plazi/treatments-xml
2
0
Fork 0
Code Issues Pull requests Releases Activity
treatments-xml/data/F0/6F/F3/F06FF311FF893A04FF7AFD61BD2DD295.xml
maintenance 5c7f71d8de added F0
2024-06-21 12:57:13 +02:00

734 lines
90 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="A487D66D56283AADDF63D3BB08503589" ID-CLB-Dataset="31084" ID-DOI="10.11646/zootaxa.4369.1.7" ID-GBIF-Dataset="e320619c-87b6-4f6d-a630-16fdaf828dfa" ID-ISSN="1175-5326" ID-Zenodo-Dep="1135628" ID-ZooBank="CD413E4C-0221-46AA-BD23-B733EFFAE6B6" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1515147780760" checkinUser="plazi" docAuthor="Kazemi, E., Niknam, G., Jabbari, H. &amp; Peña-Santiago, R." docDate="2018" docId="F06FF311FF893A04FF7AFD61BD2DD295" docLanguage="en" docName="zootaxa.4369.1.7.pdf" docOrigin="Zootaxa 4369 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Eudorylaimus kahaqensis Kazemi &amp; Niknam &amp; Jabbari &amp; Peña-Santiago 2018, sp. n." docType="treatment" docVersion="7" lastPageNumber="135" masterDocId="0C568B69FF883A0CFFEDFFEEBE6BD345" masterDocTitle="Eudorylaimus kahaqensis sp. n. (Nematoda: Dorylaimida: Qudsianematidae), an interesting new species from Iran" masterLastPageNumber="136" masterPageNumber="128" pageNumber="129" updateTime="1698512149061" updateUser="ExternalLinkService">
<mods:mods id="DB0B28ECE375F15A9B7FC383B9C8F042" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A4A9CE926805C74797DAD4D2A271E6D8">
<mods:title id="CFD794DBF1D2F140471A8E0681EF0043">Eudorylaimus kahaqensis sp. n. (Nematoda: Dorylaimida: Qudsianematidae), an interesting new species from Iran</mods:title>
</mods:titleInfo>
<mods:name id="3F0EDC2F2A266116A405AB38A87DE33C" type="personal">
<mods:role id="77AC486D21446D30D878245CF495D7F6">
<mods:roleTerm id="7AA105C62518153D7E8E347239AADAD0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9BFD9AB462E1BDF2280F452DFF9B1A01">Kazemi, E.</mods:namePart>
</mods:name>
<mods:name id="D1F16357759BDBF34568FF9CDE2C67C2" type="personal">
<mods:role id="9F3F3CF2C01B1A6BA4BA908709324259">
<mods:roleTerm id="90365A549719E5F388C18593DB3EF400">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3CB2B45CCCA1E8CAC6AC865F4DE4F497">Niknam, G.</mods:namePart>
</mods:name>
<mods:name id="E4B40BF823AEF1887122932857FC2790" type="personal">
<mods:role id="089BFD3A8FD4F1BA560860B21B74C88F">
<mods:roleTerm id="D1093574EA4C3E23FBCCDF72A58CD9A8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B5F47869F2CBFE8FDA3EFA49C597AE2F">Jabbari, H.</mods:namePart>
</mods:name>
<mods:name id="977FC30907E5C76942491F44279DBD8F" type="personal">
<mods:role id="66908447FFF3AB84C70E33287177419C">
<mods:roleTerm id="8E459700F0B6B0D3022B26CF9A3B4441">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8AB261FF813A76F43C4383961516A89A">Peña-Santiago, R.</mods:namePart>
</mods:name>
<mods:typeOfResource id="7C9E458F969F0ACADA01EA5CB1F26BCF">text</mods:typeOfResource>
<mods:relatedItem id="1515483AC9DA56996D2AAED64779D234" type="host">
<mods:titleInfo id="E70CC69519925A47F0876F3653517593">
<mods:title id="0872C436B8A85BD004AFC674BEB231A5">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="EF655E45AF128BFAFAFDFDAADEAEBC15">
<mods:date id="A36DCA3A40082FE11B3AEF4A98B64D8F">2018</mods:date>
<mods:detail id="ADE33DB2ADCC3BF2D7C1253C4D5121C5" type="pubDate">
<mods:number id="AFFF167B0068714C93399914CC08BC68">2018-01-03</mods:number>
</mods:detail>
<mods:detail id="28219C3BEA3F7ACC320F63F01CE9D16E" type="volume">
<mods:number id="BB52BF54C2533B15F16119B85D3F6739">4369</mods:number>
</mods:detail>
<mods:detail id="4B174AE0D17C7966E3FF81C346534970" type="issue">
<mods:number id="3B0F19F3DD1D5F1A05A4DD2B36552B98">1</mods:number>
</mods:detail>
<mods:extent id="8CFC0279EEF5C4110BB0F141BCBFEAC2" unit="page">
<mods:start id="E522525CD71D1C6AF9D69AF9BCD635E7">128</mods:start>
<mods:end id="1C2BEDCFCA63F64C08AEC63E321D84B1">136</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="BEF257DBCC7A5AA19B0C427683D7ACDB">journal article</mods:classification>
<mods:identifier id="BFEC309F4AC21003D68813BD1DE99667" type="CLB-Dataset">31084</mods:identifier>
<mods:identifier id="A908FED26993B346F2AA8E8FB6C98744" type="DOI">10.11646/zootaxa.4369.1.7</mods:identifier>
<mods:identifier id="D0406C790B5DD54F7995C1EE43403F38" type="GBIF-Dataset">e320619c-87b6-4f6d-a630-16fdaf828dfa</mods:identifier>
<mods:identifier id="6C3778ED85F6D231842A723D1A20A51A" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="E998848B4FF1E1BC72FB2D58335416B0" type="Zenodo-Dep">1135628</mods:identifier>
<mods:identifier id="8D2484C0ACE7F5F2C493CEA6BEC984CA" type="ZooBank">CD413E4C-0221-46AA-BD23-B733EFFAE6B6</mods:identifier>
</mods:mods>
<treatment id="F06FF311FF893A04FF7AFD61BD2DD295" ID-DOI="http://doi.org/10.5281/zenodo.5950704" ID-GBIF-Taxon="139582143" ID-Zenodo-Dep="5950704" LSID="urn:lsid:plazi:treatment:F06FF311FF893A04FF7AFD61BD2DD295" httpUri="http://treatment.plazi.org/id/F06FF311FF893A04FF7AFD61BD2DD295" lastPageId="8" lastPageNumber="135" pageId="1" pageNumber="129">
<subSubSection id="30DC118CFF893A0DFF7AFD61BF60D18E" pageId="1" pageNumber="129" type="nomenclature">
<paragraph id="78794207FF893A0DFF7AFD61BC71D1EF" blockId="1.[151,538,655,715]" box="[151,538,655,682]" pageId="1" pageNumber="129">
<heading id="2331F56BFF893A0DFF7AFD61BC71D1EF" bold="true" box="[151,538,655,682]" fontSize="11" level="1" pageId="1" pageNumber="129" reason="1">
<emphasis id="4AB29E15FF893A0DFF7AFD61BC71D1EF" bold="true" box="[151,538,655,682]" pageId="1" pageNumber="129">
<taxonomicName id="BFC63984FF893A0DFF7AFD61BFBBD1EC" authority="Kazemi &amp; Niknam &amp; Jabbari &amp; Peña-Santiago, 2018" authorityName="Kazemi &amp; Niknam &amp; Jabbari &amp; Peña-Santiago" authorityYear="2018" box="[151,464,655,681]" class="Adenophorea" family="Qudsianematidae" genus="Eudorylaimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="1" pageNumber="129" phylum="Nematoda" rank="species" species="kahaqensis" status="sp. nov.">
<emphasis id="4AB29E15FF893A0DFF7AFD61BFBBD1EC" bold="true" box="[151,464,655,681]" italics="true" pageId="1" pageNumber="129">Eudorylaimus kahaqensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5181236EFF893A0DFE3AFD7EBC71D1EF" box="[471,538,656,682]" pageId="1" pageNumber="129" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="78794207FF893A0DFF7AFD5CBF60D18E" blockId="1.[151,538,655,715]" box="[151,267,690,715]" pageId="1" pageNumber="129">
(
<figureCitation id="E0FD5E82FF893A0DFF72FD5CBF69D18E" box="[159,258,690,715]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE5" captionStart-5="FIGURE 6" captionStartId-0="3.[151,250,1938,1960]" captionStartId-1="4.[151,250,1496,1518]" captionStartId-2="5.[151,250,1526,1548]" captionStartId-3="6.[151,250,1845,1867]" captionStartId-4="7.[151,250,1011,1033]" captionStartId-5="7.[151,250,1396,1418]" captionTargetBox-0="[364,1228,639,1891]" captionTargetBox-1="[374,1225,210,1449]" captionTargetBox-2="[344,1244,451,1501]" captionTargetBox-3="[345,1239,705,1816]" captionTargetBox-4="[344,1244,199,985]" captionTargetBox-5="[348,1239,1119,1370]" captionTargetId-0="figure@3.[340,1247,625,1917]" captionTargetId-1="figure@4.[340,1247,193,1475]" captionTargetId-2="figure@5.[340,1247,445,1505]" captionTargetId-3="figure@6.[340,1247,697,1824]" captionTargetId-4="figure@7.[340,1247,193,989]" captionTargetId-5="figure@7.[340,1247,1113,1375]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="7" captionText-0="FIGURE 1. Eudorylaimus kahaqensis sp. n. (Female, LM). A: Entire. B: Lip region in surface lateral VieW.C: Lip region in median lateral VieW.D: Anterior genital branch. E: Posteriorbody region." captionText-1="FIGURE 2. Eudorylaimus kahaqensis sp. n. (Male). A: Neck, posterior region. B: Entire. C,D: Posterior body region. E, F: Lateral guiding piece.G: Spicule." captionText-2="FIGURE 3. Eudorylaimus kahaqensis sp. n. (Female, LM). A, B: Entire. C, D: Anterior genital branch. E: Posterior body region. F: Caudal region. G: OViduct-uterus junction. H, I: Vagina. (Scale bars: A, B = 200 µm; C, D = 50 µm; E–G = 20 µm; H, I = 10 µm.)" captionText-3="FIGURE 4. Eudorylaimus kahaqensis sp. n. (Anterior and neck regions, LM). A, B: Anterior region in lateral, median VieW. C: Pharyngeal expansion. D: Neck region. E: Lip region and amphid in lateral, surface VieW. F: Coelomocyte a short distance behind thenerVe ring.G:Posteriorhalf of pharyngeal expansion andpharyngo-intestinal junction shoWing S1N2, S2N and dorsal cell mass. H: Posterior part of pharyngeal expansion and pharyngo-intestinal junction shoWing S2N and dorsal cell mass.I, J: Dorsal cell mass atleVel of pharyngo-intestinal junction.(Scale bars: A, B,F, I, J = 10 µm; C = 50 µm; D = 100 µm; E= 5µm; G, H = 20 µm.)" captionText-4="FIGURE5. Eudorylaimus kahaqensissp. n. (Male, LM).A: Entire.B–D:Spicules.E, H:Caudal region.F, G:Posteriorbody region.(Scale bars:A = 200 µm; B–D, E, H = 10 µm; F, G = 50 µm.)" captionText-5="FIGURE 6. Lip region of Eudorylaimus kahaqensis sp. n. (SEM). A: SubVentral VieW. B: Sublateral VieW.C: En face VieW. (Scale bar = 2 µm.)" httpUri-0="https://zenodo.org/record/1135630/files/figure.png" httpUri-1="https://zenodo.org/record/1135632/files/figure.png" httpUri-2="https://zenodo.org/record/1135634/files/figure.png" httpUri-3="https://zenodo.org/record/1135636/files/figure.png" httpUri-4="https://zenodo.org/record/1135638/files/figure.png" httpUri-5="https://zenodo.org/record/1135640/files/figure.png" pageId="1" pageNumber="129">Figs 1–6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="30DC118CFF893A0DFF7AFD14BC7BD072" pageId="1" pageNumber="129" type="materials_examined">
<paragraph id="78794207FF893A0DFF7AFD14BC7BD072" blockId="1.[151,1427,762,823]" pageId="1" pageNumber="129">
<emphasis id="4AB29E15FF893A0DFF7AFD14BFE9D056" bold="true" box="[151,386,762,787]" pageId="1" pageNumber="129">Material examined:</emphasis>
Twenty females, twelve males and several juveniles, in acceptable (mostly good) condition.
<emphasis id="4AB29E15FF893A0DFF2AFCF0BF14D072" bold="true" box="[199,383,798,823]" pageId="1" pageNumber="129">Measurements:</emphasis>
See
<tableCitation id="354477BCFF893A0DFE5BFCF0BC61D072" box="[438,522,798,823]" captionStart="TABLE 1" captionStartId="1.[151,239,870,893]" captionText="TABLE 1. Morphometricdatafor Eudorylaimuskahaqensis sp. n. All measurements are inµm(except L, in mm)and in theform: mean ± SD (range)." pageId="1" pageNumber="129">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
<caption id="2CB9128FFF893A0DFF7AFC88BFB9D0D8" pageId="1" pageNumber="129">
<paragraph id="78794207FF893A0DFF7AFC88BFB9D0D8" blockId="1.[151,1436,869,925]" pageId="1" pageNumber="129">
<emphasis id="4AB29E15FF893A0DFF7AFC88BF62D038" bold="true" box="[151,265,870,893]" pageId="1" pageNumber="129">TABLE 1.</emphasis>
Morphometric data for
<taxonomicName id="BFC63984FF893A0DFDE5FC8BBD72D038" authority="Kazemi &amp; Niknam &amp; Jabbari &amp; Peña-Santiago, 2018" authorityName="Kazemi &amp; Niknam &amp; Jabbari &amp; Peña-Santiago" authorityYear="2018" box="[520,793,869,893]" class="Adenophorea" family="Qudsianematidae" genus="Eudorylaimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="1" pageNumber="129" phylum="Nematoda" rank="species" species="kahaqensis" status="sp. nov.">
<emphasis id="4AB29E15FF893A0DFDE5FC8BBD72D038" box="[520,793,869,893]" italics="true" pageId="1" pageNumber="129">Eudorylaimus kahaqensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5181236EFF893A0DFCCDFC88BD30D038" box="[800,859,870,893]" pageId="1" pageNumber="129" rank="species">
<emphasis id="4AB29E15FF893A0DFCCDFC88BD30D038" bold="true" box="[800,859,870,893]" pageId="1" pageNumber="129">sp. n.</emphasis>
</taxonomicNameLabel>
All measurements are in µm (except L, in mm) and in the form: mean ± SD (range).
</paragraph>
</caption>
<paragraph id="78794207FF893A0DFDB2FC57BA1CD476" pageId="1" pageNumber="129">
<table id="0AC6B0A7FF89C5F3FF72FC57BB5BD476" box="[159,1328,953,1843]" gridcols="5" gridrows="22" pageId="1" pageNumber="129">
<tr id="C6F64045FF89C5F3FF72FC57BB5BD08A" box="[159,1328,953,975]" gridrow="0" pageId="1" pageNumber="129" rowspan-0="1" rowspan-1="1" rowspan-4="1">
<th id="85272939FF89C5F3FDB2FC57BCD6D08A" box="[607,701,953,975]" gridcol="2" gridrow="0" pageId="1" pageNumber="129">Holotype</th>
<th id="85272939FF89C5F3FD1DFC57BDB4D08A" box="[752,991,953,975]" gridcol="3" gridrow="0" pageId="1" pageNumber="129">Paratypes</th>
</tr>
<tr id="C6F64045FF89C5F3FF72FC0CBB5BD0BD" box="[159,1328,994,1016]" gridrow="1" pageId="1" pageNumber="129">
<th id="85272939FF89C5F3FF72FC0CBFB8D0BD" box="[159,467,994,1016]" gridcol="0" gridrow="1" pageId="1" pageNumber="129">Characters</th>
<td id="85272939FF89C5F3FDD7FC0CBC2CD0BD" box="[570,583,994,1016]" gridcol="1" gridrow="1" pageId="1" pageNumber="129">n</td>
<td id="85272939FF89C5F3FDB2FC0CBCD6D0BD" box="[607,701,994,1016]" gridcol="2" gridrow="1" pageId="1" pageNumber="129">♀</td>
<td id="85272939FF89C5F3FD1DFC0CBDB4D0BD" box="[752,991,994,1016]" gridcol="3" gridrow="1" pageId="1" pageNumber="129">8♀</td>
<td id="85272939FF89C5F3FBADFC0CBB5BD0BD" box="[1088,1328,994,1016]" gridcol="4" gridrow="1" pageId="1" pageNumber="129">7♂</td>
</tr>
<tr id="C6F64045FF89C5F3FF72FBE2BB5BD767" box="[159,1328,1036,1058]" gridrow="2" pageId="1" pageNumber="129" rowspan-1="1">
<th id="85272939FF89C5F3FF72FBE2BFB8D767" box="[159,467,1036,1058]" gridcol="0" gridrow="2" pageId="1" pageNumber="129">L</th>
<td id="85272939FF89C5F3FDB2FBE2BCD6D767" box="[607,701,1036,1058]" gridcol="2" gridrow="2" pageId="1" pageNumber="129">1.52</td>
<td id="85272939FF89C5F3FD1DFBE2BDB4D767" box="[752,991,1036,1058]" gridcol="3" gridrow="2" pageId="1" pageNumber="129">1.61 ± 0.13 (1.40–1.75)</td>
<td id="85272939FF89C5F3FBADFBE2BB5BD767" box="[1088,1328,1036,1058]" gridcol="4" gridrow="2" pageId="1" pageNumber="129">1.56 ± 0.15 (1.38–1.86)</td>
</tr>
<tr id="C6F64045FF89C5F3FF72FBDBBB5BD70E" box="[159,1328,1077,1099]" gridrow="3" pageId="1" pageNumber="129" rowspan-1="1">
<th id="85272939FF89C5F3FF72FBDBBFB8D70E" box="[159,467,1077,1099]" gridcol="0" gridrow="3" pageId="1" pageNumber="129">a</th>
<td id="85272939FF89C5F3FDB2FBDBBCD6D70E" box="[607,701,1077,1099]" gridcol="2" gridrow="3" pageId="1" pageNumber="129">34</td>
<td id="85272939FF89C5F3FD1DFBDBBDB4D70E" box="[752,991,1077,1099]" gridcol="3" gridrow="3" pageId="1" pageNumber="129">32.5 ± 2.5 (29–39)</td>
<td id="85272939FF89C5F3FBADFBDBBB5BD70E" box="[1088,1328,1077,1099]" gridcol="4" gridrow="3" pageId="1" pageNumber="129">35.4 ±2.9 (32–39)</td>
</tr>
<tr id="C6F64045FF89C5F3FF72FBB0BB5BD731" box="[159,1328,1118,1140]" gridrow="4" pageId="1" pageNumber="129" rowspan-1="1">
<th id="85272939FF89C5F3FF72FBB0BFB8D731" box="[159,467,1118,1140]" gridcol="0" gridrow="4" pageId="1" pageNumber="129">b</th>
<td id="85272939FF89C5F3FDB2FBB0BCD6D731" box="[607,701,1118,1140]" gridcol="2" gridrow="4" pageId="1" pageNumber="129">4</td>
<td id="85272939FF89C5F3FD1DFBB0BDB4D731" box="[752,991,1118,1140]" gridcol="3" gridrow="4" pageId="1" pageNumber="129">4.2 ± 0.2 (4.0–5.0)</td>
<td id="85272939FF89C5F3FBADFBB0BB5BD731" box="[1088,1328,1118,1140]" gridcol="4" gridrow="4" pageId="1" pageNumber="129">4.1 ± 0.3 (4.0–4.5)</td>
</tr>
<tr id="C6F64045FF89C5F3FF72FB66BB5BD7DB" box="[159,1328,1160,1182]" gridrow="5" pageId="1" pageNumber="129" rowspan-1="1">
<th id="85272939FF89C5F3FF72FB66BFB8D7DB" box="[159,467,1160,1182]" gridcol="0" gridrow="5" pageId="1" pageNumber="129">c</th>
<td id="85272939FF89C5F3FDB2FB66BCD6D7DB" box="[607,701,1160,1182]" gridcol="2" gridrow="5" pageId="1" pageNumber="129">24</td>
<td id="85272939FF89C5F3FD1DFB66BDB4D7DB" box="[752,991,1160,1182]" gridcol="3" gridrow="5" pageId="1" pageNumber="129">26.0 ± 2.4 (24–30)</td>
<td id="85272939FF89C5F3FBADFB66BB5BD7DB" box="[1088,1328,1160,1182]" gridcol="4" gridrow="5" pageId="1" pageNumber="129">26.5 ± 2.8 (22–32)</td>
</tr>
<tr id="C6F64045FF89C5F3FF72FB5FBB5BD782" box="[159,1328,1201,1223]" gridrow="6" pageId="1" pageNumber="129" rowspan-1="1">
<th id="85272939FF89C5F3FF72FB5FBFB8D782" box="[159,467,1201,1223]" gridcol="0" gridrow="6" pageId="1" pageNumber="129">c΄</th>
<td id="85272939FF89C5F3FDB2FB5FBCD6D782" box="[607,701,1201,1223]" gridcol="2" gridrow="6" pageId="1" pageNumber="129">2.5</td>
<td id="85272939FF89C5F3FD1DFB5FBDB4D782" box="[752,991,1201,1223]" gridcol="3" gridrow="6" pageId="1" pageNumber="129">2.2 ± 0.3 (1.9–2.6)</td>
<td id="85272939FF89C5F3FBADFB5FBB5BD782" box="[1088,1328,1201,1223]" gridcol="4" gridrow="6" pageId="1" pageNumber="129">1.9 ± 0.2 (1.7–2.3)</td>
</tr>
<tr id="C6F64045FF89C5F3FF72FB34BB5BD7B5" box="[159,1328,1242,1264]" gridrow="7" pageId="1" pageNumber="129" rowspan-1="1">
<th id="85272939FF89C5F3FF72FB34BFB8D7B5" box="[159,467,1242,1264]" gridcol="0" gridrow="7" pageId="1" pageNumber="129">V</th>
<td id="85272939FF89C5F3FDB2FB34BCD6D7B5" box="[607,701,1242,1264]" gridcol="2" gridrow="7" pageId="1" pageNumber="129">50</td>
<td id="85272939FF89C5F3FD1DFB34BDB4D7B5" box="[752,991,1242,1264]" gridcol="3" gridrow="7" pageId="1" pageNumber="129">50.6 ± 1.4 (48–52)</td>
<td id="85272939FF89C5F3FBADFB34BB5BD7B5" box="[1088,1328,1242,1264]" gridcol="4" gridrow="7" pageId="1" pageNumber="129">-</td>
</tr>
<tr id="C6F64045FF89C5F3FF72FAEABB5BD65F" box="[159,1328,1284,1306]" gridrow="8" pageId="1" pageNumber="129" rowspan-1="1">
<th id="85272939FF89C5F3FF72FAEABFB8D65F" box="[159,467,1284,1306]" gridcol="0" gridrow="8" pageId="1" pageNumber="129">Lip region diam.</th>
<td id="85272939FF89C5F3FDB2FAEABCD6D65F" box="[607,701,1284,1306]" gridcol="2" gridrow="8" pageId="1" pageNumber="129">19</td>
<td id="85272939FF89C5F3FD1DFAEABDB4D65F" box="[752,991,1284,1306]" gridcol="3" gridrow="8" pageId="1" pageNumber="129">18.4 ± 0.7 (17.0–21.0)</td>
<td id="85272939FF89C5F3FBADFAEABB5BD65F" box="[1088,1328,1284,1306]" gridcol="4" gridrow="8" pageId="1" pageNumber="129">18.5 ± 0.5 (18.0–19.0)</td>
</tr>
<tr id="C6F64045FF89C5F3FF72FAC3BB5BD606" box="[159,1328,1325,1347]" gridrow="9" pageId="1" pageNumber="129" rowspan-1="1">
<th id="85272939FF89C5F3FF72FAC3BFB8D606" box="[159,467,1325,1347]" gridcol="0" gridrow="9" pageId="1" pageNumber="129">Odontostyle length</th>
<td id="85272939FF89C5F3FDB2FAC3BCD6D606" box="[607,701,1325,1347]" gridcol="2" gridrow="9" pageId="1" pageNumber="129">21</td>
<td id="85272939FF89C5F3FD1DFAC3BDB4D606" box="[752,991,1325,1347]" gridcol="3" gridrow="9" pageId="1" pageNumber="129">22.0 ± 1.4 (21–23)</td>
<td id="85272939FF89C5F3FBADFAC3BB5BD606" box="[1088,1328,1325,1347]" gridcol="4" gridrow="9" pageId="1" pageNumber="129">21.0 ± 0.8 (21–23)</td>
</tr>
<tr id="C6F64045FF89C5F3FF72FAB8BB5BD629" box="[159,1328,1366,1388]" gridrow="10" pageId="1" pageNumber="129" rowspan-1="1">
<th id="85272939FF89C5F3FF72FAB8BFB8D629" box="[159,467,1366,1388]" gridcol="0" gridrow="10" pageId="1" pageNumber="129">Odontophore length</th>
<td id="85272939FF89C5F3FDB2FAB8BCD6D629" box="[607,701,1366,1388]" gridcol="2" gridrow="10" pageId="1" pageNumber="129">38</td>
<td id="85272939FF89C5F3FD1DFAB8BDB4D629" box="[752,991,1366,1388]" gridcol="3" gridrow="10" pageId="1" pageNumber="129">35.6 ± 3.8(31–42)</td>
<td id="85272939FF89C5F3FBADFAB8BB5BD629" box="[1088,1328,1366,1388]" gridcol="4" gridrow="10" pageId="1" pageNumber="129">34.0 ± 3.4 (29–38)</td>
</tr>
<tr id="C6F64045FF89C5F3FF72FA6EBB5BD6D3" box="[159,1328,1408,1430]" gridrow="11" pageId="1" pageNumber="129" rowspan-1="1">
<th id="85272939FF89C5F3FF72FA6EBFB8D6D3" box="[159,467,1408,1430]" gridcol="0" gridrow="11" pageId="1" pageNumber="129">Guiding ring from ant. end</th>
<td id="85272939FF89C5F3FDB2FA6EBCD6D6D3" box="[607,701,1408,1430]" gridcol="2" gridrow="11" pageId="1" pageNumber="129">8.0</td>
<td id="85272939FF89C5F3FD1DFA6EBDB4D6D3" box="[752,991,1408,1430]" gridcol="3" gridrow="11" pageId="1" pageNumber="129">9.1±0.9 (8.0–11.0)</td>
<td id="85272939FF89C5F3FBADFA6EBB5BD6D3" box="[1088,1328,1408,1430]" gridcol="4" gridrow="11" pageId="1" pageNumber="129">9.8 ± 1.1 (9.0–12.0)</td>
</tr>
<tr id="C6F64045FF89C5F3FF72FA47BB5BD6FA" box="[159,1328,1449,1471]" gridrow="12" pageId="1" pageNumber="129" rowspan-1="1">
<th id="85272939FF89C5F3FF72FA47BFB8D6FA" box="[159,467,1449,1471]" gridcol="0" gridrow="12" pageId="1" pageNumber="129">Neck length</th>
<td id="85272939FF89C5F3FDB2FA47BCD6D6FA" box="[607,701,1449,1471]" gridcol="2" gridrow="12" pageId="1" pageNumber="129">375</td>
<td id="85272939FF89C5F3FD1DFA47BDB4D6FA" box="[752,991,1449,1471]" gridcol="3" gridrow="12" pageId="1" pageNumber="129">377 ± 17.4 (347–397)</td>
<td id="85272939FF89C5F3FBADFA47BB5BD6FA" box="[1088,1328,1449,1471]" gridcol="4" gridrow="12" pageId="1" pageNumber="129">372 ± 23 (344–400)</td>
</tr>
<tr id="C6F64045FF89C5F3FF72FA3CBB5BD6AD" box="[159,1328,1490,1512]" gridrow="13" pageId="1" pageNumber="129" rowspan-1="1">
<th id="85272939FF89C5F3FF72FA3CBFB8D6AD" box="[159,467,1490,1512]" gridcol="0" gridrow="13" pageId="1" pageNumber="129">Pharyngeal expansion length</th>
<td id="85272939FF89C5F3FDB2FA3CBCD6D6AD" box="[607,701,1490,1512]" gridcol="2" gridrow="13" pageId="1" pageNumber="129">201</td>
<td id="85272939FF89C5F3FD1DFA3CBDB4D6AD" box="[752,991,1490,1512]" gridcol="3" gridrow="13" pageId="1" pageNumber="129">192 ± 7.1(162–205)</td>
<td id="85272939FF89C5F3FBADFA3CBB5BD6AD" box="[1088,1328,1490,1512]" gridcol="4" gridrow="13" pageId="1" pageNumber="129">187 ±13.6 (170–203)</td>
</tr>
<tr id="C6F64045FF89C5F3FF72FA12BB5BD557" box="[159,1328,1532,1554]" gridrow="14" pageId="1" pageNumber="129" rowspan-1="1">
<th id="85272939FF89C5F3FF72FA12BFB8D557" box="[159,467,1532,1554]" gridcol="0" gridrow="14" pageId="1" pageNumber="129">Diam. at neck base</th>
<td id="85272939FF89C5F3FDB2FA12BCD6D557" box="[607,701,1532,1554]" gridcol="2" gridrow="14" pageId="1" pageNumber="129">42</td>
<td id="85272939FF89C5F3FD1DFA12BDB4D557" box="[752,991,1532,1554]" gridcol="3" gridrow="14" pageId="1" pageNumber="129">45.2 ± 6.4(37–53)</td>
<td id="85272939FF89C5F3FBADFA12BB5BD557" box="[1088,1328,1532,1554]" gridcol="4" gridrow="14" pageId="1" pageNumber="129">41.9 ± 5.2 (37–50)</td>
</tr>
<tr id="C6F64045FF89C5F3FF72F9CBBB5BD57E" box="[159,1328,1573,1595]" gridrow="15" pageId="1" pageNumber="129" rowspan-1="1">
<th id="85272939FF89C5F3FF72F9CBBFB8D57E" box="[159,467,1573,1595]" gridcol="0" gridrow="15" pageId="1" pageNumber="129">at mid- body</th>
<td id="85272939FF89C5F3FDB2F9CBBCD6D57E" box="[607,701,1573,1595]" gridcol="2" gridrow="15" pageId="1" pageNumber="129">43</td>
<td id="85272939FF89C5F3FD1DF9CBBDB4D57E" box="[752,991,1573,1595]" gridcol="3" gridrow="15" pageId="1" pageNumber="129">49.6 ± 5.9 (43–57)</td>
<td id="85272939FF89C5F3FBADF9CBBB5BD57E" box="[1088,1328,1573,1595]" gridcol="4" gridrow="15" pageId="1" pageNumber="129">44.0 ± 4.6 (39–51)</td>
</tr>
<tr id="C6F64045FF89C5F3FF72F9A0BB5BD521" box="[159,1328,1614,1636]" gridrow="16" pageId="1" pageNumber="129" rowspan-1="1">
<th id="85272939FF89C5F3FF72F9A0BFB8D521" box="[159,467,1614,1636]" gridcol="0" gridrow="16" pageId="1" pageNumber="129">at anus/cloaca aperture</th>
<td id="85272939FF89C5F3FDB2F9A0BCD6D521" box="[607,701,1614,1636]" gridcol="2" gridrow="16" pageId="1" pageNumber="129">25</td>
<td id="85272939FF89C5F3FD1DF9A0BDB4D521" box="[752,991,1614,1636]" gridcol="3" gridrow="16" pageId="1" pageNumber="129">27.6 ± 2.5 (25–33)</td>
<td id="85272939FF89C5F3FBADF9A0BB5BD521" box="[1088,1328,1614,1636]" gridcol="4" gridrow="16" pageId="1" pageNumber="129">31.1 ± 2.0 (29–34)</td>
</tr>
<tr id="C6F64045FF89C5F3FF72F996BB5BD5CB" box="[159,1328,1656,1678]" gridrow="17" pageId="1" pageNumber="129" rowspan-1="1">
<th id="85272939FF89C5F3FF72F996BFB8D5CB" box="[159,467,1656,1678]" gridcol="0" gridrow="17" pageId="1" pageNumber="129">Prerectum length</th>
<td id="85272939FF89C5F3FDB2F996BCD6D5CB" box="[607,701,1656,1678]" gridcol="2" gridrow="17" pageId="1" pageNumber="129">48</td>
<td id="85272939FF89C5F3FD1DF996BDB4D5CB" box="[752,991,1656,1678]" gridcol="3" gridrow="17" pageId="1" pageNumber="129">46.2 ± 3.0 (43–50)</td>
<td id="85272939FF89C5F3FBADF996BB5BD5CB" box="[1088,1328,1656,1678]" gridcol="4" gridrow="17" pageId="1" pageNumber="129">62.5</td>
</tr>
<tr id="C6F64045FF89C5F3FF72F94FBB5BD5F2" box="[159,1328,1697,1719]" gridrow="18" pageId="1" pageNumber="129" rowspan-1="1">
<th id="85272939FF89C5F3FF72F94FBFB8D5F2" box="[159,467,1697,1719]" gridcol="0" gridrow="18" pageId="1" pageNumber="129">Rectum/cloaca length</th>
<td id="85272939FF89C5F3FDB2F94FBCD6D5F2" box="[607,701,1697,1719]" gridcol="2" gridrow="18" pageId="1" pageNumber="129">33</td>
<td id="85272939FF89C5F3FD1DF94FBDB4D5F2" box="[752,991,1697,1719]" gridcol="3" gridrow="18" pageId="1" pageNumber="129">30.4 ± 4.0 (26–37)</td>
<td id="85272939FF89C5F3FBADF94FBB5BD5F2" box="[1088,1328,1697,1719]" gridcol="4" gridrow="18" pageId="1" pageNumber="129">63.0 ± 4.4 (57–68)</td>
</tr>
<tr id="C6F64045FF89C5F3FF72F924BB5BD5A5" box="[159,1328,1738,1760]" gridrow="19" pageId="1" pageNumber="129" rowspan-1="1">
<th id="85272939FF89C5F3FF72F924BFB8D5A5" box="[159,467,1738,1760]" gridcol="0" gridrow="19" pageId="1" pageNumber="129">Tail length</th>
<td id="85272939FF89C5F3FDB2F924BCD6D5A5" box="[607,701,1738,1760]" gridcol="2" gridrow="19" pageId="1" pageNumber="129">62</td>
<td id="85272939FF89C5F3FD1DF924BDB4D5A5" box="[752,991,1738,1760]" gridcol="3" gridrow="19" pageId="1" pageNumber="129">62.2 ± 7.4 (50–73)</td>
<td id="85272939FF89C5F3FBADF924BB5BD5A5" box="[1088,1328,1738,1760]" gridcol="4" gridrow="19" pageId="1" pageNumber="129">60.4 ± 6.7 (45–68)</td>
</tr>
<tr id="C6F64045FF89C5F3FF72F91ABB5BD44F" box="[159,1328,1780,1802]" gridrow="20" pageId="1" pageNumber="129" rowspan-1="1">
<th id="85272939FF89C5F3FF72F91ABFB8D44F" box="[159,467,1780,1802]" gridcol="0" gridrow="20" pageId="1" pageNumber="129">Spicule length</th>
<td id="85272939FF89C5F3FDB2F91ABCD6D44F" box="[607,701,1780,1802]" gridcol="2" gridrow="20" pageId="1" pageNumber="129">-</td>
<td id="85272939FF89C5F3FD1DF91ABDB4D44F" box="[752,991,1780,1802]" gridcol="3" gridrow="20" pageId="1" pageNumber="129">-</td>
<td id="85272939FF89C5F3FBADF91ABB5BD44F" box="[1088,1328,1780,1802]" gridcol="4" gridrow="20" pageId="1" pageNumber="129">50.2 ± 2.5 (48–55)</td>
</tr>
<tr id="C6F64045FF89C5F3FF72F8F3BB5BD476" box="[159,1328,1821,1843]" gridrow="21" pageId="1" pageNumber="129" rowspan-1="1">
<th id="85272939FF89C5F3FF72F8F3BFB8D476" box="[159,467,1821,1843]" gridcol="0" gridrow="21" pageId="1" pageNumber="129">Ventromedian supplements</th>
<td id="85272939FF89C5F3FDB2F8F3BCD6D476" box="[607,701,1821,1843]" gridcol="2" gridrow="21" pageId="1" pageNumber="129">-</td>
<td id="85272939FF89C5F3FD1DF8F3BDB4D476" box="[752,991,1821,1843]" gridcol="3" gridrow="21" pageId="1" pageNumber="129">-</td>
<td id="85272939FF89C5F3FBADF8F3BB5BD476" box="[1088,1328,1821,1843]" gridcol="4" gridrow="21" pageId="1" pageNumber="129">(7–9)</td>
</tr>
</table>
</paragraph>
<subSubSection id="30DC118CFF893A0EFF2AF899BE8ED4E5" lastPageId="2" lastPageNumber="130" pageId="1" pageNumber="129" type="description">
<paragraph id="78794207FF893A0EFF2AF899BCEDD091" blockId="1.[151,1436,1911,2009]" lastBlockId="2.[151,1437,151,2024]" lastPageId="2" lastPageNumber="130" pageId="1" pageNumber="129">
<emphasis id="4AB29E15FF893A0DFF2AF899BF32D4D5" bold="true" box="[199,345,1911,1936]" pageId="1" pageNumber="129">Description.</emphasis>
<emphasis id="4AB29E15FF893A0DFE8FF896BFF5D4D5" box="[354,414,1912,1936]" italics="true" pageId="1" pageNumber="129">Adult</emphasis>
: Slender nematodes (
<emphasis id="4AB29E15FF893A0DFD77F897BCC3D4D5" box="[666,680,1913,1936]" italics="true" pageId="1" pageNumber="129">a</emphasis>
= 29–39) of medium size,
<quantity id="BF3EEFE2FF893A0DFC08F899BAE0D4CA" box="[997,1163,1911,1936]" metricMagnitude="-3" metricUnit="m" metricValue="1.575" metricValueMax="1.75" metricValueMin="1.4" pageId="1" pageNumber="129" unit="mm" value="1.575" valueMax="1.75" valueMin="1.4">1.40–1.75 mm</quantity>
long. Body cylindrical, tapering towards both ends as the tail is conical. Habitus regularly curved ventrad, an open ‘C’ or ‘J’ shape in females, ‘J’ or ‘G’ shaped in males. Cuticle 2.0–2.5 µm thick in the anterior region, 2.0–3.5 µm at mid-body and 3.0–4.0 µm on tail; outer layer with fine transverse striation throughout the body, perceptible even under LM; inner layer thicker than the outer one, especially visible at caudal region. Lip region anteriorly truncate, nearly continuous with the adjacent body, 2.8–3.2 times as wide as high, and
<emphasis id="4AB29E15FF8A3A0EFC50FF0FBDBCD3BD" box="[957,983,225,248]" italics="true" pageId="2" pageNumber="130">ca</emphasis>
one half (45–50%) of body diameter at neck base; anterior margin visibly corrugated or wrinkled. SEM observations: lips totally amalgamated; labial and cephalic papillae distinct but weakly protruding, button-like; oral field broad, with coarse radial striation—visibly stronger than the transverse striation of the adjacent body, and responsible for the corrugated anterior marginrunning from the oral aperture to the posterior margin of lip region; oral aperture apparently small. A moderately sclerotized but perceptible cephalic framework is present. Amphidial fovea stirrup-shaped, aperture a transverse slit 9–11 µm wide and occupying
<emphasis id="4AB29E15FF8A3A0EFDFAFE57BC5AD295" box="[535,561,441,464]" italics="true" pageId="2" pageNumber="130">ca</emphasis>
one-half (50–56%) of lip region diameter. Cheilostom wider than usual, with convex walls. Odontostyle equal to lip region diameter long, comparatively slender (9–12 times as long as wide), 1.1–1.3% of total body length, and with aperture 6–8 µm long or occupying less than one-half (33–44%) of total length. Guiding ring simple, somewhat plicate, and located at 8–11 µm or 0.4–0.5 times the lip region diameter from the anterior end. Odontophore rod-like, 1.7–2.0 times the odontostyle in length, its junction with the pharyngeal lining rather inconspicuous. Anterior portion of pharynx slender but muscular, enlarging very gradually into the basal expansion that is 7.4–9.0 times as long as wide, 4.0–4.6 times as long as the body diameter at neck base and occupies 45–53% of total neck length; gland nuclei located as follows: DO = 60–66, DN = 66–70, S1N1 = 78–83, S1N2 = 82–88, S2N = 90–94, S2O = 94; DN remarkably posterior in position, far from pharyngeal enlargement, S1N1 conspicuously longer than wide, with its nucleolus much larger than that of S1N2 and even slightly larger than those of S2N, and S2O visibly behind S
<specimenCount id="6EC0898EFF8A3A0EFCDEFCD9BD3DD005" box="[819,854,809,837]" pageId="2" pageNumber="130" type="generic">2N.</specimenCount>
Nerve ring at 128–145 µm or 35–37% of the neck length from the anterior end. One coelomocyte 7.5–
<date id="0C7864C7FF8A3A0EFCEDFCA1BD58D022" box="[768,819,847,872]" pageId="2" pageNumber="130" value="2013-10">13 x</date>
5.5–10.0 µm is present in dorsal position in all the specimens examined a short distance behind the nerve ring, at 145–162 µm from the anterior end. Cardia cylindroid, 13.5–19.0 x 7.0–8.5 µm. A dorsal cell mass is always present at the pharyngo-intestinal junction. Intestine without any special differentiation.
</paragraph>
<paragraph id="78794207FF8A3A0EFF2AFC31BA49D6B5" blockId="2.[151,1437,151,2024]" pageId="2" pageNumber="130">
<emphasis id="4AB29E15FF8A3A0EFF2AFC31BF4FD0B2" box="[199,292,991,1015]" italics="true" pageId="2" pageNumber="130">Female:</emphasis>
Genital system didelphic-amphidelphic, with both branches of similar morphology and length, the anterior 192–264 µm or 11–14% of total body length, the posterior 182–273 µm or 12–14% of total body length. Ovaries reflexed, 77–101 µm long, not reaching the oviduct-uterus junction; oocytes in several rows in the germinative zone, then in one row. Oviduct 84–120 µm or 2.0–2.8 times the corresponding body diameter long, consisting of a slender portion made of prismatic cells and an appreciably longer than wide
<emphasis id="4AB29E15FF8A3A0EFB21FB9EBB0BD7C2" box="[1228,1376,1135,1159]" italics="true" pageId="2" pageNumber="130">pars dilatata</emphasis>
with visible lumen and often containing sperm cells. A marked narrowing surrounded by a weak muscular ring is present separating oviduct and uterus. Uterus a simple tube-like structure 62–125 µm or 1.5–2.3 times the corresponding body diameter long, often containing sperm cells as well. Vagina 26–33 µm long, extending to 63– 70% of body diameter:
<emphasis id="4AB29E15FF8A3A0EFE42FAEEBC08D652" box="[431,611,1279,1303]" italics="true" pageId="2" pageNumber="130">pars proximalis</emphasis>
<date id="0C7864C7FF8A3A0EFD80FB11BC9AD65D" box="[621,753,1279,1304]" pageId="2" pageNumber="130" value="2016-10-18" valueMax="2016-10-24" valueMin="2016-10-18">18–24 x 16</date>
–22 µm, with somewhat sigmoid walls and surrounded by moderately developed musculature;
<emphasis id="4AB29E15FF8A3A0EFDC3FACABCB2D67E" box="[558,729,1315,1339]" italics="true" pageId="2" pageNumber="130">pars refringens</emphasis>
consisting of two trapezoidal pieces 3.0–3.5 x 7.0–7.5 µm and a combined width of 15–18 µm;
<emphasis id="4AB29E15FF8A3A0EFDE1FAA6BCFDD61A" box="[524,662,1351,1375]" italics="true" pageId="2" pageNumber="130">pars distalis</emphasis>
5.5–6.5 µm long. One gland cell (granular in appearance) is present close the vagina both anteriorly and posteriorly. Vulva a transverse slit. Intestine-prerectum junction especially granular and colored in all the specimens examined. Prerectum 1.4–1.6, rectum 1.1–1.2 times the anal body diameter in length. Caudal region regularly curved ventrad and conical (somewhat) elongate with finely rounded tip; hyaline portion well developed, 17–30 µm or 28–45% of the total tail length.
</paragraph>
<paragraph id="78794207FF8A3A0EFF2AFA15BE8ED4E5" blockId="2.[151,1437,151,2024]" pageId="2" pageNumber="130">
<emphasis id="4AB29E15FF8A3A0EFF2AFA15BF62D556" box="[199,265,1531,1555]" italics="true" pageId="2" pageNumber="130">Male:</emphasis>
Genital system diorchic, with opposite testes. In addition to the ad-cloacal pair situated at 11–15 µm from the cloacal aperture, there is a series of 7–9 irregularly spaced (12–15 µm apart), ventromedian supplements, the most posterior of which is located at 22–28 µm from the ad-cloacal pair, at the level of or a short distance behind the anterior end of spicules. A distinct pre-cloacal space (hiatus) is therefore not present. Spicules dorylaimid, 1.7–1.9 times longer than the body diameter at level of cloacal aperture, 3.9–4.2 times longer than wide, and curved ventrad 124–130º; head 12–16 µm long, occupying 23–30% of total spicule length, 2.0–2.3 times as long as wide, with its dorsal side distinctly longer than the ventral one and slightly curved; median piece 35–41 µm long, 6.4–7.6 times longer than wide, occupying 37–46% of spicule width, and reaching the spicule posterior tip. Lateral guiding piece 13–17 µm long, 5.2–6.8 times longer than wide, with slightly bifid end. Prerectum 1.8– 2.5, cloaca 1.4–1.7 times the body diameter at level of cloacal aperture. Caudal region similar to that of female, its hyaline portion 22–25 (
<quantity id="BF3EEFE2FF8A3A0EFE4BF88DBF9CD439" box="[422,503,1891,1916]" metricMagnitude="-1" metricUnit="m" metricValue="2.921" pageId="2" pageNumber="130" unit="in" value="11.5">11.5 in</quantity>
one specimen, probably aberrant) µm long, occupying 39–40 (26)% of total tail length.
</paragraph>
</subSubSection>
<subSubSection id="30DC118CFF8A3A0FFF2AF845BF93D178" lastPageId="3" lastPageNumber="131" pageId="2" pageNumber="130" type="diagnosis">
<paragraph id="78794207FF8A3A0FFF2AF845BB78D220" blockId="2.[151,1437,151,2024]" lastBlockId="3.[151,1436,151,573]" lastPageId="3" lastPageNumber="131" pageId="2" pageNumber="130">
<emphasis id="4AB29E15FF8A3A0EFF2AF845BF2AD481" bold="true" box="[199,321,1963,1988]" pageId="2" pageNumber="130">Diagnosis.</emphasis>
The new species is characterized by its
<quantity id="BF3EEFE2FF8A3A0EFCEAF845BDC1D486" box="[775,938,1963,1988]" metricMagnitude="-3" metricUnit="m" metricValue="1.575" metricValueMax="1.75" metricValueMin="1.4" pageId="2" pageNumber="130" unit="mm" value="1.575" valueMax="1.75" valueMin="1.4">1.40–1.75 mm</quantity>
long body, lip region nearly continuous and 17–21 µm wide and bearing a weakly sclerotized but distinct cephalic framework, cheilostom broad and with convex walls, odontostyle 21–23 µm long with aperture occupying 33–44% of its length, presence of a dorsal coelomocyte a short distance behind the nerve ring, neck 347–397 µm long, DN rather posterior (66–70%), pharyngeal expansion 162–205 µm or occupying 45–53% of total neck length, presence a dorsal cell mass at level of pharyngo-intestinal junction,
<emphasis id="4AB29E15FF8B3A0FFE12FEEBBC7BD259" box="[511,528,261,284]" italics="true" pageId="3" pageNumber="131">V</emphasis>
= 48–52, caudal region conical (somewhat) elongate (50–73 µm,
<emphasis id="4AB29E15FF8B3A0FFB1FFEEBBA95D259" box="[1266,1278,261,284]" italics="true" pageId="3" pageNumber="131">c</emphasis>
= 24–30,
<emphasis id="4AB29E15FF8B3A0FFA82FEEABBEED25E" box="[1391,1413,260,283]" italics="true" pageId="3" pageNumber="131">c’</emphasis>
=
<quantity id="BF3EEFE2FF8B3A0FFF7AFEC9BF62D205" box="[151,265,295,320]" metricMagnitude="-2" metricUnit="m" metricValue="5.715" metricValueMax="6.604" metricValueMin="4.826" pageId="3" pageNumber="131" unit="in" value="2.25" valueMax="2.6" valueMin="1.9">1.9–2.6 in</quantity>
females, 45–68 µm,
<emphasis id="4AB29E15FF8B3A0FFE16FEC7BC6CD205" box="[507,519,297,320]" italics="true" pageId="3" pageNumber="131">c</emphasis>
= 22–32,
<emphasis id="4AB29E15FF8B3A0FFD90FEC6BCF8D27A" box="[637,659,296,319]" italics="true" pageId="3" pageNumber="131">c’</emphasis>
=
<quantity id="BF3EEFE2FF8B3A0FFD59FEC6BD4DD205" box="[692,806,295,320]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" metricValueMax="5.842" metricValueMin="4.318" pageId="3" pageNumber="131" unit="in" value="2.0" valueMax="2.3" valueMin="1.7">1.7–2.3 in</quantity>
males) and regularly curved ventrad with large hyaline portion, spicules 48–55 µm long and 7–9 irregularly spaced ventromedian supplements lacking hiatus.
</paragraph>
<paragraph id="78794207FF8B3A0FFF2AFE81BF93D178" blockId="3.[151,1436,151,573]" pageId="3" pageNumber="131">
<emphasis id="4AB29E15FF8B3A0FFF2AFE81BF1BD2CD" bold="true" box="[199,368,367,392]" pageId="3" pageNumber="131">Relationships.</emphasis>
<taxonomicName id="BFC63984FF8B3A0FFE95FE9EBCF1D2CD" authority="Kazemi &amp; Niknam &amp; Jabbari &amp; Peña-Santiago, 2018" authorityName="Kazemi &amp; Niknam &amp; Jabbari &amp; Peña-Santiago" authorityYear="2018" box="[376,666,368,392]" class="Adenophorea" family="Qudsianematidae" genus="Eudorylaimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="3" pageNumber="131" phylum="Nematoda" rank="species" species="kahaqensis" status="sp. nov.">
<emphasis id="4AB29E15FF8B3A0FFE95FE9EBCF1D2CD" box="[376,666,368,392]" italics="true" pageId="3" pageNumber="131">Eudorylaimus kahaqensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5181236EFF8B3A0FFD4FFE9EBCB4D2C2" box="[674,735,368,391]" pageId="3" pageNumber="131" rank="species">sp. n.</taxonomicNameLabel>
is easily recognizable by an unusual combination of features, but its conical, somewhat elongate, caudal region in both sexes and the position of its most posterior ventromedian supplement at the level of the anterior end of the spicules and not too far anterior to the pre-cloacal pair (
<emphasis id="4AB29E15FF8B3A0FFA84FE56BBFFD295" box="[1385,1428,440,464]" italics="true" pageId="3" pageNumber="131">i. e.</emphasis>
, lacking a distinct hiatus) raise doubts about its true generic identity (see further discussion below), since it possesses characteristics of the genera
<taxonomicName id="BFC63984FF8B3A0FFD9BFDEEBDB7D15D" authority="Andrassy, 1959" authorityName="Andrassy" authorityYear="1959" box="[630,988,511,536]" class="Adenophorea" family="Qudsianematidae" genus="Eudorylaimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="3" pageNumber="131" phylum="Nematoda" rank="genus">
<emphasis id="4AB29E15FF8B3A0FFD9BFDEEBD78D15D" box="[630,787,512,536]" italics="true" pageId="3" pageNumber="131">Eudorylaimus</emphasis>
<bibRefCitation id="1C573FF6FF8B3A0FFCCEFE11BDB7D15D" author="Andrassy" box="[803,988,511,536]" pageId="3" pageNumber="131" refString="Andrassy, I. (1959) Taxonomische Ubersicht der Dorylaimen (Nematoda). I. Acta Zoologica Universitatis Budapestinensis, 5, 191 - 240." type="journal article" year="1959">Andrássy, 1959</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="BFC63984FF8B3A0FFC1DFDEEBB0FD15D" authority="Andrassy, 1986" authorityName="Andrassy" authorityYear="1986" box="[1008,1380,511,536]" class="Adenophorea" family="Qudsianematidae" genus="Allodorylaimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="3" pageNumber="131" phylum="Nematoda" rank="genus">
<emphasis id="4AB29E15FF8B3A0FFC1DFDEEBAF6D15D" box="[1008,1181,512,536]" italics="true" pageId="3" pageNumber="131">Allodorylaimus</emphasis>
<bibRefCitation id="1C573FF6FF8B3A0FFB40FE11BB0FD15D" author="Andrassy" box="[1197,1380,511,536]" pageId="3" pageNumber="131" refString="Andrassy, I. (1986) The genus Eudorylaimus Andrassy, 1959 and the present status of its species (Nematoda: Qudsianematidae). Opuscula Zoologica Budapestinensis, 22, 1 - 42." type="journal article" year="1986">Andrássy, 1986</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="BFC63984FF8B3A0FFF7AFDCABF9FD178" authority="Andrassy, 1986" authorityName="Andrassy" authorityYear="1986" box="[151,500,548,573]" class="Adenophorea" family="Qudsianematidae" genus="Epidorylaimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="3" pageNumber="131" phylum="Nematoda" rank="genus">
<emphasis id="4AB29E15FF8B3A0FFF7AFDCABF57D179" box="[151,316,548,572]" italics="true" pageId="3" pageNumber="131">Epidorylaimus</emphasis>
<bibRefCitation id="1C573FF6FF8B3A0FFEAEFDCABF9FD178" author="Andrassy" box="[323,500,548,573]" pageId="3" pageNumber="131" refString="Andrassy, I. (1986) The genus Eudorylaimus Andrassy, 1959 and the present status of its species (Nematoda: Qudsianematidae). Opuscula Zoologica Budapestinensis, 22, 1 - 42." type="journal article" year="1986">Andrássy, 1986</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<caption id="2CB9128FFF8B3A0FFF7AF87CBDEAD482" httpUri="https://zenodo.org/record/1135630/files/figure.png" pageId="3" pageNumber="131" targetBox="[364,1228,639,1891]" targetPageId="3">
<paragraph id="78794207FF8B3A0FFF7AF87CBDEAD482" blockId="3.[151,1436,1938,1991]" pageId="3" pageNumber="131">
<emphasis id="4AB29E15FF8B3A0FFF7AF87CBD60D4ED" bold="true" box="[151,779,1938,1960]" pageId="3" pageNumber="131">
FIGURE 1.
<taxonomicName id="BFC63984FF8B3A0FFEF7F87CBC42D4ED" authority="Kazemi &amp; Niknam &amp; Jabbari &amp; Peña-Santiago, 2018" authorityName="Kazemi &amp; Niknam &amp; Jabbari &amp; Peña-Santiago" authorityYear="2018" box="[282,553,1938,1960]" class="Adenophorea" family="Qudsianematidae" genus="Eudorylaimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="3" pageNumber="131" phylum="Nematoda" rank="species" species="kahaqensis" status="sp. nov.">
<emphasis id="4AB29E15FF8B3A0FFEF7F87CBC42D4ED" bold="true" box="[282,553,1938,1960]" italics="true" pageId="3" pageNumber="131">Eudorylaimus kahaqensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5181236EFF8B3A0FFDC2F87CBC02D4ED" box="[559,617,1938,1960]" pageId="3" pageNumber="131" rank="species">sp. n.</taxonomicNameLabel>
(Female, LM).
</emphasis>
A: Entire. B: Lip region in surface lateral VieW. C: Lip region in median lateral VieW. D: Anterior genital branch. E: Posterior body region.
</paragraph>
</caption>
<caption id="2CB9128FFF8C3A08FF7AFA36BF86D548" httpUri="https://zenodo.org/record/1135632/files/figure.png" pageId="4" pageNumber="132" targetBox="[374,1225,210,1449]" targetPageId="4">
<paragraph id="78794207FF8C3A08FF7AFA36BF86D548" blockId="4.[151,1436,1496,1549]" pageId="4" pageNumber="132">
<emphasis id="4AB29E15FF8C3A08FF7AFA36BCA9D6AB" bold="true" box="[151,706,1496,1518]" pageId="4" pageNumber="132">
FIGURE 2.
<taxonomicName id="BFC63984FF8C3A08FEF6FA36BC40D6AB" authority="Kazemi &amp; Niknam &amp; Jabbari &amp; Peña-Santiago, 2018" authorityName="Kazemi &amp; Niknam &amp; Jabbari &amp; Peña-Santiago" authorityYear="2018" box="[283,555,1496,1518]" class="Adenophorea" family="Qudsianematidae" genus="Eudorylaimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="4" pageNumber="132" phylum="Nematoda" rank="species" species="kahaqensis" status="sp. nov.">
<emphasis id="4AB29E15FF8C3A08FEF6FA36BC40D6AB" bold="true" box="[283,555,1496,1518]" italics="true" pageId="4" pageNumber="132">Eudorylaimus kahaqensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5181236EFF8C3A08FDDFFA36BC07D6AB" box="[562,620,1496,1518]" pageId="4" pageNumber="132" rank="species">sp. n.</taxonomicNameLabel>
(Male).
</emphasis>
A: Neck, posterior region. B: Entire. C,D: Posterior body region. E, F: Lateral guiding piece. G: Spicule.
</paragraph>
</caption>
<subSubSection id="30DC118CFF8C3A09FF2AF9DABDBCD4B7" lastPageId="5" lastPageNumber="133" pageId="4" pageNumber="132" type="reference_group">
<paragraph id="78794207FF8C3A09FF2AF9DABF9FD2CD" blockId="4.[151,1436,1588,2009]" lastBlockId="5.[151,1436,151,392]" lastPageId="5" lastPageNumber="133" pageId="4" pageNumber="132">
Within
<taxonomicName id="BFC63984FF8C3A08FECDF9DBBFD6D508" authorityName="Andrassy" authorityYear="1959" box="[288,445,1589,1613]" class="Adenophorea" family="Qudsianematidae" genus="Eudorylaimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="4" pageNumber="132" phylum="Nematoda" rank="genus">
<emphasis id="4AB29E15FF8C3A08FECDF9DBBFD6D508" box="[288,445,1589,1613]" italics="true" pageId="4" pageNumber="132">Eudorylaimus</emphasis>
</taxonomicName>
, it resembles several species (some without known males), in particular
<taxonomicName id="BFC63984FF8C3A08FACFF9DBBFC5D534" authority="Tjepkema et al., (1971)" authorityName="Tjepkema et al." authorityYear="1971" class="Adenophorea" family="Qudsianematidae" genus="Eudorylaimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="4" pageNumber="132" phylum="Nematoda" rank="species" species="altherri">
<emphasis id="4AB29E15FF8C3A08FACFF9DBBBF0D508" box="[1314,1435,1589,1613]" italics="true" pageId="4" pageNumber="132">E. altherri</emphasis>
<bibRefCitation id="1C573FF6FF8C3A08FF7AF9B6BFC5D534" box="[151,430,1624,1649]" pageId="4" pageNumber="132" refString="Tjepkema, J. P., Ferris, V. R. &amp; Ferris, J. M. (1971) ReVieW of the genus Aporcelaimellus Heyns, 1965 and six species groups of the genus Eudorylaimus Andrassy, 1959 (Nematoda: Dorylaimida). Purdue University Agriculture Experimental Station Research Bulletin, 882, 52 pp." type="book">
Tjepkema
<emphasis id="4AB29E15FF8C3A08FEFFF9B7BF24D535" box="[274,335,1624,1648]" italics="true" pageId="4" pageNumber="132">et al.</emphasis>
, (1971)
</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="BFC63984FF8C3A08FE53F9B7BD7ED534" authority="Heyns, 1993" authorityName="Heyns" authorityYear="1993" box="[446,789,1624,1649]" class="Adenophorea" family="Qudsianematidae" genus="Eudorylaimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="4" pageNumber="132" phylum="Nematoda" rank="species" species="nudicaudatus">
<emphasis id="4AB29E15FF8C3A08FE53F9B7BC13D535" box="[446,632,1624,1648]" italics="true" pageId="4" pageNumber="132">E. nudicaudatus</emphasis>
<bibRefCitation id="1C573FF6FF8C3A08FD6EF9B7BD7ED534" author="Heyns" box="[643,789,1625,1649]" pageId="4" pageNumber="132" refString="Heyns, J. (1993) Eudorylaimus nudicaudatus sp. n. from Antarctica (Nematoda: Dorylaimoidea). South African Journal of Antarctic Research, 23, 33 - 36." type="journal article" year="1993">Heyns, 1993</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="BFC63984FF8C3A08FCB9F9B7BAA8D534" authority="Khan &amp; Araki, 2000" authorityName="Khan &amp; Araki" authorityYear="2000" box="[852,1219,1624,1649]" class="Adenophorea" family="Qudsianematidae" genus="Eudorylaimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="4" pageNumber="132" phylum="Nematoda" rank="species" species="unicus">
<emphasis id="4AB29E15FF8C3A08FCB9F9B7BDAAD535" box="[852,961,1625,1648]" italics="true" pageId="4" pageNumber="132">E. unicus</emphasis>
<bibRefCitation id="1C573FF6FF8C3A08FC21F9B6BAA8D534" author="Khan" box="[972,1219,1624,1649]" pageId="4" pageNumber="132" refString="Khan, Z. &amp; Araki, M. (2000) Eudorylaimus unicus sp. n., E. japonicus sp. n. and E. fransus Heyns, (Nematoda: Qudsianematidae) from Tsukuba, Japan. International Journal of Nematology, 10, 135 - 142." type="journal article" year="2000">Khan &amp; Araki, 2000</bibRefCitation>
</taxonomicName>
. It differs from
<taxonomicName id="BFC63984FF8C3A08FA69F9B7BE86D5D0" class="Adenophorea" family="Qudsianematidae" genus="Eudorylaimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="4" pageNumber="132" phylum="Nematoda" rank="species" species="altherri">
<emphasis id="4AB29E15FF8C3A08FA69F9B7BE86D5D0" italics="true" pageId="4" pageNumber="132">E. altherri</emphasis>
</taxonomicName>
in its nearly continuous (
<emphasis id="4AB29E15FF8C3A08FDF7F993BC5AD5D1" box="[538,561,1661,1684]" italics="true" pageId="4" pageNumber="132">vs</emphasis>
offset by constriction and visibly wider than the adjacent body) lip region, longer odontostyle (21–23
<emphasis id="4AB29E15FF8C3A08FE21F94FBF88D5FD" box="[460,483,1697,1720]" italics="true" pageId="4" pageNumber="132">vs</emphasis>
19.3 ± 1.2 µm), shorter female prerectum (43–50
<emphasis id="4AB29E15FF8C3A08FBC6F94FBA29D5FD" box="[1067,1090,1697,1720]" italics="true" pageId="4" pageNumber="132">vs</emphasis>
90 ± 22 µm),
<emphasis id="4AB29E15FF8C3A08FB02F94FBE9BD599" italics="true" pageId="4" pageNumber="132">pars refringens vaginae</emphasis>
with trapezoidal (
<emphasis id="4AB29E15FF8C3A08FE23F92BBF8ED599" box="[462,485,1733,1756]" italics="true" pageId="4" pageNumber="132">vs</emphasis>
triangular) pieces, and male as frequent as female (
<emphasis id="4AB29E15FF8C3A08FB8DF92BBA1CD599" box="[1120,1143,1733,1756]" italics="true" pageId="4" pageNumber="132">vs</emphasis>
male absent). From
<taxonomicName id="BFC63984FF8C3A08FA69F92BBF45D445" authorityName="Heyns" authorityYear="1993" class="Adenophorea" family="Qudsianematidae" genus="Eudorylaimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="4" pageNumber="132" phylum="Nematoda" rank="species" species="nudicaudatus">
<emphasis id="4AB29E15FF8C3A08FA69F92BBF45D445" italics="true" pageId="4" pageNumber="132">E. nudicaudatus</emphasis>
</taxonomicName>
, it differs in its nearly continuous (
<emphasis id="4AB29E15FF8C3A08FD51F907BCB8D445" box="[700,723,1769,1792]" italics="true" pageId="4" pageNumber="132">vs</emphasis>
offset by constriction) lip region, longer odontostyle (21–23
<emphasis id="4AB29E15FF8C3A08FA69F907BBF0D445" box="[1412,1435,1769,1792]" italics="true" pageId="4" pageNumber="132">vs</emphasis>
13–15 µm, equal to
<emphasis id="4AB29E15FF8C3A08FE94F8E3BFFBD461" box="[377,400,1805,1828]" italics="true" pageId="4" pageNumber="132">vs</emphasis>
distinctly shorter than lip region diameter), more posterior vulva (
<emphasis id="4AB29E15FF8C3A08FB96F8E3BAE7D461" box="[1147,1164,1805,1828]" italics="true" pageId="4" pageNumber="132">V</emphasis>
= 48–52
<emphasis id="4AB29E15FF8C3A08FB15F8E3BB4CD461" box="[1272,1319,1805,1828]" italics="true" pageId="4" pageNumber="132">vs V</emphasis>
= 43–48), comparatively longer female tail (
<emphasis id="4AB29E15FF8C3A08FDCDF8DFBC47D40D" box="[544,556,1841,1864]" italics="true" pageId="4" pageNumber="132">c</emphasis>
= 24–30
<emphasis id="4AB29E15FF8C3A08FD4DF8DFBCA5D40D" box="[672,718,1841,1864]" italics="true" pageId="4" pageNumber="132">vs c</emphasis>
= 32–46) with long (
<emphasis id="4AB29E15FF8C3A08FC21F8DFBD88D40D" box="[972,995,1841,1864]" italics="true" pageId="4" pageNumber="132">vs</emphasis>
very short) hyaline portion, and male lacking (
<emphasis id="4AB29E15FF8C3A08FF14F8BBBF7BD429" box="[249,272,1877,1900]" italics="true" pageId="4" pageNumber="132">vs</emphasis>
having) hiatus. It can be distinguished from
<taxonomicName id="BFC63984FF8C3A08FCECF8BBBD03D429" authorityName="Khan &amp; Araki" authorityYear="2000" box="[769,872,1877,1900]" class="Adenophorea" family="Qudsianematidae" genus="Eudorylaimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="4" pageNumber="132" phylum="Nematoda" rank="species" species="unicus">
<emphasis id="4AB29E15FF8C3A08FCECF8BBBD03D429" box="[769,872,1877,1900]" italics="true" pageId="4" pageNumber="132">E. unicus</emphasis>
</taxonomicName>
by its nearly continuous (
<emphasis id="4AB29E15FF8C3A08FB64F8BBBACBD429" box="[1161,1184,1877,1900]" italics="true" pageId="4" pageNumber="132">vs</emphasis>
offset by constriction) lip region, more posterior DN (
<emphasis id="4AB29E15FF8C3A08FE10F897BC7FD4D5" box="[509,532,1913,1936]" italics="true" pageId="4" pageNumber="132">vs</emphasis>
close the pharyngeal expansion), longer vagina (extending to 63–70%
<emphasis id="4AB29E15FF8C3A08FAAFF897BB32D4D5" box="[1346,1369,1913,1936]" italics="true" pageId="4" pageNumber="132">vs</emphasis>
up to one-half of body diameter), longer female tail (
<emphasis id="4AB29E15FF8C3A08FD48F873BCD0D4F1" box="[677,699,1949,1972]" italics="true" pageId="4" pageNumber="132">c’</emphasis>
= 1.9–2.6
<emphasis id="4AB29E15FF8C3A08FCDEF873BD0CD4F1" box="[819,871,1949,1972]" italics="true" pageId="4" pageNumber="132">vs c’</emphasis>
= 1.6–1.8) with different morphology (ending in a finely rounded
<emphasis id="4AB29E15FF8C3A08FEA9F82FBF30D49D" box="[324,347,1985,2008]" italics="true" pageId="4" pageNumber="132">vs</emphasis>
relatively coarse tip), and male present (
<emphasis id="4AB29E15FF8C3A08FCC1F82FBD28D49D" box="[812,835,1985,2008]" italics="true" pageId="4" pageNumber="132">vs</emphasis>
absent). The new species also resembles
<taxonomicName id="BFC63984FF8C3A09FACEF82FBC70D3F5" authority="(Andrassy, 1957) Andrassy, 1959" authorityName="(Andrassy, 1957) Andrassy" authorityYear="1959" baseAuthorityName="Andrassy" baseAuthorityYear="1957" class="Adenophorea" family="Qudsianematidae" genus="Eudorylaimus" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="5" lastPageNumber="133" order="Dorylaimida" pageId="4" pageNumber="132" phylum="Nematoda" rank="species" species="rugosus">
<emphasis id="4AB29E15FF8C3A08FACEF82FBBF7D49D" box="[1315,1436,1985,2008]" italics="true" pageId="4" pageNumber="132">E. rugosus</emphasis>
(
<bibRefCitation id="1C573FF6FF8D3A09FF73FF79BF3FD3F5" author="Andrassy" box="[158,340,151,176]" pageId="5" pageNumber="133" refString="Andrassy, I. (1957) Dorylaimus rugosus n. sp., ein neuer Nematode aus Ungarn. Nematologischen NotiZen 7. Opuscula Zoologica Instituti Zoosystematici Universitatis Budapestinensis, 2, 9 - 11." type="journal article" year="1957">Andrássy, 1957</bibRefCitation>
)
<bibRefCitation id="1C573FF6FF8D3A09FE85FF79BC70D3F5" author="Andrassy" box="[360,539,151,176]" pageId="5" pageNumber="133" refString="Andrassy, I. (1959) Taxonomische Ubersicht der Dorylaimen (Nematoda). I. Acta Zoologica Universitatis Budapestinensis, 5, 191 - 240." type="journal article" year="1959">
Andrássy,
<quantity id="BF3EEFE2FF8D3A09FE0EFF79BC70D3F5" box="[483,539,151,176]" metricMagnitude="1" metricUnit="m" metricValue="4.975859999999999" pageId="5" pageNumber="133" unit="in" value="1959.0">1959</quantity>
</bibRefCitation>
</taxonomicName>
in several aspects (general morphology and morphometry, lip region nearly truncate and visibly slender odontostyle) but differs in its longer tail (
<emphasis id="4AB29E15FF8D3A09FC5EFF53BDD4D391" box="[947,959,189,212]" italics="true" pageId="5" pageNumber="133">c</emphasis>
= 24–30,
<emphasis id="4AB29E15FF8D3A09FBDAFF52BA26D396" box="[1079,1101,188,211]" italics="true" pageId="5" pageNumber="133">c’</emphasis>
= 1.9–2.6
<emphasis id="4AB29E15FF8D3A09FB27FF53BA9DD391" box="[1226,1270,189,212]" italics="true" pageId="5" pageNumber="133">vs c</emphasis>
= 30–33,
<emphasis id="4AB29E15FF8D3A09FA80FF52BBE8D396" box="[1389,1411,188,211]" italics="true" pageId="5" pageNumber="133">c’</emphasis>
= 1.5–1.6). It can also be compared to
<taxonomicName id="BFC63984FF8D3A09FDD2FF0FBD13D3BD" authority="Zell, 1986" authorityName="Zell" authorityYear="1986" box="[575,888,223,248]" class="Adenophorea" family="Qudsianematidae" genus="Eudorylaimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="5" pageNumber="133" phylum="Nematoda" rank="species" species="pseudobokori">
<emphasis id="4AB29E15FF8D3A09FDD2FF0FBC92D3BD" box="[575,761,224,248]" italics="true" pageId="5" pageNumber="133">E. pseudobokori</emphasis>
<bibRefCitation id="1C573FF6FF8D3A09FCEFFF31BD13D3BD" author="Zell" box="[770,888,223,248]" pageId="5" pageNumber="133" refString="Zell, H. (1986) Nematoden eines Buchenealbodens. 6. Die Dorylaimen (Nematoda: Dorylaimida). Carolinea, 44, 91 - 118." type="journal article" year="1986">
Zell,
<quantity id="BF3EEFE2FF8D3A09FCADFF31BD13D3BD" box="[832,888,223,248]" metricMagnitude="1" metricUnit="m" metricValue="5.04444" pageId="5" pageNumber="133" unit="in" value="1986.0">1986</quantity>
</bibRefCitation>
</taxonomicName>
in its general morphology and morphometry as well its comparatively long caudal region, but can be distinguished by the morphology of lip region (truncate and continuous
<emphasis id="4AB29E15FF8D3A09FEF3FEC7BF5ED205" box="[286,309,297,320]" italics="true" pageId="5" pageNumber="133">vs</emphasis>
angular and offset) and odontostyle (slender
<emphasis id="4AB29E15FF8D3A09FCB5FEC7BD04D205" box="[856,879,297,320]" italics="true" pageId="5" pageNumber="133">vs</emphasis>
more robust), and the number (7–9
<emphasis id="4AB29E15FF8D3A09FAC1FEC7BB28D205" box="[1324,1347,297,320]" italics="true" pageId="5" pageNumber="133">vs</emphasis>
5) and arrangement of ventromedian supplements (most posterior of them at level of or slightly posterior to
<emphasis id="4AB29E15FF8D3A09FAC3FEA3BB2ED221" box="[1326,1349,333,356]" italics="true" pageId="5" pageNumber="133">vs</emphasis>
visibly anterior to the end of spicules).
</paragraph>
<caption id="2CB9128FFF8D3A09FF7AFA18BF42D50F" httpUri="https://zenodo.org/record/1135634/files/figure.png" pageId="5" pageNumber="133" targetBox="[344,1244,451,1501]" targetPageId="5">
<paragraph id="78794207FF8D3A09FF7AFA18BF42D50F" blockId="5.[151,1436,1526,1610]" pageId="5" pageNumber="133">
<emphasis id="4AB29E15FF8D3A09FF7AFA18BD7FD549" bold="true" box="[151,788,1526,1548]" pageId="5" pageNumber="133">
FIGURE 3.
<taxonomicName id="BFC63984FF8D3A09FEF1FA18BC46D549" authority="Kazemi &amp; Niknam &amp; Jabbari &amp; Peña-Santiago, 2018" authorityName="Kazemi &amp; Niknam &amp; Jabbari &amp; Peña-Santiago" authorityYear="2018" box="[284,557,1526,1548]" class="Adenophorea" family="Qudsianematidae" genus="Eudorylaimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="5" pageNumber="133" phylum="Nematoda" rank="species" species="kahaqensis" status="sp. nov.">
<emphasis id="4AB29E15FF8D3A09FEF1FA18BC46D549" bold="true" box="[284,557,1526,1548]" italics="true" pageId="5" pageNumber="133">Eudorylaimus kahaqensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5181236EFF8D3A09FDD9FA18BC04D549" box="[564,623,1526,1548]" pageId="5" pageNumber="133" rank="species">sp. n.</taxonomicNameLabel>
(Female, LM).
</emphasis>
A, B: Entire. C, D: Anterior genital branch. E: Posterior body region. F: Caudal region. G: OViduct-uterus junction. H, I: Vagina. (Scale bars: A, B = 200 µm; C, D = 50 µm; E–G = 20 µm; H, I = 10 µm.)
</paragraph>
</caption>
<paragraph id="78794207FF8D3A09FF2AF99CBDBCD4B7" blockId="5.[151,1436,1649,2034]" pageId="5" pageNumber="133">
In lacking a distinct precloacal space in males, the new species also resembles members of
<taxonomicName id="BFC63984FF8D3A09FB05F99FBBFFD5CC" authorityName="Andrassy" authorityYear="1986" box="[1256,1428,1649,1673]" class="Adenophorea" family="Qudsianematidae" genus="Allodorylaimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="5" pageNumber="133" phylum="Nematoda" rank="genus">
<emphasis id="4AB29E15FF8D3A09FB05F99FBBFFD5CC" box="[1256,1428,1649,1673]" italics="true" pageId="5" pageNumber="133">Allodorylaimus</emphasis>
</taxonomicName>
, especially
<taxonomicName id="BFC63984FF8D3A09FEE3F97BBCC5D5E8" authority="Quijano et al., (1991)" authorityName="Quijano et al." authorityYear="1991" box="[270,686,1684,1709]" class="Adenophorea" family="Qudsianematidae" genus="Allodorylaimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="5" pageNumber="133" phylum="Nematoda" rank="species" species="aljabaranus">
<emphasis id="4AB29E15FF8D3A09FEE3F97BBFDFD5E8" box="[270,436,1685,1709]" italics="true" pageId="5" pageNumber="133">A. aljabaranus</emphasis>
<bibRefCitation id="1C573FF6FF8D3A09FE51F97ABCC5D5E8" box="[444,686,1684,1709]" pageId="5" pageNumber="133" refString="Quijano, R., Pena-Santiago, R. &amp; JimeneZ-Guirado, D. (1991) Three neW species of the genus Allodorylaimus Andrassy, 1986 (Nematoda: Dorylaimida) from Spain. Afro-Asian Journal of Nematology, 1, 161 - 173." type="journal article">
Quijano
<emphasis id="4AB29E15FF8D3A09FDF0F978BC3DD5E8" box="[541,598,1685,1709]" italics="true" pageId="5" pageNumber="133">et al.</emphasis>
, (1991)
</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="BFC63984FF8D3A09FD08F97BBAB8D5E8" authority="(Thorne, 1939) Andrassy, 1986" authorityName="Andrassy" authorityYear="1986" baseAuthorityName="Thorne" baseAuthorityYear="1939" box="[741,1235,1684,1709]" class="Adenophorea" family="Qudsianematidae" genus="Allodorylaimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="5" pageNumber="133" phylum="Nematoda" rank="species" species="digiturus">
<emphasis id="4AB29E15FF8D3A09FD08F97BBD02D5E8" box="[741,873,1685,1709]" italics="true" pageId="5" pageNumber="133">A. digiturus</emphasis>
(
<bibRefCitation id="1C573FF6FF8D3A09FC95F97ABA79D5E8" author="Thorne" box="[888,1042,1684,1709]" pageId="5" pageNumber="133" refString="Thorne, G. (1939) A monograph of the nematodes of the superfamily Dorylaimoidea. Capita Zoologica, 8, 1 - 261." type="journal article" year="1939">Thorne, 1939</bibRefCitation>
)
<bibRefCitation id="1C573FF6FF8D3A09FBCFF97ABAB8D5E8" author="Andrassy" box="[1058,1235,1684,1709]" pageId="5" pageNumber="133" refString="Andrassy, I. (1986) The genus Eudorylaimus Andrassy, 1959 and the present status of its species (Nematoda: Qudsianematidae). Opuscula Zoologica Budapestinensis, 22, 1 - 42." type="journal article" year="1986">Andrássy, 1986</bibRefCitation>
</taxonomicName>
. It differs from
<taxonomicName id="BFC63984FF8D3A09FA69F97BBF75D594" class="Adenophorea" family="Qudsianematidae" genus="Allodorylaimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="5" pageNumber="133" phylum="Nematoda" rank="species" species="aljabaranus">
<emphasis id="4AB29E15FF8D3A09FA69F97BBF75D594" italics="true" pageId="5" pageNumber="133">A. aljabaranus</emphasis>
</taxonomicName>
in its broader (17.0–21
<emphasis id="4AB29E15FF8D3A09FDC0F954BC2FD594" box="[557,580,1722,1745]" italics="true" pageId="5" pageNumber="133">vs</emphasis>
14–16 µm) and amalgamated (
<emphasis id="4AB29E15FF8D3A09FC48F954BDD7D594" box="[933,956,1722,1745]" italics="true" pageId="5" pageNumber="133">vs</emphasis>
lips separated and angular) lip region, and fewer ventromedian supplements (7–9
<emphasis id="4AB29E15FF8D3A09FDA7F930BC0AD5B0" box="[586,609,1758,1781]" italics="true" pageId="5" pageNumber="133">vs</emphasis>
10–15) with different arrangement (only the most posterior ventromedian supplement located at the level of the anterior end of the spicules or a short distance posterior
<emphasis id="4AB29E15FF8D3A09FB01F8ECBB68D45C" box="[1260,1283,1794,1817]" italics="true" pageId="5" pageNumber="133">vs</emphasis>
two or three ventromedian supplements well within the range of the spicules). From
<taxonomicName id="BFC63984FF8D3A09FC06F8CBBA18D478" authorityName="Andrassy" authorityYear="1986" baseAuthorityName="Thorne" baseAuthorityYear="1939" box="[1003,1139,1829,1853]" class="Adenophorea" family="Qudsianematidae" genus="Allodorylaimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="5" pageNumber="133" phylum="Nematoda" rank="species" species="digiturus">
<emphasis id="4AB29E15FF8D3A09FC06F8CBBA18D478" box="[1003,1139,1829,1853]" italics="true" pageId="5" pageNumber="133">A. digiturus</emphasis>
</taxonomicName>
, a poorly known species described on the basis of only one male, it differs in its larger general size (body length 1.40–1.75
<emphasis id="4AB29E15FF8D3A09FAEBF8A4BB76D424" box="[1286,1309,1866,1889]" italics="true" pageId="5" pageNumber="133">vs</emphasis>
<quantity id="BF3EEFE2FF8D3A09FACBF8A7BBFBD424" box="[1318,1424,1865,1889]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="5" pageNumber="133" unit="mm" value="1.3">1.30 mm</quantity>
), broader lip region (17.0–21
<emphasis id="4AB29E15FF8D3A09FE22F880BC6CD4C0" box="[463,519,1902,1925]" italics="true" pageId="5" pageNumber="133">vs ca</emphasis>
12 µm), longer (21–23
<emphasis id="4AB29E15FF8D3A09FCFCF880BD22D4C0" box="[785,841,1902,1925]" italics="true" pageId="5" pageNumber="133">vs ca</emphasis>
12 µm) and more slender (9–12
<emphasis id="4AB29E15FF8D3A09FB5BF880BA84D4C0" box="[1206,1263,1902,1925]" italics="true" pageId="5" pageNumber="133">vs ca</emphasis>
6 times as long as wide) odontostyle, and fewer ventromedian supplements (7–9
<emphasis id="4AB29E15FF8D3A09FC95F87CBDE4D4EC" box="[888,911,1938,1961]" italics="true" pageId="5" pageNumber="133">vs</emphasis>
11) with different arrangement (only the most posterior ventromedian supplement located at the level of the anterior end of the spicules or a short distance behind it
<emphasis id="4AB29E15FF8D3A09FF40F834BEAFD4B4" box="[173,196,2010,2033]" italics="true" pageId="5" pageNumber="133">vs</emphasis>
two ventromedian supplements well within the range of the spicules).
</paragraph>
</subSubSection>
<subSubSection id="30DC118CFF8E3A0AFF2AFF76BAD7D1C0" pageId="6" pageNumber="134" type="description">
<paragraph id="78794207FF8E3A0AFF2AFF76BF7BD295" blockId="6.[151,1436,151,645]" pageId="6" pageNumber="134">
In its general morphology, and in particular in its comparatively long tail in both females and males, and males lacking hiatus, the new species resembles some
<taxonomicName id="BFC63984FF8E3A0AFD3DFF52BD1ED391" authorityName="Andrassy" authorityYear="1986" box="[720,885,188,212]" class="Adenophorea" family="Qudsianematidae" genus="Epidorylaimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="6" pageNumber="134" phylum="Nematoda" rank="genus">
<emphasis id="4AB29E15FF8E3A0AFD3DFF52BD1ED391" box="[720,885,188,212]" italics="true" pageId="6" pageNumber="134">Epidorylaimus</emphasis>
</taxonomicName>
representatives (see the updated diagnosis and compendium by
<bibRefCitation id="1C573FF6FF8E3A0AFEBFFF31BC47D3BD" author="Ahmad" box="[338,556,223,248]" pageId="6" pageNumber="134" refString="Ahmad, W., Imran, Z. &amp; Araki, M. (2016) Epidorylaimus monhystera sp. n., an atypical species of the genus Epidorylaimus Andrassy, 1986 (Dorylaimida: Qudsianematidae) from Japan. Zootaxa, 4072 (1), 90 - 100. https: // doi. org / 10.11646 / Zootaxa. 4072.1.4" type="journal article" year="2016">
Ahmad
<emphasis id="4AB29E15FF8E3A0AFE46FF0FBF88D3BD" box="[427,483,224,248]" italics="true" pageId="6" pageNumber="134">et al.</emphasis>
, 2016
</bibRefCitation>
), especially
<taxonomicName id="BFC63984FF8E3A0AFD58FF0FBAAFD3BD" authority="(de Man, 1880) Andrassy, 1986" authorityName="Andrassy" authorityYear="1986" baseAuthorityName="de Man" baseAuthorityYear="1880" box="[693,1220,223,248]" class="Adenophorea" family="Qudsianematidae" genus="Epidorylaimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="6" pageNumber="134" phylum="Nematoda" rank="species" species="consobrinus">
<emphasis id="4AB29E15FF8E3A0AFD58FF0FBD36D3BD" box="[693,861,224,248]" italics="true" pageId="6" pageNumber="134">E. consobrinus</emphasis>
(de
<bibRefCitation id="1C573FF6FF8E3A0AFC60FF0EBA6CD3BD" author="Man" box="[909,1031,224,248]" pageId="6" pageNumber="134" refString="de Man, J. G. (1880) Die einheimischen, frei in der reinen Erde und im sussen Wasser lebenden Nematoden. Tijdschrift Nederlandsche dierkundige Vereeniging, 5, 1 - 104." type="journal article" year="1880">Man, 1880</bibRefCitation>
)
<bibRefCitation id="1C573FF6FF8E3A0AFBFBFF31BAAFD3BD" author="Andrassy" box="[1046,1220,223,248]" pageId="6" pageNumber="134" refString="Andrassy, I. (1986) The genus Eudorylaimus Andrassy, 1959 and the present status of its species (Nematoda: Qudsianematidae). Opuscula Zoologica Budapestinensis, 22, 1 - 42." type="journal article" year="1986">Andrássy, 1986</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="BFC63984FF8E3A0AFB14FF0FBC00D258" authority="(Tjepkema et al., 1971) Andrassy, 1986" authorityName="Andrassy" authorityYear="1986" baseAuthorityName="Tjepkema et al." baseAuthorityYear="1971" class="Adenophorea" family="Qudsianematidae" genus="Epidorylaimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="6" pageNumber="134" phylum="Nematoda" rank="species" species="filicaudatus">
<emphasis id="4AB29E15FF8E3A0AFB14FF0FBBF7D3BD" box="[1273,1436,224,248]" italics="true" pageId="6" pageNumber="134">E. filicaudatus</emphasis>
(
<bibRefCitation id="1C573FF6FF8E3A0AFF73FEEABFC8D259" author="Tjepkema" box="[158,419,260,285]" pageId="6" pageNumber="134" refString="Tjepkema, J. P., Ferris, V. R. &amp; Ferris, J. M. (1971) ReVieW of the genus Aporcelaimellus Heyns, 1965 and six species groups of the genus Eudorylaimus Andrassy, 1959 (Nematoda: Dorylaimida). Purdue University Agriculture Experimental Station Research Bulletin, 882, 52 pp." type="book" year="1971">
Tjepkema
<emphasis id="4AB29E15FF8E3A0AFEF6FEEBBF33D259" box="[283,344,260,284]" italics="true" pageId="6" pageNumber="134">et al.</emphasis>
, 1971
</bibRefCitation>
)
<bibRefCitation id="1C573FF6FF8E3A0AFE5BFEEABC00D258" author="Andrassy" box="[438,619,260,285]" pageId="6" pageNumber="134" refString="Andrassy, I. (1986) The genus Eudorylaimus Andrassy, 1959 and the present status of its species (Nematoda: Qudsianematidae). Opuscula Zoologica Budapestinensis, 22, 1 - 42." type="journal article" year="1986">Andrássy, 1986</bibRefCitation>
</taxonomicName>
. The new species differs from
<taxonomicName id="BFC63984FF8E3A0AFC31FEEBBAE2D259" authorityName="Andrassy" authorityYear="1986" baseAuthorityName="de Man" baseAuthorityYear="1880" box="[988,1161,260,284]" class="Adenophorea" family="Qudsianematidae" genus="Epidorylaimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="6" pageNumber="134" phylum="Nematoda" rank="species" species="consobrinus">
<emphasis id="4AB29E15FF8E3A0AFC31FEEBBAE2D259" box="[988,1161,260,284]" italics="true" pageId="6" pageNumber="134">E. consobrinus</emphasis>
</taxonomicName>
, a poorly characterized species, in its weakly (
<emphasis id="4AB29E15FF8E3A0AFE76FEC7BFD9D205" box="[411,434,297,320]" italics="true" pageId="6" pageNumber="134">vs</emphasis>
distinctly) angular lip region, and shorter caudal region (
<emphasis id="4AB29E15FF8E3A0AFBADFEC6BA3DD27A" box="[1088,1110,296,319]" italics="true" pageId="6" pageNumber="134">c’</emphasis>
= 1.9–2.6
<emphasis id="4AB29E15FF8E3A0AFB3EFEC7BB62D27A" box="[1235,1289,296,320]" italics="true" pageId="6" pageNumber="134">vs c’</emphasis>
=
<quantity id="BF3EEFE2FF8E3A0AFAC7FEC6BBF7D205" box="[1322,1436,295,320]" metricMagnitude="-2" metricUnit="m" metricValue="8.89" metricValueMax="10.921999999999999" metricValueMin="6.858" pageId="6" pageNumber="134" unit="in" value="3.5" valueMax="4.3" valueMin="2.7">2.7–4.3 in</quantity>
females) with different morphology (with finely rounded
<emphasis id="4AB29E15FF8E3A0AFCD0FEA3BD3FD221" box="[829,852,333,356]" italics="true" pageId="6" pageNumber="134">vs</emphasis>
acute tip, large
<emphasis id="4AB29E15FF8E3A0AFBF6FEA3BA59D221" box="[1051,1074,333,356]" italics="true" pageId="6" pageNumber="134">vs</emphasis>
no hyaline portion). It can be distinguished from
<taxonomicName id="BFC63984FF8E3A0AFE9CFE9FBC7FD2CD" authorityName="Andrassy" authorityYear="1986" baseAuthorityName="Tjepkema et al." baseAuthorityYear="1971" box="[369,532,368,392]" class="Adenophorea" family="Qudsianematidae" genus="Epidorylaimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="6" pageNumber="134" phylum="Nematoda" rank="species" species="filicaudatus">
<emphasis id="4AB29E15FF8E3A0AFE9CFE9FBC7FD2CD" box="[369,532,368,392]" italics="true" pageId="6" pageNumber="134">E. filicaudatus</emphasis>
</taxonomicName>
by having weakly (
<emphasis id="4AB29E15FF8E3A0AFD1EFE9FBD61D2CD" box="[755,778,369,392]" italics="true" pageId="6" pageNumber="134">vs</emphasis>
distinctly) angular lip region, more attenuated odontostyle (9–12
<emphasis id="4AB29E15FF8E3A0AFF32FE7BBE9DD2E9" box="[223,246,405,428]" italics="true" pageId="6" pageNumber="134">vs</emphasis>
up to 7 times as long as wide), shorter caudal region (50–73
<emphasis id="4AB29E15FF8E3A0AFC45FE7BBDD4D2E9" box="[936,959,405,428]" italics="true" pageId="6" pageNumber="134">vs</emphasis>
81–123 µm), and frequent males (
<emphasis id="4AB29E15FF8E3A0AFAAAFE7BBB35D2E9" box="[1351,1374,405,428]" italics="true" pageId="6" pageNumber="134">vs</emphasis>
male unknown).
</paragraph>
<paragraph id="78794207FF8E3A0AFF2AFE32BAD7D1C0" blockId="6.[151,1436,151,645]" pageId="6" pageNumber="134">
Finally, some traits (continuous lip region, broad cheilostom) of the new species resemble those found in species of the genus
<emphasis id="4AB29E15FF8E3A0AFE6DFDEEBC66D15D" box="[384,525,512,536]" italics="true" pageId="6" pageNumber="134">Chrysonema</emphasis>
<bibRefCitation id="1C573FF6FF8E3A0AFDF9FE11BCC4D15D" author="Thorne" box="[532,687,511,536]" pageId="6" pageNumber="134" refString="Thorne, G. (1929) Nematodes from the Summit of Long's peak. Transactions of the American Microscopic Society, 48, 181 - 195." type="journal article" year="1929">Thorne, 1929</bibRefCitation>
, but it does not fit at all the very peculiar morphological pattern of this taxon (see
<bibRefCitation id="1C573FF6FF8E3A0AFED3FDCABF84D179" author="Andrassy" box="[318,495,548,573]" pageId="6" pageNumber="134" refString="Andrassy, I. (2009) Free-living nematodes of Hungary. III. PedoZoologica Hungarica nº 5. Hungarian Natural History Museum. Budapest, Hungary, 608 pp." type="book" year="2009">Andrássy, 2009</bibRefCitation>
), which is characterized by, among other diagnostic features, a very slender body (
<emphasis id="4AB29E15FF8E3A0AFA69FDCBBBF9D179" box="[1412,1426,549,572]" italics="true" pageId="6" pageNumber="134">a</emphasis>
- ratio very usually over 50), comparatively small and attenuate odontostyle, absence of
<emphasis id="4AB29E15FF8E3A0AFB8DFDA7BB00D11A" box="[1120,1387,584,608]" italics="true" pageId="6" pageNumber="134">pars refringens vaginae</emphasis>
, tail conical elongate (
<emphasis id="4AB29E15FF8E3A0AFEB3FD82BF1FD1C6" box="[350,372,620,643]" italics="true" pageId="6" pageNumber="134">c’</emphasis>
&gt; 4), tapering very gradually, and male bearing subventral caudal papillae.
</paragraph>
</subSubSection>
<caption id="2CB9128FFF8E3A0AFF7AF8DBBFFED4A0" httpUri="https://zenodo.org/record/1135636/files/figure.png" pageId="6" pageNumber="134" targetBox="[345,1239,705,1816]" targetPageId="6">
<paragraph id="78794207FF8E3A0AFF7AF8DBBFFED4A0" blockId="6.[151,1436,1845,2021]" pageId="6" pageNumber="134">
<emphasis id="4AB29E15FF8E3A0AFF7AF8DBBD9AD40E" bold="true" box="[151,1009,1845,1867]" pageId="6" pageNumber="134">
FIGURE 4.
<taxonomicName id="BFC63984FF8E3A0AFEF3F8DBBC5AD40E" authority="Kazemi &amp; Niknam &amp; Jabbari &amp; Peña-Santiago, 2018" authorityName="Kazemi &amp; Niknam &amp; Jabbari &amp; Peña-Santiago" authorityYear="2018" box="[286,561,1845,1867]" class="Adenophorea" family="Qudsianematidae" genus="Eudorylaimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="6" pageNumber="134" phylum="Nematoda" rank="species" species="kahaqensis" status="sp. nov.">
<emphasis id="4AB29E15FF8E3A0AFEF3F8DBBC5AD40E" bold="true" box="[286,561,1845,1867]" italics="true" pageId="6" pageNumber="134">Eudorylaimus kahaqensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5181236EFF8E3A0AFDD4F8DBBC1ED40E" box="[569,629,1845,1867]" pageId="6" pageNumber="134" rank="species">sp. n.</taxonomicNameLabel>
(Anterior and neck regions, LM).
</emphasis>
A, B: Anterior region in lateral, median VieW. C: Pharyngeal expansion. D: Neck region. E: Lip region and amphid in lateral, surface VieW. F: Coelomocyte a short distance behind the nerVe ring. G: Posterior half of pharyngeal expansion and pharyngo-intestinal junction shoWing S1N2, S2N and dorsal cell mass. H: Posterior part of pharyngeal expansion and pharyngo-intestinal junction shoWing S2N and dorsal cell mass. I, J: Dorsal cell mass at leVel of pharyngo-intestinal junction. (Scale bars: A, B, F, I, J = 10 µm; C = 50 µm; D = 100 µm; E = 5µm; G, H = 20 µm.)
</paragraph>
</caption>
<caption id="2CB9128FFF8F3A0BFF7AFC1DBD38D762" httpUri="https://zenodo.org/record/1135638/files/figure.png" pageId="7" pageNumber="135" targetBox="[344,1244,199,985]" targetPageId="7">
<paragraph id="78794207FF8F3A0BFF7AFC1DBD38D762" blockId="7.[151,1436,1010,1063]" pageId="7" pageNumber="135">
<emphasis id="4AB29E15FF8F3A0BFF7AFC1DBC85D74C" bold="true" box="[151,750,1010,1033]" pageId="7" pageNumber="135">
FIGURE 5.
<taxonomicName id="BFC63984FF8F3A0BFEF5FC1CBC4CD74D" authority="Kazemi &amp; Niknam &amp; Jabbari &amp; Peña-Santiago, 2018" authorityName="Kazemi &amp; Niknam &amp; Jabbari &amp; Peña-Santiago" authorityYear="2018" box="[280,551,1010,1032]" class="Adenophorea" family="Qudsianematidae" genus="Eudorylaimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="7" pageNumber="135" phylum="Nematoda" rank="species" species="kahaqensis" status="sp. nov.">
<emphasis id="4AB29E15FF8F3A0BFEF5FC1CBC4CD74D" bold="true" box="[280,551,1010,1032]" italics="true" pageId="7" pageNumber="135">Eudorylaimus kahaqensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5181236EFF8F3A0BFDC1FC1DBC0ED74C" box="[556,613,1011,1033]" pageId="7" pageNumber="135" rank="species">sp. n.</taxonomicNameLabel>
(Male, LM).
</emphasis>
A: Entire. B–D: Spicules. E, H: Caudal region. F, G: Posterior body region. (Scale bars: A = 200 µm; B–D, E, H = 10 µm; F, G = 50 µm.)
</paragraph>
</caption>
<caption id="2CB9128FFF8F3A0BFF7AFA9ABF30D6EC" httpUri="https://zenodo.org/record/1135640/files/figure.png" pageId="7" pageNumber="135" targetBox="[348,1239,1119,1370]" targetPageId="7">
<paragraph id="78794207FF8F3A0BFF7AFA9ABF30D6EC" blockId="7.[151,1437,1396,1449]" pageId="7" pageNumber="135">
<emphasis id="4AB29E15FF8F3A0BFF7AFA9ABD39D6CF" bold="true" box="[151,850,1396,1418]" pageId="7" pageNumber="135">
FIGURE 6. Lip region of
<taxonomicName id="BFC63984FF8F3A0BFE43FA9ABCD6D6CF" authority="Kazemi &amp; Niknam &amp; Jabbari &amp; Peña-Santiago, 2018" authorityName="Kazemi &amp; Niknam &amp; Jabbari &amp; Peña-Santiago" authorityYear="2018" box="[430,701,1396,1418]" class="Adenophorea" family="Qudsianematidae" genus="Eudorylaimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="7" pageNumber="135" phylum="Nematoda" rank="species" species="kahaqensis" status="sp. nov.">
<emphasis id="4AB29E15FF8F3A0BFE43FA9ABCD6D6CF" bold="true" box="[430,701,1396,1418]" italics="true" pageId="7" pageNumber="135">Eudorylaimus kahaqensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5181236EFF8F3A0BFD29FA9ABC96D6CF" box="[708,765,1396,1418]" pageId="7" pageNumber="135" rank="species">sp. n.</taxonomicNameLabel>
(SEM).
</emphasis>
A: SubVentral VieW. B: Sublateral VieW. C: En face VieW. (Scale bar = 2 µm.)
</paragraph>
</caption>
<subSubSection id="30DC118CFF8F3A0BFF2AFA3BBADFD5E7" pageId="7" pageNumber="135" type="materials_examined">
<paragraph id="78794207FF8F3A0BFF2AFA3BBC79D573" blockId="7.[151,1437,1493,2022]" pageId="7" pageNumber="135">
<materialsCitation id="C8AE485AFF8F3A0BFF2AFA3BBC79D573" ID-GBIF-Occurrence="1802482376" country="Iran" county="The" latitude="37.470028" location="Kahaq region" longLatPrecision="1" longitude="47.226585" municipality="Maragheh" pageId="7" pageNumber="135" specimenCount="1" stateProvince="East Azarbaijan province" typeStatus="holotype">
<emphasis id="4AB29E15FF8F3A0BFF2AFA3BBF91D6AB" bold="true" box="[199,506,1493,1518]" pageId="7" pageNumber="135">
<typeStatus id="A77DFCA5FF8F3A0BFF2AFA3BBF69D6AB" box="[199,258,1493,1518]" pageId="7" pageNumber="135">Type</typeStatus>
locality and habitat.
</emphasis>
<collectingCounty id="91183A8BFF8F3A0BFDEFFA3BBC45D6AB" box="[514,558,1493,1518]" pageId="7" pageNumber="135">The</collectingCounty>
new species has been collected from
<collectingCountry id="00D10297FF8F3A0BFC37FA39BA61D6AB" box="[986,1034,1495,1518]" name="Iran" pageId="7" pageNumber="135">Iran</collectingCountry>
,
<collectingRegion id="BA028CE5FF8F3A0BFBFBFA39BB5ED6AB" box="[1046,1333,1493,1518]" country="Iran" name="Azarbayjan-e Sharqi" pageId="7" pageNumber="135">East Azarbaijan province</collectingRegion>
, north of
<collectingMunicipality id="981DD87DFF8F3A0BFF7AFA17BF67D557" box="[151,268,1529,1554]" pageId="7" pageNumber="135">Maragheh</collectingMunicipality>
,
<location id="7D1914DCFF8F3A0BFEF2FA17BFDCD557" LSID="urn:lsid:plazi:treatment:F06FF311FF893A04FF7AFD61BD2DD295:7D1914DCFF8F3A0BFEF2FA17BFDCD557" box="[287,439,1529,1554]" country="Iran" county="The" latitude="37.470028" longLatPrecision="1" longitude="47.226585" municipality="Maragheh" name="Kahaq region" pageId="7" pageNumber="135" stateProvince="East Azarbaijan province">Kahaq region</location>
(GPS coordinate
<geoCoordinate id="1DF224C0FF8F3A0BFD6FFA17BD48D557" box="[642,803,1529,1554]" direction="east" orientation="longitude" pageId="7" pageNumber="135" precision="1" value="47.226585">47º13´35.7´´E</geoCoordinate>
,
<geoCoordinate id="1DF224C0FF8F3A0BFCC2FA14BDB7D557" box="[815,988,1530,1554]" direction="north" orientation="latitude" pageId="7" pageNumber="135" precision="1" value="37.470028">37º28´12.1´´ N</geoCoordinate>
), where it was found in the rhizosphere soil of walnut, spruce and willow.
</materialsCitation>
</paragraph>
<paragraph id="78794207FF8F3A0BFF2AF9AFBADFD5E7" blockId="7.[151,1437,1493,2022]" pageId="7" pageNumber="135">
<emphasis id="4AB29E15FF8F3A0BFF2AF9AFBF1FD51F" bold="true" box="[199,372,1601,1626]" pageId="7" pageNumber="135">
<typeStatus id="A77DFCA5FF8F3A0BFF2AF9AFBF69D51F" box="[199,258,1601,1626]" pageId="7" pageNumber="135">Type</typeStatus>
material.
</emphasis>
Female
<typeStatus id="A77DFCA5FF8F3A0BFE38F9AFBC51D51F" box="[469,570,1601,1626]" pageId="7" pageNumber="135" type="holotype">holotype</typeStatus>
,
<specimenCount id="6EC0898EFF8F3A0BFDA9F9ACBCD8D51F" box="[580,691,1601,1626]" pageId="7" pageNumber="135" type="female">10 female</specimenCount>
and eight male
<typeStatus id="A77DFCA5FF8F3A0BFC8BF9ACBDBAD51F" box="[870,977,1602,1626]" pageId="7" pageNumber="135" type="paratype">paratypes</typeStatus>
deposited in the Nematode Collection of Nematology Laboratory, Department of Plant Protection, Faculty of Agriculture, University of Tabriz, Tabriz,
<collectingCountry id="00D10297FF8F3A0BFA85F989BBF2D53B" box="[1384,1433,1639,1662]" name="Iran" pageId="7" pageNumber="135">Iran</collectingCountry>
. Two female and two male
<typeStatus id="A77DFCA5FF8F3A0BFE2DF964BC47D5E7" box="[448,556,1674,1698]" pageId="7" pageNumber="135" type="paratype">paratypes</typeStatus>
with the Nematode Collection, University of Jaén,
<collectingCountry id="00D10297FF8F3A0BFB80F967BADBD5E7" box="[1133,1200,1673,1698]" name="Spain" pageId="7" pageNumber="135">Spain</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="30DC118CFF8F3A0BFF2AF943BB19D583" box="[199,1394,1709,1734]" pageId="7" pageNumber="135" type="etymology">
<paragraph id="78794207FF8F3A0BFF2AF943BB19D583" blockId="7.[151,1437,1493,2022]" box="[199,1394,1709,1734]" pageId="7" pageNumber="135">
<emphasis id="4AB29E15FF8F3A0BFF2AF943BF25D583" bold="true" box="[199,334,1709,1734]" pageId="7" pageNumber="135">Etymology.</emphasis>
The specific epithet refers to the geographical origin of the new species in Kahaq region,
<collectingCountry id="00D10297FF8F3A0BFAD3F941BB04D583" box="[1342,1391,1711,1734]" name="Iran" pageId="7" pageNumber="135">Iran</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="30DC118CFF8F3A04FF2AF93FBD2DD295" lastPageId="8" lastPageNumber="136" pageId="7" pageNumber="135" type="discussion">
<paragraph id="78794207FF8F3A04FF2AF93FBD2DD295" blockId="7.[151,1437,1493,2022]" lastBlockId="8.[151,1437,151,464]" lastPageId="8" lastPageNumber="136" pageId="7" pageNumber="135">
<emphasis id="4AB29E15FF8F3A0BFF2AF93FBF50D5AF" bold="true" box="[199,315,1745,1770]" pageId="7" pageNumber="135">Remarks.</emphasis>
The new species herein described is an excellent example of the problematic taxonomy of the conical-tailed species of the family
<taxonomicName id="BFC63984FF8F3A0BFDCFF91BBC83D44B" box="[546,744,1781,1806]" class="Adenophorea" family="Qudsianematidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="7" pageNumber="135" phylum="Nematoda" rank="family">Qudsianematidae</taxonomicName>
. It is characterized by a combination of features that allows its relatively easy identification but that, at the same time, raises doubt about its generic identity. Its conical, somewhat elongate caudal region is intermediate between those observed in species of
<taxonomicName id="BFC63984FF8F3A0BFBCEF8D0BAABD413" authorityName="Andrassy" authorityYear="1959" box="[1059,1216,1854,1878]" class="Adenophorea" family="Qudsianematidae" genus="Eudorylaimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="7" pageNumber="135" phylum="Nematoda" rank="genus">
<emphasis id="4AB29E15FF8F3A0BFBCEF8D0BAABD413" box="[1059,1216,1854,1878]" italics="true" pageId="7" pageNumber="135">Eudorylaimus</emphasis>
</taxonomicName>
, which tend to be shorter, and those of
<taxonomicName id="BFC63984FF8F3A0BFE6DF88CBC4ED43F" authorityName="Andrassy" authorityYear="1986" box="[384,549,1890,1914]" class="Adenophorea" family="Qudsianematidae" genus="Epidorylaimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="7" pageNumber="135" phylum="Nematoda" rank="genus">
<emphasis id="4AB29E15FF8F3A0BFE6DF88CBC4ED43F" box="[384,549,1890,1914]" italics="true" pageId="7" pageNumber="135">Epidorylaimus</emphasis>
</taxonomicName>
species, which tend to be longer, but there is no distinct borderline between the two genera. The location of the posteriormost ventromedian supplement is also notable: situated at approximately the level of the anterior end of the spicules, it is not too much farther from the adcloacal pair than from the penultimate ventromedian supplement of the series, so that a pre-cloacal space (hiatus) is not distinctly perceptible, again representing an intermediate condition between
<taxonomicName id="BFC63984FF803A04FCA3FF76BD80D3F5" authorityName="Andrassy" authorityYear="1959" box="[846,1003,152,176]" class="Adenophorea" family="Qudsianematidae" genus="Eudorylaimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="8" pageNumber="136" phylum="Nematoda" rank="genus">
<emphasis id="4AB29E15FF803A04FCA3FF76BD80D3F5" box="[846,1003,152,176]" italics="true" pageId="8" pageNumber="136">Eudorylaimus</emphasis>
</taxonomicName>
(males with distinct hiatus) and
<taxonomicName id="BFC63984FF803A04FF7AFF52BF2FD391" authorityName="Andrassy" authorityYear="1986" box="[151,324,188,212]" class="Adenophorea" family="Qudsianematidae" genus="Allodorylaimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="8" pageNumber="136" phylum="Nematoda" rank="genus">
<emphasis id="4AB29E15FF803A04FF7AFF52BF2FD391" box="[151,324,188,212]" italics="true" pageId="8" pageNumber="136">Allodorylaimus</emphasis>
</taxonomicName>
and
<taxonomicName id="BFC63984FF803A04FE60FF52BC59D391" authorityName="Andrassy" authorityYear="1986" box="[397,562,188,212]" class="Adenophorea" family="Qudsianematidae" genus="Epidorylaimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="8" pageNumber="136" phylum="Nematoda" rank="genus">
<emphasis id="4AB29E15FF803A04FE60FF52BC59D391" box="[397,562,188,212]" italics="true" pageId="8" pageNumber="136">Epidorylaimus</emphasis>
</taxonomicName>
(both with males lacking hiatus,
<emphasis id="4AB29E15FF803A04FC0CFF53BA6FD391" box="[993,1028,189,212]" italics="true" pageId="8" pageNumber="136">i.e.</emphasis>
with one or more ventromedian supplements within the range of the spicules). With due caution, the new species has been provisionally classified under
<taxonomicName id="BFC63984FF803A04FF0CFEEABF15D259" authorityName="Andrassy" authorityYear="1959" box="[225,382,260,284]" class="Adenophorea" family="Qudsianematidae" genus="Eudorylaimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="8" pageNumber="136" phylum="Nematoda" rank="genus">
<emphasis id="4AB29E15FF803A04FF0CFEEABF15D259" box="[225,382,260,284]" italics="true" pageId="8" pageNumber="136">Eudorylaimus</emphasis>
</taxonomicName>
because its general morphology fits better the pattern of that genus. Nevertheless, leaving aside the tail length and the presence/absence of a hiatus, it should be emphasized that the new species displays other features (nearly truncate and continuous lip region with corrugated anterior margin, broad cheilostom, weakly sclerotized but perceptible cephalic framework, presence of a coelomocyte shortly behind the nerve ring, dorsal gland pharyngeal nucleus in rather posterior position, etc.) that conform a peculiar pattern that might deserve separate generic status if other species are found that share it.
</paragraph>
</subSubSection>
</treatment>
</document>
Reference in a new issue View git blame Copy permalink