treatments-xml/data/03/89/CB/0389CB1C9A092955FF00589CFB555E5D.xml
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<mods:title id="E37B6A1DE379A84FC23CCD750C8DE842">Revision of the dragon-head cusk eels of the genus Porogadus (Teleostei: Ophidiidae), with description of eight new species and one new genus</mods:title>
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<taxonomicName id="4C2001899A092953FF00589CFDE25D90" ID-CoL="4LY4Y" authority="Nybelin, 1957" authorityName="Nybelin" authorityYear="1957" box="[151,575,654,681]" class="Actinopterygii" family="Ophidiidae" genus="Porogadus" kingdom="Animalia" order="Ophidiiformes" pageId="59" pageNumber="60" phylum="Chordata" rank="species" species="abyssalis">
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<emphasis id="B954A6189A092953FF00589CFE505D91" bold="true" box="[151,397,654,681]" italics="true" pageId="59" pageNumber="60">Porogadus abyssalis</emphasis>
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<figureCitation id="131B668F9A092953FF0058A1FF2E5DF5" box="[151,243,691,717]" captionStart="FIGURE 29" captionStartId="60.[151,250,1699,1723]" captionTargetBox="[151,1436,453,1668]" captionTargetId="figure-143@60.[151,1436,453,1675]" captionTargetPageId="60" captionText="FIGURE 29. Porogadus abyssalis Nybelin, 1957; 29AB, DG, MNHN 1979-0240, 160 mm SL, 10°58N, 43°13W, 5100 m, (AB photographs, DE head drawings, F otolith inner face, G otolith ventral view); 29C, MNHN 19921231, 158 mm SL, 10°24N, 46°43W, 4830 m." figureDoi="http://doi.org/10.5281/zenodo.5454195" httpUri="https://zenodo.org/record/5454195/files/figure.png" pageId="59" pageNumber="60">Figs. 29</figureCitation>
,
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,
<figureCitation id="131B668F9A092953FEBE58A1FE955DF5" box="[297,328,691,717]" captionStart="FIGURE 49" captionStartId="88.[151,250,1943,1967]" captionTargetBox="[178,1409,194,1915]" captionTargetId="figure-26@88.[175,1412,181,1919]" captionTargetPageId="88" captionText="FIGURE 49. Depth versus distance from shelf plots for specimens of the Porogadus miles group (upper panel) and the Porogadus gracilis group (lower panel)." figureDoi="http://doi.org/10.5281/zenodo.5454235" httpUri="https://zenodo.org/record/5454235/files/figure.png" pageId="59" pageNumber="60">49</figureCitation>
, Tab. 17
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<taxonomicName id="4C2001899A092953FF0058E5FDE85C37" ID-CoL="4LY4Y" authority="Nybelin, 1957: 288" authorityName="Nybelin" authorityPageNumber="288" authorityYear="1957" box="[151,565,759,783]" class="Actinopterygii" family="Ophidiidae" genus="Porogadus" kingdom="Animalia" order="Ophidiiformes" pageId="59" pageNumber="60" phylum="Chordata" rank="species" species="abyssalis">
<emphasis id="B954A6189A092953FF0058E5FEB45C37" box="[151,361,759,783]" italics="true" pageId="59" pageNumber="60">Porogadus abyssalis</emphasis>
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, figs. 1920, pl.6 figs. 24;
<treatmentCitation id="0A815C1B9A092953FCC558E5FBE75C37" author="Nielsen, J. G. &amp; Cohen, D. M. &amp; Markle, D. F. &amp; vRobins, C. R." box="[850,1082,759,783]" page="86" pageId="59" pageNumber="60" year="1999">
<bibRefCitation id="EFB107FB9A092953FCC558E5FBE75C37" author="Nielsen, J. G. &amp; Cohen, D. M. &amp; Markle, D. F. &amp; vRobins, C. R." box="[850,1082,759,783]" pageId="59" pageNumber="60" pagination="1 - 178" refId="ref64460" refString="Nielsen, J. G., Cohen, D. M., Markle, D. F. &amp; vRobins, C. R. (1999) Ophidiiform fishes of the world (Order Ophidiiformes). FAO Fisheries Synopsis, 125, FAO Species Catalogue, 18, 1 - 178." type="journal article" year="1999">
Nielsen
<emphasis id="B954A6189A092953FC3258EAFC045C37" box="[933,985,759,783]" italics="true" pageId="59" pageNumber="60">et al.</emphasis>
1999: 86
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.
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<emphasis id="B954A6189A092953FF00592FFEA45C6F" bold="true" box="[151,377,829,855]" pageId="59" pageNumber="60">Material examined</emphasis>
(
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):
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<specimenCode id="DB86D2719A092953FDBE592CFCDF5C6F" box="[553,770,829,856]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="59" pageNumber="60">MNHN 1979-0239</specimenCode>
,
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SL,
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,
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,
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a6/cp01, collected by
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,
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<collectingDate id="EFDAA5229A092953FDE45973FC915C43" box="[627,844,865,891]" pageId="59" pageNumber="60" value="1977-11-13">13 November 1977</collectingDate>
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;
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<specimenCode id="DB86D2719A092953FCC05970FBF25C43" box="[855,1071,865,892]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="59" pageNumber="60">MNHN 1979-0240</specimenCode>
,
<quantity id="4CD8D7EF9A092953FBAD5973FB4B5C44" box="[1082,1174,865,892]" metricMagnitude="-1" metricUnit="m" metricValue="1.6" pageId="59" pageNumber="60" unit="mm" value="160.0">160 mm</quantity>
SL,
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,
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</quantity>
,
<collectorName id="26D51FDC9A092953FF615997FE035CA7" box="[246,478,901,927]" pageId="59" pageNumber="60">Biovema Expedition</collectorName>
,
<collectorName id="26D51FDC9A092953FE705997FDCF5CA7" box="[487,530,901,927]" pageId="59" pageNumber="60">Sta.</collectorName>
a6/cp01, collected by
<collectorName id="26D51FDC9A092953FC915994FCA45CA7" box="[774,889,901,928]" pageId="59" pageNumber="60">J. Charcot</collectorName>
,
<date id="FF9E5CCA9A092953FC165997FB855CA7" box="[897,1112,901,927]" pageId="59" pageNumber="60" value="1977-11-13">
<collectingDate id="EFDAA5229A092953FC165997FB855CA7" box="[897,1112,901,927]" pageId="59" pageNumber="60" value="1977-11-13">13 November 1977</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B4870579A092953FBF55994FAB45CFB" collectingDate="1980-09-28" collectionCode="MNHN" collectorName="Demeraby Expedition &amp; Sta. &amp; J. Charcot" elevation="4830" latitude="10.4" longLatPrecision="1298" longitude="-46.716667" pageId="59" pageNumber="60" specimenCount="1">
<collectionCode id="ED31E2CF9A092953FBF55994FB655C98" box="[1122,1208,902,928]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="59" pageNumber="60">MNHN</collectionCode>
<quantity id="4CD8D7EF9A092953FB2B5997FA405C98" box="[1212,1437,901,928]" metricMagnitude="2" metricUnit="m" metricValue="6.16575" metricValueMax="12.31158" metricValueMin="0.01992" pageId="59" pageNumber="60" unit="mm" value="616575.0" valueMax="1231158.0" valueMin="1992.0">1992-1231, 158 mm</quantity>
SL,
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,
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,
<collectorName id="26D51FDC9A092953FD9459BBFD215CFB" box="[515,764,937,963]" pageId="59" pageNumber="60">Demeraby Expedition</collectorName>
,
<collectorName id="26D51FDC9A092953FC9059BBFCEF5CFB" box="[775,818,937,963]" pageId="59" pageNumber="60">Sta.</collectorName>
cp14, collected by
<collectorName id="26D51FDC9A092953FB9A59B8FB5C5CFB" box="[1037,1153,937,964]" pageId="59" pageNumber="60">J. Charcot</collectorName>
,
<date id="FF9E5CCA9A092953FB1B59BBFAB45CFB" box="[1164,1385,937,963]" pageId="59" pageNumber="60" value="1980-09-28">
<collectingDate id="EFDAA5229A092953FB1B59BBFAB45CFB" box="[1164,1385,937,963]" pageId="59" pageNumber="60" value="1980-09-28">28 September 1980</collectingDate>
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.
</paragraph>
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<subSubSection id="C33A29819A092953FF5059DFFF1C5B87" pageId="59" pageNumber="60" type="diagnosis">
<paragraph id="8B9F7A0A9A092953FF5059DFFF1C5B87" blockId="59.[151,1437,829,2008]" pageId="59" pageNumber="60">
<emphasis id="B954A6189A092953FF5059DFFE9F5CDF" bold="true" box="[199,322,973,999]" pageId="59" pageNumber="60">Diagnosis.</emphasis>
Precaudal vertebrae 1819; long gill rakers on lower part of first gill arch 1314; pectoral-fin rays 1923; HL:HD 1.852.09; most head spines moderately strong or weak, present on ethmoidal, prefrontal, interorbital, supraorbital, sphenotic, inner preopercular rim; absent on lacrimal, 5th infraorbital, supratemporal, inner and outer posttemporal, outer preopercular rim; opercular spine sharp, extruding; opercular flap small; lower lateral line pores until beginning of anal fin 23; vomer with moderately broad dentition patch with 23 rows of teeth; palatines with moderately broad dentition patch 24 rows of teeth; otolith with single colliculum; OL:OH = 1.17; OL:TCL = 2.3.
</paragraph>
</subSubSection>
<subSubSection id="C33A29819A092954FF505EDBFCC15E95" lastPageId="60" lastPageNumber="61" pageId="59" pageNumber="60" type="description">
<paragraph id="8B9F7A0A9A092953FF505EDBFD325AAF" blockId="59.[151,1437,829,2008]" pageId="59" pageNumber="60">
<emphasis id="B954A6189A092953FF505EDBFE8C5BDB" bold="true" box="[199,337,1225,1251]" pageId="59" pageNumber="60">Description</emphasis>
(based on non-types and
<typeStatus id="549BC4A89A092953FDFA5EDBFD125BDB" box="[621,719,1225,1251]" pageId="59" pageNumber="60" type="holotype">holotype</typeStatus>
data from
<bibRefCitation id="EFB107FB9A092953FCDF5EDBFC3C5BDC" author="Nybelin, O." box="[840,993,1225,1252]" pageId="59" pageNumber="60" pagination="247 - 345" refId="ref64632" refString="Nybelin, O. (1957) Deep-sea bottom fishes. Reports of the Swedish Deep-Sea Expedition 1947 - 1948, 2, 247 - 345." type="journal article" year="1957">Nybelin 1957</bibRefCitation>
). Meristics: precaudal vertebrae 1819, 1 last vertebrae without ribs; pectoral-fin rays 19 (1923); D/V = 6; D/A = 2427; V/A = 1921; long gill rakers on lower gill arch 14 (1314). Gill rakers in a typical specimen (MNHN 1979-0240) on lower first gill arch with six short rakers, followed by a series of 16 long rakers. The lower 8 of those intercept by narrow plate shaped rakers. These are smaller than in most species. Upper gill raker with a series of three semi long rakers and three plate like rakers intercepted, followed by two very short rakers.
</paragraph>
<paragraph id="8B9F7A0A9A092953FF505FB3FB6D5910" blockId="59.[151,1437,829,2008]" pageId="59" pageNumber="60">Morphometrics: in % of SL: HL 16.5 (16.517.0); maximal HD 8.29.0; HD through center of eye 5.26.1; bony interorbital width 3.43.6; snout length 6.1; upper jaw length 9.710.1; predorsal 17.418.4; preanal 33.9 (29.333.9); prepelvic 12.213.4; prepectoral 16.617.4; pectoral length 10.4. Relations: HL:HD = 1.852.09; HL to snout length 2.722.77; preanal to predorsal 1.681.69; predorsal to prepectoral 1.051.06.</paragraph>
<paragraph id="8B9F7A0A9A092953FF505C23FB3958B7" blockId="59.[151,1437,829,2008]" pageId="59" pageNumber="60">
Slender fish with long tapering tail and long, pointed snout. Maximal size of fishes investigated
<quantity id="4CD8D7EF9A092953FA815C23FAA95974" box="[1302,1396,1585,1612]" metricMagnitude="-1" metricUnit="m" metricValue="2.09" pageId="59" pageNumber="60" unit="mm" value="209.0">209 mm</quantity>
SL (
<typeStatus id="549BC4A89A092953FF085C47FED55948" box="[159,264,1621,1648]" pageId="59" pageNumber="60" type="holotype">holotype</typeStatus>
). Head long and very slender, with flat dorsal profile, with moderate to weak spines as follows: ethmoidal (1), prefrontal (1 weak), interorbital (3), supraorbital (2 weak), sphenotic (23), inner preopercular rim (34); no spines on lacrimal, 5th infraorbital, supratemporal, inner and outer posttemporal, outer preopercular rim. Opercle with sharp, short spine. Eye small located in strongly asymmetric orbit. Maxilla extending far beyond eye, strongly widened posteriorly and with distinct supramaxilla. Infra-/postorbital and mandibular-preopercular pores wide, 1 pore on occiput in front of nape. Head squamation on opercle, cheeks and occiput; absent on frontal, around eyes, and on maxilla. Opercle with two distinct large neuromasts behind preopercular edge; opercular flap small, with few radial ridges along lower margin. Lateral line rows rarely well visible and therefore number of pores countable only in rare instances. Lower lateral line row with 23 pores until beginning of anal fin in
<specimenCount id="9D26B1839A092953FBD45D64FB3D58B7" box="[1091,1248,1909,1936]" pageId="59" pageNumber="60" type="generic">one specimen</specimenCount>
.
</paragraph>
<paragraph id="8B9F7A0A9A092954FF505D8BFB1D5FED" blockId="59.[151,1437,829,2008]" lastBlockId="60.[151,1437,151,429]" lastPageId="60" lastPageNumber="61" pageId="59" pageNumber="60">Dentition. All teeth tiny and needle like. Vomer with a narrow dentition patch with 23 rows of teeth anteriorly; palatines with a narrow dentition patch with 24 rows of teeth in middle part. Premaxilla tooth patches not fused anteriorly; ca. 4 teeth rows anteriorly and 23 rows posteriorly. Dentary tooth patches not fused anteriorly; ca. 4 teeth rows anteriorly and a single row posteriorly. Median basibranchial tooth patch very short.</paragraph>
<paragraph id="8B9F7A0A9A0E2954FF505ACDFAED5E5D" blockId="60.[151,1437,151,429]" pageId="60" pageNumber="61">
Otolith morphology (n = 1). Size
<quantity id="4CD8D7EF9A0E2954FDDB5ACDFD795FC1" box="[588,676,223,249]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="60" pageNumber="61" unit="mm" value="1.8">1.8 mm</quantity>
in length (MNHN 1979-0240); OL:OH = 1.17; OH:OT 2.8. Thin, small, compressed, roundish otolith. All rims regularly rounded and smooth. Inner face flat, smooth, with short, centrally positioned sulcus; OL:TCL = 2.3. Sulcus with shallow, undivided, uniform, oval, wide colliculum. Dorsal field with indistinct depression; ventral field with faint ventral furrow. Outer face smooth, without umbo.
</paragraph>
<paragraph id="8B9F7A0A9A0E2954FF505B7DFCC15E95" blockId="60.[151,1437,151,429]" pageId="60" pageNumber="61">
Coloration. Color of preserved specimens light brown; belly slightly darker, opercle distinctly darker. Live coloration somewhat darker according to
<bibRefCitation id="EFB107FB9A0E2954FDFE5B81FCC55E95" author="Nybelin, O." box="[617,792,402,429]" pageId="60" pageNumber="61" pagination="247 - 345" refId="ref64632" refString="Nybelin, O. (1957) Deep-sea bottom fishes. Reports of the Swedish Deep-Sea Expedition 1947 - 1948, 2, 247 - 345." type="journal article" year="1957">Nybelin (1957)</bibRefCitation>
.
</paragraph>
</subSubSection>
<caption id="DF5F2A829A0E2954FF005CB1FE6A583A" ID-DOI="http://doi.org/10.5281/zenodo.5454195" ID-Zenodo-Dep="5454195" httpUri="https://zenodo.org/record/5454195/files/figure.png" pageId="60" pageNumber="61" startId="60.[151,250,1699,1723]" targetBox="[151,1436,453,1668]" targetPageId="60">
<paragraph id="8B9F7A0A9A0E2954FF005CB1FE6A583A" blockId="60.[151,1437,1699,1795]" pageId="60" pageNumber="61">
<emphasis id="B954A6189A0E2954FF005CB1FEFF5983" bold="true" box="[151,290,1699,1723]" pageId="60" pageNumber="61">FIGURE 29.</emphasis>
<taxonomicName id="4C2001899A0E2954FEBD5CB1FD465983" authority="Nybelin, 1957" authorityName="Nybelin" authorityYear="1957" box="[298,667,1699,1723]" class="Actinopterygii" family="Ophidiidae" genus="Porogadus" kingdom="Animalia" order="Ophidiiformes" pageId="60" pageNumber="61" phylum="Chordata" rank="species" species="abyssalis">
<emphasis id="B954A6189A0E2954FEBD5CB1FE205983" box="[298,509,1699,1723]" italics="true" pageId="60" pageNumber="61">Porogadus abyssalis</emphasis>
<bibRefCitation id="EFB107FB9A0E2954FD925CB1FD465983" author="Nybelin, O." box="[517,667,1699,1723]" pageId="60" pageNumber="61" pagination="247 - 345" refId="ref64632" refString="Nybelin, O. (1957) Deep-sea bottom fishes. Reports of the Swedish Deep-Sea Expedition 1947 - 1948, 2, 247 - 345." type="journal article" year="1957">Nybelin, 1957</bibRefCitation>
</taxonomicName>
;
<emphasis id="B954A6189A0E2954FD305CB1FCEC5983" bold="true" box="[679,817,1699,1723]" pageId="60" pageNumber="61">29AB, DG</emphasis>
, MNHN 1979-0240, 160 mm SL, 10°58N, 43°13W, 5100 m, (AB photographs, DE head drawings, F otolith inner face, G otolith ventral view);
<emphasis id="B954A6189A0E2954FB825CD5FB9D59E7" bold="true" box="[1045,1088,1735,1759]" pageId="60" pageNumber="61">29C</emphasis>
, MNHN 19921231, 158 mm SL, 10°24N, 46°43W, 4830 m.
</paragraph>
</caption>
<subSubSection id="C33A29819A0E2955FF505D23FA855FC1" lastPageId="61" lastPageNumber="62" pageId="60" pageNumber="61" type="discussion">
<paragraph id="8B9F7A0A9A0E2955FF505D23FA855FC1" blockId="60.[151,1437,1841,2012]" lastBlockId="61.[151,1436,151,357]" lastPageId="61" lastPageNumber="62" pageId="60" pageNumber="61">
<emphasis id="B954A6189A0E2954FF505D23FE915873" bold="true" box="[199,332,1841,1867]" pageId="60" pageNumber="61">Discussion.</emphasis>
<taxonomicName id="4C2001899A0E2954FEC55D23FDE45873" authorityName="Nybelin" authorityYear="1957" box="[338,569,1841,1867]" class="Actinopterygii" family="Ophidiidae" genus="Porogadus" kingdom="Animalia" order="Ophidiiformes" pageId="60" pageNumber="61" phylum="Chordata" rank="species" species="abyssalis">
<emphasis id="B954A6189A0E2954FEC55D23FDE45873" box="[338,569,1841,1867]" italics="true" pageId="60" pageNumber="61">Porogadus abyssalis</emphasis>
</taxonomicName>
was described based on a unique
<typeStatus id="549BC4A89A0E2954FC385D23FBCC5873" box="[943,1041,1841,1867]" pageId="60" pageNumber="61" type="holotype">holotype</typeStatus>
by
<bibRefCitation id="EFB107FB9A0E2954FBAD5D23FB385874" author="Nybelin, O." box="[1082,1253,1841,1868]" pageId="60" pageNumber="61" pagination="247 - 345" refId="ref64632" refString="Nybelin, O. (1957) Deep-sea bottom fishes. Reports of the Swedish Deep-Sea Expedition 1947 - 1948, 2, 247 - 345." type="journal article" year="1957">Nybelin (1957)</bibRefCitation>
caught at
<quantity id="4CD8D7EF9A0E2954FAC15D23FF305848" metricMagnitude="3" metricUnit="m" metricValue="5.275" metricValueMax="5.3" metricValueMin="5.25" pageId="60" pageNumber="61" unit="m" value="5275.0" valueMax="5300.0" valueMin="5250.0">5250 5300 m</quantity>
in the central Atlantic, which then was the deepest catch of any
<taxonomicName id="4C2001899A0E2954FC505D47FB9C5857" authorityName=": Cohen &amp; Nielsen" authorityYear="1978" box="[967,1089,1877,1903]" class="Actinopterygii" family="Ophidiidae" genus="Porogadus" kingdom="Animalia" order="Ophidiiformes" pageId="60" pageNumber="61" phylum="Chordata" rank="genus">
<emphasis id="B954A6189A0E2954FC505D47FB9C5857" box="[967,1089,1877,1903]" italics="true" pageId="60" pageNumber="61">Porogadus</emphasis>
</taxonomicName>
specimen which hitherto were caught shallower than
<quantity id="4CD8D7EF9A0E2954FE0C5D6BFE2B58AC" box="[411,502,1913,1940]" metricMagnitude="3" metricUnit="m" metricValue="3.3" pageId="60" pageNumber="61" unit="m" value="3300.0">3300 m</quantity>
. The weak head ossification and armature and the small, morphologically simple otolith may reflect adaptation to a deep sea habitat below the CCD. Within the
<taxonomicName id="4C2001899A0E2954FC6E5D8FFB12588F" baseAuthorityName="Gunther" baseAuthorityYear="1878" box="[1017,1231,1949,1975]" class="Actinopterygii" family="Ophidiidae" genus="Porogadus" kingdom="Animalia" order="Ophidiiformes" pageId="60" pageNumber="61" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="B954A6189A0E2954FC6E5D8FFB12588F" box="[1017,1231,1949,1975]" italics="true" pageId="60" pageNumber="61">Porogadus gracilis</emphasis>
</taxonomicName>
group,
<taxonomicName id="4C2001899A0E2954FAB55D8FFF2358E3" authorityName="Nybelin" authorityYear="1957" class="Actinopterygii" family="Ophidiidae" genus="Porogadus" kingdom="Animalia" order="Ophidiiformes" pageId="60" pageNumber="61" phylum="Chordata" rank="species" species="abyssalis">
<emphasis id="B954A6189A0E2954FAB55D8FFF2358E3" italics="true" pageId="60" pageNumber="61">Porogadus abyssalis</emphasis>
</taxonomicName>
is best recognized by its unusual high number of pectoral-fin rays (1923 vs 1518) and some relatively strong spines on the head. With its slender and elongate head and the high number of lower lateral line pores until the beginning of the anal fin
<taxonomicName id="4C2001899A0F2955FE4C5AA9FD805FED" authorityName="Nybelin" authorityYear="1957" box="[475,605,187,213]" class="Actinopterygii" family="Ophidiidae" genus="Porogadus" kingdom="Animalia" order="Ophidiiformes" pageId="61" pageNumber="62" phylum="Chordata" rank="species" species="abyssalis">
<emphasis id="B954A6189A0F2955FE4C5AA9FD805FED" box="[475,605,187,213]" italics="true" pageId="61" pageNumber="62">P. abyssalis</emphasis>
</taxonomicName>
resembles several species of the
<taxonomicName id="4C2001899A0F2955FC465AA9FB515FED" authorityName=", MNHN" authorityYear="1995" box="[977,1164,187,213]" class="Actinopterygii" family="Ophidiidae" genus="Porogadus" kingdom="Animalia" order="Ophidiiformes" pageId="61" pageNumber="62" phylum="Chordata" rank="species" species="miles">
<emphasis id="B954A6189A0F2955FC465AA9FB515FED" box="[977,1164,187,213]" italics="true" pageId="61" pageNumber="62">Porogadus miles</emphasis>
</taxonomicName>
group, but differs in the generally much weaker head armature, weaker head ossification and the very small and compressed otoliths.
</paragraph>
</subSubSection>
<subSubSection id="C33A29819A0F2955FF505B10FB555E5D" pageId="61" pageNumber="62" type="distribution">
<paragraph id="8B9F7A0A9A0F2955FF505B10FB555E5D" blockId="61.[151,1436,151,357]" pageId="61" pageNumber="62">
<emphasis id="B954A6189A0F2955FF505B10FEBC5E24" bold="true" box="[199,353,258,284]" pageId="61" pageNumber="62">Distribution.</emphasis>
The specimens described here are the only other specimens beside the unique
<typeStatus id="549BC4A89A0F2955FB5E5B11FAF65E25" box="[1225,1323,259,285]" pageId="61" pageNumber="62" type="holotype">holotype</typeStatus>
described by
<bibRefCitation id="EFB107FB9A0F2955FF2E5B35FEBA5E79" author="Nybelin, O." box="[185,359,294,321]" pageId="61" pageNumber="62" pagination="247 - 345" refId="ref64632" refString="Nybelin, O. (1957) Deep-sea bottom fishes. Reports of the Swedish Deep-Sea Expedition 1947 - 1948, 2, 247 - 345." type="journal article" year="1957">Nybelin (1957)</bibRefCitation>
. They confirm the deep sea dwelling of the species in the tropical Atlantic and far away from any shelf break (
<quantity id="4CD8D7EF9A0F2955FEB65B59FE1C5E5D" box="[289,449,331,357]" metricMagnitude="5" metricUnit="m" metricValue="8.5" metricValueMax="10.0" metricValueMin="7.0" pageId="61" pageNumber="62" unit="km" value="850.0" valueMax="1000.0" valueMin="700.0">7001000 km</quantity>
,
<figureCitation id="131B668F9A0F2955FE5B5B59FDC15E5D" box="[460,540,331,357]" captionStart="FIGURE 49" captionStartId="88.[151,250,1943,1967]" captionTargetBox="[178,1409,194,1915]" captionTargetId="figure-26@88.[175,1412,181,1919]" captionTargetPageId="88" captionText="FIGURE 49. Depth versus distance from shelf plots for specimens of the Porogadus miles group (upper panel) and the Porogadus gracilis group (lower panel)." figureDoi="http://doi.org/10.5281/zenodo.5454235" httpUri="https://zenodo.org/record/5454235/files/figure.png" pageId="61" pageNumber="62">Fig. 49</figureCitation>
) on the abyssal plane near to the middle Atlantic ridge.
</paragraph>
</subSubSection>
</treatment>
</document>