefCitation>
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, figs. 925939;
<treatmentCitation id="B97910D8A876FFF1FDC9FDFC9CCEFDFF" author="Fet" box="[521,684,555,578]" page="325" pageId="97" pageNumber="96" year="2000">
<bibRefCitation id="5C494B38A876FFF1FDC9FDFC9CCEFDFF" author="Fet" box="[521,684,555,578]" firstAuthor="Fet" page="325" pageId="97" pageNumber="96" refId="ref40873" refString="FET, V., W. D. SISSOM, G. LOWE &amp; M. E. BRAUNWALDER. 2000. Catalog of the Scorpions of the World (1758 - 1998). New York: The New York Entomological Society, 689 pp." type="book" year="2000">Fet, 2000: 325</bibRefCitation>
</treatmentCitation>
(complete reference list until 1998); Kovařík, 2000: 4445, fig. 14;
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<bibRefCitation id="5C494B38A876FFF1FED9FDBF9C02FDC2" author="Lourenco &amp; Huber" box="[281,608,616,639]" firstAuthor="Lourenco" journalOrPublisher="Revue suisse de Zoologie" page="273" pageId="97" pageNumber="96" pagination="265 - 275" part="109" refId="ref43004" refString="LOURENCO, W. R. &amp; D. HUBER. 2002. New addition to the scorpion fauna (Arachnida: Scorpiones) of Sri Lanka. Revue suisse de Zoologie, 109 (2): 265 - 275." title="New addition to the scorpion fauna (Arachnida: Scorpiones) of Sri Lanka" type="journal article" year="2002">Lourenço &amp; Huber, 2002: 273</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="B97910D8A876FFF1FDABFDBF9FCAFD23" author="KOVARIK, F &amp; A. A. OJANGUREN AFFILASTRO" page="135" pageId="97" pageNumber="96" year="2013">
<bibRefCitation id="5C494B38A876FFF1FDABFDBF9FCAFD23" author="Kovarik &amp; Ojanguren" bookContentInfo="400 pp." firstAuthor="Kovarik" journalOrPublisher="Clairon Production, Prague" page="135" pageId="97" pageNumber="96" refId="ref42080" refString="KOVARIK, F &amp; A. A. OJANGUREN AFFILASTRO. 2013. Illustrated catalog of scorpions Part II. Bothriuridae; Chaerilidae; Buthidae I., genera Compsobuthus, Hottentotta, Isometrus, Lychas, and Sassanidotus. Clairon Production, Prague, 400 pp." title="Illustrated catalog of scorpions Part II. Bothriuridae; Chaerilidae; Buthidae I., genera Compsobuthus, Hottentotta, Isometrus, Lychas, and Sassanidotus" type="book" year="2013">Kovařík &amp; Ojanguren, 2013: 135</bibRefCitation>
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, 284, figs. 732733.
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LOCALITY AND
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REPOSITORY.
<collectingCountry id="40CF7659A876FFF1FDB3FD1D9CBFFD5C" box="[627,733,714,737]" name="Sri Lanka" pageId="97" pageNumber="96">Sri Lanka</collectingCountry>
(Ceylon),
<location id="3D076012A876FFF1FF38FD3E9FE0FCBD" LSID="urn:lsid:plazi:treatment:B07187DFA876FFF1FEECFE5D9FB4F8E4:3D076012A876FFF1FF38FD3E9FE0FCBD" box="[248,386,745,768]" country="Sri Lanka" name="Trincomalee" pageId="97" pageNumber="96">Trincomalee</location>
;
<collectionCode id="5EC9AE0CA876FFF1FE4DFD3E9F83FCBD" box="[397,481,745,768]" pageId="97" pageNumber="96">BMNH</collectionCode>
.
</paragraph>
<paragraph id="386736C9A876FFF1FF00FCFB9C74FC3C" blockId="97.[192,802,812,897]" pageId="97" pageNumber="96">
TYPE MATERIAL EXAMINED.
<materialsCitation id="88B03C94A876FFF1FE33FCFB9C74FC3C" ID-GBIF-Occurrence="3913992310" collectionCode="BMNH" country="Sri Lanka" location="Trincomalee" pageId="97" pageNumber="96" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<collectingCountry id="40CF7659A876FFF1FE33FCFB9C3DFCFE" box="[499,607,812,835]" name="Sri Lanka" pageId="97" pageNumber="96">Sri Lanka</collectingCountry>
,
<location id="3D076012A876FFF1FDABFCFB9C96FCFE" LSID="urn:lsid:plazi:treatment:B07187DFA876FFF1FEECFE5D9FB4F8E4:3D076012A876FFF1FDABFCFB9C96FCFE" box="[619,756,812,835]" country="Sri Lanka" municipality="Ojanguren" name="Trincomalee" pageId="97" pageNumber="96">Trincomalee</location>
,
<specimenCount id="2EDEFD40A876FFF1FCC0FCFB9D43FCFE" box="[768,801,812,835]" pageId="97" pageNumber="96" type="male">1♂</specimenCount>
(
<typeStatus id="E763886BA876FFF1FF07FC9C9F44FCDF" box="[199,294,843,866]" pageId="97" pageNumber="96" type="holotype">holotype</typeStatus>
, figs.
<quantity id="FF209B2CA876FFF1FEAAFC9C9F86FCDF" box="[362,484,843,866]" metricMagnitude="1" metricUnit="m" metricValue="1.86055" metricValueMax="1.8618199999999998" metricValueMin="1.85928" pageId="97" pageNumber="96" unit="in" value="732.5" valueMax="733.0" valueMin="732.0">732733 in</quantity>
<bibRefCitation id="5C494B38A876FFF1FE2BFC9C9E88FC3C" author="Kovarik &amp; Ojanguren" bookContentInfo="400 pp." firstAuthor="Kovarik" journalOrPublisher="Clairon Production, Prague" page="284" pageId="97" pageNumber="96" refId="ref42080" refString="KOVARIK, F &amp; A. A. OJANGUREN AFFILASTRO. 2013. Illustrated catalog of scorpions Part II. Bothriuridae; Chaerilidae; Buthidae I., genera Compsobuthus, Hottentotta, Isometrus, Lychas, and Sassanidotus. Clairon Production, Prague, 400 pp." title="Illustrated catalog of scorpions Part II. Bothriuridae; Chaerilidae; Buthidae I., genera Compsobuthus, Hottentotta, Isometrus, Lychas, and Sassanidotus" type="book" year="2013">Kovařík &amp; Ojanguren, 2013: 284</bibRefCitation>
),
<collectionCode id="5EC9AE0CA876FFF1FF3EFCBD9F2CFC3C" box="[254,334,874,897]" pageId="97" pageNumber="96">BMNH</collectionCode>
No. 1893.10.20.4.
</materialsCitation>
</paragraph>
<paragraph id="386736C9A876FFF1FF00FC7A9EB8FACC" blockId="97.[192,802,941,1393]" pageId="97" pageNumber="96">
OTHER MATERIAL EXAMINED.
<materialsCitation id="88B03C94A876FFF1FDCEFC7A9C10FBC1" ID-GBIF-Occurrence="3913992335" collectingDate="2015-04-22" collectionCode="FKCP, UPSL" collectorName="Kovarik" country="Sri Lanka" elevation="132" latitude="7.954194" location="Dambula" longLatPrecision="1" longitude="80.91261" municipality="Polonnaruwa District" pageId="97" pageNumber="96" specimenCount="6" specimenCount-female="2" specimenCount-male="3" stateProvince="North Central">
<collectingCountry id="40CF7659A876FFF1FDCEFC7A9C18FC79" box="[526,634,941,964]" name="Sri Lanka" pageId="97" pageNumber="96">Sri Lanka</collectingCountry>
,
<collectingRegion id="FA1CF82BA876FFF1FD4BFC7A9F40FC5E" country="Sri Lanka" name="North Central" pageId="97" pageNumber="96">North Central Province</collectingRegion>
,
<collectingMunicipality id="D803ACB3A876FFF1FEF2FC1B9C7FFC5E" box="[306,541,972,995]" pageId="97" pageNumber="96">Polonnaruwa District</collectingMunicipality>
, ca
<quantity id="FF209B2CA876FFF1FD91FC1B9CF8FC5E" box="[593,666,972,995]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="97" pageNumber="96" unit="km" value="35.0">35 km</quantity>
from
<location id="3D076012A876FFF1FD25FC1B9E93FBBC" LSID="urn:lsid:plazi:treatment:B07187DFA876FFF1FEECFE5D9FB4F8E4:3D076012A876FFF1FD25FC1B9E93FBBC" country="Sri Lanka" latitude="7.954194" longLatPrecision="1" longitude="80.91261" municipality="Polonnaruwa District" name="Dambula" pageId="97" pageNumber="96" stateProvince="North Central">Dambula</location>
,
<geoCoordinate id="5DEC500EA876FFF1FF3BFC3D9FEEFBBC" box="[251,396,1002,1025]" degrees="07" direction="north" minutes="57" orientation="latitude" pageId="97" pageNumber="96" precision="1" seconds="15.1" value="7.954194">07°57'15.1&quot;N</geoCoordinate>
<geoCoordinate id="5DEC500EA876FFF1FE53FC3D9C51FBBC" box="[403,563,1002,1025]" degrees="080" direction="east" minutes="54" orientation="longitude" pageId="97" pageNumber="96" precision="1" seconds="45,4" value="80.91261">080°54'45,4&quot;E</geoCoordinate>
,
<elevation id="B3F5D1FAA876FFF1FDFDFC3D9CD8FBBC" box="[573,698,1002,1025]" metricMagnitude="2" metricUnit="m" metricValue="1.3199999999999998" pageId="97" pageNumber="96" unit="m" value="132.0">
<quantity id="FF209B2CA876FFF1FDFDFC3D9CE3FBBC" box="[573,641,1002,1025]" metricMagnitude="2" metricUnit="m" metricValue="1.3199999999999998" pageId="97" pageNumber="96" unit="m" value="132.0">132 m</quantity>
a.s.l.
</elevation>
(Locality
<emphasis id="0AACEADBA876FFF1FF00FBDF9F63FB9D" bold="true" box="[192,257,1032,1056]" pageId="97" pageNumber="96">15CD</emphasis>
,
<figureCitation id="A0E32A4CA876FFF1FED1FBDE9F0EFB9D" box="[273,364,1033,1056]" captionStart="Figures 577578" captionStartId="120.[192,278,1768,1792]" captionTargetBox="[191,1440,191,1739]" captionTargetId="figure-109@120.[192,1439,192,1738]" captionTargetPageId="120" captionText="Figures 577578: Figure 577. Locality 15CC, Sri Lanka, Central Province, Kandy District, 20 km S Kandy, Meegammana, Wattegama, 07°20'41.9&quot;N 080°39'30.6&quot;E, 534 m a.s.l.. We collected a female of Heterometrus indus close to a house on 21.IV.2015 evening. There also lives Isometrus sp. Figure 578. Locality 15CD, Sri Lanka, North Central Province, Polonnaruwa District, ca 35 km from Dambula, 07°57'15.1&quot;N 080°54'45,4&quot;E, 132 m a.s.l. During the day 22.IV.2015 under stones we recorded Reddyanus besucheti (close to the type locality) and Chaerilus ceylonensis." figureDoi="http://doi.org/10.5281/zenodo.7124775" httpUri="https://zenodo.org/record/7124775/files/figure.png" pageId="97" pageNumber="96">Fig. 578</figureCitation>
),
<date id="4C661009A876FFF1FE44FBDE9C62FB9D" box="[388,512,1033,1056]" pageId="97" pageNumber="96" value="2015-04-22">
<collectingDate id="5C22E9E1A876FFF1FE44FBDE9C62FB9D" box="[388,512,1033,1056]" pageId="97" pageNumber="96" value="2015-04-22">22.IV.2015</collectingDate>
</date>
,
<specimenCount id="2EDEFD40A876FFF1FDCDFBDE9C4DFB9D" box="[525,559,1033,1056]" pageId="97" pageNumber="96" type="male">2♂</specimenCount>
(
<figureCitation id="A0E32A4CA876FFF1FDFFFBDE9CCAFB9D" box="[575,680,1033,1056]" captionStart="Figures 193200" captionStartId="48.[192,278,1070,1094]" captionTargetBox="[213,1446,171,1037]" captionTargetId="figure-11@48.[205,1452,168,1041]" captionTargetPageId="48" captionText="Figures 193200: Distal segments of leg IV, retroventral view of Sri Lankan Buthidae and Chaerilidae genera. Buthoscorpio sarasinorum, male from locality 15CF (193), Charmus laneus, male from locality 15CO (194), Hottentotta tamulus, male from locality 15CK (195), Isometrus maculatus, female from locality 15CI (196), Isometrus thwaitesi, female from locality 15CO (197), Lychas srilankensis, male from locality 15CJ (198), Reddyanus basilicus, male from locality 15CS (199), and Chaerilus ceylonensis, male from locality 15CD (200)." figureDoi="http://doi.org/10.5281/zenodo.7124608" httpUri="https://zenodo.org/record/7124608/files/figure.png" pageId="97" pageNumber="96">Figs. 200</figureCitation>
,
<figureCitation id="A0E32A4CA876FFF1FD75FBDE9C82FB9D" box="[693,736,1033,1056]" captionStart="Figures 403429" captionStartId="85.[192,278,744,768]" captionTargetId="figure-11@84.[172,1464,185,1878]" captionTargetPageId="84" captionText="Figures 403429: Telson, lateral view of Sri Lankan Buthidae and Chaerilidae species. Figures 403404. Isometrus maculatus, male from locality 15CP (403) and female from locality 15CI (404). Figures 405406. I. thwaitesi, male from locality 15CH (405) and female from locality 15CO (406). Figures 407408. Lychas srilankensis, male (407) and female (408) from locality 15CJ. Figures 409410. Reddyanus basilicus, male (409) and female (410) from locality 15CR. Figures 411412. R. besucheti, male holotype (411) and female from locality 15CG (412). Figures 413414. R. ceylonensis sp. n., male holotype (413) and female paratype (414) from locality 15CI. Figures 415416. R. jayarathnei sp. n., male (415) and female (416) paratypes. Figures 417418. R. loebli, male (417) and female (418) from locality 15CG. Figures 419420. R. ranawanai sp. n., male holotype (419) and female paratype (420). Figures 421422: Buthoscorpio sarasinorum, male from locality 15CF (421) and female from locality 15CG (422). Figures 423424. Charmus laneus, male (423) and female (424) from locality 15CO. Figures 425426. Charmus saradieli sp. n. male paratype (425) (MHNG) and female holotype (426). Figurs 427. Hottentotta tamulus, male from locality 15CK (427). Figures 428429. Chaerilus ceylonensis, male (428) and female (429) from locality 15CD." figureDoi="http://doi.org/10.5281/zenodo.7124696" httpUri="https://zenodo.org/record/7124696/files/figure.png" pageId="97" pageNumber="96">428</figureCitation>
, 430 431, 436, 438, 440442, 446449, 454)
<specimenCount id="2EDEFD40A876FFF1FD40FBFF9CC0FB82" box="[640,674,1064,1087]" pageId="97" pageNumber="96" type="female">1♀</specimenCount>
(
<figureCitation id="A0E32A4CA876FFF1FD73FBFF9D7CFB82" box="[691,798,1064,1087]" captionStart="Figures 403429" captionStartId="85.[192,278,744,768]" captionTargetId="figure-11@84.[172,1464,185,1878]" captionTargetPageId="84" captionText="Figures 403429: Telson, lateral view of Sri Lankan Buthidae and Chaerilidae species. Figures 403404. Isometrus maculatus, male from locality 15CP (403) and female from locality 15CI (404). Figures 405406. I. thwaitesi, male from locality 15CH (405) and female from locality 15CO (406). Figures 407408. Lychas srilankensis, male (407) and female (408) from locality 15CJ. Figures 409410. Reddyanus basilicus, male (409) and female (410) from locality 15CR. Figures 411412. R. besucheti, male holotype (411) and female from locality 15CG (412). Figures 413414. R. ceylonensis sp. n., male holotype (413) and female paratype (414) from locality 15CI. Figures 415416. R. jayarathnei sp. n., male (415) and female (416) paratypes. Figures 417418. R. loebli, male (417) and female (418) from locality 15CG. Figures 419420. R. ranawanai sp. n., male holotype (419) and female paratype (420). Figures 421422: Buthoscorpio sarasinorum, male from locality 15CF (421) and female from locality 15CG (422). Figures 423424. Charmus laneus, male (423) and female (424) from locality 15CO. Figures 425426. Charmus saradieli sp. n. male paratype (425) (MHNG) and female holotype (426). Figurs 427. Hottentotta tamulus, male from locality 15CK (427). Figures 428429. Chaerilus ceylonensis, male (428) and female (429) from locality 15CD." figureDoi="http://doi.org/10.5281/zenodo.7124696" httpUri="https://zenodo.org/record/7124696/files/figure.png" pageId="97" pageNumber="96">Figs. 429</figureCitation>
, 432435, 437, 439, 443445, 450453, 455),
<collectionCode id="5EC9AE0CA876FFF1FD18FB919D7FFBE0" box="[728,797,1094,1117]" pageId="97" pageNumber="96">FKCP</collectionCode>
,
<specimenCount id="2EDEFD40A876FFF1FF00FBB29E83FBC1" box="[192,225,1125,1148]" pageId="97" pageNumber="96" type="male">1♂</specimenCount>
<specimenCount id="2EDEFD40A876FFF1FF21FBB29F61FBC1" box="[225,259,1125,1148]" pageId="97" pageNumber="96" type="female">1♀</specimenCount>
<specimenCount id="2EDEFD40A876FFF1FEC3FBB29F5AFBC1" box="[259,312,1125,1148]" pageId="97" pageNumber="96" type="generic">1juv.</specimenCount>
,
<collectionCode id="5EC9AE0CA876FFF1FE8CFBB29FF2FBC1" box="[332,400,1125,1148]" pageId="97" pageNumber="96">UPSL</collectionCode>
, leg.
<collectorName id="952D531FA876FFF1FE15FBB29C4FFBC1" box="[469,557,1125,1148]" pageId="97" pageNumber="96">Kovařík</collectorName>
et al
</materialsCitation>
.;
<materialsCitation id="88B03C94A876FFF1FD48FBB29C23FB4A" ID-GBIF-Occurrence="3913992329" collectingDate="2015-04-23" collectionCode="FKCP, UPSL" collectorName="Kovarik" country="Sri Lanka" elevation="101" latitude="8.144611" location="Kaudulla National Park" longLatPrecision="15" longitude="80.85111" municipality="Polonnaruwa District" pageId="97" pageNumber="96" specimenCount="5" specimenCount-female="1" specimenCount-male="2" stateProvince="North Central">
<collectingRegion id="FA1CF82BA876FFF1FD48FBB29F40FB26" country="Sri Lanka" name="North Central" pageId="97" pageNumber="96">North Central Province</collectingRegion>
,
<collectingMunicipality id="D803ACB3A876FFF1FEEFFB539C75FB26" box="[303,535,1156,1179]" pageId="97" pageNumber="96">Polonnaruwa District</collectingMunicipality>
, near
<location id="3D076012A876FFF1FD9BFB539E93FB04" LSID="urn:lsid:plazi:treatment:B07187DFA876FFF1FEECFE5D9FB4F8E4:3D076012A876FFF1FD9BFB539E93FB04" country="Sri Lanka" latitude="8.144611" longLatPrecision="15" longitude="80.85111" municipality="Polonnaruwa District" name="Kaudulla National Park" pageId="97" pageNumber="96" stateProvince="North Central">Kaudulla National Park</location>
,
<geoCoordinate id="5DEC500EA876FFF1FEC1FB759FF1FB04" box="[257,403,1186,1209]" degrees="08" direction="north" minutes="08" orientation="latitude" pageId="97" pageNumber="96" precision="1" seconds="40.6" value="8.144611">08°08'40.6&quot;N</geoCoordinate>
<geoCoordinate id="5DEC500EA876FFF1FE5DFB759C45FB04" box="[413,551,1186,1209]" degrees="080" direction="east" minutes="51" orientation="longitude" pageId="97" pageNumber="96" precision="15" seconds="04" value="80.85111">080°51'04&quot;E</geoCoordinate>
,
<elevation id="B3F5D1FAA876FFF1FDF5FB759CD4FB04" box="[565,694,1186,1209]" metricMagnitude="2" metricUnit="m" metricValue="1.01" pageId="97" pageNumber="96" unit="m" value="101.0">
<quantity id="FF209B2CA876FFF1FDF5FB759C1EFB04" box="[565,636,1186,1209]" metricMagnitude="2" metricUnit="m" metricValue="1.01" pageId="97" pageNumber="96" unit="m" value="101.0">101 m</quantity>
a.s.l.
</elevation>
(Locality
<emphasis id="0AACEADBA876FFF1FF00FB179E9DFB65" bold="true" box="[192,255,1216,1240]" pageId="97" pageNumber="96">15CF</emphasis>
,
<figureCitation id="A0E32A4CA876FFF1FEDCFB169FE7FB65" box="[284,389,1217,1240]" captionStart="Figures 579581" captionStartId="121.[192,278,1661,1685]" captionTargetBox="[213,1417,167,1633]" captionTargetId="figure-89@121.[214,1416,168,1632]" captionTargetPageId="121" captionText="Figures 579581: Figures 579580. Locality 15CE, Sri Lanka, North Central Province, Polonnaruwa District, Giritale, 08°01'26.0&quot;N 080°54'37.2&quot;E, 233 m a.s.l. During night collecting on 22.23.IV.2015 in the hotel garden (UV detection) we recorded Heterometrus swammerdami (inside the termite constructions, Fig. 580) and Heterometrus gravimanus inside burrows in open terrain. The mothers had 30 to 60 cm long oblique burrows usually once curved from which several narrow burrows created by their juveniles separated under surface. Every burrow of juveniles has a proper exit 15 to 90 cm from the main exit of the motherś burrow. On the locality we recorded night temperature 32.8 ºC 26.5 ºC (minimum temperature) and night humidity varied between 51% and 79%. Figure 581. Locality 15CF, Sri Lanka, North Central Province, Polonnaruwa District, near Kaudulla National Park, 08°08'40.6&quot;N 080°51'04&quot;E, 101 m a.s.l. During a day 23.IV.2015 we recorded Buthoscorpio sarasinorum (under stones or woods), Isometrus thwaitesi (under dry bark of standing trees), Reddyanus besucheti (under stones), and Chaerilus ceylonensis (under stones)." figureDoi="http://doi.org/10.5281/zenodo.7124777" httpUri="https://zenodo.org/record/7124777/files/figure.png" pageId="97" pageNumber="96">Fig. 581</figureCitation>
),
<date id="4C661009A876FFF1FE6BFB169C46FB65" box="[427,548,1217,1240]" pageId="97" pageNumber="96" value="2015-04-23">
<collectingDate id="5C22E9E1A876FFF1FE6BFB169C46FB65" box="[427,548,1217,1240]" pageId="97" pageNumber="96" value="2015-04-23">23.IV.2015</collectingDate>
</date>
,
<specimenCount id="2EDEFD40A876FFF1FD82FB169C01FB65" box="[578,611,1217,1240]" pageId="97" pageNumber="96" type="male">1♂</specimenCount>
<specimenCount id="2EDEFD40A876FFF1FDA3FB169CE7FB65" box="[611,645,1217,1240]" pageId="97" pageNumber="96" type="female">1♀</specimenCount>
<specimenCount id="2EDEFD40A876FFF1FD45FB169CDFFB65" box="[645,701,1217,1240]" pageId="97" pageNumber="96" type="generic">1juv.</specimenCount>
,
<collectionCode id="5EC9AE0CA876FFF1FD18FB169D79FB65" box="[728,795,1217,1240]" pageId="97" pageNumber="96">FKCP</collectionCode>
,
<specimenCount id="2EDEFD40A876FFF1FF00FB379E83FB4A" box="[192,225,1248,1271]" pageId="97" pageNumber="96" type="male">1♂</specimenCount>
<specimenCount id="2EDEFD40A876FFF1FF21FB379F75FB4A" box="[225,279,1248,1271]" pageId="97" pageNumber="96" type="generic">1juv.</specimenCount>
,
<collectionCode id="5EC9AE0CA876FFF1FEE8FB379F0BFB4A" box="[296,361,1248,1271]" pageId="97" pageNumber="96">UPSL</collectionCode>
, leg.
<collectorName id="952D531FA876FFF1FE6AFB379C60FB4A" box="[426,514,1248,1271]" pageId="97" pageNumber="96">Kovařík</collectorName>
et al
</materialsCitation>
.;
<materialsCitation id="88B03C94A876FFF1FD94FB379EB4FACC" ID-GBIF-Occurrence="3913992322" collectingDate="2015-04-28" collectionCode="FKCP, UPSL" collectorName="Kovarik" country="Sri Lanka" elevation="38" latitude="8.2875" location="Eluwankulam" longLatPrecision="15" longitude="79.844086" municipality="Puttalam District" pageId="97" pageNumber="96" specimenCount="2" specimenCount-male="2" stateProvince="North Central">
<collectingRegion id="FA1CF82BA876FFF1FD94FB379E9FFAA8" country="Sri Lanka" name="North Central" pageId="97" pageNumber="96">North Central Province</collectingRegion>
,
<collectingMunicipality id="D803ACB3A876FFF1FED5FB299FBBFAA8" box="[277,473,1278,1301]" pageId="97" pageNumber="96">Puttalam District</collectingMunicipality>
,
<location id="3D076012A876FFF1FE35FB299CEBFAA8" LSID="urn:lsid:plazi:treatment:B07187DFA876FFF1FEECFE5D9FB4F8E4:3D076012A876FFF1FE35FB299CEBFAA8" box="[501,649,1278,1301]" country="Sri Lanka" latitude="8.2875" longLatPrecision="15" longitude="79.844086" municipality="Puttalam District" name="Eluwankulam" pageId="97" pageNumber="96" stateProvince="North Central">Eluwankulam</location>
,
<geoCoordinate id="5DEC500EA876FFF1FD64FB299D43FAA8" box="[676,801,1278,1301]" degrees="08" direction="north" minutes="17" orientation="latitude" pageId="97" pageNumber="96" precision="15" seconds="15" value="8.2875">08°17'15&quot;N</geoCoordinate>
<geoCoordinate id="5DEC500EA876FFF1FF00FACA9F3DFA89" box="[192,351,1309,1332]" degrees="079" direction="east" minutes="50" orientation="longitude" pageId="97" pageNumber="96" precision="1" seconds="38. 7" value="79.844086">079°50'38.7&quot;E</geoCoordinate>
,
<elevation id="B3F5D1FAA876FFF1FEAEFACA9F84FA89" box="[366,486,1309,1332]" metricMagnitude="1" metricUnit="m" metricValue="3.8" pageId="97" pageNumber="96" unit="m" value="38.0">
<quantity id="FF209B2CA876FFF1FEAEFACA9FC8FA89" box="[366,426,1309,1332]" metricMagnitude="1" metricUnit="m" metricValue="3.8" pageId="97" pageNumber="96" unit="m" value="38.0">38 m</quantity>
a.s.l.
</elevation>
(Locality
<emphasis id="0AACEADBA876FFF1FD9FFACB9CC0FA89" bold="true" box="[607,674,1308,1332]" pageId="97" pageNumber="96">15CO</emphasis>
,
<figureCitation id="A0E32A4CA876FFF1FD74FACA9D76FA89" box="[692,788,1309,1332]" captionStart="Figures 592594" captionStartId="126.[192,278,1617,1641]" captionTargetBox="[213,1417,167,1588]" captionTargetId="figure-251@126.[214,1416,168,1587]" captionTargetPageId="126" captionText="Figures 592594: Figure 592. Locality 15CO, Sri Lanka, North Central Province, Puttalam District, Eluwankulam, 08°17'15&quot;N 079°50'38.7&quot;E, 38 m a.s.l. During night collecting from 10.30 p.m. on 28.IV.2015 to 4.00 a.m. on 29.IV.2015 (UV detection) we recorded Charmus laneus (running on the ground among leaves), Isometrus thwaitesii (running on branches of bushes, also sitting on leaves), Lychas srilankensis (on the ground under branches, leaves or in their vicinity), Reddyanus ceylonensis sp. n. (on the ground), Chaerilus ceylonensis (inside oblique usually not curved burrows 15 to 30 cm long) and Heterometrus swammerdami (inside long burrows). On the locality we recorded between 10.00 p.m. and 4.00 a.m. temperature 25.9 ºC and humidity between 58% and 70%. On the locality we recorded 6 scorpion species and genera of all three Sri Lankan families. Figures 593594. Locality 15CP, Sri Lanka, Southern Province, Matara District, Kekanadura village, 05°58'28.2&quot;N 080°36'20.5&quot;E, 40 m a.s.l. During night collecting on 30.IV.2015 (UV detection) we recorded Isometrus maculatus (inside a house, the species occupy the roof of the house, Fig. 593) and Heterometrus serratus (inside almost straight burrows in a ground wall). On the locality we recorded night temperature 28.4 ºC 25.2 ºC (minimum temperature) and night humidity varied between 63% and 89%." figureDoi="http://doi.org/10.5281/zenodo.7124791" httpUri="https://zenodo.org/record/7124791/files/figure.png" pageId="97" pageNumber="96">Fig. 592</figureCitation>
),
<date id="4C661009A876FFF1FF00FAEB9F5EFAEE" box="[192,316,1340,1363]" pageId="97" pageNumber="96" value="2015-04-28">
<collectingDate id="5C22E9E1A876FFF1FF00FAEB9F5EFAEE" box="[192,316,1340,1363]" pageId="97" pageNumber="96" value="2015-04-28">28.IV.2015</collectingDate>
</date>
,
<specimenCount id="2EDEFD40A876FFF1FE8CFAEB9F0FFAEE" box="[332,365,1340,1363]" pageId="97" pageNumber="96" type="male">2♂</specimenCount>
,
<collectionCode id="5EC9AE0CA876FFF1FEBFFAEB9FA7FAEE" box="[383,453,1340,1363]" pageId="97" pageNumber="96">FKCP</collectionCode>
, 3juvs.,
<collectionCode id="5EC9AE0CA876FFF1FDE6FAEB9C05FAEE" box="[550,615,1340,1363]" pageId="97" pageNumber="96">UPSL</collectionCode>
, leg.
<collectorName id="952D531FA876FFF1FD6CFAEB9D61FAEE" box="[684,771,1340,1363]" pageId="97" pageNumber="96">Kovařík</collectorName>
et al
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="70C26542A876FFF1FF00FA429F82F8A1" pageId="97" pageNumber="96" type="diagnosis">
<paragraph id="386736C9A876FFF1FF00FA429F82F8A1" blockId="97.[192,803,1429,1820]" pageId="97" pageNumber="96">
DIAGNOSIS. Total length
<quantity id="FF209B2CA876FFF1FE0EFA429C27FA11" box="[462,581,1429,1452]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" metricValueMax="4.5" metricValueMin="2.7" pageId="97" pageNumber="96" unit="mm" value="36.0" valueMax="45.0" valueMin="27.0">2745 mm</quantity>
. Two pairs of lateral eyes and one pair of median eyes. Movable fingers of pedipalps with 1112 rows of granules. Fingers straight and short. Trichobothrium
<emphasis id="0AACEADBA876FFF1FE24FA269F93F9B5" box="[484,497,1521,1544]" italics="true" pageId="97" pageNumber="96">d</emphasis>
<subScript id="A45C348CA876FFF1FE32FA2C9F99F9B7" attach="left" box="[498,507,1531,1546]" fontSize="6" pageId="97" pageNumber="96">2</subScript>
of pedipalp patella located on dorsal surface, trichobothrium
<emphasis id="0AACEADBA876FFF1FDF3F9D89C2AF99B" box="[563,584,1551,1574]" italics="true" pageId="97" pageNumber="96">id</emphasis>
on internal surface. Pedipalp chela with 9 carinae. Pectinal teeth number 4 6. Entire carapace evenly covered by granules. Ratio of median eye distance from anterior and posterior margins of carapace = 1: 1.551.60. Mesosoma without carinae. Sternites smooth. First and second metasomal segments with 10 carinae, third and fourth segments with 8 carinae, fifth segment with 7 carinae. All carinae consist of large, denticulate granules.
</paragraph>
</subSubSection>
<subSubSection id="70C26542A876FFF1FF00F8959FB4F8E4" box="[192,470,1858,1881]" pageId="97" pageNumber="96" type="distribution">
<paragraph id="386736C9A876FFF1FF00F8959FB4F8E4" blockId="97.[192,470,1858,1881]" box="[192,470,1858,1881]" pageId="97" pageNumber="96">
DISTRIBUTION.
<collectingCountry id="40CF7659A876FFF1FEA7F8959FB0F8E4" box="[359,466,1858,1881]" name="Sri Lanka" pageId="97" pageNumber="96">Sri Lanka</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>