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<document id="99BAB7DA8235C9C6C23D9B8ECDDD4C40" ID-DOI="10.5252/z2009n3a12" ID-ISSN="1638-9387" ID-Zenodo-Dep="5398343" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630328017152" checkinUser="marcus" docAuthor="Desutter-Grandcolas, Laure" docDate="2009" docId="EB4887D3FFDA6739FD50FAE1FEC458B0" docLanguage="en" docName="Zoosystema.2009.31.3.619-659.pdf" docOrigin="Zoosystema 31 (3)" docSource="http://www.bioone.org/doi/abs/10.5252/z2009n3a12" docStyle="DocumentStyle:E638A3C99FA728B529662B46F84D75FD.3:Zoosystema.2009-2010.journal_article" docStyleId="E638A3C99FA728B529662B46F84D75FD" docStyleName="Zoosystema.2009-2010.journal_article" docStyleVersion="3" docTitle="Cophonemobius Chopard 1929" docType="treatment" docVersion="3" lastPageNumber="648" masterDocId="1771FFABFFC66724FFB0FFADFFAB5E14" masterDocTitle="New and little known crickets from Espiritu Santo Island, Vanuatu (Insecta, Orthoptera, Grylloidea, Pseudotrigonidium Chopard, 1915, Phaloriinae and Nemobiinae p. p.)" masterLastPageNumber="659" masterPageNumber="619" pageNumber="647" updateTime="1699229333769" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="A92133A0D6D076888A51867379154D27">New and little known crickets from Espiritu Santo Island, Vanuatu (Insecta, Orthoptera, Grylloidea, Pseudotrigonidium Chopard, 1915, Phaloriinae and Nemobiinae p. p.)</mods:title>
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<mods:name id="4541706446EBD32759F91AF6F2D48197" type="personal">
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<mods:namePart id="A1F9C4BA851B7628B9779ED330981510">Desutter-Grandcolas, Laure</mods:namePart>
<mods:affiliation id="4AFF75CBF6CCC7EE5B2E4DF5C300E830">Muséum national dHistoire naturelle, Département Systématique et Évolution, UMR 7205 CNRS, case postale 50, 57 rue Cuvier, F- 75231 Paris cedex 05 (France) desutter @ mnhn. fr</mods:affiliation>
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<mods:title id="CCE4BF631FC7CE877B2B521E8EE76B84">Zoosystema</mods:title>
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<mods:date id="A33AF97E53B5E5C0609443961A116843">2009</mods:date>
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<mods:number id="55D182164F7F5794B20E2FC389FE1B1C">2009-09-30</mods:number>
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<mods:number id="AFA46F19095DB6C6EAEDC417EAB31303">31</mods:number>
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<mods:classification id="430D9A4FACFF3D4CF7CEC8890FAD9AC5">journal article</mods:classification>
<mods:identifier id="756794E4A9AF017813FD126386216EF7" type="DOI">10.5252/z2009n3a12</mods:identifier>
<mods:identifier id="56E746739496E73C0D65D423F621BBB8" type="ISSN">1638-9387</mods:identifier>
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<treatment id="EB4887D3FFDA6739FD50FAE1FEC458B0" LSID="urn:lsid:plazi:treatment:EB4887D3FFDA6739FD50FAE1FEC458B0" httpUri="http://treatment.plazi.org/id/EB4887D3FFDA6739FD50FAE1FEC458B0" lastPageId="29" lastPageNumber="648" pageId="28" pageNumber="647">
<subSubSection id="2BFB654EFFDA6738FD50FAE1FBD55B73" box="[736,1150,1356,1383]" pageId="28" pageNumber="647" type="nomenclature">
<paragraph id="635E36C5FFDA6738FD50FAE1FBD55B73" blockId="28.[736,1150,1356,1383]" box="[736,1150,1356,1383]" pageId="28" pageNumber="647">
<heading id="381681A9FFDA6738FD50FAE1FBD55B73" box="[736,1150,1356,1383]" centered="true" fontSize="11" level="2" pageId="28" pageNumber="647" reason="2">
Genus
<taxonomicName id="A4E14D46FFDA6738FC9DFAE1FBD55B73" ID-CoL="8NPZD" authority="Chopard, 1929" authorityName="Chopard" authorityYear="1929" box="[813,1150,1356,1383]" class="Insecta" family="Trigonidiidae" genus="Cophonemobius" kingdom="Animalia" order="Orthoptera" pageId="28" pageNumber="647" phylum="Arthropoda" rank="genus">
<emphasis id="5195EAD7FFDA6738FC9DFAE1FC7A5B72" box="[813,977,1356,1382]" italics="true" pageId="28" pageNumber="647">Cophonemobius</emphasis>
Chopard, 1929
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="2BFB654EFFDA6739FD15FA2AFE805CA8" lastPageId="29" lastPageNumber="648" pageId="28" pageNumber="647" type="materials_examined">
<paragraph id="635E36C5FFDA6738FD15FA2AFB115B8B" blockId="28.[677,1210,1414,1439]" box="[677,1210,1414,1439]" pageId="28" pageNumber="647">
TYPE SPECIES. —
<taxonomicName id="A4E14D46FFDA6738FCF6FA2BFB1C5B8B" authority="Chopard, 1929" authorityName="Chopard" authorityYear="1929" box="[838,1207,1414,1439]" class="Insecta" family="Trigonidiidae" genus="Cophonemobius" kingdom="Animalia" order="Orthoptera" pageId="28" pageNumber="647" phylum="Arthropoda" rank="species" species="buxtoni">
<emphasis id="5195EAD7FFDA6738FCF6FA2BFBB45B8A" box="[838,1055,1414,1438]" italics="true" pageId="28" pageNumber="647">Cophonemobius buxtoni</emphasis>
Chopard, 1929
</taxonomicName>
.
</paragraph>
<paragraph id="635E36C5FFDA6738FD15FA15FBE7587A" blockId="28.[675,1211,1463,1699]" pageId="28" pageNumber="647">EMENDED DIAGNOSIS. — TI without tympana.TIII with 3 pairs of long subapical spurs; 3 outer apical spurs; 3 inner apical spurs, the dorsal 1.5 times as long as the median, and longer than half basitarsus III. FWs present in both males and females, hardened, very short, and only slightly overlapping; lateral and dorsal fields with longitudinal, parallel veins. Palpi very long.</paragraph>
<paragraph id="635E36C5FFDA6738FD0BF9DCFB1358B7" blockId="28.[675,1211,1463,1699]" pageId="28" pageNumber="647">
In Vanuatuan species, male genitalia wide and distinctly curved dorso-ventrally (
<figureCitation id="FBDA2A40FFDA6738FC34F926FC6058B7" box="[900,971,1675,1699]" captionStart="FIG" captionStartId="34.[113,124,1347,1364]" captionTargetBox="[114,1212,222,1299]" captionTargetId="figure-19@34.[96,1228,213,1306]" captionTargetPageId="34" captionText="FIG. 14. — Cophonemobius faustini n. sp., male genitalia in dorsal (A, C) and lateral (B) views: A, B, C. faustini faustini n. subsp.; C, C. faustini funafus n. subsp. Abbreviations: see text. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5398375" httpUri="https://zenodo.org/record/5398375/files/figure.png" pageId="28" pageNumber="647">Fig. 14</figureCitation>
). Pseudepiphallic sclerite
</paragraph>
<caption id="379E664DFFDB6739FFC1FF71FE9D5F10" pageId="29" pageNumber="648" startId="29.[113,124,220,237]">
<paragraph id="635E36C5FFDB6739FFC1FF71FE9D5F10" blockId="29.[113,1210,220,260]" pageId="29" pageNumber="648">
TABLE 11. — Measurements (in mm) of
<taxonomicName id="A4E14D46FFDB6739FE72FF71FDCC5EF9" authority="Desutter-Grandcolas, 2009" authorityName="Desutter-Grandcolas" authorityYear="2009" box="[450,615,220,237]" class="Insecta" family="Phalangopsidae" genus="Phaloria" kingdom="Animalia" order="Orthoptera" pageId="29" pageNumber="648" phylum="Arthropoda" rank="species" species="faponensis" status="sp. nov.">
<emphasis id="5195EAD7FFDB6739FE72FF71FDCC5EF9" box="[450,615,220,237]" italics="true" pageId="29" pageNumber="648">Phaloria faponensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4AA657ACFFDB6739FDDEFF71FD0B5EF9" box="[622,672,220,237]" pageId="29" pageNumber="648" rank="species">n. sp.</taxonomicNameLabel>
holotype. Abbreviations:
<emphasis id="5195EAD7FFDB6739FCCAFF71FC3A5EF9" bold="true" box="[890,913,220,237]" pageId="29" pageNumber="648">TL</emphasis>
, total length (body + wings); other abbreviations, see text.
</paragraph>
</caption>
<paragraph id="635E36C5FFDB6739FE98FE93FB085F46" blockId="29.[113,1187,318,373]" box="[296,1187,318,338]" pageId="29" pageNumber="648">
<emphasis id="5195EAD7FFDB6739FE98FE93FB085F46" bold="true" box="[296,1187,318,338]" pageId="29" pageNumber="648">TL Lpron lpron LFW lFW LW LFIII LTIII Lovip.</emphasis>
</paragraph>
<paragraph id="635E36C5FFDB6739FFC1FECFFB385F61" blockId="29.[113,1187,318,373]" box="[113,1171,354,373]" pageId="29" pageNumber="648">
<typeStatus id="BC5A8867FFDB6739FFC1FECFFF6C5F61" box="[113,199,354,373]" pageId="29" pageNumber="648" type="holotype">Holotype</typeStatus>
25.5 2.4 4.1 16.9 4.1 21.1 11.1 10.1 4.7
</paragraph>
<paragraph id="635E36C5FFDB6739FFC1FE1BFE805CA8" blockId="29.[111,649,438,700]" pageId="29" pageNumber="648">round and convex, its anterior margin deeply indented; 4 more sclerotized longitudinal areas, each bearing long and strong setae; pseudepiphallic “lophi”, parameres and ectophallic fold gathered under pseudepiphallus apex, with a 90° angle from pseudepiphallic sclerite. Pseudepiphallic parameres having the shape of vertical plates, hardly sclerotized. Female ovipositor with acute apex, slightly raised dorsally; dorsal valves with longitudinal carinae and inconspicuous lateral teeth. No distinct copulatory papilla.</paragraph>
</subSubSection>
<subSubSection id="2BFB654EFFDB6739FFC1FD79FF7D5D64" pageId="29" pageNumber="648" type="distribution">
<paragraph id="635E36C5FFDB6739FFC1FD79FF7D5D64" blockId="29.[112,647,723,880]" pageId="29" pageNumber="648">
DISTRIBUTION. — Until now
<taxonomicName id="A4E14D46FFDB6739FE38FD7EFDB25CFF" authorityName="Chopard" authorityYear="1929" box="[392,537,723,747]" class="Insecta" family="Trigonidiidae" genus="Cophonemobius" kingdom="Animalia" order="Orthoptera" pageId="29" pageNumber="648" phylum="Arthropoda" rank="genus">
<emphasis id="5195EAD7FFDB6739FE38FD7EFDB25CFF" box="[392,537,723,747]" italics="true" pageId="29" pageNumber="648">Cophonemobius</emphasis>
</taxonomicName>
was known from
<collectingCountry id="1BF67655FFDB6739FF15FD43FF4D5D12" box="[165,230,750,774]" name="Samoa" pageId="29" pageNumber="648">Samoa</collectingCountry>
islands only. Its presence is largely attested in
<collectingCountry id="1BF67655FFDB6739FF3DFCA5FF4F5D34" box="[141,228,776,800]" name="Vanuatu" pageId="29" pageNumber="648">Vanuatu</collectingCountry>
, where it is abundant in cave habitats. It is not present in
<collectingCountry id="1BF67655FFDB6739FE9EFC8EFE785D2F" box="[302,467,803,827]" name="New Caledonia" pageId="29" pageNumber="648">New Caledonia</collectingCountry>
(Chopard 1915;
<bibRefCitation id="07704B34FFDB6739FFC1FC90FEA55D41" author="GOROCHOV A. V." box="[113,270,829,853]" pageId="29" pageNumber="648" pagination="692 - 708" refId="ref21910" refString="GOROCHOV A. V. 1986. - New and little known crickets (Orthoptera, Gryllidae) from Australia and Oceania. Revue d'Entomologie de l'URSS 65: 692 - 708." type="journal article" year="1986">Gorochov 1986</bibRefCitation>
;
<bibRefCitation id="07704B34FFDB6739FEA7FC90FE195D41" author="OTTE D. &amp; ALEXANDER R. D. &amp; CADE W." box="[279,434,829,853]" pageId="29" pageNumber="648" pagination="375 - 457" refId="ref22573" refString="OTTE D., ALEXANDER R. D. &amp; CADE W. 1987. - The crickets of New Caledonia (Gryllidae). Proceedings of the Academy of Natural Science of Philadelphia 139: 375 - 457." type="journal article" year="1987">
Otte
<emphasis id="5195EAD7FFDB6739FEFAFC93FED35D41" box="[330,376,829,853]" italics="true" pageId="29" pageNumber="648">et al.</emphasis>
1987
</bibRefCitation>
; Desutter-Grandcolas 1997a, b).
</paragraph>
</subSubSection>
<subSubSection id="2BFB654EFFDB6739FFC1FC3CFEC458B0" pageId="29" pageNumber="648" type="discussion">
<paragraph id="635E36C5FFDB6739FFC1FC3CFF7E5DBD" blockId="29.[112,649,913,1700]" box="[113,213,913,939]" pageId="29" pageNumber="648">REMARKS</paragraph>
<paragraph id="635E36C5FFDB6739FFC1FC1DFE6C5B3C" blockId="29.[112,649,913,1700]" pageId="29" pageNumber="648">
<taxonomicName id="A4E14D46FFDB6739FFC1FC1DFEB65DDE" authorityName="Chopard" authorityYear="1929" box="[113,285,944,970]" class="Insecta" family="Trigonidiidae" genus="Cophonemobius" kingdom="Animalia" order="Orthoptera" pageId="29" pageNumber="648" phylum="Arthropoda" rank="genus">
<emphasis id="5195EAD7FFDB6739FFC1FC1DFEB65DDE" box="[113,285,944,970]" italics="true" pageId="29" pageNumber="648">Cophonemobius</emphasis>
</taxonomicName>
is widely distributed in Espiritu Santo. It occurs in many caves, often with great abundance and deep inside galleries, and could be one of the most characteristic animal of Vanuatuan karstic environment. It has never been observed outside caves, neither by day nor by night (pers. obs.), which would indicate that
<taxonomicName id="A4E14D46FFDB6739FF1CFB23FEC95ABC" authorityName="Chopard" authorityYear="1929" box="[172,354,1166,1192]" class="Insecta" family="Trigonidiidae" genus="Cophonemobius" kingdom="Animalia" order="Orthoptera" pageId="29" pageNumber="648" phylum="Arthropoda" rank="genus">
<emphasis id="5195EAD7FFDB6739FF1CFB23FEC95ABC" box="[172,354,1166,1192]" italics="true" pageId="29" pageNumber="648">Cophonemobius</emphasis>
</taxonomicName>
is truly troglobitic, i.e. caves may constitute its natural habitat, even though it does not present the morphological syndrome most often associated with cave life (Desutter-Grandcolas 1997c).
</paragraph>
<paragraph id="635E36C5FFDB6739FF38FA80FEC458B0" blockId="29.[112,649,913,1700]" pageId="29" pageNumber="648">
The specimens studied here form quite a homogeneous stock, which supports the description of a new species,
<taxonomicName id="A4E14D46FFDB6739FE88FAC1FD965B92" authority="Desutter-Grandcolas, 2009" authorityName="Desutter-Grandcolas" authorityYear="2009" box="[312,573,1388,1414]" class="Insecta" family="Trigonidiidae" genus="Cophonemobius" kingdom="Animalia" order="Orthoptera" pageId="29" pageNumber="648" phylum="Arthropoda" rank="species" species="faustini" status="sp. nov.">
<emphasis id="5195EAD7FFDB6739FE88FAC1FD965B92" box="[312,573,1388,1414]" italics="true" pageId="29" pageNumber="648">Cophonemobius faustini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4AA657ACFFDB6739FDF6FAC1FD2E5B93" box="[582,645,1388,1415]" pageId="29" pageNumber="648" rank="species">n. sp.</taxonomicNameLabel>
Observed variation in male genitalia and colouration is however suffi ciently stable and coherent to allow the definitions of at least two subspecies,
<taxonomicName id="A4E14D46FFDB6739FFC3FA46FEEF5811" authority="faustini" box="[115,324,1515,1541]" class="Insecta" family="Trigonidiidae" genus="Cophonemobius" kingdom="Animalia" order="Orthoptera" pageId="29" pageNumber="648" phylum="Arthropoda" rank="subSpecies" species="faustini" subSpecies="faustini">
<emphasis id="5195EAD7FFDB6739FFC3FA46FEEF5811" box="[115,324,1515,1541]" italics="true" pageId="29" pageNumber="648">C. faustini faustini</emphasis>
</taxonomicName>
and
<taxonomicName id="A4E14D46FFDB6739FE30FA46FDE55811" authorityName="Desutter-Grandcolas" authorityYear="2009" box="[384,590,1515,1541]" class="Insecta" family="Trigonidiidae" genus="Cophonemobius" kingdom="Animalia" order="Orthoptera" pageId="29" pageNumber="648" phylum="Arthropoda" rank="subSpecies" species="faustini" subSpecies="funafus">
<emphasis id="5195EAD7FFDB6739FE30FA46FDE55811" box="[384,590,1515,1541]" italics="true" pageId="29" pageNumber="648">C. faustini funafus</emphasis>
</taxonomicName>
. Additional juvenile specimens have been collected in different caves during SANTO 2006, including Amarirua on Malo island, which could not be identified at species level (material in alcohol, MNHN-ENSIF2151).
</paragraph>
</subSubSection>
</treatment>
</document>