234 lines
28 KiB
XML
234 lines
28 KiB
XML
<document id="CF656EAA90DF15393CD0E5F13CE1DCD6" ID-CLB-Dataset="2961" ID-DOI="10.11646/palaeoentomology.4.6.5" ID-GBIF-Dataset="73a0820d-7503-472f-8c32-3836cf5fe857" ID-ISSN="2624-2834" ID-Zenodo-Dep="5778576" ID-ZooBank="95AA3380-CA95-46E8-B233-E9F0E8D60603" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1639469484134" checkinUser="plazi" docAuthor="Béthoux, Olivier & Anderson, John M." docDate="2021" docId="039DFA6CFFDAFFD96EC7FBDAC941FD1B" docLanguage="en" docName="palaeoentomology.4.6.5.pdf" docOrigin="Palaeoentomology 4 (6)" docStyle="DocumentStyle:4EE6FD780FF4521D541D0D0600705D0E.4:Palaeoentomology.2018-.journal_article" docStyleId="4EE6FD780FF4521D541D0D0600705D0E" docStyleName="Palaeoentomology.2018-.journal_article" docStyleVersion="4" docTitle="Polycytella rasnitsyni Béthoux & Anderson 2021, sp. nov." docType="treatment" docUuid="0D93F67B-6562-44BE-8E7B-8BC4B3A314B8" docUuidSource="ZooBank" docVersion="7" lastPageNumber="555" masterDocId="FFA48214FFDFFFDF6E4FFF9CC82EFFF8" masterDocTitle="The Polycytellidae viewed as Gondwanan Glosselytrodea" masterLastPageNumber="558" masterPageNumber="550" pageNumber="555" updateTime="1699472406386" updateUser="ExternalLinkService">
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<mods:mods id="4CC46D1B9642446D54D0CD18114D771E" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="5D2317AF4728062CFD75F118867BA54D">
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<mods:title id="BBF7DC4D356AB5DC48142B3767CC5B5E">The Polycytellidae viewed as Gondwanan Glosselytrodea</mods:title>
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</mods:titleInfo>
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<mods:name id="370A97CDC0A1E496125CA619BA05AA7A" type="personal">
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<mods:roleTerm id="23AFCEB4BE877B35A037F15784D7BA6D">Author</mods:roleTerm>
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<mods:namePart id="9D6764AD60E82D8274130740722F89F2">Béthoux, Olivier</mods:namePart>
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<mods:affiliation id="4B865DDE3A05691A115B4A689F86B106">Centre de Recherche sur la Paléontologie - Paris (CR 2 P), Sorbonne Université, MNHN, CNRS, Paris, France</mods:affiliation>
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</mods:name>
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<mods:name id="0FC2EC09687FFD2C6E2C3B91DF550B84" type="personal">
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<mods:roleTerm id="38A734C5B1FC55333CE84203CCE5FCF1">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="37288511E7E828F5619214CCA36FA0E5">Anderson, John M.</mods:namePart>
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<mods:affiliation id="A0C56B49AE709CA77135D992F297AC85">Evolutionary Studies Institute, Witwatersrand University, Johannesburg, South Africa</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="47CA7EC67BFE568DD0D19636F282DAFE">text</mods:typeOfResource>
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<mods:title id="C5323E7D5857FA3474A01034D2F2B610">Palaeoentomology</mods:title>
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</mods:titleInfo>
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<mods:part id="9A6C10DDFCBAD7493BB9F8A2E9F28126">
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<mods:date id="84EEFF065947B055F89B70AAA6415DD3">2021</mods:date>
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<mods:detail id="75A461F7771F80FCFB36666FCF1E2021" type="pubDate">
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<mods:number id="AF3FB60AD795CFDE4088D082B85EC94E">2021-12-14</mods:number>
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<mods:number id="8E6D57FE52018987D04B4E973C5F1FDE">4</mods:number>
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<mods:number id="4F246C195345611FFA1C422E3240DC19">6</mods:number>
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<mods:classification id="F46739B9604F068D56FCB0470EA09544">journal article</mods:classification>
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<mods:identifier id="BDED71FE8F66D863A6DCBF92F614885C" type="CLB-Dataset">2961</mods:identifier>
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<mods:identifier id="88AA2C8274DF1E21C9544D371AF0B8DF" type="DOI">10.11646/palaeoentomology.4.6.5</mods:identifier>
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<treatment id="039DFA6CFFDAFFD96EC7FBDAC941FD1B" ID-DOI="http://doi.org/10.5281/zenodo.5780414" ID-GBIF-Taxon="191571261" ID-Zenodo-Dep="5780414" LSID="urn:lsid:zoobank.org:act:0D93F67B-6562-44BE-8E7B-8BC4B3A314B8" httpUri="http://treatment.plazi.org/id/039DFA6CFFDAFFD96EC7FBDAC941FD1B" lastPageId="6" lastPageNumber="555" pageId="5" pageNumber="555">
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<subSubSection id="C32E18F1FFDAFFDA6EC7FBDAC9F1FB98" box="[136,479,1094,1121]" pageId="5" pageNumber="555" type="nomenclature">
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<paragraph id="8B8B4B7AFFDAFFDA6EC7FBDAC9F1FB98" blockId="5.[136,777,1094,1228]" box="[136,479,1094,1121]" pageId="5" pageNumber="555">
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<heading id="D0C3FC16FFDAFFDA6EC7FBDAC9F1FB98" bold="true" box="[136,479,1094,1121]" fontSize="10" level="2" pageId="5" pageNumber="555" reason="3">
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<emphasis id="B9409768FFDAFFDA6EC7FBDAC9F1FB98" bold="true" box="[136,479,1094,1121]" pageId="5" pageNumber="555">
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<taxonomicName id="4C3430F9FFDAFFDA6EC7FBDAC951FB98" authority="Béthoux & Anderson, 2021" authorityName="Béthoux & Anderson" authorityYear="2021" box="[136,383,1094,1121]" class="Insecta" family="Glosselytridae" genus="Polycytella" kingdom="Animalia" order="Glosselytrodea" pageId="5" pageNumber="555" phylum="Arthropoda" rank="species" species="rasnitsyni" status="sp. nov.">
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<emphasis id="B9409768FFDAFFDA6EC7FBDAC951FB98" bold="true" box="[136,383,1094,1121]" italics="true" pageId="5" pageNumber="555">Polycytella rasnitsyni</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2732A13FFDAFFDA6FC9FBDAC9F1FB98" box="[390,479,1094,1120]" pageId="5" pageNumber="555" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C32E18F1FFDAFFDA6EC7FBF6C967FB34" pageId="5" pageNumber="555" type="description">
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<paragraph id="8B8B4B7AFFDAFFDA6EC7FBF6C8D2FB7C" blockId="5.[136,777,1094,1228]" box="[136,252,1130,1156]" pageId="5" pageNumber="555">
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(
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<figureCitation id="130F57FFFFDAFFDA6EDFFBF6C8F7FB7C" box="[144,217,1130,1156]" captionStart="FIGURE 2" captionStartId="3.[136,235,1918,1942]" captionTargetBox="[189,1398,197,1894]" captionTargetId="figure-20@3.[189,1398,189,1894]" captionTargetPageId="3" captionText="FIGURE 2. Polycytella rasnitsyni sp. nov., holotype specimen (PRE/F/10559), forewing pair. A, Interpretative drawing (RFW, right forewing; LFW, left forewing). B–D, Photograph and RTI extracts, part. B, Photograph (light-mirrored; ‘cf’, claval furrow). C, RTI extract, specular enhancement mode. D, RTI extract, normals visualisation mode." figureDoi="http://doi.org/10.5281/zenodo.5778582" httpUri="https://zenodo.org/record/5778582/files/figure.png" pageId="5" pageNumber="555">Figs 2</figureCitation>
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,
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<figureCitation id="130F57FFFFDAFFDA6EAAFBF6C8DDFB7C" box="[229,243,1130,1156]" captionStart="FIGURE 3" captionStartId="4.[136,235,1039,1063]" captionTargetBox="[192,1373,208,1007]" captionTargetId="figure-435@4.[183,1404,204,1016]" captionTargetPageId="4" captionText="FIGURE 3. Polycytella rasnitsyni sp. nov., holotype specimen (PRE/F/10559), interpretative drawings, separated. A, Left forewing, as in Fig. 2A. B, Right forewing, as in Fig. 2A and with reconstructed parts." figureDoi="http://doi.org/10.5281/zenodo.5778586" httpUri="https://zenodo.org/record/5778586/files/figure.png" pageId="5" pageNumber="555">3</figureCitation>
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)
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</paragraph>
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<paragraph id="8B8B4B7AFFDAFFDA6EC7FB12C967FB34" blockId="5.[136,777,1094,1228]" pageId="5" pageNumber="555">
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<uri id="FFA54778FFDAFFDA6EC7FB12C967FB34" pageId="5" pageNumber="555">
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urn:lsid:zoobank.org:act:
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<uuid id="FF9271AFFFDAFFDA6FECFB12C967FB34" pageId="5" pageNumber="555">0D93F67B-6562-44BE-8E7B-8BC4B3A314B8</uuid>
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</uri>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C32E18F1FFDAFFDA6EC7FB66CA59FA78" pageId="5" pageNumber="555" type="materials_examined">
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||
<paragraph id="8B8B4B7AFFDAFFDA6EC7FB66CA59FA78" blockId="5.[136,778,1274,2093]" pageId="5" pageNumber="555">
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<materialsCitation id="3B5C4127FFDAFFDA6EC7FB66CA59FA78" ID-GBIF-Occurrence="3418318301" country="South Africa" county="Johannesburg" location="Specimen" municipality="Witwatersrand" pageId="5" pageNumber="555" specimenCount="1" typeStatus="holotype">
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<emphasis id="B9409768FFDAFFDA6EC7FB66C8D2FAEC" bold="true" box="[136,252,1274,1300]" pageId="5" pageNumber="555">
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<typeStatus id="548FF5D8FFDAFFDA6EC7FB66C8D9FAEC" box="[136,247,1274,1300]" pageId="5" pageNumber="555" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<location id="8EEB1DA1FFDAFFDA6F42FB66C955FAEC" LSID="urn:lsid:plazi:treatment:039DFA6CFFDAFFD96EC7FBDAC941FD1B:8EEB1DA1FFDAFFDA6F42FB66C955FAEC" box="[269,379,1274,1300]" country="South Africa" county="Johannesburg" municipality="Witwatersrand" name="Specimen" pageId="5" pageNumber="555">Specimen</location>
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PRE/F/10559 (preserving a left forewing in ventral aspect and a right forewing in dorsal aspect),
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<location id="8EEB1DA1FFDAFFDA6EA8FADECA12FAA4" LSID="urn:lsid:plazi:treatment:039DFA6CFFDAFFD96EC7FBDAC941FD1B:8EEB1DA1FFDAFFDA6EA8FADECA12FAA4" box="[231,572,1346,1372]" country="South Africa" county="Johannesburg" municipality="Witwatersrand" name="Evolutionary Studies Institute" pageId="5" pageNumber="555">Evolutionary Studies Institute</location>
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,
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<location id="8EEB1DA1FFDAFFDA6C06FADECA91FAA4" LSID="urn:lsid:plazi:treatment:039DFA6CFFDAFFD96EC7FBDAC941FD1B:8EEB1DA1FFDAFFDA6C06FADECA91FAA4" box="[585,703,1346,1372]" country="South Africa" county="Johannesburg" municipality="Witwatersrand" name="University" pageId="5" pageNumber="555">University</location>
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of the
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<collectingMunicipality id="6BEFD100FFDAFFDA6EC7FAFAC901FA78" box="[136,303,1382,1408]" pageId="5" pageNumber="555">Witwatersrand</collectingMunicipality>
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,
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||
<collectingCounty id="62EA33F6FFDAFFDA6F75FAFAC9F8FA78" box="[314,470,1382,1408]" pageId="5" pageNumber="555">Johannesburg</collectingCounty>
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,
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<collectingCountry id="F3230BEAFFDAFFDA6FAEFAFACA5DFA78" box="[481,627,1382,1408]" name="South Africa" pageId="5" pageNumber="555">South Africa</collectingCountry>
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.
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</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C32E18F1FFDAFFDA6EF2FA16CAAEFA14" pageId="5" pageNumber="555" type="etymology">
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<paragraph id="8B8B4B7AFFDAFFDA6EF2FA16CAAEFA14" blockId="5.[136,778,1274,2093]" pageId="5" pageNumber="555">
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<emphasis id="B9409768FFDAFFDA6EF2FA16C96AFA5C" bold="true" box="[189,324,1418,1444]" pageId="5" pageNumber="555">Etymology.</emphasis>
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The specific epithet honours Alexandr P. Rasnitsyn, for his wide-ranging contribution to the fields of evolutionary entomology and systematics.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32E18F1FFDAFFDA6EF2FA6AC8D7F911" pageId="5" pageNumber="555" type="diagnosis">
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<paragraph id="8B8B4B7AFFDAFFDA6EF2FA6AC8D7F911" blockId="5.[136,778,1274,2093]" pageId="5" pageNumber="555">
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<emphasis id="B9409768FFDAFFDA6EF2FA6AC916F9E8" bold="true" box="[189,312,1526,1552]" pageId="5" pageNumber="555">Diagnosis.</emphasis>
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Forewing: length
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<quantity id="4CCCE69FFFDAFFDA6C59FA6ACA5DF9E9" box="[534,627,1526,1553]" metricMagnitude="-3" metricUnit="m" metricValue="9.4" pageId="5" pageNumber="555" unit="mm" value="9.4">9.4 mm</quantity>
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, aspect ratio 3.6; in the 3
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<superScript id="7C41E632FFDAFFDA6F59F986C90AF9D0" attach="left" box="[278,292,1562,1576]" fontSize="6" pageId="5" pageNumber="555">rd</superScript>
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quarter of wing length, cells between RA and CuA
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<emphasis id="B9409768FFDAFFDA6EA4F9A3C934F9A0" box="[235,282,1599,1624]" italics="true" pageId="5" pageNumber="555">post</emphasis>
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wider than long; no intercalary vein between CuA
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<emphasis id="B9409768FFDAFFDA6EF2F9FFC8C2F984" box="[189,236,1635,1660]" italics="true" pageId="5" pageNumber="555">post</emphasis>
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and CuP (but see above); clavus large (
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<quantity id="4CCCE69FFFDAFFDA6CFEF9FECB24F985" box="[689,778,1634,1661]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="555" unit="mm" value="0.7">0.7 mm</quantity>
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wide opposite clavus mid-length; probably indicative of a large thorax; incompletely known in
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<taxonomicName id="4C3430F9FFDAFFDA6CC3F936C8C6F910" authorityName="Tillyard" authorityYear="1922" class="Insecta" family="Glosselytridae" genus="Polycytella" kingdom="Animalia" order="Glosselytrodea" pageId="5" pageNumber="555" phylum="Arthropoda" rank="species" species="triassica">
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<emphasis id="B9409768FFDAFFDA6CC3F936C8C6F910" italics="true" pageId="5" pageNumber="555">Polycytella triassica</emphasis>
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</taxonomicName>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C32E18F1FFDAFFDA6EF2F96EC9BAF880" pageId="5" pageNumber="555" type="materials_examined">
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<paragraph id="8B8B4B7AFFDAFFDA6EF2F96EC9BAF880" blockId="5.[136,778,1274,2093]" pageId="5" pageNumber="555">
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||
<emphasis id="B9409768FFDAFFDA6EF2F96ECA26F8F4" bold="true" box="[189,520,1778,1804]" pageId="5" pageNumber="555">Type locality and horizon.</emphasis>
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Birds River (locality code ‘Bir 111’; see
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<bibRefCitation id="EFA5368BFFDAFFDA6F27F88ACA9DF8C8" author="Anderson, J. M. & Anderson, H. M." box="[360,691,1814,1840]" pageId="5" pageNumber="555" pagination="39 - 59" refId="ref5642" refString="Anderson, J. M. & Anderson, H. M. (1984) The fossil content of the Upper Triassic Molteno Formation, South Africa. Palaeontologia Africana, 25, 39 - 59." type="journal article" year="1984">Anderson & Anderson, 1984</bibRefCitation>
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||
),
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<collectingCountry id="F3230BEAFFDAFFDA6C87F88AC8FDF8AC" name="South Africa" pageId="5" pageNumber="555">South Africa</collectingCountry>
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||
; Molteno Formation; lower Carnian, Upper Triassic (
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||
<bibRefCitation id="EFA5368BFFDAFFDA6EC0F8C2C9A9F880" author="Anderson, J. M. & Anderson, H. M. & Cruickshank, A. R. I." box="[143,391,1886,1913]" pageId="5" pageNumber="555" pagination="387 - 421" refId="ref5754" refString="Anderson, J. M., Anderson, H. M. & Cruickshank, A. R. I. (1998) Late Triassic ecosystems of the Molteno / Lower Elliot Biome of Southern Africa. Palaeontology, 41 (3), 387 - 421." type="journal article" year="1998">
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||
Anderson
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||
<emphasis id="B9409768FFDAFFDA6F4AF8C3C918F880" box="[261,310,1886,1912]" italics="true" pageId="5" pageNumber="555">et al</emphasis>
|
||
., 1998
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32E18F1FFDAFFDA6EF2F81ECC2AFAEF" pageId="5" pageNumber="555" type="description">
|
||
<paragraph id="8B8B4B7AFFDAFFDA6EF2F81ECC2AFAEF" blockId="5.[136,778,1274,2093]" lastBlockId="5.[810,1452,158,2093]" pageId="5" pageNumber="555">
|
||
<emphasis id="B9409768FFDAFFDA6EF2F81EC961F864" bold="true" box="[189,335,1922,1948]" pageId="5" pageNumber="555">Description.</emphasis>
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A pair of forewings, presumably from the same individual, with left forewing very well preserved, and outline and main veins of right forewing more or less discernible; forewing length
|
||
<quantity id="4CCCE69FFFDAFFDA6F84F872CA0AF7F1" box="[459,548,2030,2057]" metricMagnitude="-3" metricUnit="m" metricValue="9.4" pageId="5" pageNumber="555" unit="mm" value="9.4">9.4 mm</quantity>
|
||
, width
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||
<quantity id="4CCCE69FFFDAFFDA6C39F872CAE1F7F1" box="[630,719,2030,2057]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="5" pageNumber="555" unit="mm" value="2.6">2.6 mm</quantity>
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||
; area between anterior wing margin and ScP broad, with at least 4 rows of small cells; in this area, occurrence of an ambient vein parallel to the anterior wing margin; just distal to the end of ScP, area between anterior wing margin and RA broad (
|
||
<quantity id="4CCCE69FFFDAFFDA6D35FE96CBFDFEDD" box="[890,979,266,293]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="555" unit="mm" value="0.6">0.6 mm</quantity>
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||
wide opposite the end of ScP), with 4 or 5 rows of cells; area between anterior wing margin and RA gradually tapering, with 2 rows of cells until shortly before the apex; RA simple; MP+CuA
|
||
<emphasis id="B9409768FFDAFFDA6AA4FEE4CD21FE69" box="[1259,1295,376,401]" italics="true" pageId="5" pageNumber="555">ant</emphasis>
|
||
rectilinear in its basal half, then slightly deflected posteriorly; area between RA and MP+CuA
|
||
<emphasis id="B9409768FFDAFFDA6A20FE5CCCBDFE21" box="[1135,1171,448,473]" italics="true" pageId="5" pageNumber="555">ant</emphasis>
|
||
broadening from wing base to the
|
||
<emphasis id="B9409768FFDAFFDA6DF9FE78CBFEFE05" box="[950,976,484,509]" italics="true" pageId="5" pageNumber="555">ca</emphasis>
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||
. 3/5 of wing length, then tapering, with 4 veins at its broadest; area between MP+CuA
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||
<emphasis id="B9409768FFDAFFDA6B1CFD94CD59FDD9" box="[1363,1399,520,545]" italics="true" pageId="5" pageNumber="555">ant</emphasis>
|
||
and CuA
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||
<emphasis id="B9409768FFDAFFDA6D10FDB0CBA0FDBD" box="[863,910,556,581]" italics="true" pageId="5" pageNumber="555">post</emphasis>
|
||
broadening from the wing base to the
|
||
<emphasis id="B9409768FFDAFFDA6B0FFDB0CD74FDBD" box="[1344,1370,556,581]" italics="true" pageId="5" pageNumber="555">ca</emphasis>
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||
. 3/5 of wing length, then tapering, with 3 veins at its broadest; in the 3
|
||
<superScript id="7C41E632FFDAFFDA6DCBFDEFCBBCFD79" attach="left" box="[900,914,627,641]" fontSize="6" pageId="5" pageNumber="555">rd</superScript>
|
||
quarter of wing length, cells between RA and CuA
|
||
<emphasis id="B9409768FFDAFFDA6D10FD04CBA0FD49" box="[863,910,664,689]" italics="true" pageId="5" pageNumber="555">post</emphasis>
|
||
wider than long; CuA
|
||
<emphasis id="B9409768FFDAFFDA6AC2FD04CC92FD49" box="[1165,1212,664,689]" italics="true" pageId="5" pageNumber="555">post</emphasis>
|
||
simple, slightly bent opposite the end of CuP; RA, MP+CuA
|
||
<emphasis id="B9409768FFDAFFDA6AA3FD21CD3EFD2E" box="[1260,1296,701,726]" italics="true" pageId="5" pageNumber="555">ant</emphasis>
|
||
and CuA
|
||
<emphasis id="B9409768FFDAFFDA6B32FD21CD82FD2E" box="[1405,1452,701,726]" italics="true" pageId="5" pageNumber="555">post</emphasis>
|
||
joining at their apices and delimiting a single marginal band; claval furrow concave, rectilinear, closely bordered by two convex vein-like elements, the anterior one herein regarded as CuP, the posterior one as AA1 or a derivative of cross-venation (hereafter ‘AA1’); distally, claval furrow and both of these elements barely distinguishable from a single vein, ending
|
||
<quantity id="4CCCE69FFFDAFFDA6A29FC24CC91FC2B" box="[1126,1215,952,979]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="5" pageNumber="555" unit="mm" value="5.4">5.4 mm</quantity>
|
||
distal to wing base; clavus large (
|
||
<quantity id="4CCCE69FFFDAFFDA6D8CFC41CC33FC0F" box="[963,1053,989,1015]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="555" unit="mm" value="0.7">0.7 mm</quantity>
|
||
wide opposite clavus mid-length); 2 well-defined AA veins (in addition to AA1), converging towards the end of CuP, with two rows of cells between them, and between AA1 and the first of these veins; area between the second of these AA veins and posterior wing margin with 5 rows of cells, progressively longer than wide towards the margin, and organized into vein-like elements parallel to the posterior wing margin, forming a depression along it.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32E18F1FFDAFFD96D10FABDC941FD1B" lastPageId="6" lastPageNumber="556" pageId="5" pageNumber="555" type="discussion">
|
||
<paragraph id="8B8B4B7AFFDAFFDA6D10FABDCC93F979" blockId="5.[810,1452,158,2093]" pageId="5" pageNumber="555">
|
||
<emphasis id="B9409768FFDAFFDA6D10FABDCBCAFAC3" bold="true" box="[863,996,1313,1339]" pageId="5" pageNumber="555">Discussion.</emphasis>
|
||
The new material is very similar to
|
||
<taxonomicName id="4C3430F9FFDAFFDA6D65FADACC3CFAA7" authorityName="Tillyard" authorityYear="1922" box="[810,1042,1350,1376]" class="Insecta" family="Glosselytridae" genus="Polycytella" kingdom="Animalia" order="Glosselytrodea" pageId="5" pageNumber="555" phylum="Arthropoda" rank="species" species="triassica">
|
||
<emphasis id="B9409768FFDAFFDA6D65FADACC3CFAA7" box="[810,1042,1350,1376]" italics="true" pageId="5" pageNumber="555">Polycytella triassica</emphasis>
|
||
</taxonomicName>
|
||
, particularly in the relatively distal location of the end of CuP (unknown in
|
||
<taxonomicName id="4C3430F9FFDAFFDA6B4FFAF7CD59FA7C" authorityName="Béthoux & Anderson" authorityYear="2021" box="[1280,1399,1386,1412]" class="Insecta" family="Polycytellidae" genus="Moltenojurina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Glosselytrodea" pageId="5" pageNumber="555" phylum="Arthropoda" rank="species" species="parva">
|
||
<emphasis id="B9409768FFDAFFDA6B4FFAF7CD0BFA7C" box="[1280,1317,1387,1412]" italics="true" pageId="5" pageNumber="555">Mo</emphasis>
|
||
.
|
||
<emphasis id="B9409768FFDAFFDA6B79FAF7CD59FA7C" box="[1334,1399,1387,1412]" italics="true" pageId="5" pageNumber="555">parva</emphasis>
|
||
</taxonomicName>
|
||
, see below). Differences mainly regard aspect ratio, the shape of cells in the 3
|
||
<superScript id="7C41E632FFDAFFDA6D85FA2ECBF6FA38" attach="left" box="[970,984,1458,1472]" fontSize="6" pageId="5" pageNumber="555">rd</superScript>
|
||
quarter of wing length, and the occurrence, or lack thereof, of an intercalary vein between CuA
|
||
<emphasis id="B9409768FFDAFFDA6B32FA4BCD82FA08" box="[1405,1452,1495,1520]" italics="true" pageId="5" pageNumber="555">post</emphasis>
|
||
and CuP (basal to the end of the latter). Considering that it is more informative to emphasize similarities rather than differences, we propose to assign the new material to the same genus as
|
||
<taxonomicName id="4C3430F9FFDAFFDA6D9EF9FACC98F978" authorityName="Tillyard" authorityYear="1922" box="[977,1206,1638,1664]" class="Insecta" family="Glosselytridae" genus="Polycytella" kingdom="Animalia" order="Glosselytrodea" pageId="5" pageNumber="555" phylum="Arthropoda" rank="species" species="triassica">
|
||
<emphasis id="B9409768FFDAFFDA6D9EF9FACC98F978" box="[977,1206,1638,1664]" italics="true" pageId="5" pageNumber="555">Polycytella triassica</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8B4B7AFFDAFFD96D10F916C941FD1B" blockId="5.[810,1452,158,2093]" lastBlockId="6.[136,778,158,740]" lastPageId="6" lastPageNumber="556" pageId="5" pageNumber="555">
|
||
The occurrence of intercalary veins running very close to the posterior wing margin, basal to the end of CuP, is probably part of a forewing interlocking mechanism. Indeed, the occurrence of one or several vein-like structures (main and/or intercalary vein(s)) bordering the actual posterior wing margin, basal to the end of CuP or along its entire length, is common in insects with elytrized forewings, such as Dermaptera and some Dictyoptera (
|
||
<bibRefCitation id="EFA5368BFFDAFFDA6D8DF836CC45F83C" author="Tillyard, R. J." box="[962,1131,1962,1988]" pageId="5" pageNumber="555" pagination="232 - 266" refId="ref7179" refString="Tillyard, R. J. (1931) Kansas Permian insects. Part 13. The new order Protelytroptera, with a discussion of its relationships. American Journal of Science (5), 21, 232 - 266. https: // doi. org / 10.2475 / ajs. s 5 - 21.123.232" type="journal article" year="1931">Tillyard, 1931</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA5368BFFDAFFDA6A33F836CD13F83C" author="Nel, A. & Prokop, J. & Grandcolas, P. & Garrouste, R. & Lapeyrie, J. & Legendre, F. & Anisyutkin, L. N. & Kirejtshuk, A. G." box="[1148,1341,1962,1989]" pageId="5" pageNumber="555" pagination="545 - 554" refId="ref6800" refString="Nel, A., Prokop, J., Grandcolas, P., Garrouste, R., Lapeyrie, J., Legendre, F., Anisyutkin, L. N. & Kirejtshuk, A. G. (2014) The beetle-like Palaeozoic and Mesozoic roachoids of the so- called " umenocoleoid " lineage (Dictyoptera: Ponopterixidae fam. nov.). Comptes Rendus Palevol, 13 (7), 545 - 554. https: // doi. org / 10.1016 / j. crpv. 2014.05.002" type="journal article" year="2014">
|
||
Nel
|
||
<emphasis id="B9409768FFDAFFDA6AFFF837CCC9F83C" box="[1200,1255,1962,1988]" italics="true" pageId="5" pageNumber="555">et al</emphasis>
|
||
., 2014
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA5368BFFDAFFDA6B03F836CB9EF81F" author="Bethoux, O. & Llamosi, A. & Toussaint, S." pageId="5" pageNumber="555" pagination="1 - 7" refId="ref5967" refString="Bethoux, O., Llamosi, A. & Toussaint, S. (2016) Reinvestigation of Protelytron permianum (Insecta; Early Permian; USA) as an example for applying reflectance transformation imaging to insect imprint fossils. Fossil Record, 20, 1 - 7. https: // doi. org / 10.5194 / fr- 20 - 1 - 2016" type="journal article" year="2016">
|
||
Béthoux
|
||
<emphasis id="B9409768FFDAFFDA6D65F852CB73F81F" box="[810,861,1997,2023]" italics="true" pageId="5" pageNumber="555">et al</emphasis>
|
||
., 2016
|
||
</bibRefCitation>
|
||
; Luo
|
||
<emphasis id="B9409768FFDAFFDA6DBCF852CC09F81F" box="[1011,1063,1997,2023]" italics="true" pageId="5" pageNumber="555">et al</emphasis>
|
||
., in press). Most elaborated cases involve a three-dimensional tongue-and-groove structure, as in Coleoptera (
|
||
<bibRefCitation id="EFA5368BFFDAFFDA6DA2F78ECCE5F7D4" author="Breed, R. S. & Ball, E. F." box="[1005,1227,2066,2092]" pageId="5" pageNumber="555" pagination="289 - 303" refId="ref6033" refString="Breed, R. S. & Ball, E. F. (1908) The interlocking mechanisms which are found in connection with the elytra of Coleoptera. Biological Bulletin, 15 (6), 289 - 303. https: // doi. org / 10.2307 / 1535924" type="journal article" year="1908">Breed & Ball, 1908</bibRefCitation>
|
||
; and see
|
||
<bibRefCitation id="EFA5368BFFDAFFD96B7FF78EC8EAFF40" author="Nachtigall, W." lastPageId="6" lastPageNumber="556" pageId="5" pageNumber="555" refId="ref6763" refString="Nachtigall, W. (1974) Biological mechanisms of attachment. Springer-Verlag, Berlin, 194 pp. https: // doi. org / 10.1007 / 978 - 3 - 642 - 85775 - 1" type="book" year="1974">Nachtigall, 1974</bibRefCitation>
|
||
), which appears as composed of multiple, parallel vein-like elements when viewed dorsally. The bordering of the claval furrow by two vein-like elements (one of the two being CuP) is a configuration that can be observed in elytrized forewings of grasshoppers (OB, pers. observ.), among others. In
|
||
<taxonomicName id="4C3430F9FFD9FFD96F1AFED0CA65FE9D" authorityName="Béthoux & Anderson" authorityYear="2021" box="[341,587,332,358]" class="Insecta" family="Glosselytridae" genus="Polycytella" kingdom="Animalia" order="Glosselytrodea" pageId="6" pageNumber="556" phylum="Arthropoda" rank="species" species="rasnitsyni">
|
||
<emphasis id="B9409768FFD9FFD96F1AFED0CA65FE9D" box="[341,587,332,358]" italics="true" pageId="6" pageNumber="556">Polycytella rasnitsyni</emphasis>
|
||
</taxonomicName>
|
||
the interlocking mechanism may have involved the posterior margin of the clavus on one hand, and the area of the claval furrow on the other. Indeed, when folded along the claval furrow, the posterior margin of the clavus of the overlapping forewing overlaps the claval furrow of the overlapped forewing. To best appreciate this the reader is encouraged to cut wings illustrated on
|
||
<figureCitation id="130F57FFFFD9FFD96F67FDA2C942FDA1" box="[296,364,574,601]" captionStart="FIGURE 3" captionStartId="4.[136,235,1039,1063]" captionTargetBox="[192,1373,208,1007]" captionTargetId="figure-435@4.[183,1404,204,1016]" captionTargetPageId="4" captionText="FIGURE 3. Polycytella rasnitsyni sp. nov., holotype specimen (PRE/F/10559), interpretative drawings, separated. A, Left forewing, as in Fig. 2A. B, Right forewing, as in Fig. 2A and with reconstructed parts." figureDoi="http://doi.org/10.5281/zenodo.5778586" httpUri="https://zenodo.org/record/5778586/files/figure.png" pageId="6" pageNumber="556">Fig. 3</figureCitation>
|
||
along their margin, use the tip of a pointed blade (or a needle) and a ruler to gently imprint a line of weakness along the claval furrow, and assemble the wing pair in resting position, with the claval furrow forming a 90° angle.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |