treatments-xml/data/EB/32/87/EB3287FD0277E70B54841FB7FBC9B586.xml
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<document id="9ED83BAA77B7439146DFBFE48EAF90A4" ID-DOI="10.25221/fee.496.1" ID-ISSN="2713-2196" ID-Zenodo-Dep="10928400" ID-ZooBank="43C2B903-897A-40B0-8100-8BDA3B771D14" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1712258596968" checkinUser="felipe" docAuthor="Matsuura, K., Hosoishi, S. &amp; Sokh, H." docDate="2024" docId="EB3287FD0277E70B54841FB7FBC9B586" docLanguage="en" docName="FarEastEntomol.496.1-9.pdf.imf" docOrigin="Far Eastern Entomologist 496" docSource="http://dx.doi.org/10.25221/fee.496.1" docStyle="DocumentStyle:68641B8F4A21450A5F9A74164B730F50.12:FarEastEntomol.2017-.journal_article" docStyleId="68641B8F4A21450A5F9A74164B730F50" docStyleName="FarEastEntomol.2017-.journal_article" docStyleVersion="12" docTitle="Carebara bokorensis Matsuura et Hosoishi 2024, sp. n." docType="treatment" docVersion="4" lastPageNumber="8" masterDocId="170BFF850275E70C54671C78FFA9B43F" masterDocTitle="A remarkable new species from the pygmaea species group of the genus Carebara Westwood, 1840 (Hymenoptera: Formicidae: Myrmicinae) from Cambodia" masterLastPageNumber="9" masterPageNumber="1" pageNumber="3" updateTime="1712599832898" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="CCADA3DA94074262914E3E87E1E28404">A remarkable new species from the pygmaea species group of the genus Carebara Westwood, 1840 (Hymenoptera: Formicidae: Myrmicinae) from Cambodia</mods:title>
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<mods:namePart id="5092A00E3FA468DD602D077E56567518">Matsuura, K.</mods:namePart>
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<mods:namePart id="60B3AED933BECEED420BC6103832E496">Hosoishi, S.</mods:namePart>
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<mods:namePart id="6B343E16B868B7D6D1B8BBBBF696AFEF">Sokh, H.</mods:namePart>
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<mods:date id="EE04B3BC41D3FC2B980AEC179D30211F">2024</mods:date>
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<mods:identifier id="86011C287C6B4D9385E0F6BC7DD10987" type="DOI">10.25221/fee.496.1</mods:identifier>
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<treatment id="EB3287FD0277E70B54841FB7FBC9B586" LSID="urn:lsid:plazi:treatment:EB3287FD0277E70B54841FB7FBC9B586" httpUri="http://treatment.plazi.org/id/EB3287FD0277E70B54841FB7FBC9B586" lastPageId="7" lastPageNumber="8" pageId="2" pageNumber="3">
<subSubSection id="2B8165600277E70E54841FB7FB90B037" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="632436EB0277E70E54841FB7FCA5B7D8" blockId="2.[227,780,975,999]" box="[227,780,975,999]" pageId="2" pageNumber="3">
<heading id="386C81870277E70E54841FB7FCA5B7D8" box="[227,780,975,999]" centered="true" fontSize="9" level="2" pageId="2" pageNumber="3" reason="2">
<emphasis id="51EFEAF90277E70E54841FB7FCA5B7D8" bold="true" box="[227,780,975,999]" pageId="2" pageNumber="3">
<taxonomicName id="A49B4D680277E70E54841FB7FD6DB7D8" authority="Matsuura et Hosoishi" authorityName="Matsuura et Hosoishi" authorityYear="2024" box="[227,708,975,999]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="bokorensis" status="sp. nov.">
<emphasis id="51EFEAF90277E70E54841FB7FE61B7D8" bold="true" box="[227,456,975,999]" italics="true" pageId="2" pageNumber="3">Carebara bokorensis</emphasis>
Matsuura et Hosoishi
</taxonomicName>
,
<taxonomicNameLabel id="4ADC57820277E70E56B71FB7FCA5B7D8" box="[720,780,975,999]" pageId="2" pageNumber="3" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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https://zoobank.org/NomenclaturalActs/
<uuid id="173D0C3E0277E70E56171F8AFB90B037" box="[624,1081,1010,1032]" pageId="2" pageNumber="3">9F1A8383-F9EE-4C51-9D94-BFD5AC006CA9</uuid>
</paragraph>
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<subSubSection id="2B8165600277E70E5484186CFE96B012" box="[227,319,1044,1069]" pageId="2" pageNumber="3" type="description">
<paragraph id="632436EB0277E70E5484186CFE96B012" blockId="2.[227,319,1044,1069]" box="[227,319,1044,1069]" pageId="2" pageNumber="3">
<figureCitation id="FBA02A6E0277E70E5484186CFE8CB012" box="[227,293,1044,1069]" captionStart="Fig" captionStartId="4.[265,300,1393,1415]" captionTargetBox="[236,1117,257,1358]" captionTargetId="figure-0@4.[236,1117,257,1359]" captionTargetPageId="4" captionText="Fig. 1. AC. Carebara bokorensis sp. n., holotype major worker (KUECCRB001). A body in profile; B dorsal view of body; C head in full-face view." figureDoi="http://doi.org/10.5281/zenodo.10945886" httpUri="https://zenodo.org/record/10945886/files/figure.png" pageId="2" pageNumber="3">Figs 1</figureCitation>
,
<figureCitation id="FBA02A6E0277E70E5555186DFE96B012" box="[306,319,1045,1069]" captionStart="Fig" captionStartId="5.[265,300,1368,1390]" captionTargetBox="[246,1108,257,1335]" captionTargetId="figure-0@5.[246,1108,257,1335]" captionTargetPageId="5" captionText="Fig. 2. AC. Carebara bokorensis sp. n., paratype minor worker (KUECCRB004). A body in profile; B dorsal view of body; C head in full-face view." figureDoi="http://doi.org/10.5281/zenodo.10945888" httpUri="https://zenodo.org/record/10945888/files/figure.png" pageId="2" pageNumber="3">2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="2B8165600277E70E556E182BFDE4B11B" pageId="2" pageNumber="3" type="materials_examined">
<paragraph id="632436EB0277E70E556E182BFDE4B11B" blockId="2.[227,1127,1106,1594]" pageId="2" pageNumber="3">
<materialsCitation id="D3F33CB60277E70E556E182BFBCAB097" collectionCode="TYPE, MATERIAL, KUEC" collectorName="S. Hosoishi" country="Cambodia" elevation="663" latitude="10.6" location="Bokor National Park" longLatPrecision="925" longitude="104.103165" pageId="2" pageNumber="3" specimenCount="1" stateProvince="Kampot" typeStatus="holotype">
<collectionCode id="058AAE2E0277E70E556E182BFEE5B055" box="[265,332,1107,1130]" pageId="2" pageNumber="3">TYPE</collectionCode>
<collectionCode id="058AAE2E0277E70E553C182AFE41B055" box="[347,488,1106,1130]" pageId="2" pageNumber="3">MATERIAL</collectionCode>
.
<typeStatus id="BC2088490277E70E559D182AFDF4B055" box="[506,605,1106,1130]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
(major worker):
<emphasis id="51EFEAF90277E70E574B182AFC0AB055" bold="true" box="[812,931,1106,1130]" pageId="2" pageNumber="3">
<collectingCountry id="1B8C767B0277E70E574B182AFC0AB055" box="[812,931,1106,1130]" name="Cambodia" pageId="2" pageNumber="3">Cambodia</collectingCountry>
</emphasis>
:
<location id="664460300277E70E57DE182AFEBEB0B6" LSID="urn:lsid:plazi:treatment:EB3287FD0277E70B54841FB7FBC9B586:664460300277E70E57DE182AFEBEB0B6" country="Cambodia" latitude="10.6" longLatPrecision="925" longitude="104.103165" name="Bokor National Park" pageId="2" pageNumber="3" stateProvince="Kampot">Bokor National Park</location>
,
<quantity id="A4639B0E0277E70E55431809FEC9B0B6" box="[292,352,1137,1161]" metricMagnitude="2" metricUnit="m" metricValue="6.63" pageId="2" pageNumber="3" unit="m" value="663.0">
<elevation id="E8B6D1D80277E70E55431809FEC9B0B6" box="[292,352,1137,1161]" metricMagnitude="2" metricUnit="m" metricValue="6.63" pageId="2" pageNumber="3" unit="m" value="663.0">663m</elevation>
</quantity>
alt.,
<collectingRegion id="A15FF8090277E70E55FB180AFDF7B0B6" box="[412,606,1137,1161]" country="Cambodia" name="Kampot" pageId="2" pageNumber="3">Kampot Province</collectingRegion>
,
<geoCoordinate id="06AF502C0277E70E560C180AFD61B0B6" box="[619,712,1137,1161]" degrees="10" direction="north" minutes="36" orientation="latitude" pageId="2" pageNumber="3" precision="925" value="10.6">N 10°36</geoCoordinate>
<geoCoordinate id="06AF502C0277E70E56B5180AFCCEB0B6" box="[722,871,1137,1161]" degrees="104" direction="east" minutes="06, 19" orientation="longitude" pageId="2" pageNumber="3" precision="9" value="104.103165">E 104°06, 19</geoCoordinate>
. XII 2011,
<collectorName id="CE6E533D0277E70E578F1809FBCFB0B6" box="[1000,1126,1137,1161]" pageId="2" pageNumber="3">S. Hosoishi</collectorName>
leg. SH11-Cam-55, decayed wood (
<collectionCode id="058AAE2E0277E70E56E818E8FD75B097" box="[655,732,1168,1192]" country="Japan" httpUri="http://grbio.org/cool/txbq-e9hf" name="Kyushu University Entomology Collection" pageId="2" pageNumber="3">KUEC</collectionCode>
, collection code KUECCRB001)
</materialsCitation>
.
<materialsCitation id="D3F33CB60277E70E548418D7FDE1B11B" collectionCode="KUEC" collectorName="S. Hosoishi" country="Cambodia" elevation="663" latitude="10.6" location="Bokor National Park" longLatPrecision="925" longitude="104.103165" pageId="2" pageNumber="3" specimenCount="1" stateProvince="Kampot" typeStatus="paratype">
<typeStatus id="BC2088490277E70E548418D7FEF8B0F8" box="[227,337,1199,1223]" pageId="2" pageNumber="3" type="paratype">Paratypes</typeStatus>
: same data as holotype, 2 major workers (
<collectionCode id="058AAE2E0277E70E573F18D7FC0FB0F8" box="[856,934,1199,1223]" country="Japan" httpUri="http://grbio.org/cool/txbq-e9hf" name="Kyushu University Entomology Collection" pageId="2" pageNumber="3">KUEC</collectionCode>
, collection code KUECCRB002, KUECCRB003) and 7 minor workers (
<collectionCode id="058AAE2E0277E70E573818B6FC05B0D9" box="[863,940,1230,1254]" country="Japan" httpUri="http://grbio.org/cool/txbq-e9hf" name="Kyushu University Entomology Collection" pageId="2" pageNumber="3">KUEC</collectionCode>
, collection code KUECCRB004, KUECCRB005, KUECCRB006, KUECCRB007, KUECCRB008, KUECCRB009, KUECCRB010)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="2B8165600277E70A556E1953FECBB716" lastPageId="6" lastPageNumber="7" pageId="2" pageNumber="3" type="description">
<paragraph id="632436EB0277E70E556E1953FE27B205" blockId="2.[227,1127,1106,1594]" pageId="2" pageNumber="3">
<materialsCitation id="D3F33CB60277E70E556E1953FE27B205" collectionCode="ML, MW, PSL, PSD, PL" country="Philippines" location="Major" municipality="Minor" pageId="2" pageNumber="3" specimenCode="PI1, PI2" specimenCount="9" typeStatus="holotype">
MEASUREMENTS.
<typeStatus id="BC2088490277E70E55961953FDF0B17C" box="[497,601,1323,1347]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
: TL 4.23, HL 0.97, HW 0.95, SL 0.44,
<collectionCode id="058AAE2E0277E70E50651953FB83B17D" box="[1026,1066,1323,1346]" country="France" name="Musee de Lectoure" pageId="2" pageNumber="3">ML</collectionCode>
0.89,
<collectionCode id="058AAE2E0277E70E54841932FEB8B15E" box="[227,273,1354,1377]" country="Netherlands" name="Museum Wasmann" pageId="2" pageNumber="3">MW</collectionCode>
0.42,
<collectionCode id="058AAE2E0277E70E553A1932FE21B15D" box="[349,392,1354,1378]" pageId="2" pageNumber="3">PSL</collectionCode>
0.17,
<collectionCode id="058AAE2E0277E70E55B31932FDABB15D" box="[468,514,1354,1378]" pageId="2" pageNumber="3">PSD</collectionCode>
0.07,
<collectionCode id="058AAE2E0277E70E56291932FDC3B15E" box="[590,618,1354,1377]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="2" pageNumber="3" type="Museum">PL</collectionCode>
0.37,
<collectingCountry id="1B8C767B0277E70E56D11932FD7CB15E" box="[694,725,1354,1377]" name="Philippines" pageId="2" pageNumber="3">PH</collectingCountry>
0.35, DPW 0.18, CI 98, SI 47,
<specimenCode id="333D9E900277E70E50261932FBCAB15D" box="[1089,1123,1354,1378]" collectionCode="PI" country="Russia" name="Paleontological Institute" pageId="2" pageNumber="3">PI1</specimenCode>
106,
<specimenCode id="333D9E900277E70E557D1911FEE9B1BE" box="[282,320,1385,1409]" collectionCode="PI" country="Russia" name="Paleontological Institute" pageId="2" pageNumber="3">PI2</specimenCode>
48.
<location id="664460300277E70E55161910FE1BB1BF" LSID="urn:lsid:plazi:treatment:EB3287FD0277E70B54841FB7FBC9B586:664460300277E70E55161910FE1BB1BF" box="[369,434,1384,1408]" country="Philippines" municipality="Minor" name="Major" pageId="2" pageNumber="3">Major</location>
worker (
<specimenCount id="759DFD620277E70E56701911FD33B1BE" box="[535,666,1385,1409]" count="2" pageId="2" pageNumber="3" type="generic" typeStatus="paratype">2 paratypes</specimenCount>
): TL 4.204.26, HL 0.921.25, HW 0.93 1.02, SL 0.430.44,
<collectionCode id="058AAE2E0277E70E55D919F0FE4FB1A0" box="[446,486,1416,1439]" country="France" name="Musee de Lectoure" pageId="2" pageNumber="3">ML</collectionCode>
0.890.92,
<collectionCode id="058AAE2E0277E70E560119F0FD3EB1A0" box="[614,663,1416,1439]" country="Netherlands" name="Museum Wasmann" pageId="2" pageNumber="3">MW</collectionCode>
0.420.50,
<collectionCode id="058AAE2E0277E70E577119F0FCEDB19F" box="[790,836,1416,1440]" pageId="2" pageNumber="3">PSL</collectionCode>
0.160.18,
<collectionCode id="058AAE2E0277E70E57A319F0FC5CB19F" box="[964,1013,1416,1440]" pageId="2" pageNumber="3">PSD</collectionCode>
0.060.07
<collectionCode id="058AAE2E0277E70E548419DFFEABB181" box="[227,258,1447,1470]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="2" pageNumber="3" type="Museum">PL</collectionCode>
0.340.37,
<collectingCountry id="1B8C767B0277E70E55E419DFFE0CB181" box="[387,421,1447,1470]" name="Philippines" pageId="2" pageNumber="3">PH</collectingCountry>
0.330.35, DPW 0.180.23, CI 98101, SI 4647,
<specimenCode id="333D9E900277E70E57B219DEFC52B181" box="[981,1019,1446,1470]" collectionCode="PI" country="Russia" name="Paleontological Institute" pageId="2" pageNumber="3">PI1</specimenCode>
102106,
<specimenCode id="333D9E900277E70E548419BDFEA1B1E2" box="[227,264,1477,1501]" collectionCode="PI" country="Russia" name="Paleontological Institute" pageId="2" pageNumber="3">PI2</specimenCode>
4868.
<collectingMunicipality id="8340AC910277E70E550219BDFE01B1E2" box="[357,424,1477,1501]" pageId="2" pageNumber="3">Minor</collectingMunicipality>
worker (
<specimenCount id="759DFD620277E70E566819BDFD3BB1E2" box="[527,658,1477,1501]" count="7" pageId="2" pageNumber="3" type="generic" typeStatus="paratype">7 paratypes</specimenCount>
): TL 1.431.68, HL 0.330.41, HW 0.33 0.38, SL 0.230.26,
<collectionCode id="058AAE2E0277E70E55AB199DFE5DB1C3" box="[460,500,1509,1532]" country="France" name="Musee de Lectoure" pageId="2" pageNumber="3">ML</collectionCode>
0.390.45,
<collectionCode id="058AAE2E0277E70E561A199DFD07B1C3" box="[637,686,1509,1532]" country="Netherlands" name="Museum Wasmann" pageId="2" pageNumber="3">MW</collectionCode>
0.160.22,
<collectionCode id="058AAE2E0277E70E5750199CFCCCB1C3" box="[823,869,1508,1532]" pageId="2" pageNumber="3">PSL</collectionCode>
0.040.05,
<collectionCode id="058AAE2E0277E70E5789199CFBB6B1C3" box="[1006,1055,1508,1532]" pageId="2" pageNumber="3">PSD</collectionCode>
0.03 0.04,
<collectionCode id="058AAE2E0277E70E55781A7CFE97B224" box="[287,318,1540,1563]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="2" pageNumber="3" type="Museum">PL</collectionCode>
0.170.22,
<collectingCountry id="1B8C767B0277E70E55DA1A7CFE76B224" box="[445,479,1540,1563]" name="Philippines" pageId="2" pageNumber="3">PH</collectingCountry>
0.140.15, DPW 0.080.10, CI 90100, SI 6771,
<specimenCode id="333D9E900277E70E50621A7BFB83B224" box="[1029,1066,1539,1563]" collectionCode="PI" country="Russia" name="Paleontological Institute" pageId="2" pageNumber="3">PI1</specimenCode>
117 152,
<specimenCode id="333D9E900277E70E557F1A5AFE97B205" box="[280,318,1570,1594]" collectionCode="PI" country="Russia" name="Paleontological Institute" pageId="2" pageNumber="3">PI2</specimenCode>
4354.
</materialsCitation>
</paragraph>
<paragraph id="632436EB0276E70F556E1D70FE9FB606" blockId="3.[227,1127,264,1634]" pageId="3" pageNumber="4">
DESCRIPTION. Dimorphism in worker caste.
<emphasis id="51EFEAF90276E70F57461D70FC63B51F" bold="true" box="[801,970,264,288]" pageId="3" pageNumber="4">Major worker</emphasis>
(
<figureCitation id="FBA02A6E0276E70F57B91D70FBFCB51F" box="[990,1109,264,288]" captionStart="Fig" captionStartId="4.[265,300,1393,1415]" captionTargetBox="[236,1117,257,1358]" captionTargetId="figure-0@4.[236,1117,257,1359]" captionTargetPageId="4" captionText="Fig. 1. AC. Carebara bokorensis sp. n., holotype major worker (KUECCRB001). A body in profile; B dorsal view of body; C head in full-face view." figureDoi="http://doi.org/10.5281/zenodo.10945886" httpUri="https://zenodo.org/record/10945886/files/figure.png" pageId="3" pageNumber="4">Fig. 1AC</figureCitation>
). Head, in full-face view, longer than wide, almost square (CI 98101); posterior cephalic border deeply concave in the middle, posterolateral corners rounded, posterior lateral margins weakly convex; anterior clypeal margin entirely convex with median concave portion; frontal carinae inconspicuous; frontal triangle welldefined; mandibles short, almost triangular, masticatory margins with 5 teeth; ocelli present; eyes with around 12 ommatidia, situated at anterior one-third of lateral head; antennae 11-segmented with 2-segmented club; scapes thin and clavate, its apical portion same as antennal segment X, short, less than even half of head (SI 4647).
</paragraph>
<paragraph id="632436EB0276E70F556E1E39FEB7B70B" blockId="3.[227,1127,264,1634]" pageId="3" pageNumber="4">
Mesosoma smaller than head (HL 0.921.25,
<collectionCode id="058AAE2E0276E70F576A1E39FC9CB667" box="[781,821,577,600]" country="France" name="Musee de Lectoure" pageId="3" pageNumber="4">ML</collectionCode>
0.890.92); promesonotum weakly convex in profile view; promesonotal suture present; scutellum present; mesopleuron clearly demarcated from pronotum and metapleuron by deep groove; metanotal groove present and separated by elevated band; metapleuron clearly demarcated from propodeum by deep groove; propodeum declining backwards; propodeal spiracle large (
<collectionCode id="058AAE2E0276E70F55911EA5FD80B6CA" box="[502,553,733,757]" pageId="3" pageNumber="4">PSD</collectionCode>
0.060.07) and rounded; propodeal spines relatively long (
<collectionCode id="058AAE2E0276E70F557C1E85FEE2B72A" box="[283,331,765,789]" pageId="3" pageNumber="4">PSL</collectionCode>
0.160.18); promesonotum distinctly higher than propodeum in profile view.
</paragraph>
<paragraph id="632436EB0276E70F556E1F43FDE7B7AD" blockId="3.[227,1127,264,1634]" pageId="3" pageNumber="4">Petiole with long peduncle and high node, broader posteriorly in dorsal view; subpetiolar process absent; postpetiole rounded and shorter than petiole, lower than petiole node, trapezoidal dorsally.</paragraph>
<paragraph id="632436EB0276E70F556E1FE1FD57B78D" blockId="3.[227,1127,264,1634]" box="[265,766,921,946]" pageId="3" pageNumber="4">Gastral tergite larger than remaining segments.</paragraph>
<paragraph id="632436EB0276E70F556E1FC0FDA8B136" blockId="3.[227,1127,264,1634]" pageId="3" pageNumber="4">Sculptures: Head, posterior one-third of head punctate anterior two-third with longitudinal striation, lateral (area below eye) face striated; clypeus smooth and shiny medially, lateral sides longitudinally striated; frontal triangle smooth and shiny; frontal lobes with fine longitudinal striation; mandibles smooth and shiny; anterior dorsal surface of pronotum with faintly reticulations, smooth and shiny laterally; mesonotum punctate; mesopleuron separated by deep groove, both higher and lower mesopleuron smooth and shiny except for surroundings, under part of lower mesopleuron punctate; propodeum entirely punctate, but lower lateral face longitudinally striated; legs smooth and shiny; petiolar peduncle punctate, petiolar node micropunctate laterally, anterior declivity of node with reticulation; gastral tergite I smooth and shiny.</paragraph>
<paragraph id="632436EB0276E70F556E1969FEF0B21C" blockId="3.[227,1127,264,1634]" pageId="3" pageNumber="4">Pilosity: Head, in full-face view, posterior cephalic border with short decumbent hairs medially and long suberect hairs laterally, short decumbent hairs over surface; frontal carinae with 3 pairs of erect hairs; anterior margin of clypeus with 2 pairs of erect hairs; mandible, masticatory margins with long decumbent hairs, short decumbent hairs over surface; scapes with dense short decumbent hairs (slightly shorter than width of apical portion of scapes); mesosoma with erect hairs and decumbent hairs; legs with decumbent hairs; petiolar node and postpetiole with erect hairs; gastral tergite I with abundant short decumbent hairs and entirely scattered long erect hairs.</paragraph>
<paragraph id="632436EB0276E70F556E1A52FD1AB25D" blockId="3.[227,1127,264,1634]" pageId="3" pageNumber="4">Color: Head and gaster dark brown; mesosoma, petiole and postpetiole reddish brown; antennae and legs yellowish brown.</paragraph>
<caption id="37E466630271E708556E1909FCDAB19D" ID-DOI="http://doi.org/10.5281/zenodo.10945886" ID-Zenodo-Dep="10945886" httpUri="https://zenodo.org/record/10945886/files/figure.png" pageId="4" pageNumber="5" startId="4.[265,300,1393,1415]" targetBox="[236,1117,257,1358]" targetPageId="4" targetType="figure">
<paragraph id="632436EB0271E708556E1909FCDAB19D" blockId="4.[227,1127,1393,1442]" pageId="4" pageNumber="5">
Fig. 1. AC.
<taxonomicName id="A49B4D680271E70855EA1909FDF3B1B9" authority="Matsuura &amp; Hosoishi &amp; Sokh, 2024" authorityName="Matsuura &amp; Hosoishi &amp; Sokh" authorityYear="2024" box="[397,602,1393,1414]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="bokorensis" status="sp. nov.">
<emphasis id="51EFEAF90271E70855EA1909FDF3B1B9" box="[397,602,1393,1414]" italics="true" pageId="4" pageNumber="5">Carebara bokorensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4ADC57820271E70856051909FD30B1B9" box="[610,665,1393,1414]" pageId="4" pageNumber="5" rank="species">sp. n.</taxonomicNameLabel>
, holotype major worker (KUECCRB001). A body in profile; B dorsal view of body; C head in full-face view.
</paragraph>
</caption>
<paragraph id="632436EB0271E709556E19BFFE15B21C" blockId="4.[227,1127,1479,1627]" lastBlockId="5.[227,1127,1455,1632]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
<emphasis id="51EFEAF90271E708556E19BFFE06B1DF" bold="true" box="[265,431,1479,1504]" pageId="4" pageNumber="5">Minor worker</emphasis>
. (
<figureCitation id="FBA02A6E0271E70855AF19B0FDE8B1DF" box="[456,577,1480,1504]" captionStart="Fig" captionStartId="5.[265,300,1368,1390]" captionTargetBox="[246,1108,257,1335]" captionTargetId="figure-0@5.[246,1108,257,1335]" captionTargetPageId="5" captionText="Fig. 2. AC. Carebara bokorensis sp. n., paratype minor worker (KUECCRB004). A body in profile; B dorsal view of body; C head in full-face view." figureDoi="http://doi.org/10.5281/zenodo.10945888" httpUri="https://zenodo.org/record/10945888/files/figure.png" pageId="4" pageNumber="5">Fig. 2AC</figureCitation>
). Head, in full-face view, wide same as long (CI 90100), almost rectangle; posterior cephalic border slightly concave in the middle, posterolateral corners rounded, posterior lateral margins weakly convex; anterior clypeal margin convex, medially concave, frontal lobe present; frontal triangle welldefined; mandibles short, almost triangular, masticatory margins with 5 teeth; eyes relatively small, present as dots, situated at anterior one-fourth of lateral head; antennae 11-segmented with 2-segmented club; scapes thin and clavate, width of apical portion same as antennal segment X, relatively long, reaching posterior one-third of head length.
</paragraph>
<caption id="37E466630270E709556E1920FCDAB1B5" ID-DOI="http://doi.org/10.5281/zenodo.10945888" ID-Zenodo-Dep="10945888" httpUri="https://zenodo.org/record/10945888/files/figure.png" pageId="5" pageNumber="6" startId="5.[265,300,1368,1390]" targetBox="[246,1108,257,1335]" targetPageId="5" targetType="figure">
<paragraph id="632436EB0270E709556E1920FCDAB1B5" blockId="5.[227,1102,1368,1418]" pageId="5" pageNumber="6">
Fig. 2. AC.
<taxonomicName id="A49B4D680270E70955E11920FDFBB152" authority="Matsuura &amp; Hosoishi &amp; Sokh, 2024" authorityName="Matsuura &amp; Hosoishi &amp; Sokh" authorityYear="2024" box="[390,594,1368,1389]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="bokorensis" status="sp. nov.">
<emphasis id="51EFEAF90270E70955E11920FDFBB152" box="[390,594,1368,1389]" italics="true" pageId="5" pageNumber="6">Carebara bokorensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4ADC57820270E70956301921FD25B151" box="[599,652,1369,1390]" pageId="5" pageNumber="6" rank="species">sp. n.</taxonomicNameLabel>
, paratype minor worker (KUECCRB004). A body in profile; B dorsal view of body; C head in full-face view.
</paragraph>
</caption>
<paragraph id="632436EB0270E70A556E1A52FE6DB5A4" blockId="5.[227,1127,1455,1632]" lastBlockId="6.[227,1127,264,1606]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
Promesonotum convex and entirely dome-shaped in profile view; promesonotal suture absent; mesopleuron clearly demarcated from metapleuron and pronotum; metanotal groove distinct; in profile view, declivity of propodeum with lamellae, maximum length of lamellae same as a diameter of propodeal spiracle; propodeal spines absent; propodeal spiracle large (
<collectionCode id="058AAE2E0273E70A56E91D3DFD69B562" box="[654,704,325,349]" pageId="6" pageNumber="7">PSD</collectionCode>
0.030.04) and rounded; dorsal outline of propodeum entirely rounded; mesosoma almost same length as head (
<collectionCode id="058AAE2E0273E70A579F1D1CFB89B543" box="[1016,1056,356,380]" country="France" name="Musee de Lectoure" pageId="6" pageNumber="7">ML</collectionCode>
0.39 0.45, HL 0.330.41).
</paragraph>
<paragraph id="632436EB0273E70A556E1DD9FE9AB62A" blockId="6.[227,1127,264,1606]" pageId="6" pageNumber="7">Petiole with relatively short peduncle and high node, spread posteriorly in dorsal view; subpetiolar process developed like triangle; postpetiole rounded and lower than petiolar node in profile view, rectangle in dorsal view; postpetiole smaller than petiole.</paragraph>
<paragraph id="632436EB0273E70A556E1E64FD57B60B" blockId="6.[227,1127,264,1606]" box="[265,766,540,564]" pageId="6" pageNumber="7">Gastral tergite larger than remaining segments.</paragraph>
<paragraph id="632436EB0273E70A556E1E42FC43B64E" blockId="6.[227,1127,264,1606]" pageId="6" pageNumber="7">Sculptures: Head, mandibles, promesonotum, legs, petiolar node and gaster smooth and shiny; mesopleuron, propodeum, petiolar peduncle punctate.</paragraph>
<paragraph id="632436EB0273E70A556E1E00FBF2B6D3" blockId="6.[227,1127,264,1606]" pageId="6" pageNumber="7">Pilosity: Head and mandibles with sparse decumbent hairs; scapes with sparse decumbent hairs (slightly shorter than width of apical portion of scapes); anterior margin of clypeus with 2 pairs of erect hairs; mesosoma, petiole and postpetiole without erect hairs; legs with decumbent hairs; gastral tergite I with decumbent hairs.</paragraph>
<paragraph id="632436EB0273E70A556E1E8AFECBB716" blockId="6.[227,1127,264,1606]" pageId="6" pageNumber="7">Color: Head, postpetiole and gaster brown; mesosoma, petiole, legs and antennae pale brown.</paragraph>
</subSubSection>
<subSubSection id="2B8165600273E70A556E1F48FE74B03F" pageId="6" pageNumber="7" type="diagnosis">
<paragraph id="632436EB0273E70A556E1F48FE74B03F" blockId="6.[227,1127,264,1606]" pageId="6" pageNumber="7">
DIAGNOSIS.
<taxonomicName id="A49B4D680273E70A55CF1F48FD97B777" authority="Matsuura &amp; Hosoishi &amp; Sokh, 2024" authorityName="Matsuura &amp; Hosoishi &amp; Sokh" authorityYear="2024" box="[424,574,816,840]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="bokorensis" status="sp. nov.">
<emphasis id="51EFEAF90273E70A55CF1F48FE13B778" box="[424,442,816,839]" italics="true" pageId="6" pageNumber="7">C</emphasis>
.
<emphasis id="51EFEAF90273E70A55AD1F48FD97B777" box="[458,574,816,840]" italics="true" pageId="6" pageNumber="7">bokorensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4ADC57820273E70A56201F48FD2BB778" box="[583,642,816,839]" pageId="6" pageNumber="7" rank="species">sp. n.</taxonomicNameLabel>
can be distinguished from the other species in the genus by the following characters: 11-segmented antennae, punctation on posterior surface of head, long petiolar pedicel, well-developed scutellum and metanotal groove separated by elevated band in the major worker subcaste; 11- segmented antennae, median concavity of anterior clypeal margin, entirely rounded promesonotum, short petiolar pedicel and presence of subpetiolar process in the minor worker subcaste.
</paragraph>
</subSubSection>
<subSubSection id="2B8165600273E70B556E187EFE23B562" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="discussion">
<paragraph id="632436EB0273E70A556E187EFDE2B1B1" blockId="6.[227,1127,264,1606]" pageId="6" pageNumber="7">
REMARKS.
<taxonomicName id="A49B4D680273E70A55FF187EFDD3B021" authority="Matsuura &amp; Hosoishi &amp; Sokh, 2024" authorityName="Matsuura &amp; Hosoishi &amp; Sokh" authorityYear="2024" box="[408,634,1030,1054]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="bokorensis" status="sp. nov.">
<emphasis id="51EFEAF90273E70A55FF187EFDD3B021" box="[408,634,1030,1054]" italics="true" pageId="6" pageNumber="7">Carebara bokorensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4ADC57820273E70A56E5187FFD14B021" box="[642,701,1031,1054]" pageId="6" pageNumber="7" rank="species">sp. n.</taxonomicNameLabel>
shares most features except for promesonotal suture with the
<emphasis id="51EFEAF90273E70A558A185DFE56B003" box="[493,511,1061,1084]" italics="true" pageId="6" pageNumber="7">C</emphasis>
.
<emphasis id="51EFEAF90273E70A5677185EFDD9B002" box="[528,624,1062,1085]" italics="true" pageId="6" pageNumber="7">pygmaea</emphasis>
species group.
<bibRefCitation id="070A4B1A0273E70A5744185DFBA6B002" author="Jaitrong, W. &amp; Pitaktunsakul, P. &amp; Jantarit, S." box="[803,1039,1061,1085]" pageId="6" pageNumber="7" pagination="7 - 20" refId="ref3512" refString="Jaitrong, W., Pitaktunsakul, P. &amp; Jantarit, S. 2021. A new species of the genus Carebara Westwood, 1840 (Hymenoptera: Formicidae: Myrmicinae) inhabits a cave in Thailand. Far Eastern Entomologist, 425: 7 - 20. DOI: 10.25221 / fee. 425.2" type="journal article" year="2021">
Jaitrong
<emphasis id="51EFEAF90273E70A57E4185EFC1FB002" box="[899,950,1061,1085]" italics="true" pageId="6" pageNumber="7">et al</emphasis>
. (2021)
</bibRefCitation>
defines promesonotal suture of major workers as indistinct in definition of
<emphasis id="51EFEAF90273E70A5785183CFC5DB064" box="[994,1012,1092,1115]" italics="true" pageId="6" pageNumber="7">C</emphasis>
.
<emphasis id="51EFEAF90273E70A5061183DFBCFB063" box="[1030,1126,1093,1116]" italics="true" pageId="6" pageNumber="7">pygmaea</emphasis>
species group, and this is thought to be due to the exceptionally partial presence of promesonotal suture in
<taxonomicName id="A49B4D680273E70A558B18F9FDF0B0A6" authorityName="Jaitrong, Pitaktunsakul et Jantarit" authorityYear="2021" box="[492,601,1153,1177]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="panhai">
<emphasis id="51EFEAF90273E70A558B18F9FE57B0A7" box="[492,510,1153,1176]" italics="true" pageId="6" pageNumber="7">C</emphasis>
.
<emphasis id="51EFEAF90273E70A566818F9FDF0B0A6" box="[527,601,1153,1177]" italics="true" pageId="6" pageNumber="7">panhai</emphasis>
</taxonomicName>
. This new species has a distinct promesonotal suture in major workers, but other morphological characters are consistent with the definition of the species group. Therefore, it is appropriate to include this new species in the species group. Furthermore, the major workers of this species also have scutellum, which is not found in the species group. Developed scutellum in major workers are found in some species such as
<emphasis id="51EFEAF90273E70A57431963FC9FB10D" box="[804,822,1307,1330]" italics="true" pageId="6" pageNumber="7">C</emphasis>
.
<emphasis id="51EFEAF90273E70A572E1963FCC2B10D" box="[841,875,1307,1330]" italics="true" pageId="6" pageNumber="7">oni</emphasis>
(Terayama, 1996),
<taxonomicName id="A49B4D680273E70A50291963FC85B16E" authority="(Fischer, Azorsa &amp; Hita Garcia, 2015)" baseAuthorityName="Fischer, Azorsa &amp; Hita Garcia" baseAuthorityYear="2015" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="phragmotica">
<emphasis id="51EFEAF90273E70A50291963FBC9B10D" box="[1102,1120,1307,1330]" italics="true" pageId="6" pageNumber="7">C</emphasis>
.
<emphasis id="51EFEAF90273E70A54841941FEC2B16E" box="[227,363,1337,1361]" italics="true" pageId="6" pageNumber="7">phragmotica</emphasis>
(
<bibRefCitation id="070A4B1A0273E70A55181941FC8CB16E" author="Fischer, G. &amp; Azorsa, F. &amp; Hita Garcia, F. &amp; Mikheyev, A. S. &amp; Economo, E. P." box="[383,805,1337,1361]" pageId="6" pageNumber="7" pagination="77 - 105" refId="ref3387" refString="Fischer, G., Azorsa, F., Hita Garcia, F., Mikheyev, A. S. &amp; Economo, E. P. 2015. Two new phragmotic ant species from Africa: morphology and next-generation sequencing solve a caste association problem in the genus Carebara Westwood. ZooKeys, 525, 77 - 105. DOI: 10.3897 / zookeys. 525.6057" type="journal article" year="2015">Fischer, Azorsa &amp; Hita Garcia, 2015</bibRefCitation>
)
</taxonomicName>
and
<emphasis id="51EFEAF90273E70A570B1941FCD7B16F" box="[876,894,1337,1360]" italics="true" pageId="6" pageNumber="7">C</emphasis>
.
<emphasis id="51EFEAF90273E70A57F71941FC59B16E" box="[912,1008,1337,1361]" italics="true" pageId="6" pageNumber="7">aberrans</emphasis>
(
<bibRefCitation id="070A4B1A0273E70A50631941FEB5B14F" author="Santschi, F." pageId="6" pageNumber="7" pagination="211 - 250" refId="ref3617" refString="Santschi, F. 1937. Resultats de la Mission scientifique suisse en Angola (2 me voyage) 1932 - 1933. Fourmis angolaises. Revue Suisse de Zoologie, 44: 211 - 250." type="journal article" year="1937">Santschi, 1937</bibRefCitation>
). Although it is beyond this study to infer it, such character state can be found in different lineages of
<taxonomicName id="A49B4D680273E70A55BB190EFDEDB1B1" authorityName="WESTWOOD" authorityYear="1840" box="[476,580,1398,1422]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="51EFEAF90273E70A55BB190EFDEDB1B1" box="[476,580,1398,1422]" italics="true" pageId="6" pageNumber="7">Carebara</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="632436EB0273E70B556E19EDFE23B562" blockId="6.[227,1127,264,1606]" lastBlockId="7.[227,1127,264,441]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
The
<emphasis id="51EFEAF90273E70A555B19EDFDA4B192" box="[316,525,1429,1453]" italics="true" pageId="6" pageNumber="7">
<taxonomicName id="A49B4D680273E70A555B19EDFE0DB192" authorityName="WESTWOOD" authorityYear="1840" box="[316,420,1429,1453]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">Carebara</taxonomicName>
pygmaea
</emphasis>
species group includes five species,
<taxonomicName id="A49B4D680273E70A57CE19EDFB96B192" authority="Matsuura &amp; Hosoishi &amp; Sokh, 2024" authorityName="Matsuura &amp; Hosoishi &amp; Sokh" authorityYear="2024" box="[937,1087,1429,1453]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="bokorensis" status="sp. nov.">
<emphasis id="51EFEAF90273E70A57CE19EDFC12B193" box="[937,955,1429,1452]" italics="true" pageId="6" pageNumber="7">C</emphasis>
.
<emphasis id="51EFEAF90273E70A57AC19EDFB96B192" box="[971,1087,1429,1453]" italics="true" pageId="6" pageNumber="7">bokorensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4ADC57820273E70A502F19EEFF51B1F4" pageId="6" pageNumber="7" rank="species">sp. n.</taxonomicNameLabel>
,
<emphasis id="51EFEAF90273E70A556119CCFE2FB1F3" box="[262,390,1460,1484]" italics="true" pageId="6" pageNumber="7">C. pygmaea</emphasis>
,
<emphasis id="51EFEAF90273E70A55F219CCFE5BB1F3" box="[405,498,1460,1484]" italics="true" pageId="6" pageNumber="7">C. rubra</emphasis>
,
<emphasis id="51EFEAF90273E70A566619CCFD02B1F3" box="[513,683,1460,1484]" italics="true" pageId="6" pageNumber="7">C. transversalis</emphasis>
and
<taxonomicName id="A49B4D680273E70A568619CCFCE5B1F3" authorityName="Jaitrong, Pitaktunsakul et Jantarit" authorityYear="2021" box="[737,844,1460,1484]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="panhai">
<emphasis id="51EFEAF90273E70A568619CCFCE5B1F3" box="[737,844,1460,1484]" italics="true" pageId="6" pageNumber="7">C. panhai</emphasis>
</taxonomicName>
. The major workers of
<taxonomicName id="A49B4D680273E70A502919CCFEFEB1D5" authorityName="Matsuura &amp; Hosoishi &amp; Sokh" authorityYear="2024" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="bokorensis">
<emphasis id="51EFEAF90273E70A502919CCFBC9B1F4" box="[1102,1120,1460,1483]" italics="true" pageId="6" pageNumber="7">C</emphasis>
.
<emphasis id="51EFEAF90273E70A548419AAFEFEB1D5" box="[227,343,1490,1514]" italics="true" pageId="6" pageNumber="7">bokorensis</emphasis>
</taxonomicName>
are distinguished from the other four species in the species group by the presence of punctation on posterior surface of head, presence of scutellum and projection in the metanotal groove. The minor workers of
<taxonomicName id="A49B4D680273E70A57031A68FC51B217" authority="Matsuura &amp; Hosoishi &amp; Sokh, 2024" authorityName="Matsuura &amp; Hosoishi &amp; Sokh" authorityYear="2024" box="[868,1016,1552,1576]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="bokorensis" status="sp. nov.">
<emphasis id="51EFEAF90273E70A57031A68FCDFB218" box="[868,886,1552,1575]" italics="true" pageId="6" pageNumber="7">C</emphasis>
.
<emphasis id="51EFEAF90273E70A57E21A68FC51B217" box="[901,1016,1552,1576]" italics="true" pageId="6" pageNumber="7">bokorensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4ADC57820273E70A50651A68FB92B218" box="[1026,1083,1552,1575]" pageId="6" pageNumber="7" rank="species">sp. n.</taxonomicNameLabel>
are distinguished from the three species,
<emphasis id="51EFEAF90273E70A56E31A57FCACB279" box="[644,773,1583,1606]" italics="true" pageId="6" pageNumber="7">C. pygmaea</emphasis>
,
<emphasis id="51EFEAF90273E70A57741A57FCD8B279" box="[787,881,1582,1606]" italics="true" pageId="6" pageNumber="7">C. rubra</emphasis>
and
<taxonomicName id="A49B4D680273E70A57C01A57FBBBB279" authorityName="Jaitrong, Pitaktunsakul et Jantarit" authorityYear="2021" box="[935,1042,1582,1606]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="panhai">
<emphasis id="51EFEAF90273E70A57C01A57FBBBB279" box="[935,1042,1582,1606]" italics="true" pageId="6" pageNumber="7">C. panhai</emphasis>
</taxonomicName>
(minor worker of
<emphasis id="51EFEAF90272E70B55301D70FEC0B520" box="[343,361,264,287]" italics="true" pageId="7" pageNumber="8">C</emphasis>
.
<emphasis id="51EFEAF90272E70B551E1D70FDAAB51F" box="[377,515,264,288]" italics="true" pageId="7" pageNumber="8">transversalis</emphasis>
is unknown), in the species group by the presence of a well-developed subtriangular subpetiolar process and medial concavity of anterior clypeal margin.
</paragraph>
</subSubSection>
<subSubSection id="2B8165600272E70B556E1D1CFBFDB543" box="[265,1108,356,380]" pageId="7" pageNumber="8" type="description">
<paragraph id="632436EB0272E70B556E1D1CFBFDB543" blockId="7.[227,1127,264,441]" box="[265,1108,356,380]" pageId="7" pageNumber="8">
HABITAT.
<typeStatus id="BC2088490272E70B55EC1D1CFE6BB543" box="[395,450,356,380]" pageId="7" pageNumber="8">Type</typeStatus>
series specimens were collected from decayed wood (
<figureCitation id="FBA02A6E0272E70B50611D1CFBEEB543" box="[1030,1095,356,380]" captionStart="Fig" captionStartId="7.[321,356,1054,1076]" captionTargetBox="[246,1109,457,1032]" captionTargetId="figure-60@7.[246,1109,457,1032]" captionTargetPageId="7" captionText="Fig. 3. Carebara bokorensis sp. n. nesting in decayed wood (S. Hosoishi)." figureDoi="http://doi.org/10.5281/zenodo.10928406" httpUri="https://zenodo.org/record/10928406/files/figure.png" pageId="7" pageNumber="8">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="2B8165600272E70B556E1DFBFC37B5A4" box="[265,926,386,411]" pageId="7" pageNumber="8" type="distribution">
<paragraph id="632436EB0272E70B556E1DFBFC37B5A4" blockId="7.[227,1127,264,441]" box="[265,926,386,411]" pageId="7" pageNumber="8">
DISTRIBUTION.
<collectingCountry id="1B8C767B0272E70B55B61DFAFD97B5A5" box="[465,574,386,410]" name="Cambodia" pageId="7" pageNumber="8">Cambodia</collectingCountry>
(Bokor National Park,
<collectingRegion id="A15FF8090272E70B575F1DFBFC3AB5A4" box="[824,915,387,411]" country="Cambodia" name="Kampot" pageId="7" pageNumber="8">Kampot</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="2B8165600272E70B556E1DD9FBC9B586" box="[265,1120,417,441]" pageId="7" pageNumber="8" type="etymology">
<paragraph id="632436EB0272E70B556E1DD9FBC9B586" blockId="7.[227,1127,264,441]" box="[265,1120,417,441]" pageId="7" pageNumber="8">ETYMOLOGY. The species is named after the name of the Bokor National Park.</paragraph>
</subSubSection>
</treatment>
</document>