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<document id="7D4BD5C54D0BFC44FC34769E69F4D827" ID-CLB-Dataset="153629" ID-DOI="10.18590/euscorpius.2016.vol2016.iss220.1" ID-GBIF-Dataset="03196918-10d5-44d2-ab42-d38f88314c7b" ID-ISSN="1536-9307" ID-Zenodo-Dep="7124466" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638249082081" checkinUser="felipe" docAuthor="Kovařík, František, Lowe, Graeme, Ranawana, Kithsiri B., Hoferek, David &amp; Š, V. A." docDate="2016" docId="B07187DFA802FF8DFE8CF9B69FB4FDB7" docLanguage="en" docName="Euscorpius.220.1-133.pdf" docOrigin="Euscorpius 220" docSource="https://doi.org/10.18590/euscorpius.2016.vol2016.iss220.1" docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.3:Euscorpius.2002-2019.journal_article" docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A" docStyleName="Euscorpius.2002-2019.journal_article" docStyleVersion="3" docTitle="Charmus laneus Karsch 1879" docType="treatment" docVersion="8" lastPageNumber="28" masterDocId="4C48FFA7A817FF90FFC0FFD79E62FFBD" masterDocTitle="Scorpions of Sri Lanka (Scorpiones Buthidae, Chaerilidae, Scorpionidae) with description of four new species of the genera Charmus Karsch, 1879 and Reddyanus Vachon, 1972, stat n" masterLastPageNumber="133" masterPageNumber="1" pageNumber="20" updateTime="1699471154095" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="2DB6737D36C4FCC0CDDB1AD66E1DDA2C">Scorpions of Sri Lanka (Scorpiones Buthidae, Chaerilidae, Scorpionidae) with description of four new species of the genera Charmus Karsch, 1879 and Reddyanus Vachon, 1972, stat n</mods:title>
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<mods:namePart id="940F0DC50E1017A5A7CB251A62A221BE">Kovařík, František</mods:namePart>
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<mods:namePart id="3DAF9A5D5DF938360BFAA0517A091D4E">Lowe, Graeme</mods:namePart>
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<mods:namePart id="7047AD03DF41D5A05D8DD2BE56500960">Ranawana, Kithsiri B.</mods:namePart>
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<mods:namePart id="ED0905E7A5126292B419E050726F349C">Hoferek, David</mods:namePart>
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<treatment id="B07187DFA802FF8DFE8CF9B69FB4FDB7" ID-DOI="http://doi.org/10.5281/zenodo.7124485" ID-GBIF-Taxon="201134896" ID-Zenodo-Dep="7124485" LSID="urn:lsid:plazi:treatment:B07187DFA802FF8DFE8CF9B69FB4FDB7" httpUri="http://treatment.plazi.org/id/B07187DFA802FF8DFE8CF9B69FB4FDB7" lastPageId="29" lastPageNumber="28" pageId="21" pageNumber="20">
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<heading id="632F81A5A802FF85FE8CF9B69CF4F9C4" box="[332,662,1633,1657]" centered="true" fontSize="11" level="2" pageId="21" pageNumber="20" reason="2">
<taxonomicName id="FFD84D4AA802FF85FE8CF9B69CF4F9C4" ID-CoL="69PQ4" authority="Karsch, 1879" authorityName="Karsch" authorityYear="1879" box="[332,662,1633,1657]" class="Arachnida" family="Buthidae" genus="Charmus" kingdom="Animalia" order="Scorpiones" pageId="21" pageNumber="20" phylum="Arthropoda" rank="species" species="laneus">
<emphasis id="0AACEADBA802FF85FE8CF9B69C60F9C4" bold="true" box="[332,514,1633,1657]" italics="true" pageId="21" pageNumber="20">Charmus laneus</emphasis>
<bibRefCitation id="5C494B38A802FF85FDC8F9B59CF4F9C4" author="Karsch" box="[520,662,1634,1657]" firstAuthor="Karsch" pageId="21" pageNumber="20" pagination="97 - 136" refId="ref41417" refString="KARSCH, F. 1879. Skorpionologische Beitrage I. and II. Mitteilungen des Munchener Entomologischen Vereins, 3: 6 - 22, 97 - 136." type="journal article" year="1879">Karsch, 1879</bibRefCitation>
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(
<figureCitation id="A0E32A4CA802FF85FF17F9569F4FF925" box="[215,301,1665,1688]" captionStart="Figures 12–15" captionStartId="6.[192,278,1885,1909]" captionTargetBox="[191,1440,167,1856]" captionTargetId="figure-11@6.[192,1439,168,1855]" captionTargetPageId="6" captionText="Figures 12–15: Maps showing the known geographical distribution of the Sri Lankan species." figureDoi="http://doi.org/10.5281/zenodo.7124476" httpUri="https://zenodo.org/record/7124476/files/figure.png" pageId="21" pageNumber="20">Figs. 12</figureCitation>
,
<figureCitation id="A0E32A4CA802FF85FEFAF9569F1FF925" box="[314,381,1665,1688]" captionStart="Figures 39–46" captionStartId="13.[192,278,946,970]" captionTargetBox="[196,1427,173,927]" captionTargetId="figure-6@13.[192,1439,168,933]" captionTargetPageId="13" captionText="Figures 39–46: Chela of pedipalps (39–45) and movable finger of chela (46). Figures 39–40. Buthoscorpio sarasinorum, male from locality 15CF (39) and female from locality 15CG (40). Figures 41–43. Charmus laneus, male (41) and female (42) from locality 15CO, and female holotype (43). Figure 44. Charmus saradieli, sp. n., female holotype. Figures 45–46. Hottentotta tamulus, male from locality 15CK." figureDoi="http://doi.org/10.5281/zenodo.7124493" httpUri="https://zenodo.org/record/7124493/files/figure.png" pageId="21" pageNumber="20">41–43</figureCitation>
,
<figureCitation id="A0E32A4CA802FF85FE4AF9569FAEF925" box="[394,460,1665,1688]" captionStart-0="Figures 47–53" captionStart-1="Figures 54–59" captionStart-2="Figures 60–63" captionStart-3="Figures 64–65" captionStart-4="Figures 66–67" captionStart-5="Figures 68–70" captionStart-6="Figures 71–73" captionStart-7="Figures 74–79" captionStart-8="Figures 80–84" captionStartId-0="14.[192,278,1770,1794]" captionStartId-1="15.[192,278,1589,1613]" captionStartId-2="16.[192,278,1844,1868]" captionStartId-3="17.[192,278,1310,1334]" captionStartId-4="18.[192,278,909,933]" captionStartId-5="19.[192,278,1708,1732]" captionStartId-6="20.[192,278,1872,1896]" captionStartId-7="21.[192,278,1257,1281]" captionStartId-8="22.[192,278,1533,1557]" captionTargetBox-0="[207,1426,157,1733]" captionTargetBox-1="[203,1426,153,1559]" captionTargetBox-2="[213,1417,167,1815]" captionTargetBox-3="[191,1440,215,1282]" captionTargetBox-4="[204,1433,173,864]" captionTargetBox-5="[192,1439,168,1678]" captionTargetBox-6="[191,1440,167,1843]" captionTargetBox-7="[199,1428,176,1222]" captionTargetBox-8="[206,1423,177,1494]" captionTargetId-0="figure-11@14.[192,1439,144,1741]" captionTargetId-1="figure-40@15.[192,1439,144,1560]" captionTargetId-2="figure-70@16.[214,1416,168,1814]" captionTargetId-3="figure-631@17.[192,1439,216,1281]" captionTargetId-4="figure-11@18.[192,1439,168,880]" captionTargetId-5="figure-7@19.[192,1439,168,1678]" captionTargetId-6="figure-69@20.[192,1439,168,1842]" captionTargetId-7="figure-654@21.[192,1439,168,1228]" captionTargetId-8="figure-11@22.[192,1439,168,1504]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionTargetPageId-2="16" captionTargetPageId-3="17" captionTargetPageId-4="18" captionTargetPageId-5="19" captionTargetPageId-6="20" captionTargetPageId-7="21" captionTargetPageId-8="22" captionText-0="Figures 47–53: Figure 47. Left hemispermatophore of Buthoscorpio sarasinorum from locality 15CF. Figures 48–53. Charmus laneus from locality 15CO. Figure 48. Left hemispermatophore. Figures 49–53. Capsule region and flagellum. External (49), dorsal (50), internal (51) and ventral (52) views of left hemispermatophore, and externo-dorsal (53) view of right hemispermatophore presented as mirror image for comparison. Scale bars: 1 mm for 47, 500 µm for 48, 250 µm for 49–52, and 200 µm for 53. Abbreviations: bl, basal lobe; f, flagellum; ml, median lobe; mlc, median lobe carina; pr, pars recta of flagellum; prf, pars reflecta of flagellum; t, trunk." captionText-1="Figures 54–59: Charmus laneus from locality 15CO. Figures 54–55. Male 14.7 mm long in dorsal (54) and ventral (55) views. Figures 56–57. Juvenile male 7.5 mm long in dorsal (56) and ventral (57) views. Figures 58–59. Female 20.1 mm long in dorsal (58) and ventral (59) views." captionText-2="Figures 60–63: Charmus laneus from locality 15CO. Figures 60, 62. Male, chelicerae, carapace and tergites I–III (60) and sternopectinal region and sternites III–VII (62). Figures 61, 63. Female, chelicerae, carapace and tergites I–III (61) and sternopectinal region and sternite III (63)." captionText-3="Figures 64–65: Charmus laneus from locality 15CO, showing granulation and color pattern of male. Chelicerae, carapace and tergites (64). Coxosternal region and sternites (65). Scale bar: 500 µm." captionText-4="Figures 66–67: Left chelicera of male Charmus laneus from locality 15CO. Figures 66–67. Dorsal (66) and ventral (67a) view (white light + UV), and ventroexternal (67b) aspect (UV) showing 2 denticles on ventral surface of fixed finger (d1, d2). Scale bar: 200 µm." captionText-5="Figures 68–70: Charmus laneus, male from locality 15CO. Metasoma V and telson under UV fluorescence, showing granulation, punctuation and setation in dorsal (68), lateral (69) and ventral (70) views. Scale bar: 500 µm." captionText-6="Figures 71–73: Metasoma and telson of Charmus laneus male from locality 15CO, showing granulation, punctuation, setation and color pattern in dorsal (71), lateral (72) and ventral (73) views. Scale bar: 1 mm." captionText-7="Figures 74–79: Charmus laneus from locality 15CO. Figures 74–76. Male, metasoma and telson, lateral (74), ventral (75), and dorsal (76) views. Figures 77–79. Female, metasoma and telson, lateral (77), ventral (78), and dorsal (79) views." captionText-8="Figures 80–84: Comparing Sri Lankan Charmus female metasomas. Figures 80–82. Female holotype of Charmus laneus, metasoma and telson, lateral (80), ventral (81), and dorsal (82) views. Figures 83. Female of Charmus laneus from locality 15CO, metasoma and telson dorsal. Figures 84. Female holotype of Charmus saradieli sp. n. (MHNG) metasoma and telson dorsal." figureDoi-0="http://doi.org/10.5281/zenodo.7124495" figureDoi-1="http://doi.org/10.5281/zenodo.7124497" figureDoi-2="http://doi.org/10.5281/zenodo.7124499" figureDoi-3="http://doi.org/10.5281/zenodo.7124503" figureDoi-4="http://doi.org/10.5281/zenodo.7124507" figureDoi-5="http://doi.org/10.5281/zenodo.7124511" figureDoi-6="http://doi.org/10.5281/zenodo.7124513" figureDoi-7="http://doi.org/10.5281/zenodo.7124521" figureDoi-8="http://doi.org/10.5281/zenodo.7124523" httpUri-0="https://zenodo.org/record/7124495/files/figure.png" httpUri-1="https://zenodo.org/record/7124497/files/figure.png" httpUri-2="https://zenodo.org/record/7124499/files/figure.png" httpUri-3="https://zenodo.org/record/7124503/files/figure.png" httpUri-4="https://zenodo.org/record/7124507/files/figure.png" httpUri-5="https://zenodo.org/record/7124511/files/figure.png" httpUri-6="https://zenodo.org/record/7124513/files/figure.png" httpUri-7="https://zenodo.org/record/7124521/files/figure.png" httpUri-8="https://zenodo.org/record/7124523/files/figure.png" pageId="21" pageNumber="20">47–83</figureCitation>
,
<figureCitation id="A0E32A4CA802FF85FE1AF9569C7FF925" box="[474,541,1665,1688]" captionStart="Figures 85–87" captionStartId="23.[192,278,1859,1883]" captionTargetBox="[191,1440,167,1830]" captionTargetId="figure-69@23.[192,1439,168,1829]" captionTargetPageId="23" captionText="Figures 85–87: Charmus. Figures 85–86. Male (85) and female (86) of Charmus laneus from locality 15CO. Figure 87. Male paratype of Charmus saradieli sp. n. (UPSL)." figureDoi="http://doi.org/10.5281/zenodo.7124529" httpUri="https://zenodo.org/record/7124529/files/figure.png" pageId="21" pageNumber="20">85–86</figureCitation>
,
<figureCitation id="A0E32A4CA802FF85FDEAF9569C0FF925" box="[554,621,1665,1688]" captionStart="Figures 92–100" captionStartId="25.[192,278,1693,1717]" captionTargetBox="[191,1440,167,1664]" captionTargetId="figure-217@25.[192,1439,168,1663]" captionTargetPageId="25" captionText="Figures 92–100: Charmus. Figures 92–95, 99–100. Charmus saradieli sp. n., male paratype (MHNG), chelicerae, carapace and tergites I–IV (92), sternopectinal region and sternites III–IV (94), and telson lateral (99); female holotype, chelicerae, carapace and tergites I–III (93), sternopectinal region and sternite III (95), and telson lateral (100). Figures 96–98. Charmus laneus, female holotype, chelicera and carapace (96), original label (97), and sternopectinal region and sternites III–IV (98)." figureDoi="http://doi.org/10.5281/zenodo.7124537" httpUri="https://zenodo.org/record/7124537/files/figure.png" pageId="21" pageNumber="20">96–98</figureCitation>
,
<figureCitation id="A0E32A4CA802FF85FDBAF9569CBAF925" box="[634,728,1665,1688]" captionStart="Figures 107–119" captionStartId="27.[192,278,1733,1757]" captionTargetBox="[211,1428,190,1691]" captionTargetId="figure-6@27.[192,1439,181,1704]" captionTargetPageId="27" captionText="Figures 107–119: Charmus. Figures 107–114. Charmus saradieli sp. n., female holotype. Pedipalp movable finger (107). Pedipalp chela, dorsal (108), external (109), and ventral (110) views. Pedipalp patella, dorsal (111) and external (112) views. Pedipalp femur and trochanter, internal (113) and dorsal (114) views. The trichobothrial pattern is indicated in Figures 108a– 114a. Figures 115–117. Charmus saradieli sp. n., male paratype (MHNG), chelicerae (115), pedipalp dorsal (116), and fourth leg dorsal (117). Figures 118–119. Charmus laneus from locality 15CO. Pedipalp dorsal of male (118) and female (119)." figureDoi="http://doi.org/10.5281/zenodo.7124543" httpUri="https://zenodo.org/record/7124543/files/figure.png" pageId="21" pageNumber="20">118–119</figureCitation>
,
<figureCitation id="A0E32A4CA802FF85FD25F9569D6FF925" box="[741,781,1665,1688]" captionStart="Figures 193–200" captionStartId="48.[192,278,1070,1094]" captionTargetBox="[213,1446,171,1037]" captionTargetId="figure-11@48.[205,1452,168,1041]" captionTargetPageId="48" captionText="Figures 193–200: Distal segments of leg IV, retroventral view of Sri Lankan Buthidae and Chaerilidae genera. Buthoscorpio sarasinorum, male from locality 15CF (193), Charmus laneus, male from locality 15CO (194), Hottentotta tamulus, male from locality 15CK (195), Isometrus maculatus, female from locality 15CI (196), Isometrus thwaitesi, female from locality 15CO (197), Lychas srilankensis, male from locality 15CJ (198), Reddyanus basilicus, male from locality 15CS (199), and Chaerilus ceylonensis, male from locality 15CD (200)." figureDoi="http://doi.org/10.5281/zenodo.7124608" httpUri="https://zenodo.org/record/7124608/files/figure.png" pageId="21" pageNumber="20">194</figureCitation>
,
<figureCitation id="A0E32A4CA802FF85FEA1F9779FDDF90A" box="[353,447,1696,1719]" captionStart="Figures 403–429" captionStartId="85.[192,278,744,768]" captionTargetId="figure-11@84.[172,1464,185,1878]" captionTargetPageId="84" captionText="Figures 403–429: Telson, lateral view of Sri Lankan Buthidae and Chaerilidae species. Figures 403–404. Isometrus maculatus, male from locality 15CP (403) and female from locality 15CI (404). Figures 405–406. I. thwaitesi, male from locality 15CH (405) and female from locality 15CO (406). Figures 407–408. Lychas srilankensis, male (407) and female (408) from locality 15CJ. Figures 409–410. Reddyanus basilicus, male (409) and female (410) from locality 15CR. Figures 411–412. R. besucheti, male holotype (411) and female from locality 15CG (412). Figures 413–414. R. ceylonensis sp. n., male holotype (413) and female paratype (414) from locality 15CI. Figures 415–416. R. jayarathnei sp. n., male (415) and female (416) paratypes. Figures 417–418. R. loebli, male (417) and female (418) from locality 15CG. Figures 419–420. R. ranawanai sp. n., male holotype (419) and female paratype (420). Figures 421–422: Buthoscorpio sarasinorum, male from locality 15CF (421) and female from locality 15CG (422). Figures 423–424. Charmus laneus, male (423) and female (424) from locality 15CO. Figures 425–426. Charmus saradieli sp. n. male paratype (425) (MHNG) and female holotype (426). Figurs 427. Hottentotta tamulus, male from locality 15CK (427). Figures 428–429. Chaerilus ceylonensis, male (428) and female (429) from locality 15CD." figureDoi="http://doi.org/10.5281/zenodo.7124696" httpUri="https://zenodo.org/record/7124696/files/figure.png" pageId="21" pageNumber="20">423–424</figureCitation>
, 548,
<tableCitation id="755A0372A802FF85FDC1F9779C1AF90A" box="[513,632,1696,1719]" captionStart-0="Table 1" captionStart-1="Table 2" captionStartId-0="28.[192,257,853,877]" captionStartId-1="28.[192,257,1382,1406]" captionTargetPageId-0="28" captionTargetPageId-1="28" captionText-0="Table 1: Comparative measurements of adults of Charmus laneus and C. saradieli sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." captionText-1="Table 2: Comparison among Sri Lankan Charmus species (specimens), based upon selected morphometric ratios of adult females. Abbreviations: length (L), width (W), depth (D)." httpUri-0="http://table.plazi.org/id/6CA76641A80BFF8CFF00FC829C80FC3B" httpUri-1="http://table.plazi.org/id/6CA76641A80BFF8CFF00FAB19C88FA2B" pageId="21" pageNumber="20" tableUuid-0="6CA76641A80BFF8CFF00FC829C80FC3B" tableUuid-1="6CA76641A80BFF8CFF00FAB19C88FA2B">Tables 1–2</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="70C26542A802FF85FF00F90A9B1FFA42" pageId="21" pageNumber="20" type="reference_group">
<paragraph id="386736C9A802FF85FF00F90A9FA2F8D3" blockId="21.[192,802,1757,1902]" pageId="21" pageNumber="20">
<treatmentCitationGroup id="18C811E7A802FF85FF00F90A9FA2F8D3" pageId="21" pageNumber="20">
<treatmentCitation id="B97910D8A802FF85FF00F90A9CFEF949" author="KARSCH" box="[192,668,1757,1780]" page="105" pageId="21" pageNumber="20" year="1879">
<taxonomicName id="FFD84D4AA802FF85FF00F90A9CFEF949" ID-CoL="69PQ4" authority="Karsch, 1879: 104 - 105" authorityName="Karsch" authorityPageNumber="104 - 105" authorityYear="1879" box="[192,668,1757,1780]" class="Arachnida" family="Buthidae" genus="Charmus" kingdom="Animalia" order="Scorpiones" pageId="21" pageNumber="20" phylum="Arthropoda" rank="species" species="laneus">
<emphasis id="0AACEADBA802FF85FF00F90A9F1BF949" box="[192,377,1757,1780]" italics="true" pageId="21" pageNumber="20">Charmus laneus</emphasis>
<bibRefCitation id="5C494B38A802FF85FE4AF90A9CFEF949" author="Karsch" box="[394,668,1757,1780]" firstAuthor="Karsch" page="104 - 105" pageId="21" pageNumber="20" pagination="97 - 136" refId="ref41417" refString="KARSCH, F. 1879. Skorpionologische Beitrage I. and II. Mitteilungen des Munchener Entomologischen Vereins, 3: 6 - 22, 97 - 136." type="journal article" year="1879">Karsch, 1879: 104–105</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="B97910D8A802FF85FD72F90A9F33F8AE" author="KRAEPELIN" page="39" pageId="21" pageNumber="20" year="1899">
<bibRefCitation id="5C494B38A802FF85FD72F90A9F33F8AE" author="Kraepelin" firstAuthor="Kraepelin" page="39" pageId="21" pageNumber="20" refId="ref42361" refString="KRAEPELIN, K. 1899. Scorpiones und Pedipalpi. In: F. DAHL (ed.), Das Tierreich. Herausgegeben von der Deutschen Zoologischen Gesellschaft. Berlin: R. Friedlander und Sohn Verlag, 8. Lieferung. 265 pp." type="book" year="1899">Kraepelin, 1899: 39</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="B97910D8A802FF85FEA2F92B9C40F8AE" author="POCOCK, R. I." box="[354,546,1788,1811]" page="32" pageId="21" pageNumber="20" year="1900">
<bibRefCitation id="5C494B38A802FF85FEA2F92B9C40F8AE" author="Pocock" bookContentInfo="279 pp" box="[354,546,1788,1811]" firstAuthor="Pocock" journalOrPublisher="London: W. T. Blandford" page="32" pageId="21" pageNumber="20" part="xii" refId="ref43790" refString="POCOCK, R. I. 1900. Arachnida. The fauna of British India, including Ceylon and Burma. Published un- der the authority of the Secretary of State for India in Council. London: W. T. Blandford, xii, 279 pp" title="Arachnida. The fauna of British India, including Ceylon and Burma. Published un- der the authority of the Secretary of State for India in Council" type="book" year="1900">Pocock, 1900: 32</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="B97910D8A802FF85FDF3F92B9D7FF8AE" author="KRAEPELIN" box="[563,797,1788,1811]" page="131" pageId="21" pageNumber="20" year="1913">
<bibRefCitation id="5C494B38A802FF85FDF3F92B9D7FF8AE" author="Kraepelin" box="[563,797,1788,1811]" firstAuthor="Kraepelin" page="131" pageId="21" pageNumber="20" pagination="123 - 196" refId="ref42407" refString="KRAEPELIN, K. 1913. Neue Beitrage zur Systematik der Gliederspinnen. III. A. Bemerkungen zur Skorpionenfauna Indiens. B. Die Skorpione, Pedipalpen und Solifugen Deutsch - Ostafrikas. Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten, 30: 123 - 196." type="journal article" year="1913">Kraepelin, 1913: 131</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="B97910D8A802FF85FF30F8CD9C12F88C" author="SREENIVASA-REDDY, R. P." box="[240,624,1818,1841]" page="253" pageId="21" pageNumber="20" year="1966">
<bibRefCitation id="5C494B38A802FF85FF30F8CD9C12F88C" author="Sreenivasa-Reddy" box="[240,624,1818,1841]" firstAuthor="Sreenivasa-Reddy" journalOrPublisher="Bulletin du Museum d'Histoire Naturelle, (Paris)" page="253 - 254" pageId="21" pageNumber="20" pagination="247 - 256" part="38" refId="ref44396" refString="SREENIVASA-REDDY, R. P. 1966. Contribution a la connaissance des scorpions de l'Inde. I. Charmus indicus Hirst 1915 (fam. Buthidae). Bulletin du Museum d'Histoire Naturelle, (Paris), 38 (3): 247 - 256." title="Contribution a la connaissance des scorpions de l'Inde. I. Charmus indicus Hirst 1915 (fam. Buthidae)" type="journal article" year="1966">Sreenivasa-Reddy, 1966: 253–254</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="B97910D8A802FF85FD41F8CD9FFEF8ED" author="MORITZ, M. &amp; S. - CH. FISCHER" page="317" pageId="21" pageNumber="20" year="1980">
<bibRefCitation id="5C494B38A802FF85FD41F8CD9FFEF8ED" author="Moritz &amp; Fischer" firstAuthor="Moritz" journalOrPublisher="Mitteilungen aus dem Zoologischen Museum in Berlin" page="317" pageId="21" pageNumber="20" pagination="309 - 326" part="56" refId="ref43512" refString="MORITZ, M. &amp; S. - CH. FISCHER 1980. Die typen der Arachniden-Sammlung des Zoologischen Museums Berlin. III. Scorpiones. Mitteilungen aus dem Zoologischen Museum in Berlin, 56: 309 - 326." title="Die typen der Arachniden-Sammlung des Zoologischen Museums Berlin. III. Scorpiones" type="journal article" year="1980">Moritz &amp; Fischer, 1980: 317</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="B97910D8A802FF85FE6DF8EE9CE4F8ED" author="KOVARIK" box="[429,646,1849,1872]" page="108" pageId="21" pageNumber="20" year="1998">
<bibRefCitation id="5C494B38A802FF85FE6DF8EE9CE4F8ED" author="Kovarik" box="[429,646,1849,1872]" firstAuthor="Kovarik" page="108" pageId="21" pageNumber="20" refId="ref41687" refString="KOVARIK, F. 1998. Stiri [Scorpiones]. Publishing House &quot; Madagaskar &quot;, Jihlava (Czech Republic). 175 pp. (in Czech)." type="book" year="1998">Kovařík, 1998: 108</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="B97910D8A802FF85FD54F8EE9F39F8D3" author="FET, V. &amp; W. D. SISSOM &amp; G. LOWE &amp; M. E. BRAUNWALDER" page="123" pageId="21" pageNumber="20" year="2000">
<bibRefCitation id="5C494B38A802FF85FD54F8EE9F39F8D3" author="Fet &amp; Lowe" bookContentInfo="689 pp." firstAuthor="Fet" journalOrPublisher="New York: The New York Entomological Society" page="123" pageId="21" pageNumber="20" refId="ref40873" refString="FET, V., W. D. SISSOM, G. LOWE &amp; M. E. BRAUNWALDER. 2000. Catalog of the Scorpions of the World (1758 - 1998). New York: The New York Entomological Society, 689 pp." title="Catalog of the Scorpions of the World (1758 - 1998)" type="book" year="2000">Fet &amp; Lowe, 2000: 123</bibRefCitation>
</treatmentCitation>
(in part).
</treatmentCitationGroup>
</paragraph>
<paragraph id="386736C9A802FF85FCFEFA999AC0FA1E" blockId="21.[830,1440,1358,1535]" pageId="21" pageNumber="20">
<treatmentCitationGroup id="18C811E7A802FF85FCFEFA999AC0FA1E" pageId="21" pageNumber="20">
=
<treatmentCitation id="B97910D8A802FF85FC9CFA999B12FAD8" author="POCOCK, R. I." box="[860,1392,1358,1381]" page="48" pageId="21" pageNumber="20" year="1892">
<taxonomicName id="FFD84D4AA802FF85FC9CFA999B12FAD8" authority="Pocock, 1892: 47 - 48" authorityName="Pocock" authorityPageNumber="47 - 48" authorityYear="1892" box="[860,1392,1358,1381]" class="Arachnida" family="Buthidae" genus="Heterocharmus" kingdom="Animalia" order="Scorpiones" pageId="21" pageNumber="20" phylum="Arthropoda" rank="species" species="cinctipes">
<emphasis id="0AACEADBA802FF85FC9CFA999A0DFAD8" box="[860,1135,1358,1381]" italics="true" pageId="21" pageNumber="20">Heterocharmus cinctipes</emphasis>
<bibRefCitation id="5C494B38A802FF85FBBEFA999B12FAD8" author="Pocock" box="[1150,1392,1358,1381]" firstAuthor="Pocock" journalOrPublisher="Annals and Magazine of Natural History" page="47 - 48" pageId="21" pageNumber="20" pagination="38 - 49" part="6" refId="ref43682" refString="POCOCK, R. I. 1892. Descriptions of two new genera of scorpions, with notes upon some species of Palamnaeus. Annals and Magazine of Natural History, 6 (9): 38 - 49." title="Descriptions of two new genera of scorpions, with notes upon some species of Palamnaeus" type="journal article" year="1892">Pocock, 1892: 47–48</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, pl. IIIB, fig. 2, 2a–b (TL:
<collectingCountry id="40CF7659A802FF85FBAAFABA9AFDFA39" box="[1130,1183,1389,1412]" name="India" pageId="21" pageNumber="20">India</collectingCountry>
or
<collectingCountry id="40CF7659A802FF85FB08FABA9B56FA39" box="[1224,1332,1389,1412]" name="Sri Lanka" pageId="21" pageNumber="20">Sri Lanka</collectingCountry>
; BMNH). Syn. by
<treatmentCitation id="B97910D8A802FF85FC07FA5B9AF9FA1E" author="KRAEPELIN" box="[967,1179,1420,1443]" page="39" pageId="21" pageNumber="20" year="1899">
<bibRefCitation id="5C494B38A802FF85FC07FA5B9AF9FA1E" author="Kraepelin" box="[967,1179,1420,1443]" firstAuthor="Kraepelin" page="39" pageId="21" pageNumber="20" refId="ref42361" refString="KRAEPELIN, K. 1899. Scorpiones und Pedipalpi. In: F. DAHL (ed.), Das Tierreich. Herausgegeben von der Deutschen Zoologischen Gesellschaft. Berlin: R. Friedlander und Sohn Verlag, 8. Lieferung. 265 pp." type="book" year="1899">Kraepelin, 1899: 39</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="386736C9A802FF85FCFEFA7D9B1FFA42" blockId="21.[830,1440,1358,1535]" pageId="21" pageNumber="20">
<treatmentCitationGroup id="18C811E7A802FF85FCFEFA7D9B1FFA42" pageId="21" pageNumber="20">
=
<treatmentCitation id="B97910D8A802FF85FC9AFA7D9B7CFA7C" author="LOURENCO, W. R." box="[858,1310,1450,1473]" page="24" pageId="21" pageNumber="20" year="2002">
<taxonomicName id="FFD84D4AA802FF85FC9AFA7D9B7CFA7C" authority="Lourenco, 2002: 19 - 24" authorityName="Lourenco" authorityPageNumber="19 - 24" authorityYear="2002" box="[858,1310,1450,1473]" class="Arachnida" family="Buthidae" genus="Charmus" kingdom="Animalia" order="Scorpiones" pageId="21" pageNumber="20" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="0AACEADBA802FF85FC9AFA7D9A6BFA7C" box="[858,1033,1450,1473]" italics="true" pageId="21" pageNumber="20">Charmus minor</emphasis>
<bibRefCitation id="5C494B38A802FF85FBD6FA7D9B7CFA7C" author="Lourenco" box="[1046,1310,1450,1473]" firstAuthor="Lourenco" journalOrPublisher="Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg" page="19 - 24" pageId="21" pageNumber="20" pagination="17 - 25" part="14" refId="ref42746" refString="LOURENCO, W. R. 2002. Further taxonomic considerations about the genus Charmus Karsch, 1879 (Scorpiones, Buthidae), with the description of a new species from Sri Lanka. Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg, 14 (165): 17 - 25" title="Further taxonomic considerations about the genus Charmus Karsch, 1879 (Scorpiones, Buthidae), with the description of a new species from Sri Lanka" type="journal article" year="2002">Lourenço, 2002: 19–24</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs. 1–14 (TL:
<collectingCountry id="40CF7659A802FF85FC68FA1E9A77FA5D" box="[936,1045,1481,1504]" name="Sri Lanka" pageId="21" pageNumber="20">Sri Lanka</collectingCountry>
, Mannar District, Wilpattu National Park, 0.5 miles NE Cockmuttai; ZMUH).
<emphasis id="0AACEADBA802FF85FAEFFA309B38FA42" bold="true" box="[1327,1370,1511,1535]" pageId="21" pageNumber="20">Syn</emphasis>
.
<emphasis id="0AACEADBA802FF85FAA7FA309B14FA42" bold="true" box="[1383,1398,1511,1535]" pageId="21" pageNumber="20">n</emphasis>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="70C26542A802FF86FCFEF9F29CF6F8AB" lastPageId="22" lastPageNumber="21" pageId="21" pageNumber="20" type="materials_examined">
<paragraph id="386736C9A802FF85FCFEF9F29BF9F981" blockId="21.[830,1435,1573,1596]" box="[830,1435,1573,1596]" pageId="21" pageNumber="20">
<typeStatus id="E763886BA802FF85FCFEF9F29D15F986" box="[830,887,1573,1596]" pageId="21" pageNumber="20">TYPE</typeStatus>
LOCALITY AND
<typeStatus id="E763886BA802FF85FBE0F9FF9A37F986" box="[1056,1109,1576,1595]" pageId="21" pageNumber="20">TYPE</typeStatus>
REPOSITORY.
<collectingCountry id="40CF7659A802FF85FB2BF9F29B5EF981" box="[1259,1340,1573,1596]" name="Sri Lanka" pageId="21" pageNumber="20">Ceylon</collectingCountry>
;
<collectionCode id="5EC9AE0CA802FF85FA87F9F29BF6F981" box="[1351,1428,1573,1596]" pageId="21" pageNumber="20">ZMHB</collectionCode>
.
</paragraph>
<paragraph id="386736C9A802FF85FCFEF9B59D9FF90A" blockId="21.[830,1440,1634,1719]" pageId="21" pageNumber="20">
TYPE MATERIAL EXAMINED.
<materialsCitation id="88B03C94A802FF85FB43F9B59D9BF90A" ID-GBIF-Occurrence="3913992313" collectionCode="ZMHB" collectorName="Hoffmeister" country="Sri Lanka" location="Ceylon" pageId="21" pageNumber="20" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<collectingCountry id="40CF7659A802FF85FB43F9B59AB2F9C4" box="[1155,1232,1634,1657]" name="Sri Lanka" pageId="21" pageNumber="20">Ceylon</collectingCountry>
(now
<collectingCountry id="40CF7659A802FF85FAE1F9B59BF0F9C4" box="[1313,1426,1634,1657]" name="Sri Lanka" pageId="21" pageNumber="20">Sri Lanka</collectingCountry>
), leg.
<collectorName id="952D531FA802FF85FCAEF9569D90F925" box="[878,1010,1665,1688]" pageId="21" pageNumber="20">Hoffmeister</collectorName>
,
<specimenCount id="2EDEFD40A802FF85FC3EF9569A7DF925" box="[1022,1055,1665,1688]" pageId="21" pageNumber="20" type="female">1♀</specimenCount>
<typeStatus id="E763886BA802FF85FBE9F9569AE7F925" box="[1065,1157,1665,1688]" pageId="21" pageNumber="20" type="holotype">holotype</typeStatus>
(
<figureCitation id="A0E32A4CA802FF85FB56F9569A8FF925" box="[1174,1261,1665,1688]" captionStart="Figures 39–46" captionStartId="13.[192,278,946,970]" captionTargetBox="[196,1427,173,927]" captionTargetId="figure-6@13.[192,1439,168,933]" captionTargetPageId="13" captionText="Figures 39–46: Chela of pedipalps (39–45) and movable finger of chela (46). Figures 39–40. Buthoscorpio sarasinorum, male from locality 15CF (39) and female from locality 15CG (40). Figures 41–43. Charmus laneus, male (41) and female (42) from locality 15CO, and female holotype (43). Figure 44. Charmus saradieli, sp. n., female holotype. Figures 45–46. Hottentotta tamulus, male from locality 15CK." figureDoi="http://doi.org/10.5281/zenodo.7124493" httpUri="https://zenodo.org/record/7124493/files/figure.png" pageId="21" pageNumber="20">Figs. 43</figureCitation>
,
<figureCitation id="A0E32A4CA802FF85FB3CF9569B5DF925" box="[1276,1343,1665,1688]" captionStart="Figures 80–84" captionStartId="22.[192,278,1533,1557]" captionTargetBox="[206,1423,177,1494]" captionTargetId="figure-11@22.[192,1439,168,1504]" captionTargetPageId="22" captionText="Figures 80–84: Comparing Sri Lankan Charmus female metasomas. Figures 80–82. Female holotype of Charmus laneus, metasoma and telson, lateral (80), ventral (81), and dorsal (82) views. Figures 83. Female of Charmus laneus from locality 15CO, metasoma and telson dorsal. Figures 84. Female holotype of Charmus saradieli sp. n. (MHNG) metasoma and telson dorsal." figureDoi="http://doi.org/10.5281/zenodo.7124523" httpUri="https://zenodo.org/record/7124523/files/figure.png" pageId="21" pageNumber="20">80–82</figureCitation>
,
<figureCitation id="A0E32A4CA802FF85FA8EF9569BF0F925" box="[1358,1426,1665,1688]" captionStart="Figures 92–100" captionStartId="25.[192,278,1693,1717]" captionTargetBox="[191,1440,167,1664]" captionTargetId="figure-217@25.[192,1439,168,1663]" captionTargetPageId="25" captionText="Figures 92–100: Charmus. Figures 92–95, 99–100. Charmus saradieli sp. n., male paratype (MHNG), chelicerae, carapace and tergites I–IV (92), sternopectinal region and sternites III–IV (94), and telson lateral (99); female holotype, chelicerae, carapace and tergites I–III (93), sternopectinal region and sternite III (95), and telson lateral (100). Figures 96–98. Charmus laneus, female holotype, chelicera and carapace (96), original label (97), and sternopectinal region and sternites III–IV (98)." figureDoi="http://doi.org/10.5281/zenodo.7124537" httpUri="https://zenodo.org/record/7124537/files/figure.png" pageId="21" pageNumber="20">96–98</figureCitation>
),
<collectionCode id="5EC9AE0CA802FF85FC80F9779DE9F90A" box="[832,907,1696,1719]" pageId="21" pageNumber="20">ZMHB</collectionCode>
No. 3051
</materialsCitation>
.
</paragraph>
<paragraph id="386736C9A802FF86FCFEF90A9CF6F8AB" blockId="21.[830,1441,1757,1902]" lastBlockId="22.[192,802,1668,1814]" lastPageId="22" lastPageNumber="21" pageId="21" pageNumber="20">
OTHER MATERIAL EXAMINED.
<materialsCitation id="88B03C94A802FF85FB4CF90A9B70F8ED" ID-GBIF-Occurrence="3913992350" collectingDate="2015-04-28" collectionCode="FKCP" collectorName="Kovarik" country="Sri Lanka" elevation="52" latitude="8.20975" location="Eluwankulam" longLatPrecision="15" longitude="79.85889" municipality="Puttalam District" pageId="21" pageNumber="20" specimenCount="1" specimenCount-male="1" stateProvince="North Central">
<collectingCountry id="40CF7659A802FF85FB4CF90A9A9BF949" box="[1164,1273,1757,1780]" name="Sri Lanka" pageId="21" pageNumber="20">Sri Lanka</collectingCountry>
,
<collectingRegion id="FA1CF82BA802FF85FAC9F90A9DFDF8AE" country="Sri Lanka" name="North Central" pageId="21" pageNumber="20">North Central Province</collectingRegion>
,
<collectingMunicipality id="D803ACB3A802FF85FC72F92B9A13F8AE" box="[946,1137,1788,1811]" pageId="21" pageNumber="20">Puttalam District</collectingMunicipality>
,
<location id="3D076012A802FF85FB43F92B9B78F8AE" LSID="urn:lsid:plazi:treatment:B07187DFA802FF8DFE8CF9B69FB4FDB7:3D076012A802FF85FB43F92B9B78F8AE" box="[1155,1306,1788,1811]" country="Sri Lanka" latitude="8.20975" longLatPrecision="15" longitude="79.85889" municipality="Puttalam District" name="Eluwankulam" pageId="21" pageNumber="20" stateProvince="North Central">Eluwankulam</location>
,
<geoCoordinate id="5DEC500EA802FF85FAECF92B9D3EF88C" degrees="08" direction="north" minutes="12" orientation="latitude" pageId="21" pageNumber="20" precision="1" seconds="35.1" value="8.20975">08°12'35.1 &quot;N</geoCoordinate>
<geoCoordinate id="5DEC500EA802FF85FCA6F8CD9D92F88C" box="[870,1008,1818,1841]" degrees="079" direction="east" minutes="51" orientation="longitude" pageId="21" pageNumber="20" precision="15" seconds="32" value="79.85889">079°51'32&quot;E</geoCoordinate>
,
<elevation id="B3F5D1FAA802FF85FC3DF8CD9A12F88C" box="[1021,1136,1818,1841]" metricMagnitude="1" metricUnit="m" metricValue="5.2" pageId="21" pageNumber="20" unit="m" value="52.0">
<quantity id="FF209B2CA802FF85FC3DF8CD9A54F88C" box="[1021,1078,1818,1841]" metricMagnitude="1" metricUnit="m" metricValue="5.2" pageId="21" pageNumber="20" unit="m" value="52.0">52 m</quantity>
a.s.l.
</elevation>
(Locality
<emphasis id="0AACEADBA802FF85FB24F8CE9B47F88C" bold="true" box="[1252,1317,1817,1841]" pageId="21" pageNumber="20">15CN</emphasis>
,
<figureCitation id="A0E32A4CA802FF85FAF5F8CD9BF3F88C" box="[1333,1425,1818,1841]" captionStart="Figures 590–591" captionStartId="125.[192,278,1721,1745]" captionTargetBox="[191,1440,167,1692]" captionTargetId="figure-179@125.[192,1439,168,1691]" captionTargetPageId="125" captionText="Figures 590–591: Figure 590. Locality 15CM, Sri Lanka, Central Province, Matale District, Pallegama, 07°32'49.5&quot;N 080°47'50&quot;E, 434 m a.s.l. During a day 28.IV.2015 we recorded Reddyanus loebli (under bark). Figure 591. Locality 15CN, Sri Lanka, North Central Province, Puttalam District, Eluwankulam, 08°12'35.1&quot;N 079°51'32&quot;E, 52 m a.s.l. During night collecting on 28.IV.2015 (UV detection) between 9.00 and 10.00 p.m. we recorded Charmus laneus (running on the ground among leaves), Lychas srilankensis (on the ground under branches or leaves or in their vicinity), Reddyanus ceylonensis sp. n. (on the ground and inside old moulder stump), Reddyanus loebli (hidden under scales of bark of standing tree trunks) and Heterometrus swammerdami (inside burrows). On the locality we recorded between 9.00 and 10.00 p.m. temperature 28.5 ºC– 25.9 ºC and humidity between 61% and 78%." figureDoi="http://doi.org/10.5281/zenodo.7124787" httpUri="https://zenodo.org/record/7124787/files/figure.png" pageId="21" pageNumber="20">Fig. 591</figureCitation>
),
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<collectingDate id="5C22E9E1A802FF85FCFEF8EE9DD9F8ED" box="[830,955,1849,1872]" pageId="21" pageNumber="20" value="2015-04-28">28.IV.2015</collectingDate>
</date>
,
<specimenCount id="2EDEFD40A802FF85FC08F8EE9D8BF8ED" box="[968,1001,1849,1872]" pageId="21" pageNumber="20" type="male">1♂</specimenCount>
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<collectionCode id="5EC9AE0CA802FF85FC39F8EE9A5DF8ED" box="[1017,1087,1849,1872]" pageId="21" pageNumber="20">FKCP</collectionCode>
, leg.
<collectorName id="952D531FA802FF85FBBDF8EE9AB6F8ED" box="[1149,1236,1849,1872]" pageId="21" pageNumber="20">Kovařík</collectorName>
et al
</materialsCitation>
.;
<materialsCitation id="88B03C94A802FF86FAE5F8EE9CEFF8AB" ID-GBIF-Occurrence="3913992377" collectingDate="2015-04-28" collectionCode="FKCP, UPSL" collectorName="Kovarik" country="Sri Lanka" elevation="38" lastPageId="22" lastPageNumber="21" latitude="8.2875" location="Eluwankulam" longLatPrecision="15" longitude="79.844086" municipality="Puttalam District" pageId="21" pageNumber="20" specimenCount="6" specimenCount-female="2" specimenCount-male="3" stateProvince="North Central">
<collectingRegion id="FA1CF82BA802FF85FAE5F8EE9DB3F8D3" country="Sri Lanka" name="North Central" pageId="21" pageNumber="20">North Central Province</collectingRegion>
,
<collectingMunicipality id="D803ACB3A802FF85FC22F8809AC3F8D3" box="[994,1185,1879,1902]" pageId="21" pageNumber="20">Puttalam District</collectingMunicipality>
,
<location id="3D076012A802FF85FB72F8809B24F8D3" LSID="urn:lsid:plazi:treatment:B07187DFA802FF8DFE8CF9B69FB4FDB7:3D076012A802FF85FB72F8809B24F8D3" box="[1202,1350,1879,1902]" country="Sri Lanka" latitude="8.2875" longLatPrecision="15" longitude="79.844086" municipality="Puttalam District" name="Eluwankulam" pageId="21" pageNumber="20" stateProvince="North Central">Eluwankulam</location>
,
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<geoCoordinate id="5DEC500EA801FF86FEC4F9529FC1F921" box="[260,419,1669,1692]" degrees="079" direction="east" minutes="50" orientation="longitude" pageId="22" pageNumber="21" precision="1" seconds="38.7" value="79.844086">079°50'38.7&quot;E</geoCoordinate>
,
<elevation id="B3F5D1FAA801FF86FE72F9529C48F921" box="[434,554,1669,1692]" metricMagnitude="1" metricUnit="m" metricValue="3.8" pageId="22" pageNumber="21" unit="m" value="38.0">
<quantity id="FF209B2CA801FF86FE72F9529F8CF921" box="[434,494,1669,1692]" metricMagnitude="1" metricUnit="m" metricValue="3.8" pageId="22" pageNumber="21" unit="m" value="38.0">38 m</quantity>
a.s.l.
</elevation>
(Locality
<emphasis id="0AACEADBA801FF86FD63F9539C87F921" bold="true" box="[675,741,1668,1692]" pageId="22" pageNumber="21">15CO</emphasis>
,
<figureCitation id="A0E32A4CA801FF86FD37F9529E8AF907" captionStart-0="Figures 5–11" captionStart-1="Figures 12–15" captionStart-10="Figures 60–63" captionStart-11="Figures 64–65" captionStart-12="Figures 66–67" captionStart-13="Figures 68–70" captionStart-14="Figures 71–73" captionStart-15="Figures 74–79" captionStart-16="Figures 80–84" captionStart-17="Figures 85–87" captionStart-18="Figures 88–91" captionStart-19="Figures 92–100" captionStart-2="Figures 16–19" captionStart-3="Figures 20–23" captionStart-4="Figures 24–29" captionStart-5="Figures 30–33" captionStart-6="Figures 34–38" captionStart-7="Figures 39–46" captionStart-8="Figures 47–53" captionStart-9="Figures 54–59" captionStartId-0="4.[192,278,1745,1769]" captionStartId-1="6.[192,278,1885,1909]" captionStartId-10="16.[192,278,1844,1868]" captionStartId-11="17.[192,278,1310,1334]" captionStartId-12="18.[192,278,909,933]" captionStartId-13="19.[192,278,1708,1732]" captionStartId-14="20.[192,278,1872,1896]" captionStartId-15="21.[192,278,1257,1281]" captionStartId-16="22.[192,278,1533,1557]" captionStartId-17="23.[192,278,1859,1883]" captionStartId-18="24.[192,278,1804,1828]" captionStartId-19="25.[192,278,1693,1717]" captionStartId-2="8.[192,278,1853,1877]" captionStartId-3="9.[192,278,1829,1853]" captionStartId-4="10.[192,278,1378,1402]" captionStartId-5="11.[192,278,1229,1253]" captionStartId-6="12.[192,278,1853,1877]" captionStartId-7="13.[192,278,946,970]" captionStartId-8="14.[192,278,1770,1794]" captionStartId-9="15.[192,278,1589,1613]" captionTargetBox-0="[227,1387,167,1716]" captionTargetBox-1="[191,1440,167,1856]" captionTargetBox-10="[213,1417,167,1815]" captionTargetBox-11="[191,1440,215,1282]" captionTargetBox-12="[204,1433,173,864]" captionTargetBox-13="[192,1439,168,1678]" captionTargetBox-14="[191,1440,167,1843]" captionTargetBox-15="[199,1428,176,1222]" captionTargetBox-16="[206,1423,177,1494]" captionTargetBox-17="[191,1440,167,1830]" captionTargetBox-18="[200,1418,242,1791]" captionTargetBox-19="[191,1440,167,1664]" captionTargetBox-2="[256,1384,176,1817]" captionTargetBox-3="[233,1393,199,1801]" captionTargetBox-4="[199,1433,201,1332]" captionTargetBox-5="[196,1432,202,1188]" captionTargetBox-6="[213,1417,192,1825]" captionTargetBox-7="[196,1427,173,927]" captionTargetBox-8="[207,1426,157,1733]" captionTargetBox-9="[203,1426,153,1559]" captionTargetId-0="figure-11@4.[228,1386,168,1716]" captionTargetId-1="figure-11@6.[192,1439,168,1855]" captionTargetId-10="figure-70@16.[214,1416,168,1814]" captionTargetId-11="figure-631@17.[192,1439,216,1281]" captionTargetId-12="figure-11@18.[192,1439,168,880]" captionTargetId-13="figure-7@19.[192,1439,168,1678]" captionTargetId-14="figure-69@20.[192,1439,168,1842]" captionTargetId-15="figure-654@21.[192,1439,168,1228]" captionTargetId-16="figure-11@22.[192,1439,168,1504]" captionTargetId-17="figure-69@23.[192,1439,168,1829]" captionTargetId-18="figure-11@24.[192,1439,223,1800]" captionTargetId-19="figure-217@25.[192,1439,168,1663]" captionTargetId-2="figure-11@8.[192,1439,170,1824]" captionTargetId-3="figure-6@9.[234,1392,200,1800]" captionTargetId-4="figure-11@10.[192,1439,192,1349]" captionTargetId-5="figure-6@11.[192,1439,191,1200]" captionTargetId-6="figure-11@12.[214,1416,193,1824]" captionTargetId-7="figure-6@13.[192,1439,168,933]" captionTargetId-8="figure-11@14.[192,1439,144,1741]" captionTargetId-9="figure-40@15.[192,1439,144,1560]" captionTargetPageId-0="4" captionTargetPageId-1="6" captionTargetPageId-10="16" captionTargetPageId-11="17" captionTargetPageId-12="18" captionTargetPageId-13="19" captionTargetPageId-14="20" captionTargetPageId-15="21" captionTargetPageId-16="22" captionTargetPageId-17="23" captionTargetPageId-18="24" captionTargetPageId-19="25" captionTargetPageId-2="8" captionTargetPageId-3="9" captionTargetPageId-4="10" captionTargetPageId-5="11" captionTargetPageId-6="12" captionTargetPageId-7="13" captionTargetPageId-8="14" captionTargetPageId-9="15" captionText-0="Figures 5–11: Figure 5. The reserarch team at a field station in the locality 15CB. Figure 6. Tea time in Girithale Wildlife Training Center close to the locality 15CD. From left P. Devasurendra, F. Kovařík and K. B. Ranawana. Figure 7. P. Rajkumar (Environmental Research, Jaffna), F. Kovařík and local people in the locality 15CK video recording Hottentotta tamulus. Figure 8. S. Jayarathne taking a picture of a dangerous snake Hypnale hypnale in the locality 15CD. Figure 9. Herpetologist S. Goonewardene and F. Kovařík in the locality 15CN. Figure 10. Meeting chief monk, in Monaragala &quot;Sri Sambodhi Viharaya&quot;, who has helped to research scorpions in the locality 15CQ. Figure 11. Profesor K. B. Ranawana in his office at University of Peradeniya." captionText-1="Figures 12–15: Maps showing the known geographical distribution of the Sri Lankan species." captionText-10="Figures 60–63: Charmus laneus from locality 15CO. Figures 60, 62. Male, chelicerae, carapace and tergites I–III (60) and sternopectinal region and sternites III–VII (62). Figures 61, 63. Female, chelicerae, carapace and tergites I–III (61) and sternopectinal region and sternite III (63)." captionText-11="Figures 64–65: Charmus laneus from locality 15CO, showing granulation and color pattern of male. Chelicerae, carapace and tergites (64). Coxosternal region and sternites (65). Scale bar: 500 µm." captionText-12="Figures 66–67: Left chelicera of male Charmus laneus from locality 15CO. Figures 66–67. Dorsal (66) and ventral (67a) view (white light + UV), and ventroexternal (67b) aspect (UV) showing 2 denticles on ventral surface of fixed finger (d1, d2). Scale bar: 200 µm." captionText-13="Figures 68–70: Charmus laneus, male from locality 15CO. Metasoma V and telson under UV fluorescence, showing granulation, punctuation and setation in dorsal (68), lateral (69) and ventral (70) views. Scale bar: 500 µm." captionText-14="Figures 71–73: Metasoma and telson of Charmus laneus male from locality 15CO, showing granulation, punctuation, setation and color pattern in dorsal (71), lateral (72) and ventral (73) views. Scale bar: 1 mm." captionText-15="Figures 74–79: Charmus laneus from locality 15CO. Figures 74–76. Male, metasoma and telson, lateral (74), ventral (75), and dorsal (76) views. Figures 77–79. Female, metasoma and telson, lateral (77), ventral (78), and dorsal (79) views." captionText-16="Figures 80–84: Comparing Sri Lankan Charmus female metasomas. Figures 80–82. Female holotype of Charmus laneus, metasoma and telson, lateral (80), ventral (81), and dorsal (82) views. Figures 83. Female of Charmus laneus from locality 15CO, metasoma and telson dorsal. Figures 84. Female holotype of Charmus saradieli sp. n. (MHNG) metasoma and telson dorsal." captionText-17="Figures 85–87: Charmus. Figures 85–86. Male (85) and female (86) of Charmus laneus from locality 15CO. Figure 87. Male paratype of Charmus saradieli sp. n. (UPSL)." captionText-18="Figures 88–91: Charmus saradieli sp. n. Figures 88–89. Male paratype (MHNG) in dorsal (88) and ventral (89) views, and original labels. Figures 90–91. Female holotype in dorsal (90) and ventral (91) views, and original labels. Scale bar: 5 mm." captionText-19="Figures 92–100: Charmus. Figures 92–95, 99–100. Charmus saradieli sp. n., male paratype (MHNG), chelicerae, carapace and tergites I–IV (92), sternopectinal region and sternites III–IV (94), and telson lateral (99); female holotype, chelicerae, carapace and tergites I–III (93), sternopectinal region and sternite III (95), and telson lateral (100). Figures 96–98. Charmus laneus, female holotype, chelicera and carapace (96), original label (97), and sternopectinal region and sternites III–IV (98)." captionText-2="Figures 16–19: Buthoscorpio sarasinorum. Figures 16–17. Male from locality 15CF in dorsal (16) and ventral (17) views. Figures 18–19. Female from locality 15CG in dorsal (18) and ventral (19) views." captionText-3="Figures 20–23: Buthoscorpio sarasinorum. Figures 20, 22. Male from locality 15CF, chelicerae, carapace and tergites I–III (20) and sternopectinal region and sternite III (22). Figures 21, 23. Female from locality 15CG, chelicerae, carapace and tergites I–III (21) and sternopectinal region and sternite III (23)." captionText-4="Figures 24–29: Buthoscorpio sarasinorum. Figures 24–26. Male from locality 15CF, metasoma and telson, lateral (24), ventral (25), and dorsal (26) views. Figures 27–29. Female from locality 15CG, metasoma and telson, lateral (27), ventral (28), and dorsal (29) views." captionText-5="Figures 30–33: Capsule region and flagellum of left hemispermatophore of Buthoscorpio sarasinorum from locality 15CF. External (30), dorsal (31), internal (32) and ventral (33) views. Note: damaged section of pars reflecta of flagellum is interpolated as a dotted outline. Scale bar: 500 µm. Abbreviations: bl, basal lobe; f, flagellum; ml, median lobe; mlc, median lobe carina; pr, pars recta of flagellum; prf, pars reflecta of flagellum; t, trunk." captionText-6="Figures 34–38: Buthoscorpio sarasinorum. Figures 34. Male at locality 15CF. Figures 35. Female at locality 15CS under UV. Figures 36–38. Female from locality 15CF (36) with newborns (37), and with juveniles after first ecdysis (38)." captionText-7="Figures 39–46: Chela of pedipalps (39–45) and movable finger of chela (46). Figures 39–40. Buthoscorpio sarasinorum, male from locality 15CF (39) and female from locality 15CG (40). Figures 41–43. Charmus laneus, male (41) and female (42) from locality 15CO, and female holotype (43). Figure 44. Charmus saradieli, sp. n., female holotype. Figures 45–46. Hottentotta tamulus, male from locality 15CK." captionText-8="Figures 47–53: Figure 47. Left hemispermatophore of Buthoscorpio sarasinorum from locality 15CF. Figures 48–53. Charmus laneus from locality 15CO. Figure 48. Left hemispermatophore. Figures 49–53. Capsule region and flagellum. External (49), dorsal (50), internal (51) and ventral (52) views of left hemispermatophore, and externo-dorsal (53) view of right hemispermatophore presented as mirror image for comparison. Scale bars: 1 mm for 47, 500 µm for 48, 250 µm for 49–52, and 200 µm for 53. Abbreviations: bl, basal lobe; f, flagellum; ml, median lobe; mlc, median lobe carina; pr, pars recta of flagellum; prf, pars reflecta of flagellum; t, trunk." captionText-9="Figures 54–59: Charmus laneus from locality 15CO. Figures 54–55. Male 14.7 mm long in dorsal (54) and ventral (55) views. Figures 56–57. Juvenile male 7.5 mm long in dorsal (56) and ventral (57) views. Figures 58–59. Female 20.1 mm long in dorsal (58) and ventral (59) views." figureDoi-0="http://doi.org/10.5281/zenodo.7124472" figureDoi-1="http://doi.org/10.5281/zenodo.7124476" figureDoi-10="http://doi.org/10.5281/zenodo.7124499" figureDoi-11="http://doi.org/10.5281/zenodo.7124503" figureDoi-12="http://doi.org/10.5281/zenodo.7124507" figureDoi-13="http://doi.org/10.5281/zenodo.7124511" figureDoi-14="http://doi.org/10.5281/zenodo.7124513" figureDoi-15="http://doi.org/10.5281/zenodo.7124521" figureDoi-16="http://doi.org/10.5281/zenodo.7124523" figureDoi-17="http://doi.org/10.5281/zenodo.7124529" figureDoi-18="http://doi.org/10.5281/zenodo.7124535" figureDoi-19="http://doi.org/10.5281/zenodo.7124537" figureDoi-2="http://doi.org/10.5281/zenodo.7124478" figureDoi-3="http://doi.org/10.5281/zenodo.7124482" figureDoi-4="http://doi.org/10.5281/zenodo.7124487" figureDoi-5="http://doi.org/10.5281/zenodo.7124489" figureDoi-6="http://doi.org/10.5281/zenodo.7124491" figureDoi-7="http://doi.org/10.5281/zenodo.7124493" figureDoi-8="http://doi.org/10.5281/zenodo.7124495" figureDoi-9="http://doi.org/10.5281/zenodo.7124497" httpUri-0="https://zenodo.org/record/7124472/files/figure.png" httpUri-1="https://zenodo.org/record/7124476/files/figure.png" httpUri-10="https://zenodo.org/record/7124499/files/figure.png" httpUri-11="https://zenodo.org/record/7124503/files/figure.png" httpUri-12="https://zenodo.org/record/7124507/files/figure.png" httpUri-13="https://zenodo.org/record/7124511/files/figure.png" httpUri-14="https://zenodo.org/record/7124513/files/figure.png" httpUri-15="https://zenodo.org/record/7124521/files/figure.png" httpUri-16="https://zenodo.org/record/7124523/files/figure.png" httpUri-17="https://zenodo.org/record/7124529/files/figure.png" httpUri-18="https://zenodo.org/record/7124535/files/figure.png" httpUri-19="https://zenodo.org/record/7124537/files/figure.png" httpUri-2="https://zenodo.org/record/7124478/files/figure.png" httpUri-3="https://zenodo.org/record/7124482/files/figure.png" httpUri-4="https://zenodo.org/record/7124487/files/figure.png" httpUri-5="https://zenodo.org/record/7124489/files/figure.png" httpUri-6="https://zenodo.org/record/7124491/files/figure.png" httpUri-7="https://zenodo.org/record/7124493/files/figure.png" httpUri-8="https://zenodo.org/record/7124495/files/figure.png" httpUri-9="https://zenodo.org/record/7124497/files/figure.png" pageId="22" pageNumber="21">Fig. 5 92</figureCitation>
),
<date id="4C661009A801FF86FEC2F9749F19F907" box="[258,379,1699,1722]" pageId="22" pageNumber="21" value="2015-04-28">
<collectingDate id="5C22E9E1A801FF86FEC2F9749F19F907" box="[258,379,1699,1722]" pageId="22" pageNumber="21" value="2015-04-28">28.IV.2015</collectingDate>
</date>
,
<specimenCount id="2EDEFD40A801FF86FE4DF9749FCCF907" box="[397,430,1699,1722]" pageId="22" pageNumber="21" type="male">1♂</specimenCount>
(
<figureCitation id="A0E32A4CA801FF86FE02F9749C7EF907" box="[450,540,1699,1722]" captionStart="Figures 39–46" captionStartId="13.[192,278,946,970]" captionTargetBox="[196,1427,173,927]" captionTargetId="figure-6@13.[192,1439,168,933]" captionTargetPageId="13" captionText="Figures 39–46: Chela of pedipalps (39–45) and movable finger of chela (46). Figures 39–40. Buthoscorpio sarasinorum, male from locality 15CF (39) and female from locality 15CG (40). Figures 41–43. Charmus laneus, male (41) and female (42) from locality 15CO, and female holotype (43). Figure 44. Charmus saradieli, sp. n., female holotype. Figures 45–46. Hottentotta tamulus, male from locality 15CK." figureDoi="http://doi.org/10.5281/zenodo.7124493" httpUri="https://zenodo.org/record/7124493/files/figure.png" pageId="22" pageNumber="21">Figs. 41</figureCitation>
,
<figureCitation id="A0E32A4CA801FF86FDEDF9749C12F907" box="[557,624,1699,1722]" captionStart-0="Figures 47–53" captionStart-1="Figures 54–59" captionStartId-0="14.[192,278,1770,1794]" captionStartId-1="15.[192,278,1589,1613]" captionTargetBox-0="[207,1426,157,1733]" captionTargetBox-1="[203,1426,153,1559]" captionTargetId-0="figure-11@14.[192,1439,144,1741]" captionTargetId-1="figure-40@15.[192,1439,144,1560]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionText-0="Figures 47–53: Figure 47. Left hemispermatophore of Buthoscorpio sarasinorum from locality 15CF. Figures 48–53. Charmus laneus from locality 15CO. Figure 48. Left hemispermatophore. Figures 49–53. Capsule region and flagellum. External (49), dorsal (50), internal (51) and ventral (52) views of left hemispermatophore, and externo-dorsal (53) view of right hemispermatophore presented as mirror image for comparison. Scale bars: 1 mm for 47, 500 µm for 48, 250 µm for 49–52, and 200 µm for 53. Abbreviations: bl, basal lobe; f, flagellum; ml, median lobe; mlc, median lobe carina; pr, pars recta of flagellum; prf, pars reflecta of flagellum; t, trunk." captionText-1="Figures 54–59: Charmus laneus from locality 15CO. Figures 54–55. Male 14.7 mm long in dorsal (54) and ventral (55) views. Figures 56–57. Juvenile male 7.5 mm long in dorsal (56) and ventral (57) views. Figures 58–59. Female 20.1 mm long in dorsal (58) and ventral (59) views." figureDoi-0="http://doi.org/10.5281/zenodo.7124495" figureDoi-1="http://doi.org/10.5281/zenodo.7124497" httpUri-0="https://zenodo.org/record/7124495/files/figure.png" httpUri-1="https://zenodo.org/record/7124497/files/figure.png" pageId="22" pageNumber="21">48–55</figureCitation>
,
<figureCitation id="A0E32A4CA801FF86FD40F9749CA5F907" box="[640,711,1699,1722]" captionStart="Figures 60–63" captionStartId="16.[192,278,1844,1868]" captionTargetBox="[213,1417,167,1815]" captionTargetId="figure-70@16.[214,1416,168,1814]" captionTargetPageId="16" captionText="Figures 60–63: Charmus laneus from locality 15CO. Figures 60, 62. Male, chelicerae, carapace and tergites I–III (60) and sternopectinal region and sternites III–VII (62). Figures 61, 63. Female, chelicerae, carapace and tergites I–III (61) and sternopectinal region and sternite III (63)." figureDoi="http://doi.org/10.5281/zenodo.7124499" httpUri="https://zenodo.org/record/7124499/files/figure.png" pageId="22" pageNumber="21">60, 62</figureCitation>
,
<figureCitation id="A0E32A4CA801FF86FD18F9749D79F907" box="[728,795,1699,1722]" captionStart="Figures 74–79" captionStartId="21.[192,278,1257,1281]" captionTargetBox="[199,1428,176,1222]" captionTargetId="figure-654@21.[192,1439,168,1228]" captionTargetPageId="21" captionText="Figures 74–79: Charmus laneus from locality 15CO. Figures 74–76. Male, metasoma and telson, lateral (74), ventral (75), and dorsal (76) views. Figures 77–79. Female, metasoma and telson, lateral (77), ventral (78), and dorsal (79) views." figureDoi="http://doi.org/10.5281/zenodo.7124521" httpUri="https://zenodo.org/record/7124521/files/figure.png" pageId="22" pageNumber="21">74–76</figureCitation>
,
<figureCitation id="A0E32A4CA801FF86FF00F9159E88F964" box="[192,234,1730,1753]" captionStart-0="Figures 1–4" captionStart-1="Figures 5–11" captionStart-2="Figures 12–15" captionStart-3="Figures 16–19" captionStartId-0="3.[192,278,1840,1864]" captionStartId-1="4.[192,278,1745,1769]" captionStartId-2="6.[192,278,1885,1909]" captionStartId-3="8.[192,278,1853,1877]" captionTargetBox-0="[191,1440,191,1811]" captionTargetBox-1="[227,1387,167,1716]" captionTargetBox-2="[191,1440,167,1856]" captionTargetBox-3="[256,1384,176,1817]" captionTargetId-0="figure-6@3.[192,1439,192,1810]" captionTargetId-1="figure-11@4.[228,1386,168,1716]" captionTargetId-2="figure-11@6.[192,1439,168,1855]" captionTargetId-3="figure-11@8.[192,1439,170,1824]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="6" captionTargetPageId-3="8" captionText-0="Figures 1–4: Figure 1. Sri Lankan scorpion expedition 2015 research team, the official photo. Figure 2. Collecting in the locality 15CA, within Peradeniya Univesity land. Figure 3. The main building of the Faculty of Science, Univesity of Peradeniya. Figure 4. F. Kovařík taking photo of Heterometrus swammerdami in the locality 15CB." captionText-1="Figures 5–11: Figure 5. The reserarch team at a field station in the locality 15CB. Figure 6. Tea time in Girithale Wildlife Training Center close to the locality 15CD. From left P. Devasurendra, F. Kovařík and K. B. Ranawana. Figure 7. P. Rajkumar (Environmental Research, Jaffna), F. Kovařík and local people in the locality 15CK video recording Hottentotta tamulus. Figure 8. S. Jayarathne taking a picture of a dangerous snake Hypnale hypnale in the locality 15CD. Figure 9. Herpetologist S. Goonewardene and F. Kovařík in the locality 15CN. Figure 10. Meeting chief monk, in Monaragala &quot;Sri Sambodhi Viharaya&quot;, who has helped to research scorpions in the locality 15CQ. Figure 11. Profesor K. B. Ranawana in his office at University of Peradeniya." captionText-2="Figures 12–15: Maps showing the known geographical distribution of the Sri Lankan species." captionText-3="Figures 16–19: Buthoscorpio sarasinorum. Figures 16–17. Male from locality 15CF in dorsal (16) and ventral (17) views. Figures 18–19. Female from locality 15CG in dorsal (18) and ventral (19) views." figureDoi-0="http://doi.org/10.5281/zenodo.7124468" figureDoi-1="http://doi.org/10.5281/zenodo.7124472" figureDoi-2="http://doi.org/10.5281/zenodo.7124476" figureDoi-3="http://doi.org/10.5281/zenodo.7124478" httpUri-0="https://zenodo.org/record/7124468/files/figure.png" httpUri-1="https://zenodo.org/record/7124472/files/figure.png" httpUri-2="https://zenodo.org/record/7124476/files/figure.png" httpUri-3="https://zenodo.org/record/7124478/files/figure.png" pageId="22" pageNumber="21">1 18</figureCitation>
,
<figureCitation id="A0E32A4CA801FF86FF38F9159F43F964" box="[248,289,1730,1753]" captionStart="Figures 193–200" captionStartId="48.[192,278,1070,1094]" captionTargetBox="[213,1446,171,1037]" captionTargetId="figure-11@48.[205,1452,168,1041]" captionTargetPageId="48" captionText="Figures 193–200: Distal segments of leg IV, retroventral view of Sri Lankan Buthidae and Chaerilidae genera. Buthoscorpio sarasinorum, male from locality 15CF (193), Charmus laneus, male from locality 15CO (194), Hottentotta tamulus, male from locality 15CK (195), Isometrus maculatus, female from locality 15CI (196), Isometrus thwaitesi, female from locality 15CO (197), Lychas srilankensis, male from locality 15CJ (198), Reddyanus basilicus, male from locality 15CS (199), and Chaerilus ceylonensis, male from locality 15CD (200)." figureDoi="http://doi.org/10.5281/zenodo.7124608" httpUri="https://zenodo.org/record/7124608/files/figure.png" pageId="22" pageNumber="21">194</figureCitation>
,
<figureCitation id="A0E32A4CA801FF86FEEDF9159F37F964" box="[301,341,1730,1753]" captionStart="Figures 39–46" captionStartId="13.[192,278,946,970]" captionTargetBox="[196,1427,173,927]" captionTargetId="figure-6@13.[192,1439,168,933]" captionTargetPageId="13" captionText="Figures 39–46: Chela of pedipalps (39–45) and movable finger of chela (46). Figures 39–40. Buthoscorpio sarasinorum, male from locality 15CF (39) and female from locality 15CG (40). Figures 41–43. Charmus laneus, male (41) and female (42) from locality 15CO, and female holotype (43). Figure 44. Charmus saradieli, sp. n., female holotype. Figures 45–46. Hottentotta tamulus, male from locality 15CK." figureDoi="http://doi.org/10.5281/zenodo.7124493" httpUri="https://zenodo.org/record/7124493/files/figure.png" pageId="22" pageNumber="21">42 3</figureCitation>
,
<figureCitation id="A0E32A4CA801FF86FEA4F9159FEEF964" box="[356,396,1730,1753]" captionStart="Figures 547–554" captionStartId="114.[192,278,810,834]" captionTargetBox="[191,1440,191,782]" captionTargetId="figure-538@114.[192,1440,192,781]" captionTargetPageId="114" captionText="Figures 547–554: The mitotic metaphases of buthid males from Sri Lanka. Buthoscorpio sarasinorum from locality 15CF (2n=14) (547), Charmus laneus from locality 15CO (2n=9) (548), Isometrus thwaitesi from locality 15CF (2n=8) (549), Lychas srilankensis from locality 15CN (2n=16) (550), Reddyanus basilicus from locality 15CS (2n=16) (551), Reddyanus basilicus from locality 15CR (2n=15) (552), Reddyanus ceylonensis from locality 15CI (2n=16) (553), Reddyanus loebli from locality 15CH (2n=17) (554). Arrowheads indicate extra large odd chromosomes. Scale bar = 5 µm for 547–554." figureDoi="http://doi.org/10.5281/zenodo.7124765" httpUri="https://zenodo.org/record/7124765/files/figure.png" pageId="22" pageNumber="21">548</figureCitation>
)
<specimenCount id="2EDEFD40A801FF86FE5EF9159FDCF964" box="[414,446,1730,1753]" pageId="22" pageNumber="21" type="female">1♀</specimenCount>
(
<figureCitation id="A0E32A4CA801FF86FE0FF9159C44F964" box="[463,550,1730,1753]" captionStart="Figures 39–46" captionStartId="13.[192,278,946,970]" captionTargetBox="[196,1427,173,927]" captionTargetId="figure-6@13.[192,1439,168,933]" captionTargetPageId="13" captionText="Figures 39–46: Chela of pedipalps (39–45) and movable finger of chela (46). Figures 39–40. Buthoscorpio sarasinorum, male from locality 15CF (39) and female from locality 15CG (40). Figures 41–43. Charmus laneus, male (41) and female (42) from locality 15CO, and female holotype (43). Figure 44. Charmus saradieli, sp. n., female holotype. Figures 45–46. Hottentotta tamulus, male from locality 15CK." figureDoi="http://doi.org/10.5281/zenodo.7124493" httpUri="https://zenodo.org/record/7124493/files/figure.png" pageId="22" pageNumber="21">Figs. 42</figureCitation>
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<figureCitation id="A0E32A4CA801FF86FDF6F9159C18F964" box="[566,634,1730,1753]" captionStart="Figures 54–59" captionStartId="15.[192,278,1589,1613]" captionTargetBox="[203,1426,153,1559]" captionTargetId="figure-40@15.[192,1439,144,1560]" captionTargetPageId="15" captionText="Figures 54–59: Charmus laneus from locality 15CO. Figures 54–55. Male 14.7 mm long in dorsal (54) and ventral (55) views. Figures 56–57. Juvenile male 7.5 mm long in dorsal (56) and ventral (57) views. Figures 58–59. Female 20.1 mm long in dorsal (58) and ventral (59) views." figureDoi="http://doi.org/10.5281/zenodo.7124497" httpUri="https://zenodo.org/record/7124497/files/figure.png" pageId="22" pageNumber="21">58–59</figureCitation>
,
<figureCitation id="A0E32A4CA801FF86FD47F9159CABF964" box="[647,713,1730,1753]" captionStart="Figures 60–63" captionStartId="16.[192,278,1844,1868]" captionTargetBox="[213,1417,167,1815]" captionTargetId="figure-70@16.[214,1416,168,1814]" captionTargetPageId="16" captionText="Figures 60–63: Charmus laneus from locality 15CO. Figures 60, 62. Male, chelicerae, carapace and tergites I–III (60) and sternopectinal region and sternites III–VII (62). Figures 61, 63. Female, chelicerae, carapace and tergites I–III (61) and sternopectinal region and sternite III (63)." figureDoi="http://doi.org/10.5281/zenodo.7124499" httpUri="https://zenodo.org/record/7124499/files/figure.png" pageId="22" pageNumber="21">61, 63</figureCitation>
,
<figureCitation id="A0E32A4CA801FF86FD19F9159D7CF964" box="[729,798,1730,1753]" captionStart="Figures 74–79" captionStartId="21.[192,278,1257,1281]" captionTargetBox="[199,1428,176,1222]" captionTargetId="figure-654@21.[192,1439,168,1228]" captionTargetPageId="21" captionText="Figures 74–79: Charmus laneus from locality 15CO. Figures 74–76. Male, metasoma and telson, lateral (74), ventral (75), and dorsal (76) views. Figures 77–79. Female, metasoma and telson, lateral (77), ventral (78), and dorsal (79) views." figureDoi="http://doi.org/10.5281/zenodo.7124521" httpUri="https://zenodo.org/record/7124521/files/figure.png" pageId="22" pageNumber="21">77–79</figureCitation>
,
<figureCitation id="A0E32A4CA801FF86FF01F9369EB8F945" box="[193,218,1761,1784]" captionStart="Figures 5–11" captionStartId="4.[192,278,1745,1769]" captionTargetBox="[227,1387,167,1716]" captionTargetId="figure-11@4.[228,1386,168,1716]" captionTargetPageId="4" captionText="Figures 5–11: Figure 5. The reserarch team at a field station in the locality 15CB. Figure 6. Tea time in Girithale Wildlife Training Center close to the locality 15CD. From left P. Devasurendra, F. Kovařík and K. B. Ranawana. Figure 7. P. Rajkumar (Environmental Research, Jaffna), F. Kovařík and local people in the locality 15CK video recording Hottentotta tamulus. Figure 8. S. Jayarathne taking a picture of a dangerous snake Hypnale hypnale in the locality 15CD. Figure 9. Herpetologist S. Goonewardene and F. Kovařík in the locality 15CN. Figure 10. Meeting chief monk, in Monaragala &quot;Sri Sambodhi Viharaya&quot;, who has helped to research scorpions in the locality 15CQ. Figure 11. Profesor K. B. Ranawana in his office at University of Peradeniya." figureDoi="http://doi.org/10.5281/zenodo.7124472" httpUri="https://zenodo.org/record/7124472/files/figure.png" pageId="22" pageNumber="21">8 3</figureCitation>
,
<figureCitation id="A0E32A4CA801FF86FF2EF9369F6BF945" box="[238,265,1761,1784]" captionStart="Figures 85–87" captionStartId="23.[192,278,1859,1883]" captionTargetBox="[191,1440,167,1830]" captionTargetId="figure-69@23.[192,1439,168,1829]" captionTargetPageId="23" captionText="Figures 85–87: Charmus. Figures 85–86. Male (85) and female (86) of Charmus laneus from locality 15CO. Figure 87. Male paratype of Charmus saradieli sp. n. (UPSL)." figureDoi="http://doi.org/10.5281/zenodo.7124529" httpUri="https://zenodo.org/record/7124529/files/figure.png" pageId="22" pageNumber="21">85</figureCitation>
,
<figureCitation id="A0E32A4CA801FF86FEDBF9369F27F945" box="[283,325,1761,1784]" captionStart="Figures 107–119" captionStartId="27.[192,278,1733,1757]" captionTargetBox="[211,1428,190,1691]" captionTargetId="figure-6@27.[192,1439,181,1704]" captionTargetPageId="27" captionText="Figures 107–119: Charmus. Figures 107–114. Charmus saradieli sp. n., female holotype. Pedipalp movable finger (107). Pedipalp chela, dorsal (108), external (109), and ventral (110) views. Pedipalp patella, dorsal (111) and external (112) views. Pedipalp femur and trochanter, internal (113) and dorsal (114) views. The trichobothrial pattern is indicated in Figures 108a– 114a. Figures 115–117. Charmus saradieli sp. n., male paratype (MHNG), chelicerae (115), pedipalp dorsal (116), and fourth leg dorsal (117). Figures 118–119. Charmus laneus from locality 15CO. Pedipalp dorsal of male (118) and female (119)." figureDoi="http://doi.org/10.5281/zenodo.7124543" httpUri="https://zenodo.org/record/7124543/files/figure.png" pageId="22" pageNumber="21">119</figureCitation>
,
<figureCitation id="A0E32A4CA801FF86FE96F9369FE2F945" box="[342,384,1761,1784]" captionStart="Figures 403–429" captionStartId="85.[192,278,744,768]" captionTargetId="figure-11@84.[172,1464,185,1878]" captionTargetPageId="84" captionText="Figures 403–429: Telson, lateral view of Sri Lankan Buthidae and Chaerilidae species. Figures 403–404. Isometrus maculatus, male from locality 15CP (403) and female from locality 15CI (404). Figures 405–406. I. thwaitesi, male from locality 15CH (405) and female from locality 15CO (406). Figures 407–408. Lychas srilankensis, male (407) and female (408) from locality 15CJ. Figures 409–410. Reddyanus basilicus, male (409) and female (410) from locality 15CR. Figures 411–412. R. besucheti, male holotype (411) and female from locality 15CG (412). Figures 413–414. R. ceylonensis sp. n., male holotype (413) and female paratype (414) from locality 15CI. Figures 415–416. R. jayarathnei sp. n., male (415) and female (416) paratypes. Figures 417–418. R. loebli, male (417) and female (418) from locality 15CG. Figures 419–420. R. ranawanai sp. n., male holotype (419) and female paratype (420). Figures 421–422: Buthoscorpio sarasinorum, male from locality 15CF (421) and female from locality 15CG (422). Figures 423–424. Charmus laneus, male (423) and female (424) from locality 15CO. Figures 425–426. Charmus saradieli sp. n. male paratype (425) (MHNG) and female holotype (426). Figurs 427. Hottentotta tamulus, male from locality 15CK (427). Figures 428–429. Chaerilus ceylonensis, male (428) and female (429) from locality 15CD." figureDoi="http://doi.org/10.5281/zenodo.7124696" httpUri="https://zenodo.org/record/7124696/files/figure.png" pageId="22" pageNumber="21">424</figureCitation>
)
<specimenCount id="2EDEFD40A801FF86FE53F9369FA8F945" box="[403,458,1761,1784]" pageId="22" pageNumber="21" type="generic">1juv.</specimenCount>
(
<figureCitation id="A0E32A4CA801FF86FE1DF9369C00F945" box="[477,610,1761,1784]" captionStart="Figures 54–59" captionStartId="15.[192,278,1589,1613]" captionTargetBox="[203,1426,153,1559]" captionTargetId="figure-40@15.[192,1439,144,1560]" captionTargetPageId="15" captionText="Figures 54–59: Charmus laneus from locality 15CO. Figures 54–55. Male 14.7 mm long in dorsal (54) and ventral (55) views. Figures 56–57. Juvenile male 7.5 mm long in dorsal (56) and ventral (57) views. Figures 58–59. Female 20.1 mm long in dorsal (58) and ventral (59) views." figureDoi="http://doi.org/10.5281/zenodo.7124497" httpUri="https://zenodo.org/record/7124497/files/figure.png" pageId="22" pageNumber="21">Figs. 56–57</figureCitation>
,
<figureCitation id="A0E32A4CA801FF86FDB5F9369CF2F945" box="[629,656,1761,1784]" captionStart="Figures 85–87" captionStartId="23.[192,278,1859,1883]" captionTargetBox="[191,1440,167,1830]" captionTargetId="figure-69@23.[192,1439,168,1829]" captionTargetPageId="23" captionText="Figures 85–87: Charmus. Figures 85–86. Male (85) and female (86) of Charmus laneus from locality 15CO. Figure 87. Male paratype of Charmus saradieli sp. n. (UPSL)." figureDoi="http://doi.org/10.5281/zenodo.7124529" httpUri="https://zenodo.org/record/7124529/files/figure.png" pageId="22" pageNumber="21">86</figureCitation>
),
<collectionCode id="5EC9AE0CA801FF86FD6BF9369C93F945" box="[683,753,1761,1784]" pageId="22" pageNumber="21">FKCP</collectionCode>
,
<specimenCount id="2EDEFD40A801FF86FCC0F9369D40F945" box="[768,802,1761,1784]" pageId="22" pageNumber="21" type="male">2♂</specimenCount>
(
<figureCitation id="A0E32A4CA801FF86FF09F9289F2AF8AB" box="[201,328,1791,1814]" captionStart-0="Figures 64–65" captionStart-1="Figures 66–67" captionStart-2="Figures 68–70" captionStart-3="Figures 71–73" captionStartId-0="17.[192,278,1310,1334]" captionStartId-1="18.[192,278,909,933]" captionStartId-2="19.[192,278,1708,1732]" captionStartId-3="20.[192,278,1872,1896]" captionTargetBox-0="[191,1440,215,1282]" captionTargetBox-1="[204,1433,173,864]" captionTargetBox-2="[192,1439,168,1678]" captionTargetBox-3="[191,1440,167,1843]" captionTargetId-0="figure-631@17.[192,1439,216,1281]" captionTargetId-1="figure-11@18.[192,1439,168,880]" captionTargetId-2="figure-7@19.[192,1439,168,1678]" captionTargetId-3="figure-69@20.[192,1439,168,1842]" captionTargetPageId-0="17" captionTargetPageId-1="18" captionTargetPageId-2="19" captionTargetPageId-3="20" captionText-0="Figures 64–65: Charmus laneus from locality 15CO, showing granulation and color pattern of male. Chelicerae, carapace and tergites (64). Coxosternal region and sternites (65). Scale bar: 500 µm." captionText-1="Figures 66–67: Left chelicera of male Charmus laneus from locality 15CO. Figures 66–67. Dorsal (66) and ventral (67a) view (white light + UV), and ventroexternal (67b) aspect (UV) showing 2 denticles on ventral surface of fixed finger (d1, d2). Scale bar: 200 µm." captionText-2="Figures 68–70: Charmus laneus, male from locality 15CO. Metasoma V and telson under UV fluorescence, showing granulation, punctuation and setation in dorsal (68), lateral (69) and ventral (70) views. Scale bar: 500 µm." captionText-3="Figures 71–73: Metasoma and telson of Charmus laneus male from locality 15CO, showing granulation, punctuation, setation and color pattern in dorsal (71), lateral (72) and ventral (73) views. Scale bar: 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7124503" figureDoi-1="http://doi.org/10.5281/zenodo.7124507" figureDoi-2="http://doi.org/10.5281/zenodo.7124511" figureDoi-3="http://doi.org/10.5281/zenodo.7124513" httpUri-0="https://zenodo.org/record/7124503/files/figure.png" httpUri-1="https://zenodo.org/record/7124507/files/figure.png" httpUri-2="https://zenodo.org/record/7124511/files/figure.png" httpUri-3="https://zenodo.org/record/7124513/files/figure.png" pageId="22" pageNumber="21">Figs. 64–73</figureCitation>
)
<specimenCount id="2EDEFD40A801FF86FE96F9289F1AF8AB" box="[342,376,1791,1814]" pageId="22" pageNumber="21" type="female">1♀</specimenCount>
,
<collectionCode id="5EC9AE0CA801FF86FE45F9289FABF8AB" box="[389,457,1791,1814]" pageId="22" pageNumber="21">UPSL</collectionCode>
, leg.
<collectorName id="952D531FA801FF86FDC2F9289C3BF8AB" box="[514,601,1791,1814]" pageId="22" pageNumber="21">Kovařík</collectorName>
et al
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="6CA76641A801FF86FF00FA2A9F60F9E2" ID-DOI="http://doi.org/10.5281/zenodo.7124523" ID-Zenodo-Dep="7124523" httpUri="https://zenodo.org/record/7124523/files/figure.png" pageId="22" pageNumber="21" startId="22.[192,278,1533,1557]" targetBox="[206,1423,177,1494]" targetPageId="22">
<paragraph id="386736C9A801FF86FF00FA2A9F60F9E2" blockId="22.[192,1440,1533,1631]" pageId="22" pageNumber="21">
<emphasis id="0AACEADBA801FF86FF00FA2A9F09F9A8" bold="true" box="[192,363,1533,1557]" pageId="22" pageNumber="21">Figures 80–84:</emphasis>
Comparing Sri Lankan
<taxonomicName id="FFD84D4AA801FF86FDA3F9D79CD9F9A8" authorityName="Karsch" authorityYear="1879" box="[611,699,1536,1557]" class="Arachnida" family="Buthidae" genus="Charmus" kingdom="Animalia" order="Scorpiones" pageId="22" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="0AACEADBA801FF86FDA3F9D79CD9F9A8" box="[611,699,1536,1557]" italics="true" pageId="22" pageNumber="21">Charmus</emphasis>
</taxonomicName>
female metasomas.
<emphasis id="0AACEADBA801FF86FC4AFA289A7FF9A9" bold="true" box="[906,1053,1535,1557]" pageId="22" pageNumber="21">Figures 80–82</emphasis>
. Female holotype of
<taxonomicName id="FFD84D4AA801FF86FB39F9D79BF8F9A8" authorityName="Karsch" authorityYear="1879" box="[1273,1434,1536,1557]" class="Arachnida" family="Buthidae" genus="Charmus" kingdom="Animalia" order="Scorpiones" pageId="22" pageNumber="21" phylum="Arthropoda" rank="species" species="laneus">
<emphasis id="0AACEADBA801FF86FB39F9D79BF8F9A8" box="[1273,1434,1536,1557]" italics="true" pageId="22" pageNumber="21">Charmus laneus</emphasis>
</taxonomicName>
, metasoma and telson, lateral (80), ventral (81), and dorsal (82) views.
<emphasis id="0AACEADBA801FF86FC4AF9CF9D9BF990" bold="true" box="[906,1017,1560,1582]" pageId="22" pageNumber="21">Figures 83</emphasis>
. Female of
<taxonomicName id="FFD84D4AA801FF86FBB6F9CE9B75F993" authorityName="Karsch" authorityYear="1879" box="[1142,1303,1561,1582]" class="Arachnida" family="Buthidae" genus="Charmus" kingdom="Animalia" order="Scorpiones" pageId="22" pageNumber="21" phylum="Arthropoda" rank="species" species="laneus">
<emphasis id="0AACEADBA801FF86FBB6F9CE9B75F993" box="[1142,1303,1561,1582]" italics="true" pageId="22" pageNumber="21">Charmus laneus</emphasis>
</taxonomicName>
from locality 15CO, metasoma and telson dorsal.
<emphasis id="0AACEADBA801FF86FDE9F9E69CF5F9FB" bold="true" box="[553,663,1585,1607]" pageId="22" pageNumber="21">Figures 84</emphasis>
. Female holotype of
<taxonomicName id="FFD84D4AA801FF86FCAFF9E59A46F9FA" authority="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š, 2016" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" box="[879,1060,1586,1607]" class="Arachnida" family="Buthidae" genus="Charmus" kingdom="Animalia" order="Scorpiones" pageId="22" pageNumber="21" phylum="Arthropoda" rank="species" species="saradieli" status="sp. nov.">
<emphasis id="0AACEADBA801FF86FCAFF9E59A46F9FA" box="[879,1060,1586,1607]" italics="true" pageId="22" pageNumber="21">Charmus saradieli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="119F57A0A801FF86FBEEF9E69A04F9FA" box="[1070,1126,1585,1607]" pageId="22" pageNumber="21" rank="species">
<emphasis id="0AACEADBA801FF86FBEEF9E69A27F9FB" bold="true" box="[1070,1093,1585,1606]" pageId="22" pageNumber="21">sp</emphasis>
.
<emphasis id="0AACEADBA801FF86FB93F9E69A02F9FB" bold="true" box="[1107,1120,1585,1606]" pageId="22" pageNumber="21">n</emphasis>
.
</taxonomicNameLabel>
(MHNG) metasoma and telson dorsal.
</paragraph>
</caption>
<subSubSection id="70C26542A801FF8AFF00F8EA9D8EF8D4" lastPageId="26" lastPageNumber="25" pageId="22" pageNumber="21" type="diagnosis">
<paragraph id="386736C9A801FF8AFF00F8EA9D67FA4D" blockId="22.[192,802,1853,1906]" lastBlockId="26.[192,802,1436,1520]" lastPageId="26" lastPageNumber="25" pageId="22" pageNumber="21">
DIAGNOSIS. Total length
<quantity id="FF209B2CA801FF86FE14F8EA9C43F8E9" box="[468,545,1853,1876]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="22" pageNumber="21" unit="mm" value="14.0">14 mm</quantity>
(male) – 21.3 mm (female). Mesosoma, carapace, metasoma and telson of adults black; pedipalp femur almost entirely black with several small yellow spots; pedipalp patella yellowish with several black spots; legs yellow with black spots; chelicerae brown, with black reticulation. Carapace granular without carinae, anterior edge with epistome present medially. Tergites I–VI granular, obviously with one carina. Sternites without carinae. Metasomal segments IV–V or III–V punctate without developed carinae. Fifth metasomal segment length / width ratio
<quantity id="FF209B2CA80EFF89FF00F8949F06F8E7" box="[192,356,1859,1882]" metricMagnitude="1" metricUnit="m" metricValue="3.44551" metricValueMax="3.6195" metricValueMin="3.2715199999999998" pageId="25" pageNumber="24" unit="in" value="1356.5" valueMax="1425.0" valueMin="1288.0">1.288–1.425 in</quantity>
female. Pectines with or without fulcra. Ratio of pedipalp chela length / fixed finger length in female 1.692–1.791. Movable and fixed fingers of pedipalps bearing 8 rows of granules, apical rows of 4–6 granules, and 3 terminal granules; each row of granules (except last) with one internal and two external accessory granules. Pectinal teeth number
<quantity id="FF209B2CA80DFF8AFDBBFA6D9C81FA6C" box="[635,739,1466,1489]" metricMagnitude="-1" metricUnit="m" metricValue="4.318" metricValueMax="4.572" metricValueMin="4.064" pageId="26" pageNumber="25" unit="in" value="17.0" valueMax="18.0" valueMin="16.0">16–18 in</quantity>
both sexes. Telson vesicle punctate, rather bulbous in male.
</paragraph>
<caption id="6CA76641A800FF87FF00F8949C18F8CE" ID-DOI="http://doi.org/10.5281/zenodo.7124529" ID-Zenodo-Dep="7124529" httpUri="https://zenodo.org/record/7124529/files/figure.png" pageId="23" pageNumber="22" startId="23.[192,278,1859,1883]" targetBox="[191,1440,167,1830]" targetPageId="23">
<paragraph id="386736C9A800FF87FF00F8949C18F8CE" blockId="23.[192,1440,1859,1907]" pageId="23" pageNumber="22">
<emphasis id="0AACEADBA800FF87FF00F8949F0BF8E6" bold="true" box="[192,361,1859,1883]" pageId="23" pageNumber="22">Figures 85–87:</emphasis>
<taxonomicName id="FFD84D4AA800FF87FEAFF8919FA5F8E6" authorityName="Karsch" authorityYear="1879" box="[367,455,1862,1883]" class="Arachnida" family="Buthidae" genus="Charmus" kingdom="Animalia" order="Scorpiones" pageId="23" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="0AACEADBA800FF87FEAFF8919FA5F8E6" box="[367,455,1862,1883]" italics="true" pageId="23" pageNumber="22">Charmus</emphasis>
</taxonomicName>
.
<emphasis id="0AACEADBA800FF87FE14F8929C06F8E7" bold="true" box="[468,612,1861,1883]" pageId="23" pageNumber="22">Figures 85–86</emphasis>
. Male (85) and female (86) of
<taxonomicName id="FFD84D4AA800FF87FC52F8919A4DF8E6" authorityName="Karsch" authorityYear="1879" box="[914,1071,1862,1883]" class="Arachnida" family="Buthidae" genus="Charmus" kingdom="Animalia" order="Scorpiones" pageId="23" pageNumber="22" phylum="Arthropoda" rank="species" species="laneus">
<emphasis id="0AACEADBA800FF87FC52F8919A4DF8E6" box="[914,1071,1862,1883]" italics="true" pageId="23" pageNumber="22">Charmus laneus</emphasis>
</taxonomicName>
from locality 15CO.
<emphasis id="0AACEADBA800FF87FAC0F8929B00F8E7" bold="true" box="[1280,1378,1861,1883]" pageId="23" pageNumber="22">Figure 87</emphasis>
. Male paratype of
<taxonomicName id="FFD84D4AA800FF87FEF3F8899F84F8CE" authority="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š, 2016" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" box="[307,486,1886,1907]" class="Arachnida" family="Buthidae" genus="Charmus" kingdom="Animalia" order="Scorpiones" pageId="23" pageNumber="22" phylum="Arthropoda" rank="species" species="saradieli" status="sp. nov.">
<emphasis id="0AACEADBA800FF87FEF3F8899F84F8CE" box="[307,486,1886,1907]" italics="true" pageId="23" pageNumber="22">Charmus saradieli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="119F57A0A800FF87FE2CF8899C40F8CE" box="[492,546,1886,1907]" pageId="23" pageNumber="22" rank="species">
<emphasis id="0AACEADBA800FF87FE2CF8899C61F8CE" bold="true" box="[492,515,1886,1907]" pageId="23" pageNumber="22">sp</emphasis>
.
<emphasis id="0AACEADBA800FF87FDCFF8899C7EF8CE" bold="true" box="[527,540,1886,1907]" pageId="23" pageNumber="22">n</emphasis>
.
</taxonomicNameLabel>
(UPSL).
</paragraph>
</caption>
<caption id="6CA76641A80FFF88FF00F8DB9B01F881" ID-DOI="http://doi.org/10.5281/zenodo.7124535" ID-Zenodo-Dep="7124535" httpUri="https://zenodo.org/record/7124535/files/figure.png" pageId="24" pageNumber="23" startId="24.[192,278,1804,1828]" targetBox="[200,1418,242,1791]" targetPageId="24">
<paragraph id="386736C9A80FFF88FF00F8DB9B01F881" blockId="24.[192,1440,1804,1853]" pageId="24" pageNumber="23">
<emphasis id="0AACEADBA80FFF88FF00F8DB9F09F899" bold="true" box="[192,363,1804,1828]" pageId="24" pageNumber="23">Figures 88–91:</emphasis>
<taxonomicName id="FFD84D4AA80FFF88FEB2F8D89C4AF899" authority="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š, 2016" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" box="[370,552,1807,1828]" class="Arachnida" family="Buthidae" genus="Charmus" kingdom="Animalia" order="Scorpiones" pageId="24" pageNumber="23" phylum="Arthropoda" rank="species" species="saradieli" status="sp. nov.">
<emphasis id="0AACEADBA80FFF88FEB2F8D89C4AF899" box="[370,552,1807,1828]" italics="true" pageId="24" pageNumber="23">Charmus saradieli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="119F57A0A80FFF88FDF2F8D99C0AF899" box="[562,616,1806,1828]" pageId="24" pageNumber="23" rank="species">
<emphasis id="0AACEADBA80FFF88FDF2F8D99C2BF89E" bold="true" box="[562,585,1806,1827]" pageId="24" pageNumber="23">sp</emphasis>
.
<emphasis id="0AACEADBA80FFF88FD94F8D99C03F89E" bold="true" box="[596,609,1806,1827]" pageId="24" pageNumber="23">n</emphasis>
.
</taxonomicNameLabel>
<emphasis id="0AACEADBA80FFF88FDB0F8D99D63F89E" bold="true" box="[624,769,1806,1828]" pageId="24" pageNumber="23">Figures 88–89</emphasis>
. Male paratype (MHNG) in dorsal (88) and ventral (89) views, and original labels.
<emphasis id="0AACEADBA80FFF88FE95F8F09F86F881" bold="true" box="[341,484,1831,1853]" pageId="24" pageNumber="23">Figures 90–91</emphasis>
. Female holotype in dorsal (90) and ventral (91) views, and original labels. Scale bar: 5 mm.
</paragraph>
</caption>
<caption id="6CA76641A80EFF89FF00F94A9B05F942" ID-DOI="http://doi.org/10.5281/zenodo.7124537" ID-Zenodo-Dep="7124537" httpUri="https://zenodo.org/record/7124537/files/figure.png" pageId="25" pageNumber="24" startId="25.[192,278,1693,1717]" targetBox="[191,1440,167,1664]" targetPageId="25">
<paragraph id="386736C9A80EFF89FF00F94A9B05F942" blockId="25.[192,1441,1693,1791]" pageId="25" pageNumber="24">
<emphasis id="0AACEADBA80EFF89FF00F94A9F1AF908" bold="true" box="[192,376,1693,1717]" pageId="25" pageNumber="24">Figures 92–100:</emphasis>
<taxonomicName id="FFD84D4AA80EFF89FE40F9489FBAF909" authorityName="Karsch" authorityYear="1879" box="[384,472,1695,1716]" class="Arachnida" family="Buthidae" genus="Charmus" kingdom="Animalia" order="Scorpiones" pageId="25" pageNumber="24" phylum="Arthropoda" rank="genus">
<emphasis id="0AACEADBA80EFF89FE40F9489FBAF909" box="[384,472,1695,1716]" italics="true" pageId="25" pageNumber="24">Charmus</emphasis>
</taxonomicName>
.
<emphasis id="0AACEADBA80EFF89FE27F9489CACF909" bold="true" box="[487,718,1695,1717]" pageId="25" pageNumber="24">Figures 92–95, 99–100</emphasis>
.
<taxonomicName id="FFD84D4AA80EFF89FD1DF9489DF1F909" authority="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š, 2016" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" box="[733,915,1695,1716]" class="Arachnida" family="Buthidae" genus="Charmus" kingdom="Animalia" order="Scorpiones" pageId="25" pageNumber="24" phylum="Arthropoda" rank="species" species="saradieli" status="sp. nov.">
<emphasis id="0AACEADBA80EFF89FD1DF9489DF1F909" box="[733,915,1695,1716]" italics="true" pageId="25" pageNumber="24">Charmus saradieli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="119F57A0A80EFF89FC5CF9489DB1F909" box="[924,979,1695,1716]" pageId="25" pageNumber="24" rank="species">
<emphasis id="0AACEADBA80EFF89FC5CF9489DD1F909" bold="true" box="[924,947,1695,1716]" pageId="25" pageNumber="24">sp</emphasis>
.
<emphasis id="0AACEADBA80EFF89FC01F9489DACF909" bold="true" box="[961,974,1695,1716]" pageId="25" pageNumber="24">n</emphasis>
.
</taxonomicNameLabel>
, male paratype (MHNG), chelicerae, carapace and tergites I–IV (92), sternopectinal region and sternites III–IV (94), and telson lateral (99); female holotype, chelicerae, carapace and tergites I–III (93), sternopectinal region and sternite III (95), and telson lateral (100).
<emphasis id="0AACEADBA80EFF89FB66F9079B5BF958" bold="true" box="[1190,1337,1744,1766]" pageId="25" pageNumber="24">Figures 96–98</emphasis>
.
<taxonomicName id="FFD84D4AA80EFF89FA88F9069E9DF942" authorityName="Karsch" authorityYear="1879" class="Arachnida" family="Buthidae" genus="Charmus" kingdom="Animalia" order="Scorpiones" pageId="25" pageNumber="24" phylum="Arthropoda" rank="species" species="laneus">
<emphasis id="0AACEADBA80EFF89FA88F9069E9DF942" italics="true" pageId="25" pageNumber="24">Charmus laneus</emphasis>
</taxonomicName>
, female holotype, chelicera and carapace (96), original label (97), and sternopectinal region and sternites III–IV (98).
</paragraph>
</caption>
<caption id="6CA76641A80DFF8AFF00FAFA9F18FACB" ID-DOI="http://doi.org/10.5281/zenodo.7124541" ID-Zenodo-Dep="7124541" httpUri="https://zenodo.org/record/7124541/files/figure.png" pageId="26" pageNumber="25" startId="26.[192,278,1325,1349]" targetBox="[190,1412,176,1289]" targetPageId="26">
<paragraph id="386736C9A80DFF8AFF00FAFA9F18FACB" blockId="26.[192,1440,1325,1398]" pageId="26" pageNumber="25">
<emphasis id="0AACEADBA80DFF8AFF00FAFA9FE7FAF8" bold="true" box="[192,389,1325,1349]" pageId="26" pageNumber="25">Figures 101–106:</emphasis>
<taxonomicName id="FFD84D4AA80DFF8AFE4EFAF89C26FAF9" authority="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š, 2016" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" box="[398,580,1327,1348]" class="Arachnida" family="Buthidae" genus="Charmus" kingdom="Animalia" order="Scorpiones" pageId="26" pageNumber="25" phylum="Arthropoda" rank="species" species="saradieli" status="sp. nov.">
<emphasis id="0AACEADBA80DFF8AFE4EFAF89C26FAF9" box="[398,580,1327,1348]" italics="true" pageId="26" pageNumber="25">Charmus saradieli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="119F57A0A80DFF8AFD8DFAF89CE4FAF9" box="[589,646,1327,1348]" pageId="26" pageNumber="25" rank="species">
<emphasis id="0AACEADBA80DFF8AFD8DFAF89C06FAF9" bold="true" box="[589,612,1327,1348]" pageId="26" pageNumber="25">sp</emphasis>
.
<emphasis id="0AACEADBA80DFF8AFDB2FAF89C1DFAF9" bold="true" box="[626,639,1327,1348]" pageId="26" pageNumber="25">n</emphasis>
.
</taxonomicNameLabel>
<emphasis id="0AACEADBA80DFF8AFD4EFAF89D5AFAF9" bold="true" box="[654,824,1327,1349]" pageId="26" pageNumber="25">Figures 101–103</emphasis>
. Male paratype (MHNG), metasoma and telson, lateral (101), ventral (102), and dorsal (103) views.
<emphasis id="0AACEADBA80DFF8AFDF6FA9F9CBFFAE0" bold="true" box="[566,733,1352,1374]" pageId="26" pageNumber="25">Figures 104–106</emphasis>
. Female holotype, metasoma and telson, lateral (104), ventral (105), and dorsal (106) views.
</paragraph>
</caption>
<paragraph id="386736C9A80DFF8AFF00F9C79FF1F966" blockId="26.[192,803,1552,1755]" pageId="26" pageNumber="25">
HEMISPERMATOPHORE (
<figureCitation id="A0E32A4CA80DFF8AFE04F9C79C2FF99A" box="[452,589,1552,1575]" captionStart="Figures 47–53" captionStartId="14.[192,278,1770,1794]" captionTargetBox="[207,1426,157,1733]" captionTargetId="figure-11@14.[192,1439,144,1741]" captionTargetPageId="14" captionText="Figures 47–53: Figure 47. Left hemispermatophore of Buthoscorpio sarasinorum from locality 15CF. Figures 48–53. Charmus laneus from locality 15CO. Figure 48. Left hemispermatophore. Figures 49–53. Capsule region and flagellum. External (49), dorsal (50), internal (51) and ventral (52) views of left hemispermatophore, and externo-dorsal (53) view of right hemispermatophore presented as mirror image for comparison. Scale bars: 1 mm for 47, 500 µm for 48, 250 µm for 49–52, and 200 µm for 53. Abbreviations: bl, basal lobe; f, flagellum; ml, median lobe; mlc, median lobe carina; pr, pars recta of flagellum; prf, pars reflecta of flagellum; t, trunk." figureDoi="http://doi.org/10.5281/zenodo.7124495" httpUri="https://zenodo.org/record/7124495/files/figure.png" pageId="26" pageNumber="25">Figs. 48–53</figureCitation>
). Trunk very narrow, elongate, capsule region short (
<figureCitation id="A0E32A4CA80DFF8AFD8EF9F99CC0F9F8" box="[590,674,1582,1605]" captionStart="Figures 47–53" captionStartId="14.[192,278,1770,1794]" captionTargetBox="[207,1426,157,1733]" captionTargetId="figure-11@14.[192,1439,144,1741]" captionTargetPageId="14" captionText="Figures 47–53: Figure 47. Left hemispermatophore of Buthoscorpio sarasinorum from locality 15CF. Figures 48–53. Charmus laneus from locality 15CO. Figure 48. Left hemispermatophore. Figures 49–53. Capsule region and flagellum. External (49), dorsal (50), internal (51) and ventral (52) views of left hemispermatophore, and externo-dorsal (53) view of right hemispermatophore presented as mirror image for comparison. Scale bars: 1 mm for 47, 500 µm for 48, 250 µm for 49–52, and 200 µm for 53. Abbreviations: bl, basal lobe; f, flagellum; ml, median lobe; mlc, median lobe carina; pr, pars recta of flagellum; prf, pars reflecta of flagellum; t, trunk." figureDoi="http://doi.org/10.5281/zenodo.7124495" httpUri="https://zenodo.org/record/7124495/files/figure.png" pageId="26" pageNumber="25">Fig. 48</figureCitation>
). Flagellum cylindriform, relatively short, robust, coiled. Median lobe broad, distally truncate, with straight dorsal carina near internal margin. Basal lobe well developed, a prominent, blunt, bilobate scoop arising dorsally near base of median lobe carina.
</paragraph>
<paragraph id="386736C9A80DFF8AFF00F92D9D8EF8D4" blockId="26.[192,802,1786,1900]" lastBlockId="26.[830,1441,1436,1897]" pageId="26" pageNumber="25">
C HELICERA (
<figureCitation id="A0E32A4CA80DFF8AFE8EF92D9FB3F8AC" box="[334,465,1786,1809]" captionStart="Figures 66–67" captionStartId="18.[192,278,909,933]" captionTargetBox="[204,1433,173,864]" captionTargetId="figure-11@18.[192,1439,168,880]" captionTargetPageId="18" captionText="Figures 66–67: Left chelicera of male Charmus laneus from locality 15CO. Figures 66–67. Dorsal (66) and ventral (67a) view (white light + UV), and ventroexternal (67b) aspect (UV) showing 2 denticles on ventral surface of fixed finger (d1, d2). Scale bar: 200 µm." figureDoi="http://doi.org/10.5281/zenodo.7124507" httpUri="https://zenodo.org/record/7124507/files/figure.png" pageId="26" pageNumber="25">Figs. 66–67</figureCitation>
). Manus with dark reticulated patterns on dorsal and ventral surfaces. Dorsal manus with 9 macrosetae near anterior margin. Movable finger with 2 dorsal macrosetae on anterior half. Ventrointernal aspect of manus and fixed finger with dense brush of microsetae, most if not all appearing fluorescent with intense terminal pinpoint fluorescence (i.e. putative chemoreceptive setae). Microsetae also present but sparse on ventral aspect of movable finger. Fingers with typical buthid dentition (
<bibRefCitation id="5C494B38A80DFF8AFB9AF9EF9A9FF9F2" author="Vachon" box="[1114,1277,1592,1615]" firstAuthor="Vachon" journalOrPublisher="Bulletin du Museum national d'histoire naturelle (Paris)" pageId="26" pageNumber="25" pagination="161 - 66" part="2" refId="ref45003" refString="VACHON, M. 1963. De l'utilite, en systematique, d'une nomenclature des dents des cheliceres chez les scorpions. Bulletin du Museum national d'histoire naturelle (Paris), ser. 2, 35: 161 - 66." title="De l'utilite, en systematique, d'une nomenclature des dents des cheliceres chez les scorpions" type="journal article" year="1963">Vachon, 1963</bibRefCitation>
). Fixed finger with distal and subdistal denticles, and 2 basal denticles fused into bicusp. Dorsal margin of movable finger with 5 denticles: 1 large distal and 1 large medial, 1 smaller subdistal, and 2 small partially fused basal denticles. Ventral margin of movable finger with 3 denticles: 1 large distal and 2 smaller medial denticles. Ventral surface of fixed finger armed medially and basally with 2 small denticles concealed by dense brush of microsetae (
<figureCitation id="A0E32A4CA80DFF8AFC42F8859D82F8D4" box="[898,992,1874,1897]" captionStart="Figures 66–67" captionStartId="18.[192,278,909,933]" captionTargetBox="[204,1433,173,864]" captionTargetId="figure-11@18.[192,1439,168,880]" captionTargetPageId="18" captionText="Figures 66–67: Left chelicera of male Charmus laneus from locality 15CO. Figures 66–67. Dorsal (66) and ventral (67a) view (white light + UV), and ventroexternal (67b) aspect (UV) showing 2 denticles on ventral surface of fixed finger (d1, d2). Scale bar: 200 µm." figureDoi="http://doi.org/10.5281/zenodo.7124507" httpUri="https://zenodo.org/record/7124507/files/figure.png" pageId="26" pageNumber="25">Fig. 67b</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="6CA76641A80CFF8BFF00F9129B32F8FD" ID-DOI="http://doi.org/10.5281/zenodo.7124543" ID-Zenodo-Dep="7124543" httpUri="https://zenodo.org/record/7124543/files/figure.png" pageId="27" pageNumber="26" startId="27.[192,278,1733,1757]" targetBox="[211,1428,190,1691]" targetPageId="27">
<paragraph id="386736C9A80CFF8BFF00F9129B32F8FD" blockId="27.[192,1441,1733,1856]" pageId="27" pageNumber="26">
<emphasis id="0AACEADBA80CFF8BFF00F9129FE4F960" bold="true" box="[192,390,1733,1757]" pageId="27" pageNumber="26">Figures 107–119:</emphasis>
<taxonomicName id="FFD84D4AA80CFF8BFE50F9109F8AF961" authorityName="Karsch" authorityYear="1879" box="[400,488,1735,1756]" class="Arachnida" family="Buthidae" genus="Charmus" kingdom="Animalia" order="Scorpiones" pageId="27" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="0AACEADBA80CFF8BFE50F9109F8AF961" box="[400,488,1735,1756]" italics="true" pageId="27" pageNumber="26">Charmus</emphasis>
</taxonomicName>
.
<emphasis id="0AACEADBA80CFF8BFE37F9109CC1F961" bold="true" box="[503,675,1735,1757]" pageId="27" pageNumber="26">Figures 107–114</emphasis>
.
<taxonomicName id="FFD84D4AA80CFF8BFD72F9109D0BF961" authority="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š, 2016" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" box="[690,873,1735,1756]" class="Arachnida" family="Buthidae" genus="Charmus" kingdom="Animalia" order="Scorpiones" pageId="27" pageNumber="26" phylum="Arthropoda" rank="species" species="saradieli" status="sp. nov.">
<emphasis id="0AACEADBA80CFF8BFD72F9109D0BF961" box="[690,873,1735,1756]" italics="true" pageId="27" pageNumber="26">Charmus saradieli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="119F57A0A80CFF8BFCB2F9109DC9F961" box="[882,939,1735,1756]" pageId="27" pageNumber="26" rank="species">
<emphasis id="0AACEADBA80CFF8BFCB2F9109DEBF961" bold="true" box="[882,905,1735,1756]" pageId="27" pageNumber="26">sp</emphasis>
.
<emphasis id="0AACEADBA80CFF8BFC58F9109DC7F961" bold="true" box="[920,933,1735,1756]" pageId="27" pageNumber="26">n</emphasis>
.
</taxonomicNameLabel>
, female holotype. Pedipalp movable finger (107). Pedipalp chela, dorsal (108), external (109), and ventral (110) views. Pedipalp patella, dorsal (111) and external (112) views. Pedipalp femur and trochanter, internal (113) and dorsal (114) views. The trichobothrial pattern is indicated in Figures 1 08a– 1 14a.
<emphasis id="0AACEADBA80CFF8BFF3DF8C69FC4F89B" bold="true" box="[253,422,1809,1831]" pageId="27" pageNumber="26">Figures 115–117</emphasis>
.
<taxonomicName id="FFD84D4AA80CFF8BFE74F8C59C0BF89A" authority="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š, 2016" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" box="[436,617,1810,1831]" class="Arachnida" family="Buthidae" genus="Charmus" kingdom="Animalia" order="Scorpiones" pageId="27" pageNumber="26" phylum="Arthropoda" rank="species" species="saradieli" status="sp. nov.">
<emphasis id="0AACEADBA80CFF8BFE74F8C59C0BF89A" box="[436,617,1810,1831]" italics="true" pageId="27" pageNumber="26">Charmus saradieli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="119F57A0A80CFF8BFDB3F8C59CCAF89A" box="[627,680,1810,1831]" pageId="27" pageNumber="26" rank="species">
<emphasis id="0AACEADBA80CFF8BFDB3F8C59CE8F89A" bold="true" box="[627,650,1810,1831]" pageId="27" pageNumber="26">sp</emphasis>
.
<emphasis id="0AACEADBA80CFF8BFD56F8C59CC1F89A" bold="true" box="[662,675,1810,1831]" pageId="27" pageNumber="26">n</emphasis>
.
</taxonomicNameLabel>
, male paratype (MHNG), chelicerae (115), pedipalp dorsal (116), and fourth leg dorsal (117).
<emphasis id="0AACEADBA80CFF8BFEA4F8FD9C6EF882" bold="true" box="[356,524,1834,1856]" pageId="27" pageNumber="26">Figures 118–119</emphasis>
.
<taxonomicName id="FFD84D4AA80CFF8BFDD7F8FC9CD6F8FD" authorityName="Karsch" authorityYear="1879" box="[535,692,1835,1856]" class="Arachnida" family="Buthidae" genus="Charmus" kingdom="Animalia" order="Scorpiones" pageId="27" pageNumber="26" phylum="Arthropoda" rank="species" species="laneus">
<emphasis id="0AACEADBA80CFF8BFDD7F8FC9CD6F8FD" box="[535,692,1835,1856]" italics="true" pageId="27" pageNumber="26">Charmus laneus</emphasis>
</taxonomicName>
from locality 15CO. Pedipalp dorsal of male (118) and female (119).
</paragraph>
</caption>
<paragraph id="386736C9A80BFF8CFE01FF1D9B19FC89" pageId="28" pageNumber="27">
<table id="4AD8C469A80B006FFF01FF1D9BF9FC88" box="[193,1435,202,821]" gridcols="6" gridrows="20" pageId="28" pageNumber="27">
<tr id="86E8348BA80B006FFF01FF1D9BF9FF5E" box="[193,1435,202,227]" gridrow="0" pageId="28" pageNumber="27" rowspan-0="1">
<th id="C5395DF7A80B006FFE4EFF1D9CB2FF5E" box="[398,720,202,227]" colspan="2" colspanRight="1" gridcol="1" gridrow="0" pageId="28" pageNumber="27">
<taxonomicName id="FFD84D4AA80BFF8CFE01FF1D9C15FF5F" authorityName="Karsch" authorityYear="1879" box="[449,631,202,226]" class="Arachnida" family="Buthidae" genus="Charmus" kingdom="Animalia" order="Scorpiones" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="laneus">
<emphasis id="0AACEADBA80BFF8CFE01FF1D9C15FF5F" bold="true" box="[449,631,202,226]" italics="true" pageId="28" pageNumber="27">Charmus laneus</emphasis>
</taxonomicName>
</th>
<th id="C5395DF7A80B006FFD2AFF1D9DA9FF5E" box="[746,971,202,227]" gridcol="3" gridrow="0" pageId="28" pageNumber="27">
<emphasis id="0AACEADBA80BFF8CFC04FF1C9DA9FF5F" bold="true" box="[964,971,203,226]" italics="true" pageId="28" pageNumber="27">.</emphasis>
</th>
<th id="C5395DF7A80B006FFC1AFF1D9BF9FF5E" box="[986,1435,202,227]" colspan="2" colspanRight="1" gridcol="4" gridrow="0" pageId="28" pageNumber="27">
<emphasis id="0AACEADBA80BFF8CFBFBFF1D9B2BFF5E" bold="true" box="[1083,1353,202,227]" pageId="28" pageNumber="27">
<taxonomicName id="FFD84D4AA80BFF8CFBFBFF1D9B65FF5F" authority="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š, 2016" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" box="[1083,1287,202,226]" class="Arachnida" family="Buthidae" genus="Charmus" kingdom="Animalia" order="Scorpiones" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="saradieli" status="sp. nov.">
<emphasis id="0AACEADBA80BFF8CFBFBFF1D9B65FF5F" bold="true" box="[1083,1287,202,226]" italics="true" pageId="28" pageNumber="27">Charmus saradieli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="119F57A0A80BFF8CFACDFF1C9B2BFF5E" box="[1293,1353,203,227]" pageId="28" pageNumber="27" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</th>
</tr>
<tr id="86E8348BA80B006FFF01FF3E9BF9FEBC" box="[193,1435,233,257]" gridrow="1" pageId="28" pageNumber="27">
<th id="C5395DF7A80B006FFF01FF3E9F8DFEBC" box="[193,495,233,257]" colspan="2" colspanRight="1" gridcol="0" gridrow="1" pageId="28" pageNumber="27">
<emphasis id="0AACEADBA80BFF8CFF39FF3E9FA1FEBC" bold="true" box="[249,451,233,257]" pageId="28" pageNumber="27">Dimensions (mm)</emphasis>
</th>
<td id="C5395DF7A80B006FFE3EFF3E9CB2FEBC" box="[510,720,233,257]" gridcol="2" gridrow="1" pageId="28" pageNumber="27">
♀
<emphasis id="0AACEADBA80BFF8CFD9AFF3E9CDEFEBC" bold="true" box="[602,700,233,257]" pageId="28" pageNumber="27">holotype</emphasis>
</td>
<td id="C5395DF7A80B006FFD2AFF3E9DA9FEBC" box="[746,971,233,257]" gridcol="3" gridrow="1" pageId="28" pageNumber="27">
♀
<emphasis id="0AACEADBA80BFF8CFC99FF3E9DFEFEBC" bold="true" box="[857,924,233,257]" pageId="28" pageNumber="27">15CO</emphasis>
</td>
<td id="C5395DF7A80B006FFC1AFF3E9ACFFEBC" box="[986,1197,233,257]" gridcol="4" gridrow="1" pageId="28" pageNumber="27">
♀
<emphasis id="0AACEADBA80BFF8CFBF9FF3E9AF9FEBC" bold="true" box="[1081,1179,233,257]" pageId="28" pageNumber="27">holotype</emphasis>
</td>
<td id="C5395DF7A80B006FFB08FF3E9BF9FEBC" box="[1224,1435,233,257]" gridcol="5" gridrow="1" pageId="28" pageNumber="27">
♂
<emphasis id="0AACEADBA80BFF8CFADFFF3E9BE7FEBC" bold="true" box="[1311,1413,233,257]" pageId="28" pageNumber="27">paratype</emphasis>
</td>
</tr>
<tr id="86E8348BA80B006FFF01FEDB9BF9FE9F" box="[193,1435,268,290]" gridrow="2" pageId="28" pageNumber="27">
<th id="C5395DF7A80B006FFF01FEDB9F1AFE9F" box="[193,376,268,290]" gridcol="0" gridrow="2" pageId="28" pageNumber="27">Carapace</th>
<td id="C5395DF7A80B006FFE4EFEDB9F8DFE9F" box="[398,495,268,290]" gridcol="1" gridrow="2" pageId="28" pageNumber="27">L / W</td>
<td id="C5395DF7A80B006FFE3EFEDB9CB2FE9F" box="[510,720,268,290]" gridcol="2" gridrow="2" pageId="28" pageNumber="27">2.125 / 2.425</td>
<td id="C5395DF7A80B006FFD2AFEDB9DA9FE9F" box="[746,971,268,290]" gridcol="3" gridrow="2" pageId="28" pageNumber="27">2.250 / 2.900</td>
<td id="C5395DF7A80B006FFC1AFEDB9ACFFE9F" box="[986,1197,268,290]" gridcol="4" gridrow="2" pageId="28" pageNumber="27">2.625 / 2.900</td>
<td id="C5395DF7A80B006FFB08FEDB9BF9FE9F" box="[1224,1435,268,290]" gridcol="5" gridrow="2" pageId="28" pageNumber="27">1.450 / 1.575</td>
</tr>
<tr id="86E8348BA80B006FFF01FEFE9BF9FE82" box="[193,1435,297,319]" gridrow="3" pageId="28" pageNumber="27">
<th id="C5395DF7A80B006FFF01FEFE9F1AFE82" box="[193,376,297,319]" gridcol="0" gridrow="3" pageId="28" pageNumber="27">Mesosoma</th>
<td id="C5395DF7A80B006FFE4EFEFE9F8DFE82" box="[398,495,297,319]" gridcol="1" gridrow="3" pageId="28" pageNumber="27">L</td>
<td id="C5395DF7A80B006FFE3EFEFE9CB2FE82" box="[510,720,297,319]" gridcol="2" gridrow="3" pageId="28" pageNumber="27">6.100</td>
<td id="C5395DF7A80B006FFD2AFEFE9DA9FE82" box="[746,971,297,319]" gridcol="3" gridrow="3" pageId="28" pageNumber="27">6.900</td>
<td id="C5395DF7A80B006FFC1AFEFE9ACFFE82" box="[986,1197,297,319]" gridcol="4" gridrow="3" pageId="28" pageNumber="27">8.800</td>
<td id="C5395DF7A80B006FFB08FEFE9BF9FE82" box="[1224,1435,297,319]" gridcol="5" gridrow="3" pageId="28" pageNumber="27">3.800</td>
</tr>
<tr id="86E8348BA80B006FFF01FE919BF9FEE1" box="[193,1435,326,348]" gridrow="4" pageId="28" pageNumber="27">
<th id="C5395DF7A80B006FFF01FE919F1AFEE1" box="[193,376,326,348]" gridcol="0" gridrow="4" pageId="28" pageNumber="27">Tergite VII</th>
<td id="C5395DF7A80B006FFE4EFE919F8DFEE1" box="[398,495,326,348]" gridcol="1" gridrow="4" pageId="28" pageNumber="27">L / W</td>
<td id="C5395DF7A80B006FFE3EFE919CB2FEE1" box="[510,720,326,348]" gridcol="2" gridrow="4" pageId="28" pageNumber="27">1.250 / 2.050</td>
<td id="C5395DF7A80B006FFD2AFE919DA9FEE1" box="[746,971,326,348]" gridcol="3" gridrow="4" pageId="28" pageNumber="27">1.550 / 2.600</td>
<td id="C5395DF7A80B006FFC1AFE919ACFFEE1" box="[986,1197,326,348]" gridcol="4" gridrow="4" pageId="28" pageNumber="27">1.600 / 3.000</td>
<td id="C5395DF7A80B006FFB08FE919BF9FEE1" box="[1224,1435,326,348]" gridcol="5" gridrow="4" pageId="28" pageNumber="27">0.825 / 1.375</td>
</tr>
<tr id="86E8348BA80B006FFF01FEB39BF9FE2A" box="[193,1435,356,407]" gridrow="5" pageId="28" pageNumber="27">
<th id="C5395DF7A80B006FFF01FEB39F1AFE2A" box="[193,376,356,407]" gridcol="0" gridrow="5" pageId="28" pageNumber="27">Metasoma &amp; telson</th>
<td id="C5395DF7A80B006FFE4EFEB39F8DFE2A" box="[398,495,356,407]" gridcol="1" gridrow="5" pageId="28" pageNumber="27">L</td>
<td id="C5395DF7A80B006FFE3EFEB39CB2FE2A" box="[510,720,356,407]" gridcol="2" gridrow="5" pageId="28" pageNumber="27">10.275</td>
<td id="C5395DF7A80B006FFD2AFEB39DA9FE2A" box="[746,971,356,407]" gridcol="3" gridrow="5" pageId="28" pageNumber="27">10.900</td>
<td id="C5395DF7A80B006FFC1AFEB39ACFFE2A" box="[986,1197,356,407]" gridcol="4" gridrow="5" pageId="28" pageNumber="27">12.070</td>
<td id="C5395DF7A80B006FFB08FEB39BF9FE2A" box="[1224,1435,356,407]" gridcol="5" gridrow="5" pageId="28" pageNumber="27">6.925</td>
</tr>
<tr id="86E8348BA80B006FFF01FE499BF9FE09" box="[193,1435,414,436]" gridrow="6" pageId="28" pageNumber="27">
<th id="C5395DF7A80B006FFF01FE499F1AFE09" box="[193,376,414,436]" gridcol="0" gridrow="6" pageId="28" pageNumber="27">Segment I</th>
<td id="C5395DF7A80B006FFE4EFE499F8DFE09" box="[398,495,414,436]" gridcol="1" gridrow="6" pageId="28" pageNumber="27">L / W / D</td>
<td id="C5395DF7A80B006FFE3EFE499CB2FE09" box="[510,720,414,436]" gridcol="2" gridrow="6" pageId="28" pageNumber="27">1.175 / 1.475 / 1.260</td>
<td id="C5395DF7A80B006FFD2AFE499DA9FE09" box="[746,971,414,436]" gridcol="3" gridrow="6" pageId="28" pageNumber="27">1.150 / 1.625 / 1.425</td>
<td id="C5395DF7A80B006FFC1AFE499ACFFE09" box="[986,1197,414,436]" gridcol="4" gridrow="6" pageId="28" pageNumber="27">1.375 / 1.700 / 1.500</td>
<td id="C5395DF7A80B006FFB08FE499BF9FE09" box="[1224,1435,414,436]" gridcol="5" gridrow="6" pageId="28" pageNumber="27">0.800 / 0.900 / 0.825</td>
</tr>
<tr id="86E8348BA80B006FFF01FE6C9BF9FE6C" box="[193,1435,443,465]" gridrow="7" pageId="28" pageNumber="27">
<th id="C5395DF7A80B006FFF01FE6C9F1AFE6C" box="[193,376,443,465]" gridcol="0" gridrow="7" pageId="28" pageNumber="27">Segment II</th>
<td id="C5395DF7A80B006FFE4EFE6C9F8DFE6C" box="[398,495,443,465]" gridcol="1" gridrow="7" pageId="28" pageNumber="27">L / W / D</td>
<td id="C5395DF7A80B006FFE3EFE6C9CB2FE6C" box="[510,720,443,465]" gridcol="2" gridrow="7" pageId="28" pageNumber="27">1.350 / 1.450 / 1.375</td>
<td id="C5395DF7A80B006FFD2AFE6C9DA9FE6C" box="[746,971,443,465]" gridcol="3" gridrow="7" pageId="28" pageNumber="27">1.500 / 1.680 / 1.480</td>
<td id="C5395DF7A80B006FFC1AFE6C9ACFFE6C" box="[986,1197,443,465]" gridcol="4" gridrow="7" pageId="28" pageNumber="27">1.575 / 1.600 / 1.525</td>
<td id="C5395DF7A80B006FFB08FE6C9BF9FE6C" box="[1224,1435,443,465]" gridcol="5" gridrow="7" pageId="28" pageNumber="27">0.875 / 0.825 / 0.825</td>
</tr>
<tr id="86E8348BA80B006FFF01FE0F9BF9FE53" box="[193,1435,472,494]" gridrow="8" pageId="28" pageNumber="27">
<th id="C5395DF7A80B006FFF01FE0F9F1AFE53" box="[193,376,472,494]" gridcol="0" gridrow="8" pageId="28" pageNumber="27">Segment III</th>
<td id="C5395DF7A80B006FFE4EFE0F9F8DFE53" box="[398,495,472,494]" gridcol="1" gridrow="8" pageId="28" pageNumber="27">L / W / D</td>
<td id="C5395DF7A80B006FFE3EFE0F9CB2FE53" box="[510,720,472,494]" gridcol="2" gridrow="8" pageId="28" pageNumber="27">1.600 / 1.550 / 1.425</td>
<td id="C5395DF7A80B006FFD2AFE0F9DA9FE53" box="[746,971,472,494]" gridcol="3" gridrow="8" pageId="28" pageNumber="27">1.625 / 1.825 / 1.600</td>
<td id="C5395DF7A80B006FFC1AFE0F9ACFFE53" box="[986,1197,472,494]" gridcol="4" gridrow="8" pageId="28" pageNumber="27">1.600 / 1.650 / 1.610</td>
<td id="C5395DF7A80B006FFB08FE0F9BF9FE53" box="[1224,1435,472,494]" gridcol="5" gridrow="8" pageId="28" pageNumber="27">0.925 / 0.850 / 0.825</td>
</tr>
<tr id="86E8348BA80B006FFF01FE229BF9FDB6" box="[193,1435,501,523]" gridrow="9" pageId="28" pageNumber="27">
<th id="C5395DF7A80B006FFF01FE229F1AFDB6" box="[193,376,501,523]" gridcol="0" gridrow="9" pageId="28" pageNumber="27">Segment IV</th>
<td id="C5395DF7A80B006FFE4EFE229F8DFDB6" box="[398,495,501,523]" gridcol="1" gridrow="9" pageId="28" pageNumber="27">L / W / D</td>
<td id="C5395DF7A80B006FFE3EFE229CB2FDB6" box="[510,720,501,523]" gridcol="2" gridrow="9" pageId="28" pageNumber="27">1.750 / 1.600 / 1.425</td>
<td id="C5395DF7A80B006FFD2AFE229DA9FDB6" box="[746,971,501,523]" gridcol="3" gridrow="9" pageId="28" pageNumber="27">1.875 / 1.875 / 1.675</td>
<td id="C5395DF7A80B006FFC1AFE229ACFFDB6" box="[986,1197,501,523]" gridcol="4" gridrow="9" pageId="28" pageNumber="27">2.100 / 1.580 / 1.630</td>
<td id="C5395DF7A80B006FFB08FE229BF9FDB6" box="[1224,1435,501,523]" gridcol="5" gridrow="9" pageId="28" pageNumber="27">1.250 / 0.825 / 0.775</td>
</tr>
<tr id="86E8348BA80B006FFF01FDC59BF9FD95" box="[193,1435,530,552]" gridrow="10" pageId="28" pageNumber="27">
<th id="C5395DF7A80B006FFF01FDC59F1AFD95" box="[193,376,530,552]" gridcol="0" gridrow="10" pageId="28" pageNumber="27">Segment V</th>
<td id="C5395DF7A80B006FFE4EFDC59F8DFD95" box="[398,495,530,552]" gridcol="1" gridrow="10" pageId="28" pageNumber="27">L / W / D</td>
<td id="C5395DF7A80B006FFE3EFDC59CB2FD95" box="[510,720,530,552]" gridcol="2" gridrow="10" pageId="28" pageNumber="27">2.300 / 1.625 / 1.375</td>
<td id="C5395DF7A80B006FFD2AFDC59DA9FD95" box="[746,971,530,552]" gridcol="3" gridrow="10" pageId="28" pageNumber="27">2.480 / 1.925 / 1.575</td>
<td id="C5395DF7A80B006FFC1AFDC59ACFFD95" box="[986,1197,530,552]" gridcol="4" gridrow="10" pageId="28" pageNumber="27">2.900 / 1.610 / 1.600</td>
<td id="C5395DF7A80B006FFB08FDC59BF9FD95" box="[1224,1435,530,552]" gridcol="5" gridrow="10" pageId="28" pageNumber="27">1.675 / 0.800 / 0.738</td>
</tr>
<tr id="86E8348BA80B006FFF01FDF89BF9FDF8" box="[193,1435,559,581]" gridrow="11" pageId="28" pageNumber="27">
<th id="C5395DF7A80B006FFF01FDF89F1AFDF8" box="[193,376,559,581]" gridcol="0" gridrow="11" pageId="28" pageNumber="27">Telson</th>
<td id="C5395DF7A80B006FFE4EFDF89F8DFDF8" box="[398,495,559,581]" gridcol="1" gridrow="11" pageId="28" pageNumber="27">L / W / D</td>
<td id="C5395DF7A80B006FFE3EFDF89CB2FDF8" box="[510,720,559,581]" gridcol="2" gridrow="11" pageId="28" pageNumber="27">2.100 / 0.910 / 0.850</td>
<td id="C5395DF7A80B006FFD2AFDF89DA9FDF8" box="[746,971,559,581]" gridcol="3" gridrow="11" pageId="28" pageNumber="27">2.275 / 1.125 / 0.950</td>
<td id="C5395DF7A80B006FFC1AFDF89ACFFDF8" box="[986,1197,559,581]" gridcol="4" gridrow="11" pageId="28" pageNumber="27">2.520 / 1.020 / 1.000</td>
<td id="C5395DF7A80B006FFB08FDF89BF9FDF8" box="[1224,1435,559,581]" gridcol="5" gridrow="11" pageId="28" pageNumber="27">1.400 / 0.525 / 0.475</td>
</tr>
<tr id="86E8348BA80B006FFF01FD9A9BF9FDDE" box="[193,1435,589,611]" gridrow="12" pageId="28" pageNumber="27">
<th id="C5395DF7A80B006FFF01FD9A9F1AFDDE" box="[193,376,589,611]" gridcol="0" gridrow="12" pageId="28" pageNumber="27">Pedipalp</th>
<td id="C5395DF7A80B006FFE4EFD9A9F8DFDDE" box="[398,495,589,611]" gridcol="1" gridrow="12" pageId="28" pageNumber="27">L</td>
<td id="C5395DF7A80B006FFE3EFD9A9CB2FDDE" box="[510,720,589,611]" gridcol="2" gridrow="12" pageId="28" pageNumber="27">7.050</td>
<td id="C5395DF7A80B006FFD2AFD9A9DA9FDDE" box="[746,971,589,611]" gridcol="3" gridrow="12" pageId="28" pageNumber="27">7.520</td>
<td id="C5395DF7A80B006FFC1AFD9A9ACFFDDE" box="[986,1197,589,611]" gridcol="4" gridrow="12" pageId="28" pageNumber="27">9.200</td>
<td id="C5395DF7A80B006FFB08FD9A9BF9FDDE" box="[1224,1435,589,611]" gridcol="5" gridrow="12" pageId="28" pageNumber="27">5.200</td>
</tr>
<tr id="86E8348BA80B006FFF01FDBD9BF9FD3D" box="[193,1435,618,640]" gridrow="13" pageId="28" pageNumber="27">
<th id="C5395DF7A80B006FFF01FDBD9F1AFD3D" box="[193,376,618,640]" gridcol="0" gridrow="13" pageId="28" pageNumber="27">Femur</th>
<td id="C5395DF7A80B006FFE4EFDBD9F8DFD3D" box="[398,495,618,640]" gridcol="1" gridrow="13" pageId="28" pageNumber="27">L / W</td>
<td id="C5395DF7A80B006FFE3EFDBD9CB2FD3D" box="[510,720,618,640]" gridcol="2" gridrow="13" pageId="28" pageNumber="27">1.800 / 0.530</td>
<td id="C5395DF7A80B006FFD2AFDBD9DA9FD3D" box="[746,971,618,640]" gridcol="3" gridrow="13" pageId="28" pageNumber="27">1.900 / 0.630</td>
<td id="C5395DF7A80B006FFC1AFDBD9ACFFD3D" box="[986,1197,618,640]" gridcol="4" gridrow="13" pageId="28" pageNumber="27">2.050 / 0.700</td>
<td id="C5395DF7A80B006FFB08FDBD9BF9FD3D" box="[1224,1435,618,640]" gridcol="5" gridrow="13" pageId="28" pageNumber="27">1.275 / 0.425</td>
</tr>
<tr id="86E8348BA80B006FFF01FD509BF9FD20" box="[193,1435,647,669]" gridrow="14" pageId="28" pageNumber="27">
<th id="C5395DF7A80B006FFF01FD509F1AFD20" box="[193,376,647,669]" gridcol="0" gridrow="14" pageId="28" pageNumber="27">Patela</th>
<td id="C5395DF7A80B006FFE4EFD509F8DFD20" box="[398,495,647,669]" gridcol="1" gridrow="14" pageId="28" pageNumber="27">L / W</td>
<td id="C5395DF7A80B006FFE3EFD509CB2FD20" box="[510,720,647,669]" gridcol="2" gridrow="14" pageId="28" pageNumber="27">2.150 / 0.750</td>
<td id="C5395DF7A80B006FFD2AFD509DA9FD20" box="[746,971,647,669]" gridcol="3" gridrow="14" pageId="28" pageNumber="27">2.325 / 0.900</td>
<td id="C5395DF7A80B006FFC1AFD509ACFFD20" box="[986,1197,647,669]" gridcol="4" gridrow="14" pageId="28" pageNumber="27">2.900 / 0.980</td>
<td id="C5395DF7A80B006FFB08FD509BF9FD20" box="[1224,1435,647,669]" gridcol="5" gridrow="14" pageId="28" pageNumber="27">1.625 / 0.525</td>
</tr>
<tr id="86E8348BA80B006FFF01FD739BF9FD07" box="[193,1435,676,698]" gridrow="15" pageId="28" pageNumber="27">
<th id="C5395DF7A80B006FFF01FD739F1AFD07" box="[193,376,676,698]" gridcol="0" gridrow="15" pageId="28" pageNumber="27">Chela</th>
<td id="C5395DF7A80B006FFE4EFD739F8DFD07" box="[398,495,676,698]" gridcol="1" gridrow="15" pageId="28" pageNumber="27">L</td>
<td id="C5395DF7A80B006FFE3EFD739CB2FD07" box="[510,720,676,698]" gridcol="2" gridrow="15" pageId="28" pageNumber="27">3.000</td>
<td id="C5395DF7A80B006FFD2AFD739DA9FD07" box="[746,971,676,698]" gridcol="3" gridrow="15" pageId="28" pageNumber="27">3.300</td>
<td id="C5395DF7A80B006FFC1AFD739ACFFD07" box="[986,1197,676,698]" gridcol="4" gridrow="15" pageId="28" pageNumber="27">4.250</td>
<td id="C5395DF7A80B006FFB08FD739BF9FD07" box="[1224,1435,676,698]" gridcol="5" gridrow="15" pageId="28" pageNumber="27">2.300</td>
</tr>
<tr id="86E8348BA80B006FFF01FD169BF9FD6A" box="[193,1435,705,727]" gridrow="16" pageId="28" pageNumber="27">
<th id="C5395DF7A80B006FFF01FD169F1AFD6A" box="[193,376,705,727]" gridcol="0" gridrow="16" pageId="28" pageNumber="27">Manus</th>
<td id="C5395DF7A80B006FFE4EFD169F8DFD6A" box="[398,495,705,727]" gridcol="1" gridrow="16" pageId="28" pageNumber="27">L / W / D</td>
<td id="C5395DF7A80B006FFE3EFD169CB2FD6A" box="[510,720,705,727]" gridcol="2" gridrow="16" pageId="28" pageNumber="27">1.050 / 0.600 / 0.575</td>
<td id="C5395DF7A80B006FFD2AFD169DA9FD6A" box="[746,971,705,727]" gridcol="3" gridrow="16" pageId="28" pageNumber="27">1.025 / 0.725 / 0.710</td>
<td id="C5395DF7A80B006FFC1AFD169ACFFD6A" box="[986,1197,705,727]" gridcol="4" gridrow="16" pageId="28" pageNumber="27">1.300 / 0.800 / 0.790</td>
<td id="C5395DF7A80B006FFB08FD169BF9FD6A" box="[1224,1435,705,727]" gridcol="5" gridrow="16" pageId="28" pageNumber="27">0.625 / 0.475 / 0.475</td>
</tr>
<tr id="86E8348BA80B006FFF01FD099BF9FD49" box="[193,1435,734,756]" gridrow="17" pageId="28" pageNumber="27">
<th id="C5395DF7A80B006FFF01FD099F1AFD49" box="[193,376,734,756]" gridcol="0" gridrow="17" pageId="28" pageNumber="27">Fixed finger</th>
<td id="C5395DF7A80B006FFE4EFD099F8DFD49" box="[398,495,734,756]" gridcol="1" gridrow="17" pageId="28" pageNumber="27">L</td>
<td id="C5395DF7A80B006FFE3EFD099CB2FD49" box="[510,720,734,756]" gridcol="2" gridrow="17" pageId="28" pageNumber="27">1.675</td>
<td id="C5395DF7A80B006FFD2AFD099DA9FD49" box="[746,971,734,756]" gridcol="3" gridrow="17" pageId="28" pageNumber="27">1.950</td>
<td id="C5395DF7A80B006FFC1AFD099ACFFD49" box="[986,1197,734,756]" gridcol="4" gridrow="17" pageId="28" pageNumber="27">2.925</td>
<td id="C5395DF7A80B006FFB08FD099BF9FD49" box="[1224,1435,734,756]" gridcol="5" gridrow="17" pageId="28" pageNumber="27">1.400</td>
</tr>
<tr id="86E8348BA80B006FFF01FD2C9BF9FCAC" box="[193,1435,763,785]" gridrow="18" pageId="28" pageNumber="27">
<th id="C5395DF7A80B006FFF01FD2C9F1AFCAC" box="[193,376,763,785]" gridcol="0" gridrow="18" pageId="28" pageNumber="27">Movable finger</th>
<td id="C5395DF7A80B006FFE4EFD2C9F8DFCAC" box="[398,495,763,785]" gridcol="1" gridrow="18" pageId="28" pageNumber="27">L</td>
<td id="C5395DF7A80B006FFE3EFD2C9CB2FCAC" box="[510,720,763,785]" gridcol="2" gridrow="18" pageId="28" pageNumber="27">2.050</td>
<td id="C5395DF7A80B006FFD2AFD2C9DA9FCAC" box="[746,971,763,785]" gridcol="3" gridrow="18" pageId="28" pageNumber="27">2.275</td>
<td id="C5395DF7A80B006FFC1AFD2C9ACFFCAC" box="[986,1197,763,785]" gridcol="4" gridrow="18" pageId="28" pageNumber="27">2.950</td>
<td id="C5395DF7A80B006FFB08FD2C9BF9FCAC" box="[1224,1435,763,785]" gridcol="5" gridrow="18" pageId="28" pageNumber="27">1.675</td>
</tr>
<tr id="86E8348BA80B006FFF01FCCB9BF9FC88" box="[193,1435,796,821]" gridrow="19" pageId="28" pageNumber="27" rowspan-1="1">
<th id="C5395DF7A80B006FFF01FCCB9F1AFC88" box="[193,376,796,821]" gridcol="0" gridrow="19" pageId="28" pageNumber="27">Total length</th>
<td id="C5395DF7A80B006FFE3EFCCB9CB2FC88" box="[510,720,796,821]" gridcol="2" gridrow="19" pageId="28" pageNumber="27">18.50</td>
<td id="C5395DF7A80B006FFD2AFCCB9DA9FC88" box="[746,971,796,821]" gridcol="3" gridrow="19" pageId="28" pageNumber="27">20.10</td>
<td id="C5395DF7A80B006FFC1AFCCB9ACFFC88" box="[986,1197,796,821]" gridcol="4" gridrow="19" pageId="28" pageNumber="27">23.50</td>
<td id="C5395DF7A80B006FFB08FCCB9BF9FC88" box="[1224,1435,796,821]" gridcol="5" gridrow="19" pageId="28" pageNumber="27">12.18</td>
</tr>
</table>
</paragraph>
<caption id="6CA76641A80BFF8CFF00FC829C80FC3B" ID-Table-UUID="6CA76641A80BFF8CFF00FC829C80FC3B" httpUri="http://table.plazi.org/id/6CA76641A80BFF8CFF00FC829C80FC3B" pageId="28" pageNumber="27" startId="28.[192,257,853,877]" targetBox="[193,1435,202,821]" targetIsTable="true" targetPageId="28">
<paragraph id="386736C9A80BFF8CFF00FC829C80FC3B" blockId="28.[192,1440,853,902]" pageId="28" pageNumber="27">
<emphasis id="0AACEADBA80BFF8CFF00FC829F42FCD0" bold="true" box="[192,288,853,877]" pageId="28" pageNumber="27">Table 1:</emphasis>
Comparative measurements of adults of
<taxonomicName id="FFD84D4AA80BFF8CFD71FC8F9D2DFCD0" authorityName="Karsch" authorityYear="1879" box="[689,847,856,877]" class="Arachnida" family="Buthidae" genus="Charmus" kingdom="Animalia" order="Scorpiones" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="laneus">
<emphasis id="0AACEADBA80BFF8CFD71FC8F9D2DFCD0" box="[689,847,856,877]" italics="true" pageId="28" pageNumber="27">Charmus laneus</emphasis>
</taxonomicName>
and
<taxonomicName id="FFD84D4AA80BFF8CFC40FC8F9D90FCD0" authority="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š, 2016" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" box="[896,1010,856,877]" class="Arachnida" family="Buthidae" genus="Charmus" kingdom="Animalia" order="Scorpiones" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="saradieli" status="sp. nov.">
<emphasis id="0AACEADBA80BFF8CFC40FC8F9D90FCD0" box="[896,1010,856,877]" italics="true" pageId="28" pageNumber="27">C. saradieli</emphasis>
</taxonomicName>
<emphasis id="0AACEADBA80BFF8CFC3AFC809A53FCD1" bold="true" box="[1018,1073,855,876]" pageId="28" pageNumber="27">
<taxonomicNameLabel id="119F57A0A80BFF8CFC3AFC809A53FCD1" box="[1018,1073,855,876]" pageId="28" pageNumber="27" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D).
</paragraph>
</caption>
<paragraph id="386736C9A80BFF8CFF33FC139A84FAF8" pageId="28" pageNumber="27">
<table id="4AD8C469A80B006FFF33FC629B4AFAF8" box="[243,1320,949,1349]" gridcols="3" gridrows="12" pageId="28" pageNumber="27">
<tr id="86E8348BA80B006FFF33FC629B4AFC51" box="[243,1320,949,1004]" gridrow="0" pageId="28" pageNumber="27">
<th id="C5395DF7A80B006FFF33FC629CFAFC51" box="[243,664,949,1004]" gridcol="0" gridrow="0" pageId="28" pageNumber="27">
<emphasis id="0AACEADBA80BFF8CFF33FC139F95FC61" bold="true" box="[243,503,964,988]" pageId="28" pageNumber="27">Ratios of adult females</emphasis>
</th>
<th id="C5395DF7A80B006FFCEFFC629DD7FC51" box="[815,949,949,1004]" gridcol="1" gridrow="0" pageId="28" pageNumber="27">
<emphasis id="0AACEADBA80BFF8CFCFEFC629DF5FC56" bold="true" pageId="28" pageNumber="27">
<taxonomicName id="FFD84D4AA80BFF8CFCFEFC629DC5FC70" authorityName="Karsch" authorityYear="1879" box="[830,935,949,973]" class="Arachnida" family="Buthidae" genus="Charmus" kingdom="Animalia" order="Scorpiones" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="laneus">
<emphasis id="0AACEADBA80BFF8CFCFEFC629DC5FC70" bold="true" box="[830,935,949,973]" italics="true" pageId="28" pageNumber="27">C. laneus</emphasis>
</taxonomicName>
(n = 4)
</emphasis>
</th>
<th id="C5395DF7A80B006FFBA8FC629B4AFC51" box="[1128,1320,949,1004]" gridcol="2" gridrow="0" pageId="28" pageNumber="27">
<emphasis id="0AACEADBA80BFF8CFBA8FC629A8CFC56" bold="true" pageId="28" pageNumber="27">
<taxonomicName id="FFD84D4AA80BFF8CFBA8FC629A84FC70" authority="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š, 2016" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" box="[1128,1254,949,973]" class="Arachnida" family="Buthidae" genus="Charmus" kingdom="Animalia" order="Scorpiones" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="saradieli" status="sp. nov.">
<emphasis id="0AACEADBA80BFF8CFBA8FC629A84FC70" bold="true" box="[1128,1254,949,973]" italics="true" pageId="28" pageNumber="27">C. saradieli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="119F57A0A80BFF8CFB2CFC629B4AFC70" box="[1260,1320,949,973]" pageId="28" pageNumber="27" rank="species">sp. n.</taxonomicNameLabel>
(n = 1)
</emphasis>
</th>
</tr>
<tr id="86E8348BA80B006FFF33FC209B4AFBB3" box="[243,1320,1015,1038]" gridrow="1" pageId="28" pageNumber="27">
<th id="C5395DF7A80B006FFF33FC209CFAFBB3" box="[243,664,1015,1038]" gridcol="0" gridrow="1" pageId="28" pageNumber="27">Metasomal segment I (L/W)</th>
<td id="C5395DF7A80B006FFCEFFC209DD7FBB3" box="[815,949,1015,1038]" gridcol="1" gridrow="1" pageId="28" pageNumber="27">0.707–0.796</td>
<td id="C5395DF7A80B006FFBA8FC209B4AFBB3" box="[1128,1320,1015,1038]" gridcol="2" gridrow="1" pageId="28" pageNumber="27">0.808</td>
</tr>
<tr id="86E8348BA80B006FFF33FBC29B4AFB91" box="[243,1320,1045,1068]" gridrow="2" pageId="28" pageNumber="27">
<th id="C5395DF7A80B006FFF33FBC29CFAFB91" box="[243,664,1045,1068]" gridcol="0" gridrow="2" pageId="28" pageNumber="27">Metasomal segment II (L/W)</th>
<td id="C5395DF7A80B006FFCEFFBC29DD7FB91" box="[815,949,1045,1068]" gridcol="1" gridrow="2" pageId="28" pageNumber="27">0.892–0.931</td>
<td id="C5395DF7A80B006FFBA8FBC29B4AFB91" box="[1128,1320,1045,1068]" gridcol="2" gridrow="2" pageId="28" pageNumber="27">0.984</td>
</tr>
<tr id="86E8348BA80B006FFF33FBE39B4AFBF6" box="[243,1320,1076,1099]" gridrow="3" pageId="28" pageNumber="27">
<th id="C5395DF7A80B006FFF33FBE39CFAFBF6" box="[243,664,1076,1099]" gridcol="0" gridrow="3" pageId="28" pageNumber="27">Metasomal segment IV (L/W)</th>
<td id="C5395DF7A80B006FFCEFFBE39DD7FBF6" box="[815,949,1076,1099]" gridcol="1" gridrow="3" pageId="28" pageNumber="27">1.000–1.093</td>
<td id="C5395DF7A80B006FFBA8FBE39B4AFBF6" box="[1128,1320,1076,1099]" gridcol="2" gridrow="3" pageId="28" pageNumber="27">1.329</td>
</tr>
<tr id="86E8348BA80B006FFF33FB849B4AFBD7" box="[243,1320,1107,1130]" gridrow="4" pageId="28" pageNumber="27">
<th id="C5395DF7A80B006FFF33FB849CFAFBD7" box="[243,664,1107,1130]" gridcol="0" gridrow="4" pageId="28" pageNumber="27">Metasomal segment IV (L/D)</th>
<td id="C5395DF7A80B006FFCEFFB849DD7FBD7" box="[815,949,1107,1130]" gridcol="1" gridrow="4" pageId="28" pageNumber="27">1.119–1.228</td>
<td id="C5395DF7A80B006FFBA8FB849B4AFBD7" box="[1128,1320,1107,1130]" gridcol="2" gridrow="4" pageId="28" pageNumber="27">1.288</td>
</tr>
<tr id="86E8348BA80B006FFF33FBA69B4AFB35" box="[243,1320,1137,1160]" gridrow="5" pageId="28" pageNumber="27">
<th id="C5395DF7A80B006FFF33FBA69CFAFB35" box="[243,664,1137,1160]" gridcol="0" gridrow="5" pageId="28" pageNumber="27">Metasomal segment V (L/W)</th>
<td id="C5395DF7A80B006FFCEFFBA69DD7FB35" box="[815,949,1137,1160]" gridcol="1" gridrow="5" pageId="28" pageNumber="27">1.288–1.425</td>
<td id="C5395DF7A80B006FFBA8FBA69B4AFB35" box="[1128,1320,1137,1160]" gridcol="2" gridrow="5" pageId="28" pageNumber="27">1.801</td>
</tr>
<tr id="86E8348BA80B006FFF33FB479B4AFB1A" box="[243,1320,1168,1191]" gridrow="6" pageId="28" pageNumber="27">
<th id="C5395DF7A80B006FFF33FB479CFAFB1A" box="[243,664,1168,1191]" gridcol="0" gridrow="6" pageId="28" pageNumber="27">Metasomal segment V (L/D)</th>
<td id="C5395DF7A80B006FFCEFFB479DD7FB1A" box="[815,949,1168,1191]" gridcol="1" gridrow="6" pageId="28" pageNumber="27">1.574–1.672</td>
<td id="C5395DF7A80B006FFBA8FB479B4AFB1A" box="[1128,1320,1168,1191]" gridcol="2" gridrow="6" pageId="28" pageNumber="27">1.812</td>
</tr>
<tr id="86E8348BA80B006FFF33FB789B4AFB7B" box="[243,1320,1199,1222]" gridrow="7" pageId="28" pageNumber="27">
<th id="C5395DF7A80B006FFF33FB789CFAFB7B" box="[243,664,1199,1222]" gridcol="0" gridrow="7" pageId="28" pageNumber="27">Telson (L/D)</th>
<td id="C5395DF7A80B006FFCEFFB789DD7FB7B" box="[815,949,1199,1222]" gridcol="1" gridrow="7" pageId="28" pageNumber="27">2.394–2.470</td>
<td id="C5395DF7A80B006FFBA8FB789B4AFB7B" box="[1128,1320,1199,1222]" gridcol="2" gridrow="7" pageId="28" pageNumber="27">2.520</td>
</tr>
<tr id="86E8348BA80B006FFF33FB1A9B4AFB59" box="[243,1320,1229,1252]" gridrow="8" pageId="28" pageNumber="27">
<th id="C5395DF7A80B006FFF33FB1A9CFAFB59" box="[243,664,1229,1252]" gridcol="0" gridrow="8" pageId="28" pageNumber="27">Pedipalp chela (L/W)</th>
<td id="C5395DF7A80B006FFCEFFB1A9DD7FB59" box="[815,949,1229,1252]" gridcol="1" gridrow="8" pageId="28" pageNumber="27">4.551–5.000</td>
<td id="C5395DF7A80B006FFBA8FB1A9B4AFB59" box="[1128,1320,1229,1252]" gridcol="2" gridrow="8" pageId="28" pageNumber="27">5.312</td>
</tr>
<tr id="86E8348BA80B006FFF33FB3B9B4AFABE" box="[243,1320,1260,1283]" gridrow="9" pageId="28" pageNumber="27">
<th id="C5395DF7A80B006FFF33FB3B9CFAFABE" box="[243,664,1260,1283]" gridcol="0" gridrow="9" pageId="28" pageNumber="27">Pedipalp chela (L) / fixed finger (L)</th>
<td id="C5395DF7A80B006FFCEFFB3B9DD7FABE" box="[815,949,1260,1283]" gridcol="1" gridrow="9" pageId="28" pageNumber="27">1.692–1.791</td>
<td id="C5395DF7A80B006FFBA8FB3B9B4AFABE" box="[1128,1320,1260,1283]" gridcol="2" gridrow="9" pageId="28" pageNumber="27">1.452</td>
</tr>
<tr id="86E8348BA80B006FFF33FADC9B4AFA9F" box="[243,1320,1291,1314]" gridrow="10" pageId="28" pageNumber="27">
<th id="C5395DF7A80B006FFF33FADC9CFAFA9F" box="[243,664,1291,1314]" gridcol="0" gridrow="10" pageId="28" pageNumber="27">Pedipalp chela (L) / movable finger (L)</th>
<td id="C5395DF7A80B006FFCEFFADC9DD7FA9F" box="[815,949,1291,1314]" gridcol="1" gridrow="10" pageId="28" pageNumber="27">1.450–1.512</td>
<td id="C5395DF7A80B006FFBA8FADC9B4AFA9F" box="[1128,1320,1291,1314]" gridcol="2" gridrow="10" pageId="28" pageNumber="27">1.440</td>
</tr>
<tr id="86E8348BA80B006FFF33FAFA9B4AFAF8" box="[243,1320,1325,1349]" gridrow="11" pageId="28" pageNumber="27">
<th id="C5395DF7A80B006FFF33FAFA9CFAFAF8" box="[243,664,1325,1349]" gridcol="0" gridrow="11" pageId="28" pageNumber="27">Total (L)</th>
<td id="C5395DF7A80B006FFCEFFAFA9DD7FAF8" box="[815,949,1325,1349]" gridcol="1" gridrow="11" pageId="28" pageNumber="27">18.50–21.30</td>
<td id="C5395DF7A80B006FFBA8FAFA9B4AFAF8" box="[1128,1320,1325,1349]" gridcol="2" gridrow="11" pageId="28" pageNumber="27">23.50</td>
</tr>
</table>
</paragraph>
<caption id="6CA76641A80BFF8CFF00FAB19C88FA2B" ID-Table-UUID="6CA76641A80BFF8CFF00FAB19C88FA2B" httpUri="http://table.plazi.org/id/6CA76641A80BFF8CFF00FAB19C88FA2B" pageId="28" pageNumber="27" startId="28.[192,257,1382,1406]" targetBox="[243,1320,949,1349]" targetIsTable="true" targetPageId="28">
<paragraph id="386736C9A80BFF8CFF00FAB19C88FA2B" blockId="28.[192,1440,1382,1430]" pageId="28" pageNumber="27">
<emphasis id="0AACEADBA80BFF8CFF00FAB19F41FAC3" bold="true" box="[192,291,1382,1406]" pageId="28" pageNumber="27">Table 2:</emphasis>
Comparison among Sri Lankan
<taxonomicName id="FFD84D4AA80BFF8CFDB3FABE9D79FAC3" box="[627,795,1385,1406]" class="Arachnida" family="Buthidae" genus="Charmus" kingdom="Animalia" order="Scorpiones" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="0AACEADBA80BFF8CFDB3FABE9CA9FAC3" box="[627,715,1385,1406]" italics="true" pageId="28" pageNumber="27">Charmus</emphasis>
species
</taxonomicName>
(specimens), based upon selected morphometric ratios of adult females. Abbreviations: length (L), width (W), depth (D).
</paragraph>
</caption>
<subSubSection id="70C26542A80BFF8DFF00FA039F2DFE6E" lastPageId="29" lastPageNumber="28" pageId="28" pageNumber="27" type="discussion">
<paragraph id="386736C9A80BFF8DFF00FA039F2DFE6E" blockId="28.[192,803,1492,1914]" lastBlockId="29.[192,802,199,467]" lastPageId="29" lastPageNumber="28" pageId="28" pageNumber="27">
COMMENTS. Lourenço did not study any specimens of
<taxonomicName id="FFD84D4AA80BFF8CFCC9FA039F64F9B7" authorityName="Karsch" authorityYear="1879" class="Arachnida" family="Buthidae" genus="Charmus" kingdom="Animalia" order="Scorpiones" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="laneus">
<emphasis id="0AACEADBA80BFF8CFCC9FA039F64F9B7" italics="true" pageId="28" pageNumber="27">C. laneus</emphasis>
</taxonomicName>
in spite of the fact that he cited six characters for distinguishing
<taxonomicName id="FFD84D4AA80BFF8CFEA9F9C59FBBF994" authorityName="Karsch" authorityYear="1879" box="[361,473,1554,1577]" class="Arachnida" family="Buthidae" genus="Charmus" kingdom="Animalia" order="Scorpiones" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="laneus">
<emphasis id="0AACEADBA80BFF8CFEA9F9C59FBBF994" box="[361,473,1554,1577]" italics="true" pageId="28" pageNumber="27">C. laneus</emphasis>
</taxonomicName>
from
<taxonomicName id="FFD84D4AA80BFF8CFDF0F9C59F50F9FA" authority="(Lourenco, 2002: 23)" baseAuthorityName="Lourenco" baseAuthorityPageNumber="23" baseAuthorityYear="2002" class="Arachnida" family="Buthidae" genus="Charmus" kingdom="Animalia" order="Scorpiones" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="0AACEADBA80BFF8CFDF0F9C59CF8F994" box="[560,666,1554,1577]" italics="true" pageId="28" pageNumber="27">C. minor</emphasis>
(
<bibRefCitation id="5C494B38A80BFF8CFD73F9C59F4BF9FA" author="Lourenco" firstAuthor="Lourenco" journalOrPublisher="Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg" page="23" pageId="28" pageNumber="27" pagination="17 - 25" part="14" refId="ref42746" refString="LOURENCO, W. R. 2002. Further taxonomic considerations about the genus Charmus Karsch, 1879 (Scorpiones, Buthidae), with the description of a new species from Sri Lanka. Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg, 14 (165): 17 - 25" title="Further taxonomic considerations about the genus Charmus Karsch, 1879 (Scorpiones, Buthidae), with the description of a new species from Sri Lanka" type="journal article" year="2002">Lourenço, 2002: 23</bibRefCitation>
)
</taxonomicName>
. Several of these 'character differences' lie within the range of intraspecific variation (e.g. fulcra of pectines), and others are not valid. Lourenço stated that the movable finger of
<taxonomicName id="FFD84D4AA80BFF8CFE0EF95B9C5EF91E" authorityName="Karsch" authorityYear="1879" box="[462,572,1676,1699]" class="Arachnida" family="Buthidae" genus="Charmus" kingdom="Animalia" order="Scorpiones" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="laneus">
<emphasis id="0AACEADBA80BFF8CFE0EF95B9C5EF91E" box="[462,572,1676,1699]" italics="true" pageId="28" pageNumber="27">C. laneus</emphasis>
</taxonomicName>
bears 7–8 rows of granules, while that of
<taxonomicName id="FFD84D4AA80BFF8CFE21F97C9C29F97F" baseAuthorityName="Lourenco" baseAuthorityYear="2002" box="[481,587,1707,1730]" class="Arachnida" family="Buthidae" genus="Charmus" kingdom="Animalia" order="Scorpiones" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="0AACEADBA80BFF8CFE21F97C9C29F97F" box="[481,587,1707,1730]" italics="true" pageId="28" pageNumber="27">C. minor</emphasis>
</taxonomicName>
bears 9 rows of granules. However, his own figure (
<bibRefCitation id="5C494B38A80BFF8CFD86F91D9D7EF95C" author="Lourenco" box="[582,796,1738,1761]" firstAuthor="Lourenco" journalOrPublisher="Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg" page="18" pageId="28" pageNumber="27" pagination="17 - 25" part="14" refId="ref42746" refString="LOURENCO, W. R. 2002. Further taxonomic considerations about the genus Charmus Karsch, 1879 (Scorpiones, Buthidae), with the description of a new species from Sri Lanka. Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg, 14 (165): 17 - 25" title="Further taxonomic considerations about the genus Charmus Karsch, 1879 (Scorpiones, Buthidae), with the description of a new species from Sri Lanka" type="journal article" year="2002">Lourenço, 2002: 18</bibRefCitation>
, fig. 1) shows that the male &quot;
<typeStatus id="E763886BA80BFF8CFE3EF93F9C3DF942" box="[510,607,1768,1791]" pageId="28" pageNumber="27" type="paratype">paratype</typeStatus>
&quot; of
<taxonomicName id="FFD84D4AA80BFF8CFD52F93F9C96F942" baseAuthorityName="Lourenco" baseAuthorityYear="2002" box="[658,756,1768,1791]" class="Arachnida" family="Buthidae" genus="Charmus" kingdom="Animalia" order="Scorpiones" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="0AACEADBA80BFF8CFD52F93F9C96F942" box="[658,756,1768,1791]" italics="true" pageId="28" pageNumber="27">C. minor</emphasis>
</taxonomicName>
has only 8 rows of granules. Another problem is that according to the type material section (
<bibRefCitation id="5C494B38A80BFF8CFDADF8F19EB9F8E6" author="Lourenco" firstAuthor="Lourenco" journalOrPublisher="Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg" page="19" pageId="28" pageNumber="27" pagination="17 - 25" part="14" refId="ref42746" refString="LOURENCO, W. R. 2002. Further taxonomic considerations about the genus Charmus Karsch, 1879 (Scorpiones, Buthidae), with the description of a new species from Sri Lanka. Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg, 14 (165): 17 - 25" title="Further taxonomic considerations about the genus Charmus Karsch, 1879 (Scorpiones, Buthidae), with the description of a new species from Sri Lanka" type="journal article" year="2002">Lourenço, 2002: 19</bibRefCitation>
) there exist only
<specimenCount id="2EDEFD40A80BFF8CFE66F8939C72F8E6" box="[422,528,1860,1883]" pageId="28" pageNumber="27" type="generic">two types</specimenCount>
of
<taxonomicName id="FFD84D4AA80BFF8CFDFDF8939CC2F8E6" baseAuthorityName="Lourenco" baseAuthorityYear="2002" box="[573,672,1860,1883]" class="Arachnida" family="Buthidae" genus="Charmus" kingdom="Animalia" order="Scorpiones" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="0AACEADBA80BFF8CFDFDF8939CC2F8E6" box="[573,672,1860,1883]" italics="true" pageId="28" pageNumber="27">C. minor</emphasis>
</taxonomicName>
– the male
<typeStatus id="E763886BA80BFF8CFF00F8B49F7FF8C7" box="[192,285,1891,1914]" pageId="28" pageNumber="27" type="holotype">holotype</typeStatus>
and a juvenile (second instar) female
<typeStatus id="E763886BA80BFF8CFD7FF8B49D7CF8C7" box="[703,798,1891,1914]" pageId="28" pageNumber="27" type="paratype">paratype</typeStatus>
. Neither a male
<typeStatus id="E763886BA80BFF8CFC31FA039A32FA56" box="[1009,1104,1492,1515]" pageId="28" pageNumber="27" type="paratype">paratype</typeStatus>
, nor an adult female
<typeStatus id="E763886BA80BFF8CFA86FA039BC0FA56" box="[1350,1442,1492,1515]" pageId="28" pageNumber="27" type="paratype">paratype</typeStatus>
were listed. Thus, it is surprising that Lourenço claimed that the female of
<taxonomicName id="FFD84D4AA80BFF8CFBD3F9C59A14F994" baseAuthorityName="Lourenco" baseAuthorityYear="2002" box="[1043,1142,1554,1577]" class="Arachnida" family="Buthidae" genus="Charmus" kingdom="Animalia" order="Scorpiones" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="0AACEADBA80BFF8CFBD3F9C59A14F994" box="[1043,1142,1554,1577]" italics="true" pageId="28" pageNumber="27">C. minor</emphasis>
</taxonomicName>
has a differently flattened sternum than the male (
<bibRefCitation id="5C494B38A80BFF8CFBFFF9E79B7BF9FA" author="Lourenco" box="[1087,1305,1584,1607]" firstAuthor="Lourenco" journalOrPublisher="Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg" page="23" pageId="28" pageNumber="27" pagination="17 - 25" part="14" refId="ref42746" refString="LOURENCO, W. R. 2002. Further taxonomic considerations about the genus Charmus Karsch, 1879 (Scorpiones, Buthidae), with the description of a new species from Sri Lanka. Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg, 14 (165): 17 - 25" title="Further taxonomic considerations about the genus Charmus Karsch, 1879 (Scorpiones, Buthidae), with the description of a new species from Sri Lanka" type="journal article" year="2002">Lourenço, 2002: 23</bibRefCitation>
). If this was a reference to the second instar juvenile female
<typeStatus id="E763886BA80BFF8CFAFDF9989BF9F9DB" box="[1341,1435,1615,1638]" pageId="28" pageNumber="27" type="paratype">paratype</typeStatus>
, the diagnostic character needs to specified for adult females. Under &quot;Ecological observations&quot;
<bibRefCitation id="5C494B38A80BFF8CFAFAF95B9DD5F97F" author="Lourenco" firstAuthor="Lourenco" journalOrPublisher="Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg" page="23" pageId="28" pageNumber="27" pagination="17 - 25" part="14" refId="ref42746" refString="LOURENCO, W. R. 2002. Further taxonomic considerations about the genus Charmus Karsch, 1879 (Scorpiones, Buthidae), with the description of a new species from Sri Lanka. Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg, 14 (165): 17 - 25" title="Further taxonomic considerations about the genus Charmus Karsch, 1879 (Scorpiones, Buthidae), with the description of a new species from Sri Lanka" type="journal article" year="2002">Lourenço (2002: 23)</bibRefCitation>
wrote that &quot;The specimens of
<taxonomicName id="FFD84D4AA80BFF8CFAF3F97C9BC2F97F" authorityName="Karsch" authorityYear="1879" box="[1331,1440,1707,1730]" class="Arachnida" family="Buthidae" genus="Charmus" kingdom="Animalia" order="Scorpiones" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="laneus">
<emphasis id="0AACEADBA80BFF8CFAF3F97C9BC2F97F" box="[1331,1440,1707,1730]" italics="true" pageId="28" pageNumber="27">C. laneus</emphasis>
</taxonomicName>
studied by
<bibRefCitation id="5C494B38A80BFF8CFC05F91D9A0DF95C" author="Vachon" box="[965,1135,1738,1761]" firstAuthor="Vachon" journalOrPublisher="Revue suisse de Zoologie" pageId="28" pageNumber="27" pagination="77 - 114" part="89" refId="ref45190" refString="VACHON, M. 1982. Les scorpions de Sri Lanka (Recherches sur les scorpions appartenant ou deposes au Museum d`Histoire naturelle de Geneve III.). Revue suisse de Zoologie, 89: 77 - 114." title="Les scorpions de Sri Lanka (Recherches sur les scorpions appartenant ou deposes au Museum d`Histoire naturelle de Geneve III.)" type="journal article" year="1982">Vachon (1982)</bibRefCitation>
were all collected in the central-south region of
<collectingCountry id="40CF7659A80BFF8CFBFDF93F9ACAF942" box="[1085,1192,1768,1791]" name="Sri Lanka" pageId="28" pageNumber="27">Sri Lanka</collectingCountry>
which is characterised by high altitudes reaching to more than
<quantity id="FF209B2CA80BFF8CFAC4F8D09B3CF8A3" box="[1284,1374,1799,1822]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="28" pageNumber="27" unit="m" value="1000.0">1000 m</quantity>
...&quot;. In fact, the
<specimenCount id="2EDEFD40A80BFF8CFC60F8F19A2CF880" box="[928,1102,1830,1853]" pageId="28" pageNumber="27" type="generic">three specimens</specimenCount>
studied by
<bibRefCitation id="5C494B38A80BFF8CFB11F8F19BC2F880" author="Vachon" box="[1233,1440,1830,1853]" firstAuthor="Vachon" journalOrPublisher="Revue suisse de Zoologie" page="81" pageId="28" pageNumber="27" pagination="77 - 114" part="89" refId="ref45190" refString="VACHON, M. 1982. Les scorpions de Sri Lanka (Recherches sur les scorpions appartenant ou deposes au Museum d`Histoire naturelle de Geneve III.). Revue suisse de Zoologie, 89: 77 - 114." title="Les scorpions de Sri Lanka (Recherches sur les scorpions appartenant ou deposes au Museum d`Histoire naturelle de Geneve III.)" type="journal article" year="1982">Vachon (1982: 81)</bibRefCitation>
were collected at altitudes of
<quantity id="FF209B2CA80BFF8CFBBBF8939AA0F8E6" box="[1147,1218,1860,1883]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="28" pageNumber="27" unit="m" value="250.0">250 m</quantity>
,
<quantity id="FF209B2CA80BFF8CFB0EF8939B76F8E6" box="[1230,1300,1860,1883]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="28" pageNumber="27" unit="m" value="600.0">600 m</quantity>
, and
<quantity id="FF209B2CA80BFF8CFA8EF8939BC2F8E6" box="[1358,1440,1860,1883]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="28" pageNumber="27" unit="m" value="1350.0">1350 m</quantity>
a.s.l. and according to Vachon’s opinion they all belong to the same species, '
<taxonomicName id="FFD84D4AA80AFF8DFE69FF109C73FF63" authorityName="Karsch" authorityYear="1879" box="[425,529,199,222]" class="Arachnida" family="Buthidae" genus="Charmus" kingdom="Animalia" order="Scorpiones" pageId="29" pageNumber="28" phylum="Arthropoda" rank="species" species="laneus">
<emphasis id="0AACEADBA80AFF8DFE69FF109C73FF63" box="[425,529,199,222]" italics="true" pageId="29" pageNumber="28">C. laneus</emphasis>
</taxonomicName>
'. Neither
<bibRefCitation id="5C494B38A80AFF8DFDBEFF109D40FF63" author="Vachon" box="[638,802,199,222]" firstAuthor="Vachon" journalOrPublisher="Revue suisse de Zoologie" pageId="29" pageNumber="28" pagination="77 - 114" part="89" refId="ref45190" refString="VACHON, M. 1982. Les scorpions de Sri Lanka (Recherches sur les scorpions appartenant ou deposes au Museum d`Histoire naturelle de Geneve III.). Revue suisse de Zoologie, 89: 77 - 114." title="Les scorpions de Sri Lanka (Recherches sur les scorpions appartenant ou deposes au Museum d`Histoire naturelle de Geneve III.)" type="journal article" year="1982">Vachon (1982)</bibRefCitation>
nor
<bibRefCitation id="5C494B38A80AFF8DFF2DFF319FC6FF40" author="Lourenco" box="[237,420,230,253]" firstAuthor="Lourenco" journalOrPublisher="Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg" pageId="29" pageNumber="28" pagination="17 - 25" part="14" refId="ref42746" refString="LOURENCO, W. R. 2002. Further taxonomic considerations about the genus Charmus Karsch, 1879 (Scorpiones, Buthidae), with the description of a new species from Sri Lanka. Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg, 14 (165): 17 - 25" title="Further taxonomic considerations about the genus Charmus Karsch, 1879 (Scorpiones, Buthidae), with the description of a new species from Sri Lanka" type="journal article" year="2002">Lourenço (2002)</bibRefCitation>
studied the
<typeStatus id="E763886BA80AFF8DFDEDFF319CE8FF40" box="[557,650,230,253]" pageId="29" pageNumber="28" type="holotype">holotype</typeStatus>
of
<taxonomicName id="FFD84D4AA80AFF8DFD74FF319D78FF40" authorityName="Karsch" authorityYear="1879" box="[692,794,230,253]" class="Arachnida" family="Buthidae" genus="Charmus" kingdom="Animalia" order="Scorpiones" pageId="29" pageNumber="28" phylum="Arthropoda" rank="species" species="laneus">
<emphasis id="0AACEADBA80AFF8DFD74FF319D78FF40" box="[692,794,230,253]" italics="true" pageId="29" pageNumber="28">C. laneus</emphasis>
</taxonomicName>
, and as we show here, these three
<taxonomicName id="FFD84D4AA80AFF8DFD85FED39CC5FEA6" authorityName="Karsch" authorityYear="1879" box="[581,679,260,283]" class="Arachnida" family="Buthidae" genus="Charmus" kingdom="Animalia" order="Scorpiones" pageId="29" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="0AACEADBA80AFF8DFD85FED39CC5FEA6" box="[581,679,260,283]" italics="true" pageId="29" pageNumber="28">Charmus</emphasis>
</taxonomicName>
specimens from central
<collectingCountry id="40CF7659A80AFF8DFE97FEF49FA4FE87" box="[343,454,291,314]" name="Sri Lanka" pageId="29" pageNumber="28">Sri Lanka</collectingCountry>
actually belong to a different species. These
<specimenCount id="2EDEFD40A80AFF8DFEA5FE959C70FEE4" box="[357,530,322,345]" pageId="29" pageNumber="28" type="generic">three specimens</specimenCount>
are herein designated as types of
<taxonomicName id="FFD84D4AA80AFF8DFEE8FEB79FCFFECA" authority="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š, 2016" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" box="[296,429,352,375]" class="Arachnida" family="Buthidae" genus="Charmus" kingdom="Animalia" order="Scorpiones" pageId="29" pageNumber="28" phylum="Arthropoda" rank="species" species="saradieli" status="sp. nov.">
<emphasis id="0AACEADBA80AFF8DFEE8FEB79FCFFECA" box="[296,429,352,375]" italics="true" pageId="29" pageNumber="28">C. saradieli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="119F57A0A80AFF8DFE7AFEB79F99FECA" box="[442,507,352,376]" pageId="29" pageNumber="28" rank="species">
<emphasis id="0AACEADBA80AFF8DFE7AFEB79FB1FEC5" bold="true" box="[442,467,352,376]" pageId="29" pageNumber="28">sp</emphasis>
.
<emphasis id="0AACEADBA80AFF8DFE26FEB79F97FEC5" bold="true" box="[486,501,352,376]" pageId="29" pageNumber="28">n</emphasis>
.
</taxonomicNameLabel>
, and
<taxonomicName id="FFD84D4AA80AFF8DFD83FEB79E97FE2B" authority="Lourenco, 2002" authorityName="Lourenco" authorityYear="2002" class="Arachnida" family="Buthidae" genus="Charmus" kingdom="Animalia" order="Scorpiones" pageId="29" pageNumber="28" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="0AACEADBA80AFF8DFD83FEB79CCAFECA" box="[579,680,352,375]" italics="true" pageId="29" pageNumber="28">C. minor</emphasis>
<bibRefCitation id="5C494B38A80AFF8DFD75FEB79E97FE2B" author="Lourenco" firstAuthor="Lourenco" journalOrPublisher="Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg" pageId="29" pageNumber="28" pagination="17 - 25" part="14" refId="ref42746" refString="LOURENCO, W. R. 2002. Further taxonomic considerations about the genus Charmus Karsch, 1879 (Scorpiones, Buthidae), with the description of a new species from Sri Lanka. Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg, 14 (165): 17 - 25" title="Further taxonomic considerations about the genus Charmus Karsch, 1879 (Scorpiones, Buthidae), with the description of a new species from Sri Lanka" type="journal article" year="2002">Lourenço, 2002</bibRefCitation>
</taxonomicName>
is synonymized with
<taxonomicName id="FFD84D4AA80AFF8DFE35FEA89C9DFE2B" authority="Karsch, 1879" authorityName="Karsch" authorityYear="1879" box="[501,767,383,406]" class="Arachnida" family="Buthidae" genus="Charmus" kingdom="Animalia" order="Scorpiones" pageId="29" pageNumber="28" phylum="Arthropoda" rank="species" species="laneus">
<emphasis id="0AACEADBA80AFF8DFE35FEA89C02FE2B" box="[501,608,383,406]" italics="true" pageId="29" pageNumber="28">C. laneus</emphasis>
<bibRefCitation id="5C494B38A80AFF8DFDACFEA89C9DFE2B" author="Karsch" box="[620,767,383,406]" firstAuthor="Karsch" pageId="29" pageNumber="28" pagination="97 - 136" refId="ref41417" refString="KARSCH, F. 1879. Skorpionologische Beitrage I. and II. Mitteilungen des Munchener Entomologischen Vereins, 3: 6 - 22, 97 - 136." type="journal article" year="1879">Karsch, 1879</bibRefCitation>
</taxonomicName>
as there are no significant differences between them at the species level.
</paragraph>
</subSubSection>
<subSubSection id="70C26542A80AFF8DFF00FE249FB4FDB7" box="[192,470,499,522]" pageId="29" pageNumber="28" type="distribution">
<paragraph id="386736C9A80AFF8DFF00FE249FB4FDB7" blockId="29.[192,470,499,522]" box="[192,470,499,522]" pageId="29" pageNumber="28">
DISTRIBUTION.
<collectingCountry id="40CF7659A80AFF8DFEA7FE249FADFDB7" box="[359,463,499,522]" name="Sri Lanka" pageId="29" pageNumber="28">Sri Lanka</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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