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<document id="B1AC954F3A0C50597ACB07ED32824C05" ID-DOI="10.11646/zootaxa.5437.4.9" ID-ISSN="1175-5326" ID-Zenodo-Dep="10985633" ID-ZooBank="7114B49F-4A44-4165-9FC6-8A52A6E898F9" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1713347432768" checkinUser="plazi" docAuthor="Kasparyan, Dmitry R. &amp; Khalaim, Andrey I." docDate="2024" docId="03A03624FFF7B10F4DA4AA3CFDDEF87C" docLanguage="en" docName="zootaxa.5437.4.9.pdf" docOrigin="Zootaxa 5437 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5437.4.9" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Glodianus rufithorax Kasparyan &amp; Khalaim 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="574" masterDocId="FF994E5CFFF5B10C4D33A81EFFA1FF95" masterDocTitle="Two new species of Glodianus Cameron and Hylophasma Townes (Hymenoptera: Ichneumonidae: Cryptinae) from Mexico" masterLastPageNumber="581" masterPageNumber="571" pageNumber="573" updateTime="1713348150872" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="25AABC249DF92EEDD07DB2CF5E5C1F8B">Two new species of Glodianus Cameron and Hylophasma Townes (Hymenoptera: Ichneumonidae: Cryptinae) from Mexico</mods:title>
</mods:titleInfo>
<mods:name id="B7DF89CFC020AC4355A26F38C5B4A9AB" type="personal">
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<mods:namePart id="FDDFA631FF11235F5ADEF2B08F233E6E">Kasparyan, Dmitry R.</mods:namePart>
<mods:affiliation id="ADF0721BFC347EEDECB74E9CACDEA802">Zoological Institute of the Russian Academy of Sciences, St. Petersburg, Russia</mods:affiliation>
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<mods:namePart id="B9190CC8B42B5142D66D76D6F5944250">Khalaim, Andrey I.</mods:namePart>
<mods:nameIdentifier id="95538656DA5682207BA9694F44A6E879" type="ORCID">0000-0003-1802-2649</mods:nameIdentifier>
<mods:affiliation id="A1043C0070C761C65FF3F5F0754F69A6">Zoological Institute of the Russian Academy of Sciences, St. Petersburg, Russia</mods:affiliation>
<mods:nameIdentifier id="97365BC35072748C5EFC20F2ED37FDFC" type="email">akhalaim@gmail.com</mods:nameIdentifier>
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<mods:title id="30841808EC5ADCAD3B183B81D7A92CF0">Zootaxa</mods:title>
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<mods:date id="4175653FC56590893FBBA254D34253A5">2024</mods:date>
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<mods:number id="80DA6FB5608CB88288F64D81BF0FF8A9">2024-04-16</mods:number>
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<mods:detail id="FCA7D13DF5933EC1DF0CDFB0B44CAA82" type="volume">
<mods:number id="8CB60E42A94915CD83685F815A10579D">5437</mods:number>
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<mods:number id="A87EC3018B954BA7429DB9FF8FBC9739">4</mods:number>
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<mods:classification id="C1CCD35E7AD3FEA0C44CF4E18179861E">journal article</mods:classification>
<mods:identifier id="BD14D7062F13371AF5AE3C040C8F56DB" type="DOI">10.11646/zootaxa.5437.4.9</mods:identifier>
<mods:identifier id="FB7E4CC4B05C519478A92099CB6244AE" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="A555C5374C7D1AB872405D81D74564B4" type="Zenodo-Dep">10985633</mods:identifier>
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<treatment id="03A03624FFF7B10F4DA4AA3CFDDEF87C" ID-DOI="http://doi.org/10.5281/zenodo.10985661" ID-Zenodo-Dep="10985661" LSID="urn:lsid:plazi:treatment:03A03624FFF7B10F4DA4AA3CFDDEF87C" httpUri="http://treatment.plazi.org/id/03A03624FFF7B10F4DA4AA3CFDDEF87C" lastPageId="3" lastPageNumber="574" pageId="2" pageNumber="573">
<subSubSection id="C313D4B9FFF7B10E4DA4AA3CFCE8FDA8" box="[151,841,546,573]" pageId="2" pageNumber="573" type="nomenclature">
<paragraph id="8BB68732FFF7B10E4DA4AA3CFCE8FDA8" blockId="2.[151,841,546,609]" box="[151,841,546,573]" pageId="2" pageNumber="573">
<heading id="D0FE305EFFF7B10E4DA4AA3CFCE8FDA8" bold="true" box="[151,841,546,573]" fontSize="11" level="1" pageId="2" pageNumber="573" reason="1">
<emphasis id="B97D5B20FFF7B10E4DA4AA3CFCE8FDA8" bold="true" box="[151,841,546,573]" pageId="2" pageNumber="573">
3.
<taxonomicName id="4C09FCB1FFF7B10E4D86AA3CFD7EFDA9" authority="Kasparyan &amp; Khalaim" authorityName="Kasparyan &amp; Khalaim" authorityYear="2024" box="[181,735,546,573]" class="Insecta" family="Ichneumonidae" genus="Glodianus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="573" phylum="Arthropoda" rank="species" species="rufithorax" status="sp. nov.">
<emphasis id="B97D5B20FFF7B10E4D86AA3CFE16FDA8" bold="true" box="[181,439,546,573]" italics="true" pageId="2" pageNumber="573">Glodianus rufithorax</emphasis>
Kasparyan &amp; Khalaim
</taxonomicName>
,
<taxonomicNameLabel id="A24EE65BFFF7B10E4FDFAA3DFCE8FDA8" box="[748,841,547,573]" pageId="2" pageNumber="573" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C313D4B9FFF7B10E4DA4AA59FEB8FDF4" box="[151,281,583,609]" pageId="2" pageNumber="573" type="description">
<paragraph id="8BB68732FFF7B10E4DA4AA59FEB8FDF4" blockId="2.[151,841,546,609]" box="[151,281,583,609]" pageId="2" pageNumber="573">
(
<figureCitation id="13329BB7FFF7B10E4DACAA59FEB0FDF4" box="[159,273,583,609]" captionStart-0="FIGURES 1–5" captionStart-1="FIGURES 6–10" captionStartId-0="3.[151,264,1497,1521]" captionStartId-1="4.[151,264,1792,1816]" captionTargetBox-0="[151,1435,181,1473]" captionTargetBox-1="[151,1435,181,1759]" captionTargetId-0="figure-19@3.[151,1435,181,1473]" captionTargetId-1="figure-26@4.[151,1435,181,1768]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURES 1–5. Glodianus rufithorax sp. nov., ♀, holotype: 1—habitus, lateral; 2—head and mesoscutum, dorsal; 3—head, antero-lateral; 4—head and mesosoma, lateral; 5—fore wing." captionText-1="FIGURES 6–10. Glodianus rufithorax sp. nov., ♀, holotype: 6—posterior part of mesosoma and base of metasoma, lateral; 7—propodeum and first tergite, dorsal; 8—apex of metasoma with ovipositor, lateral; 9—apex of ovipositor, lateral; 10—apex of ovipositor, dorsal." figureDoi-0="http://doi.org/10.5281/zenodo.10985636" figureDoi-1="http://doi.org/10.5281/zenodo.10985639" httpUri-0="https://zenodo.org/record/10985636/files/figure.png" httpUri-1="https://zenodo.org/record/10985639/files/figure.png" pageId="2" pageNumber="573">Figs 1–10</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C313D4B9FFF7B10E4DA4AA90FCF3FD64" pageId="2" pageNumber="573" type="materials_examined">
<paragraph id="8BB68732FFF7B10E4DA4AA90FDE0FD58" blockId="2.[151,1437,654,2013]" pageId="2" pageNumber="573">
<emphasis id="B97D5B20FFF7B10E4DA4AA90FE25FD3D" bold="true" box="[151,388,654,680]" pageId="2" pageNumber="573">Material examined.</emphasis>
<materialsCitation id="3B618D6FFFF7B10E4CBCAA91FDE0FD58" collectingDate="2017-10" collectionCode="UNAM" collectorName="M. Lopez-Ortega." country="Mexico" location="Tejerias" municipality="Municipio Teocelo" pageId="2" pageNumber="573" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="holotype">
<emphasis id="B97D5B20FFF7B10E4CBCAA91FE5DFD3C" bold="true" box="[399,508,655,681]" pageId="2" pageNumber="573">
<typeStatus id="54B23990FFF7B10E4CBCAA91FE5DFD3C" box="[399,508,655,681]" pageId="2" pageNumber="573" type="holotype">Holotype</typeStatus>
</emphasis>
female (
<collectionCode id="ED181FF7FFF7B10E4F56AA91FD61FD3C" box="[613,704,655,681]" country="Mexico" httpUri="http://grbio.org/cool/10ei-x9nj" name="Universidad Nacional Autonoma de Mexico" pageId="2" pageNumber="573">UNAM</collectionCode>
),
<collectingCountry id="F31EC7A2FFF7B10E4FEBAA91FC91FD3C" box="[728,816,655,681]" name="Mexico" pageId="2" pageNumber="573">Mexico</collectingCountry>
,
<collectingRegion id="49CD49D0FFF7B10E4E0CAA91FC07FD3C" box="[831,934,655,681]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="2" pageNumber="573">Veracruz</collectingRegion>
,
<collectingMunicipality id="6BD21D48FFF7B10E4E87AA91FB2FFD3C" box="[948,1166,655,681]" pageId="2" pageNumber="573">Municipio Teocelo</collectingMunicipality>
,
<location id="8ED6D1E9FFF7B10E49AFAA91FB56FD3C" LSID="urn:lsid:plazi:treatment:03A03624FFF7B10F4DA4AA3CFDDEF87C:8ED6D1E9FFF7B10E49AFAA91FB56FD3C" box="[1180,1271,655,681]" country="Mexico" municipality="Municipio Teocelo" name="Tejerias" pageId="2" pageNumber="573" stateProvince="Veracruz de Ignacio de la Llave">Tejerias</location>
,
<collectingMethod id="5248FF25FFF7B10E4835AA91FA39FD3C" box="[1286,1432,655,681]" pageId="2" pageNumber="573">Malaise trap</collectingMethod>
,
<date id="FFB7A1F2FFF7B10E4DA4AAADFE92FD58" box="[151,307,691,717]" pageId="2" pageNumber="573" value="2017-10">
<collectingDate id="EFF3581AFFF7B10E4DA4AAADFE92FD58" box="[151,307,691,717]" pageId="2" pageNumber="573" value="2017-10">October 2017</collectingDate>
</date>
, coll.
<collectorName id="26FCE2E4FFF7B10E4C45AAADFDE0FD58" box="[374,577,691,717]" pageId="2" pageNumber="573">M. López-Ortega.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8BB68732FFF7B10E4DF4AAC9FCF3FD64" blockId="2.[151,1437,654,2013]" box="[199,850,726,753]" pageId="2" pageNumber="573">
<materialsCitation id="3B618D6FFFF7B10E4DF4AAC9FCEFFD64" box="[199,846,726,753]" collectingDate="2017-10" collectionCode="ZISP" collectorName="M. Lopez-Ortega." country="Mexico" location="Tejerias" municipality="Municipio Teocelo" pageId="2" pageNumber="573" specimenCount="1" specimenCount-female="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="paratype">
<emphasis id="B97D5B20FFF7B10E4DF4AAC9FE63FD65" bold="true" box="[199,450,726,753]" pageId="2" pageNumber="573">
<typeStatus id="54B23990FFF7B10E4DF4AAC9FE97FD64" box="[199,310,727,753]" pageId="2" pageNumber="573" type="paratype">Paratype</typeStatus>
.
<collectingCountry id="F31EC7A2FFF7B10E4C71AAC8FE1CFD65" box="[322,445,726,752]" name="Mexico" pageId="2" pageNumber="573">MEXICO</collectingCountry>
:
</emphasis>
<specimenCount id="9D0F4CBBFFF7B10E4CFAAAC9FE52FD64" box="[457,499,727,753]" count="1" pageId="2" pageNumber="573" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED181FF7FFF7B10E4F31AAC9FDE0FD64" box="[514,577,727,753]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="2" pageNumber="573">ZISP</collectionCode>
), same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C313D4B9FFF7B10F4DF4AAE4FE24F914" lastPageId="3" lastPageNumber="574" pageId="2" pageNumber="573" type="description">
<paragraph id="8BB68732FFF7B10E4DF4AAE4FC4EFC80" blockId="2.[151,1437,654,2013]" box="[199,1007,762,789]" pageId="2" pageNumber="573">
<emphasis id="B97D5B20FFF7B10E4DF4AAE4FEF8FC81" bold="true" box="[199,345,762,788]" pageId="2" pageNumber="573">Description.</emphasis>
<emphasis id="B97D5B20FFF7B10E4C53AAE5FE15FC80" box="[352,436,763,789]" italics="true" pageId="2" pageNumber="573">Female</emphasis>
. Body length 15.0 mm, fore wing length 10.0 mm.
</paragraph>
<paragraph id="8BB68732FFF7B10E4DF4AB01FAE1FC14" blockId="2.[151,1437,654,2013]" pageId="2" pageNumber="573">
Antenna with about 39 flagellomeres; flagellomeres distad the white band distinctly widened, strongly flattened ventrally (
<figureCitation id="13329BB7FFF7B10E4C38AB5DFEEFFCC8" box="[267,334,835,861]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,1497,1521]" captionTargetBox="[151,1435,181,1473]" captionTargetId="figure-19@3.[151,1435,181,1473]" captionTargetPageId="3" captionText="FIGURES 1–5. Glodianus rufithorax sp. nov., ♀, holotype: 1—habitus, lateral; 2—head and mesoscutum, dorsal; 3—head, antero-lateral; 4—head and mesosoma, lateral; 5—fore wing." figureDoi="http://doi.org/10.5281/zenodo.10985636" httpUri="https://zenodo.org/record/10985636/files/figure.png" pageId="2" pageNumber="573">Fig. 1</figureCitation>
); the widest subapical flagellomeres about 3.0× as broad as two apical flagellomeres. Frons with median compressed horn (
<figureCitation id="13329BB7FFF7B10E4C8DAB79FDA5FC14" box="[446,516,871,897]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,1497,1521]" captionTargetBox="[151,1435,181,1473]" captionTargetId="figure-19@3.[151,1435,181,1473]" captionTargetPageId="3" captionText="FIGURES 1–5. Glodianus rufithorax sp. nov., ♀, holotype: 1—habitus, lateral; 2—head and mesoscutum, dorsal; 3—head, antero-lateral; 4—head and mesosoma, lateral; 5—fore wing." figureDoi="http://doi.org/10.5281/zenodo.10985636" httpUri="https://zenodo.org/record/10985636/files/figure.png" pageId="2" pageNumber="573">Fig. 2</figureCitation>
). Clypeus weakly convex, with lower margin distinctly truncated (
<figureCitation id="13329BB7FFF7B10E49DDAB79FA93FC14" box="[1262,1330,871,897]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,1497,1521]" captionTargetBox="[151,1435,181,1473]" captionTargetId="figure-19@3.[151,1435,181,1473]" captionTargetPageId="3" captionText="FIGURES 1–5. Glodianus rufithorax sp. nov., ♀, holotype: 1—habitus, lateral; 2—head and mesoscutum, dorsal; 3—head, antero-lateral; 4—head and mesosoma, lateral; 5—fore wing." figureDoi="http://doi.org/10.5281/zenodo.10985636" httpUri="https://zenodo.org/record/10985636/files/figure.png" pageId="2" pageNumber="573">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="8BB68732FFF7B10E4DF4AB95FBD2FA98" blockId="2.[151,1437,654,2013]" pageId="2" pageNumber="573">
Epomia long and strong, its upper end above transverse groove subdivided into two or three close wrinkles. Mesoscutum with central lobe evenly, densely and moderately finely punctate; lateral lobes sparsely punctate anteriorly, polished and impunctate in posterior half.Anterolateral part of mesoscutum at base of notaulus with strong transverse rugosity extending towards centre of mesoscutum (wrinkles stronger anteriorly and weaker posteriorly); posterior half of mesoscutum between notauli with strong longitudinal striae extending to scuto-scutellar groove (
<figureCitation id="13329BB7FFF7B10E4DACAC21FF44FBCC" box="[159,229,1087,1113]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,1497,1521]" captionTargetBox="[151,1435,181,1473]" captionTargetId="figure-19@3.[151,1435,181,1473]" captionTargetPageId="3" captionText="FIGURES 1–5. Glodianus rufithorax sp. nov., ♀, holotype: 1—habitus, lateral; 2—head and mesoscutum, dorsal; 3—head, antero-lateral; 4—head and mesosoma, lateral; 5—fore wing." figureDoi="http://doi.org/10.5281/zenodo.10985636" httpUri="https://zenodo.org/record/10985636/files/figure.png" pageId="2" pageNumber="573">Fig. 2</figureCitation>
). Scutellum polished with lateral carinae extending in its basal half. Mesopleuron (
<figureCitation id="13329BB7FFF7B10E49B0AC21FB67FBCC" box="[1155,1222,1087,1113]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,1497,1521]" captionTargetBox="[151,1435,181,1473]" captionTargetId="figure-19@3.[151,1435,181,1473]" captionTargetPageId="3" captionText="FIGURES 1–5. Glodianus rufithorax sp. nov., ♀, holotype: 1—habitus, lateral; 2—head and mesoscutum, dorsal; 3—head, antero-lateral; 4—head and mesosoma, lateral; 5—fore wing." figureDoi="http://doi.org/10.5281/zenodo.10985636" httpUri="https://zenodo.org/record/10985636/files/figure.png" pageId="2" pageNumber="573">Fig. 4</figureCitation>
) in upper half with oblique longitudinal wrinkles, in lower half with fine and dense punctures; speculum polished; epicnemium and mesosternum with small and dense punctures. Metapleuron entirely with dense oblique wrinkles, without distinct punctures; upper division of metapleuron with irregular wrinkles (
<figureCitation id="13329BB7FFF7B10E4EBFACB5FC72FB50" box="[908,979,1195,1221]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,1497,1521]" captionTargetBox="[151,1435,181,1473]" captionTargetId="figure-19@3.[151,1435,181,1473]" captionTargetPageId="3" captionText="FIGURES 1–5. Glodianus rufithorax sp. nov., ♀, holotype: 1—habitus, lateral; 2—head and mesoscutum, dorsal; 3—head, antero-lateral; 4—head and mesosoma, lateral; 5—fore wing." figureDoi="http://doi.org/10.5281/zenodo.10985636" httpUri="https://zenodo.org/record/10985636/files/figure.png" pageId="2" pageNumber="573">Fig. 4</figureCitation>
). Propodeum (
<figureCitation id="13329BB7FFF7B10E494FACB5FB42FB50" box="[1148,1251,1195,1221]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1792,1816]" captionTargetBox="[151,1435,181,1759]" captionTargetId="figure-26@4.[151,1435,181,1768]" captionTargetPageId="4" captionText="FIGURES 6–10. Glodianus rufithorax sp. nov., ♀, holotype: 6—posterior part of mesosoma and base of metasoma, lateral; 7—propodeum and first tergite, dorsal; 8—apex of metasoma with ovipositor, lateral; 9—apex of ovipositor, lateral; 10—apex of ovipositor, dorsal." figureDoi="http://doi.org/10.5281/zenodo.10985639" httpUri="https://zenodo.org/record/10985639/files/figure.png" pageId="2" pageNumber="573">Figs 6, 7</figureCitation>
) before anterior transverse carina polished dorsally (between long-oval spiracles), with wrinkles around spiracles; behind anterior transverse with strong, dense, parallel rugae; areola distinguished by irregular sculpture.
</paragraph>
<paragraph id="8BB68732FFF7B10E4DF4AD09FE83FA54" blockId="2.[151,1437,654,2013]" pageId="2" pageNumber="573">
Fore wing with areolet 0.7× as long as wide and about 0.55× as long as portion of vein 2m-cu above bulla (
<figureCitation id="13329BB7FFF7B10E485CAD09FF06FAC0" captionStart="FIGURES 1–5" captionStartId="3.[151,264,1497,1521]" captionTargetBox="[151,1435,181,1473]" captionTargetId="figure-19@3.[151,1435,181,1473]" captionTargetPageId="3" captionText="FIGURES 1–5. Glodianus rufithorax sp. nov., ♀, holotype: 1—habitus, lateral; 2—head and mesoscutum, dorsal; 3—head, antero-lateral; 4—head and mesosoma, lateral; 5—fore wing." figureDoi="http://doi.org/10.5281/zenodo.10985636" httpUri="https://zenodo.org/record/10985636/files/figure.png" pageId="2" pageNumber="573">Fig. 5</figureCitation>
). Hind wing with nervellus intercepted at lower 0.15 (
<figureCitation id="13329BB7FFF7B10E4E39AD25FCEEFAC0" box="[778,847,1338,1365]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,1497,1521]" captionTargetBox="[151,1435,181,1473]" captionTargetId="figure-19@3.[151,1435,181,1473]" captionTargetPageId="3" captionText="FIGURES 1–5. Glodianus rufithorax sp. nov., ♀, holotype: 1—habitus, lateral; 2—head and mesoscutum, dorsal; 3—head, antero-lateral; 4—head and mesosoma, lateral; 5—fore wing." figureDoi="http://doi.org/10.5281/zenodo.10985636" httpUri="https://zenodo.org/record/10985636/files/figure.png" pageId="2" pageNumber="573">Fig. 5</figureCitation>
). Legs slender (
<figureCitation id="13329BB7FFF7B10E4937AD25FBE4FAC0" box="[1028,1093,1339,1365]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,1497,1521]" captionTargetBox="[151,1435,181,1473]" captionTargetId="figure-19@3.[151,1435,181,1473]" captionTargetPageId="3" captionText="FIGURES 1–5. Glodianus rufithorax sp. nov., ♀, holotype: 1—habitus, lateral; 2—head and mesoscutum, dorsal; 3—head, antero-lateral; 4—head and mesosoma, lateral; 5—fore wing." figureDoi="http://doi.org/10.5281/zenodo.10985636" httpUri="https://zenodo.org/record/10985636/files/figure.png" pageId="2" pageNumber="573">Fig. 1</figureCitation>
). Hind leg with femur 8.0× as long as broad; inner tibial spur 1.9× longer than the outer spur; proportion of length of tarsomeres 1–5 of hind tarsus 8.8: 3.7: 2.6: 0.8: 3.2; inner (the longest) spur 1.25× as long as tarsomere 2; tarsomere 2 almost 1.2× as long as tarsomere 5.
</paragraph>
<paragraph id="8BB68732FFF7B10E4DF4ADD5FDD6F928" blockId="2.[151,1437,654,2013]" pageId="2" pageNumber="573">
First metasomal segment polished, long and narrow, 4.0× as long as wide at apex; second tergite about as long as its posterior width. Second and subsequent tergites smooth, with very fine and dense punctures and short and dense setae. Ovipositor sheath as long as hind tibia. Ovipositor strongly compressed laterally, with distinct teeth ventrally (
<figureCitation id="13329BB7FFF7B10E4C38AE29FEF0F9C4" box="[267,337,1591,1617]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1792,1816]" captionTargetBox="[151,1435,181,1759]" captionTargetId="figure-26@4.[151,1435,181,1768]" captionTargetPageId="4" captionText="FIGURES 6–10. Glodianus rufithorax sp. nov., ♀, holotype: 6—posterior part of mesosoma and base of metasoma, lateral; 7—propodeum and first tergite, dorsal; 8—apex of metasoma with ovipositor, lateral; 9—apex of ovipositor, lateral; 10—apex of ovipositor, dorsal." figureDoi="http://doi.org/10.5281/zenodo.10985639" httpUri="https://zenodo.org/record/10985639/files/figure.png" pageId="2" pageNumber="573">Fig. 9</figureCitation>
); its upper valve dorsally at apex bearing about 11 small oblique ridges, of them four most basal ridges are deeper and extending onto lateral sides of the upper valve (
<figureCitation id="13329BB7FFF7B10E4E9FAE45FB80F9E0" box="[940,1057,1627,1653]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1792,1816]" captionTargetBox="[151,1435,181,1759]" captionTargetId="figure-26@4.[151,1435,181,1768]" captionTargetPageId="4" captionText="FIGURES 6–10. Glodianus rufithorax sp. nov., ♀, holotype: 6—posterior part of mesosoma and base of metasoma, lateral; 7—propodeum and first tergite, dorsal; 8—apex of metasoma with ovipositor, lateral; 9—apex of ovipositor, lateral; 10—apex of ovipositor, dorsal." figureDoi="http://doi.org/10.5281/zenodo.10985639" httpUri="https://zenodo.org/record/10985639/files/figure.png" pageId="2" pageNumber="573">Figs 9, 10</figureCitation>
); portion of upper valve from the dorsal end of basal ridge (
<figureCitation id="13329BB7FFF7B10E4CF2AE61FDA6F90C" box="[449,519,1663,1689]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1792,1816]" captionTargetBox="[151,1435,181,1759]" captionTargetId="figure-26@4.[151,1435,181,1768]" captionTargetPageId="4" captionText="FIGURES 6–10. Glodianus rufithorax sp. nov., ♀, holotype: 6—posterior part of mesosoma and base of metasoma, lateral; 7—propodeum and first tergite, dorsal; 8—apex of metasoma with ovipositor, lateral; 9—apex of ovipositor, lateral; 10—apex of ovipositor, dorsal." figureDoi="http://doi.org/10.5281/zenodo.10985639" httpUri="https://zenodo.org/record/10985639/files/figure.png" pageId="2" pageNumber="573">Fig. 9</figureCitation>
, arrow) to apex 1.5× as long as hind tarsomere 2, and 1.3× as long as portion of lower valve from most basal tooth to apex.
</paragraph>
<paragraph id="8BB68732FFF7B10E4DF4AED9FF58F848" blockId="2.[151,1437,654,2013]" pageId="2" pageNumber="573">
Antenna black with scape fulvous and flagellomeres 6 to 11 white (blackish ventrally). Head predominantly red; face, clypeus, mandibles (except for blackish teeth), labrum and palpi yellow. Mesosoma entirely red. All coxae and fore and mid legs (except for brown apical tarsomeres) red. Hind legs with femur and tibia completely black, trochanter and trochantellus black with rufous markings ventrally (
<figureCitation id="13329BB7FFF7B10E4EA0AF2DFC5DF8D8" box="[915,1020,1843,1869]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1792,1816]" captionTargetBox="[151,1435,181,1759]" captionTargetId="figure-26@4.[151,1435,181,1768]" captionTargetPageId="4" captionText="FIGURES 6–10. Glodianus rufithorax sp. nov., ♀, holotype: 6—posterior part of mesosoma and base of metasoma, lateral; 7—propodeum and first tergite, dorsal; 8—apex of metasoma with ovipositor, lateral; 9—apex of ovipositor, lateral; 10—apex of ovipositor, dorsal." figureDoi="http://doi.org/10.5281/zenodo.10985639" httpUri="https://zenodo.org/record/10985639/files/figure.png" pageId="2" pageNumber="573">Figs 6, 7</figureCitation>
), spurs brown, tarsus predominantly white with basal 0.3 of basitarsus and extreme base of tarsomeres 2–4 black, and tarsomere 5 brown. Metasoma predominantly black; first segment with basal half of petiole and sternite yellowish red (
<figureCitation id="13329BB7FFF7B10E49B2AF65FB65F800" box="[1153,1220,1915,1941]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,1497,1521]" captionTargetBox="[151,1435,181,1473]" captionTargetId="figure-19@3.[151,1435,181,1473]" captionTargetPageId="3" captionText="FIGURES 1–5. Glodianus rufithorax sp. nov., ♀, holotype: 1—habitus, lateral; 2—head and mesoscutum, dorsal; 3—head, antero-lateral; 4—head and mesosoma, lateral; 5—fore wing." figureDoi="http://doi.org/10.5281/zenodo.10985636" httpUri="https://zenodo.org/record/10985636/files/figure.png" pageId="2" pageNumber="573">Fig. 1</figureCitation>
); tergites 2–7 with hind margins and epipleurae white; tergite 8 with lateral sides widely white; sternites white with black markings laterally.
</paragraph>
<caption id="DF76D7BAFFF6B10F4DA4ADC7FCA7F980" ID-DOI="http://doi.org/10.5281/zenodo.10985636" ID-Zenodo-Dep="10985636" httpUri="https://zenodo.org/record/10985636/files/figure.png" pageId="3" pageNumber="574" startId="3.[151,264,1497,1521]" targetBox="[151,1435,181,1473]" targetPageId="3" targetType="figure">
<paragraph id="8BB68732FFF6B10F4DA4ADC7FCA7F980" blockId="3.[151,1437,1497,1557]" pageId="3" pageNumber="574">
<emphasis id="B97D5B20FFF6B10F4DA4ADC7FE9DFA64" bold="true" box="[151,316,1497,1521]" pageId="3" pageNumber="574">FIGURES 1–5.</emphasis>
<taxonomicName id="4C09FCB1FFF6B10F4C77ADC7FDBCFA64" authority="Kasparyan &amp; Khalaim, 2024" authorityName="Kasparyan &amp; Khalaim" authorityYear="2024" box="[324,541,1497,1521]" class="Insecta" family="Ichneumonidae" genus="Glodianus" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="574" phylum="Arthropoda" rank="species" species="rufithorax" status="sp. nov.">
<emphasis id="B97D5B20FFF6B10F4C77ADC7FDBCFA64" box="[324,541,1497,1521]" italics="true" pageId="3" pageNumber="574">Glodianus rufithorax</emphasis>
</taxonomicName>
<emphasis id="B97D5B20FFF6B10F4F16ADC7FDD6FA64" bold="true" box="[549,631,1497,1521]" pageId="3" pageNumber="574">
<taxonomicNameLabel id="A24EE65BFFF6B10F4F16ADC7FDD6FA64" box="[549,631,1497,1521]" pageId="3" pageNumber="574" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, ♀, holotype:
<emphasis id="B97D5B20FFF6B10F4E3CADC7FCBDFA64" bold="true" box="[783,796,1497,1521]" pageId="3" pageNumber="574">1</emphasis>
—habitus, lateral;
<emphasis id="B97D5B20FFF6B10F4EE8ADC7FC49FA64" bold="true" box="[987,1000,1497,1521]" pageId="3" pageNumber="574">2</emphasis>
—head and mesoscutum, dorsal;
<emphasis id="B97D5B20FFF6B10F4873ADC7FAECFA64" bold="true" box="[1344,1357,1497,1521]" pageId="3" pageNumber="574">3</emphasis>
—head, antero-lateral;
<emphasis id="B97D5B20FFF6B10F4C1FADE3FE98F980" bold="true" box="[300,313,1533,1557]" pageId="3" pageNumber="574">4</emphasis>
—head and mesosoma, lateral;
<emphasis id="B97D5B20FFF6B10F4F44ADE3FD25F980" bold="true" box="[631,644,1533,1557]" pageId="3" pageNumber="574">5</emphasis>
—fore wing.
</paragraph>
</caption>
<paragraph id="8BB68732FFF6B10F4DF4AE79FE24F914" blockId="3.[151,1437,1639,2025]" box="[199,389,1639,1666]" pageId="3" pageNumber="574">
<emphasis id="B97D5B20FFF6B10F4DF4AE79FEA0F914" box="[199,257,1639,1665]" italics="true" pageId="3" pageNumber="574">Male</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C313D4B9FFF6B10F4DF4AE95FA37F933" box="[199,1430,1675,1702]" pageId="3" pageNumber="574" type="etymology">
<paragraph id="8BB68732FFF6B10F4DF4AE95FA37F933" blockId="3.[151,1437,1639,2025]" box="[199,1430,1675,1702]" pageId="3" pageNumber="574">
<emphasis id="B97D5B20FFF6B10F4DF4AE95FEE9F930" bold="true" box="[199,328,1675,1701]" pageId="3" pageNumber="574">Etymology</emphasis>
. The species name derives from the Latin
<emphasis id="B97D5B20FFF6B10F4E17AE95FCFFF930" box="[804,862,1675,1701]" italics="true" pageId="3" pageNumber="574">rufus</emphasis>
(red) and
<emphasis id="B97D5B20FFF6B10F4EE1AE95FBBAF930" box="[978,1051,1675,1701]" italics="true" pageId="3" pageNumber="574">thorax</emphasis>
(thorax), after the red mesosoma.
</paragraph>
</subSubSection>
<subSubSection id="C313D4B9FFF6B10F4DF4AEB1FDDEF87C" pageId="3" pageNumber="574" type="distribution">
<paragraph id="8BB68732FFF6B10F4DF4AEB1FDE3F95C" blockId="3.[151,1437,1639,2025]" box="[199,578,1711,1737]" pageId="3" pageNumber="574">
<emphasis id="B97D5B20FFF6B10F4DF4AEB1FEC0F95C" bold="true" box="[199,353,1711,1737]" pageId="3" pageNumber="574">Distribution.</emphasis>
<collectingCountry id="F31EC7A2FFF6B10F4C54AEB1FE1CF95C" box="[359,445,1711,1737]" name="Mexico" pageId="3" pageNumber="574">Mexico</collectingCountry>
(
<collectingRegion id="49CD49D0FFF6B10F4CFFAEB1FD96F95C" box="[460,567,1711,1737]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="3" pageNumber="574">Veracruz</collectingRegion>
).
</paragraph>
<paragraph id="8BB68732FFF6B10F4DF4AECDFBFFF8CF" blockId="3.[151,1437,1639,2025]" pageId="3" pageNumber="574">
<emphasis id="B97D5B20FFF6B10F4DF4AECDFEC3F978" bold="true" box="[199,354,1747,1773]" pageId="3" pageNumber="574">Comparison.</emphasis>
<taxonomicName id="4C09FCB1FFF6B10F4C59AECDFDF8F978" authority="Kasparyan &amp; Khalaim, 2024" authorityName="Kasparyan &amp; Khalaim" authorityYear="2024" box="[362,601,1747,1773]" class="Insecta" family="Ichneumonidae" genus="Glodianus" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="574" phylum="Arthropoda" rank="species" species="rufithorax" status="sp. nov.">
<emphasis id="B97D5B20FFF6B10F4C59AECDFDF8F978" box="[362,601,1747,1773]" italics="true" pageId="3" pageNumber="574">Glodianus rufithorax</emphasis>
</taxonomicName>
<emphasis id="B97D5B20FFF6B10F4F53AECDFD1AF978" bold="true" box="[608,699,1747,1773]" pageId="3" pageNumber="574">
<taxonomicNameLabel id="A24EE65BFFF6B10F4F53AECDFD1AF978" box="[608,699,1747,1773]" pageId="3" pageNumber="574" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is easily distinguished from its Mexican congeners by its mostly red head (except reddish yellow face, clypeus and lower part of gena, see
<figureCitation id="13329BB7FFF6B10F4EF9AEE9FBADF884" box="[970,1036,1783,1809]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,1497,1521]" captionTargetBox="[151,1435,181,1473]" captionTargetId="figure-19@3.[151,1435,181,1473]" captionTargetPageId="3" captionText="FIGURES 1–5. Glodianus rufithorax sp. nov., ♀, holotype: 1—habitus, lateral; 2—head and mesoscutum, dorsal; 3—head, antero-lateral; 4—head and mesosoma, lateral; 5—fore wing." figureDoi="http://doi.org/10.5281/zenodo.10985636" httpUri="https://zenodo.org/record/10985636/files/figure.png" pageId="3" pageNumber="574">Fig. 3</figureCitation>
), mesosoma, fore and mid legs, and hind coxa. Additionally, in the new species the hind tarsomere 2 is almost 1.2× as long as tarsomere 5, while in two other Mexican species the hind tarsomere 2 either 1.0× or 1.5× as long as tarsomere 5.
</paragraph>
<paragraph id="8BB68732FFF6B10F4DF4AF7AFDDEF87C" blockId="3.[151,1437,1639,2025]" pageId="3" pageNumber="574">
In its color pattern,
<taxonomicName id="4C09FCB1FFF6B10F4C98AF7DFDE0F8E8" authority="Kasparyan &amp; Khalaim, 2024" authorityName="Kasparyan &amp; Khalaim" authorityYear="2024" box="[427,577,1891,1917]" class="Insecta" family="Ichneumonidae" genus="Glodianus" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="574" phylum="Arthropoda" rank="species" species="rufithorax" status="sp. nov.">
<emphasis id="B97D5B20FFF6B10F4C98AF7DFDE0F8E8" box="[427,577,1891,1917]" italics="true" pageId="3" pageNumber="574">G. rufithorax</emphasis>
</taxonomicName>
<emphasis id="B97D5B20FFF6B10F4F7AAF7DFD04F8E8" bold="true" box="[585,677,1891,1917]" pageId="3" pageNumber="574">
<taxonomicNameLabel id="A24EE65BFFF6B10F4F7AAF7DFD04F8E8" box="[585,677,1891,1917]" pageId="3" pageNumber="574" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
resembles the Venezuelan
<taxonomicName id="4C09FCB1FFF6B10F4EE8AF7DFB5CF8E8" authority="Taschenberg" authorityName="Taschenberg" box="[987,1277,1891,1917]" class="Insecta" family="Ichneumonidae" genus="Glodianus" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="574" phylum="Arthropoda" rank="species" species="areolatus">
<emphasis id="B97D5B20FFF6B10F4EE8AF7DFBC9F8E8" box="[987,1128,1891,1917]" italics="true" pageId="3" pageNumber="574">G. areolatus</emphasis>
Taschenberg
</taxonomicName>
(
<figureCitation id="13329BB7FFF6B10F483DAF7DFAC0F8E8" box="[1294,1377,1891,1917]" captionStart="FIGURE 11" captionStartId="6.[151,250,1162,1186]" captionTargetBox="[151,1436,181,1138]" captionTargetId="figure-26@6.[151,1436,181,1138]" captionTargetPageId="6" captionText="FIGURE 11. Glodianus areolatus, ♀, holotype: habitus, lateral." figureDoi="http://doi.org/10.5281/zenodo.10985641" httpUri="https://zenodo.org/record/10985641/files/figure.png" pageId="3" pageNumber="574">Fig. 11</figureCitation>
), but differs from this species in having fore wing with areolet receiving vein 2m-cu at posterior 0.35 (
<figureCitation id="13329BB7FFF6B10F49FCAF99FAB0F837" box="[1231,1297,1927,1954]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,1497,1521]" captionTargetBox="[151,1435,181,1473]" captionTargetId="figure-19@3.[151,1435,181,1473]" captionTargetPageId="3" captionText="FIGURES 1–5. Glodianus rufithorax sp. nov., ♀, holotype: 1—habitus, lateral; 2—head and mesoscutum, dorsal; 3—head, antero-lateral; 4—head and mesosoma, lateral; 5—fore wing." figureDoi="http://doi.org/10.5281/zenodo.10985636" httpUri="https://zenodo.org/record/10985636/files/figure.png" pageId="3" pageNumber="574">Fig. 5</figureCitation>
) (at anterior
<quantity id="4CF12AD7FFF6B10F4DA4AFB5FF44F850" box="[151,229,1963,1990]" metricMagnitude="-2" metricUnit="m" metricValue="1.143" pageId="3" pageNumber="574" unit="in" value="0.45">0.45 in</quantity>
<taxonomicName id="4C09FCB1FFF6B10F4DDFAFB5FED6F850" box="[236,375,1963,1989]" class="Insecta" family="Ichneumonidae" genus="Glodianus" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="574" phylum="Arthropoda" rank="species" species="areolatus">
<emphasis id="B97D5B20FFF6B10F4DDFAFB5FED6F850" box="[236,375,1963,1989]" italics="true" pageId="3" pageNumber="574">G. areolatus</emphasis>
</taxonomicName>
), and hind basitarsus black in basal 0.2–0.3 and white in apical 0.7–0.8 (
<figureCitation id="13329BB7FFF6B10F4995AFB5FB48F850" box="[1190,1257,1963,1989]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1792,1816]" captionTargetBox="[151,1435,181,1759]" captionTargetId="figure-26@4.[151,1435,181,1768]" captionTargetPageId="4" captionText="FIGURES 6–10. Glodianus rufithorax sp. nov., ♀, holotype: 6—posterior part of mesosoma and base of metasoma, lateral; 7—propodeum and first tergite, dorsal; 8—apex of metasoma with ovipositor, lateral; 9—apex of ovipositor, lateral; 10—apex of ovipositor, dorsal." figureDoi="http://doi.org/10.5281/zenodo.10985639" httpUri="https://zenodo.org/record/10985639/files/figure.png" pageId="3" pageNumber="574">Fig. 8</figureCitation>
) (black in basal 0.7 and white in apical
<quantity id="4CF12AD7FFF6B10F4CAFAFD1FE7DF87C" box="[412,476,1999,2025]" metricMagnitude="-3" metricUnit="m" metricValue="7.62" pageId="3" pageNumber="574" unit="in" value="0.3">0.3 in</quantity>
<taxonomicName id="4C09FCB1FFF6B10F4CD0AFD1FDCFF87C" box="[483,622,1999,2025]" class="Insecta" family="Ichneumonidae" genus="Glodianus" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="574" phylum="Arthropoda" rank="species" species="areolatus">
<emphasis id="B97D5B20FFF6B10F4CD0AFD1FDCFF87C" box="[483,622,1999,2025]" italics="true" pageId="3" pageNumber="574">G. areolatus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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