233 lines
26 KiB
XML
233 lines
26 KiB
XML
<document id="79851F1D1911FF88E0063B604E8EE096" ID-CLB-Dataset="222437" ID-DOI="10.11646/zootaxa.5227.3.7" ID-GBIF-Dataset="4ceb3cf6-8307-4cee-ab01-20922b7a910f" ID-ISSN="1175-5326" ID-Zenodo-Dep="7518846" ID-ZooBank="8026B56A-60D7-447F-B9C9-CE43ABDC9D63" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1673311596627" checkinUser="plazi" docAuthor="Wallaard, Jonathan J. W., Fraaije, René H. B., Van Bakel, Barry W. M., Nance, John R., Lindholm, Adam & Jagt, John W. M." docDate="2023" docId="D462382A2133FF93FF0BD9CC0F66FED8" docLanguage="en" docName="zootaxa.5227.3.7.pdf" docOrigin="Zootaxa 5227 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paguristes timoni Wallaard & Fraaije & Van Bakel & Nance & Lindholm & Jagt 2023, n. sp." docType="treatment" docUuid="C4F34FA0-7A19-46B4-91BC-C93C621CD977" docUuidSource="ZooBank" docVersion="7" lastPageNumber="394" masterDocId="285B40522137FF96FF9CD95B0D55FFBD" masterDocTitle="New hermit crab species (Anomura, Paguroidea) from the upper Miocene St Marys Formation of Maryland (USA), preserved in their host shells" masterLastPageNumber="397" masterPageNumber="389" pageNumber="393" updateTime="1699473637289" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="80E3CE5BC130F0DB5BF2898833B97166">New hermit crab species (Anomura, Paguroidea) from the upper Miocene St Marys Formation of Maryland (USA), preserved in their host shells</mods:title>
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<mods:namePart id="167038E9096C0DDC116853D41CEAD479">Wallaard, Jonathan J. W.</mods:namePart>
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<mods:affiliation id="37707EE2BD509052DE75E0D3DB4B0682">Oertijdmuseum, Bosscheweg 80, 5283 WB Boxtel, the Netherlands. & curator @ oertijdmuseum. nl; https: // orcid. org / 0000 - 0002 - 7752 - 1598</mods:affiliation>
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<mods:nameIdentifier id="5437D68212BEDAF9F5F8BED83A0947AF" type="email">curator@oertijdmuseum.nl</mods:nameIdentifier>
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<mods:namePart id="1B4470182B968218122FE32C6CC9B15F">Fraaije, René H. B.</mods:namePart>
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<mods:nameIdentifier id="86DC59F94289887B221A27E93527FDD5" type="ORCID">0000-0002-3465-1093</mods:nameIdentifier>
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<mods:affiliation id="E7602E33829F9050A8B0B9574CFFBDEF">Oertijdmuseum, Bosscheweg 80, 5283 WB Boxtel, the Netherlands. & info @ oertijdmuseum. nl; https: // orcid. org / 0000 - 0002 - 3465 - 1093</mods:affiliation>
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<mods:nameIdentifier id="A64A79546C6B0CE5CEFC5A877AEBF348" type="email">info@oertijdmuseum.nl</mods:nameIdentifier>
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</mods:name>
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<mods:roleTerm id="8E13695DCAC538BE2234B8A0CBF4CD6C">Author</mods:roleTerm>
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<mods:namePart id="3EA677BC4812A49E990EC788191F50C0">Van Bakel, Barry W. M.</mods:namePart>
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<mods:affiliation id="100FC2BD787B70FFF03D2ACBB18C0E9B">Oertijdmuseum, Bosscheweg 80, 5283 WB Boxtel, the Netherlands. & Utrecht University, Budapestlaan 4, 3584 CD Utrecht, the Netherlands.</mods:affiliation>
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<mods:namePart id="BA7B2201C9E931DA238F8416ADAB401B">Nance, John R.</mods:namePart>
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<mods:affiliation id="33D790FF27ECA2D2B345B98F834D1C23">Calvert Marine Museum, 14200 Solomons Island Road, Solomons, MD 20688, USA.</mods:affiliation>
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<mods:namePart id="92AF6003AE61246D2712CCE4F9C6A71C">Lindholm, Adam</mods:namePart>
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<mods:affiliation id="7B805BF552A98642B605DF9989A644A3">Department of Earth Sciences, University of Oxford, South Parks Road, Oxford, UK, OX 1 3 AN,</mods:affiliation>
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<mods:namePart id="B92B55C6359D4BFA8478C73F6CE47E6A">Jagt, John W. M.</mods:namePart>
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<mods:affiliation id="06A11E1E25F25E3F5431839EC144A2C5">Natuurhistorisch Museum Maastricht, De Bosquetplein 6 - 7, 6211 KJ Maastricht, the Netherlands.</mods:affiliation>
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<mods:typeOfResource id="C063E3B804DC2A837DC1514C06B74385">text</mods:typeOfResource>
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<mods:title id="9FA75CA76346269FAE1F68BFA572B996">Zootaxa</mods:title>
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<mods:part id="389357D094D073ADE23F0A256F034EF1">
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<mods:date id="C0A1D865D4BE831931C1DD654B1B81F3">2023</mods:date>
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<mods:number id="A22868E354C0E0111411C3433DE94775">2023-01-06</mods:number>
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<mods:number id="BF7CE5FD9E46CD2B92A3F7A7984012E7">5227</mods:number>
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<mods:identifier id="C8D22CFECF6C1367250E14B7E46698BA" type="DOI">10.11646/zootaxa.5227.3.7</mods:identifier>
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<treatment id="D462382A2133FF93FF0BD9CC0F66FED8" ID-DOI="http://doi.org/10.5281/zenodo.7525500" ID-GBIF-Taxon="208791089" ID-Zenodo-Dep="7525500" LSID="urn:lsid:zoobank.org:act:C4F34FA0-7A19-46B4-91BC-C93C621CD977" httpUri="http://treatment.plazi.org/id/D462382A2133FF93FF0BD9CC0F66FED8" lastPageId="5" lastPageNumber="394" pageId="4" pageNumber="393">
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<subSubSection id="14D1DAB72133FF92FF0BD9CC0CE3FF0C" box="[151,438,151,177]" pageId="4" pageNumber="393" type="nomenclature">
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<paragraph id="5C74893C2133FF92FF0BD9CC0CE3FF0C" blockId="4.[151,1076,151,249]" box="[151,438,151,177]" pageId="4" pageNumber="393">
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<heading id="073C3E502133FF92FF0BD9CC0CE3FF0C" bold="true" box="[151,438,151,177]" fontSize="11" level="1" pageId="4" pageNumber="393" reason="1">
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<emphasis id="6EBF552E2133FF92FF0BD9CC0CE3FF0C" bold="true" box="[151,438,151,177]" pageId="4" pageNumber="393">
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<taxonomicName id="9BCBF2BF2133FF92FF0BD9CC0C39FF0C" authority="Wallaard & Fraaije & Van Bakel & Nance & Lindholm & Jagt, 2023" authorityName="Wallaard & Fraaije & Van Bakel & Nance & Lindholm & Jagt" authorityYear="2023" box="[151,364,151,177]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="393" phylum="Arthropoda" rank="species" species="timoni" status="sp. nov.">
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<emphasis id="6EBF552E2133FF92FF0BD9CC0C39FF0C" bold="true" box="[151,364,151,177]" italics="true" pageId="4" pageNumber="393">Paguristes timoni</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="758CE8552133FF92FEEFD9CC0CE3FF0C" box="[371,438,151,177]" pageId="4" pageNumber="393" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="14D1DAB72133FF92FF0BD9E00961FF44" pageId="4" pageNumber="393" type="description">
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<paragraph id="5C74893C2133FF92FF0BD9E00DB9FF68" blockId="4.[151,1076,151,249]" box="[151,236,187,213]" pageId="4" pageNumber="393">
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(
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<figureCitation id="C4F095B92133FF92FF03D9E00DB6FF68" box="[159,227,187,213]" captionStart="FIGURE 2" captionStartId="4.[151,250,1692,1716]" captionTargetBox="[151,1436,885,1668]" captionTargetId="figure-262@4.[151,1436,885,1668]" captionTargetPageId="4" captionText="FIGURE 2. Paguristes timoni n. sp., holotype, CMM-I-4600, from the upper Miocene St. Marys Formation in the cliff south of Little Cove Point, Calvert County (Maryland), in its host shell, and in ventral (A) and dorsal (B) aspects." figureDoi="http://doi.org/10.5281/zenodo.7518852" httpUri="https://zenodo.org/record/7518852/files/figure.png" pageId="4" pageNumber="393">Fig. 2</figureCitation>
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)
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</paragraph>
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<paragraph id="5C74893C2133FF92FF0BD9840961FF44" blockId="4.[151,1076,151,249]" box="[151,1076,223,249]" pageId="4" pageNumber="393">
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Zoobank:
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<uri id="285A853E2133FF92FE9AD9840961FF44" box="[262,1076,223,249]" pageId="4" pageNumber="393">
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urn:lsid:zoobank.org:act:
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<uuid id="286DB3E92133FF92FD84D9840961FF44" box="[536,1076,223,249]" pageId="4" pageNumber="393">C4F34FA0-7A19-46B4-91BC-C93C621CD977</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="14D1DAB72133FF92FF0BD87C0EA8FE34" pageId="4" pageNumber="393" type="diagnosis">
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<paragraph id="5C74893C2133FF92FF0BD87C0EA8FE34" blockId="4.[151,1437,295,862]" pageId="4" pageNumber="393">
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<emphasis id="6EBF552E2133FF92FF0BD87C0C47FEFC" bold="true" box="[151,274,295,321]" pageId="4" pageNumber="393">Diagnosis.</emphasis>
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Keeled merus with row of small teeth on distal margin; moveable finger corneous; fixed finger covered with rows of alveolate tubercles, decreasing in size on distal end. Upper margin with tubercles; cutting edge with several teeth proximally. Walking legs with keeled edge and row of tubercles.
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</paragraph>
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</subSubSection>
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<subSubSection id="14D1DAB72133FF92FF5BD8C8090CFE10" box="[199,1113,403,430]" pageId="4" pageNumber="393" type="materials_examined">
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<paragraph id="5C74893C2133FF92FF5BD8C8090CFE10" blockId="4.[151,1437,295,862]" box="[199,1113,403,430]" pageId="4" pageNumber="393">
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<emphasis id="6EBF552E2133FF92FF5BD8C80C20FE10" bold="true" box="[199,373,403,429]" pageId="4" pageNumber="393">Type material.</emphasis>
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The
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<typeStatus id="8370379E2133FF92FE32D8C80F46FE10" box="[430,531,403,429]" pageId="4" pageNumber="393" type="holotype">holotype</typeStatus>
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, and sole specimen known to date, is CMM-I-4600.
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</paragraph>
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</subSubSection>
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<subSubSection id="14D1DAB72133FF92FF5BD8EC0E07FE4B" pageId="4" pageNumber="393" type="etymology">
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<paragraph id="5C74893C2133FF92FF5BD8EC0E07FE4B" blockId="4.[151,1437,295,862]" pageId="4" pageNumber="393">
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<emphasis id="6EBF552E2133FF92FF5BD8EC0C1BFE6C" bold="true" box="[199,334,439,465]" pageId="4" pageNumber="393">Etymology.</emphasis>
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Named after the legendary misanthrope and hermit, Timon of Athens, who was popularized in the play ‘Timon of Athens’ by William Shakespeare (1564–1616).
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</paragraph>
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</subSubSection>
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<subSubSection id="14D1DAB72133FF92FF5BD8A40DBDFD38" pageId="4" pageNumber="393" type="distribution">
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<paragraph id="5C74893C2133FF92FF5BD8A40DBDFD38" blockId="4.[151,1437,295,862]" pageId="4" pageNumber="393">
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<emphasis id="6EBF552E2133FF92FF5BD8A40CAAFDA4" bold="true" box="[199,511,511,537]" pageId="4" pageNumber="393">Locality and stratigraphy.</emphasis>
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Driftwood Beach, Calvert County,
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<collectingRegion id="9E0F47DE2133FF92FC11D8A40EA8FDA4" box="[909,1021,511,537]" country="United States of America" name="Maryland" pageId="4" pageNumber="393">Maryland</collectingRegion>
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, from the upper Miocene (Tortonian) Little Cove Point Member of the St. Marys Formation in a silty lens within Bed E (
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<bibRefCitation id="385AF4CD2133FF92FBB6DB780864FD80" author="Ward, L. W. & Andrews, G. W." box="[1066,1329,547,573]" pageId="4" pageNumber="393" pagination="1 - 60" refId="ref7576" refString="Ward, L. W. & Andrews, G. W. (2008) Stratigraphy of the Calvert, Choptank, and St. Mary's formations (Miocene) in the Chesapeake Bay area, Maryland and Virginia. Virginia Museum of Natural History Memoir, 9, 1 - 60." type="journal article" year="2008">Ward & Andrews 2008</bibRefCitation>
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).
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<bibRefCitation id="385AF4CD2133FF92FADEDB780C71FDDF" author="Kidwell, S. M. & Powars, D. S. & Edwards, L. E. & Vogt, P. R." pageId="4" pageNumber="393" pagination="231 - 279" refId="ref6107" refString="Kidwell, S. M., Powars, D. S., Edwards, L. E. & Vogt, P. R. (2015) Miocene stratigraphy and paleoenvironments of the Calvert Cliffs, Maryland. Bulletin of the Geological Society of America, 40, 231 - 279." type="journal article" year="2015">
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Kidwell
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<emphasis id="6EBF552E2133FF92FF0BDB130D9FFDDC" box="[151,202,583,609]" italics="true" pageId="4" pageNumber="393">et al</emphasis>
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. (2015)
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</bibRefCitation>
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identified this level as belonging to “SM-C,” assigned to Shattuck zones 22–23 and part of dinocyst zone 8.
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</paragraph>
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</subSubSection>
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<subSubSection id="14D1DAB72133FF92FF5BDBD40E6AF831" pageId="4" pageNumber="393" type="description">
|
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<paragraph id="5C74893C2133FF92FF5BDBD40975FCE0" blockId="4.[151,1437,295,862]" pageId="4" pageNumber="393">
|
||
<emphasis id="6EBF552E2133FF92FF5BDBD40C0CFD14" bold="true" box="[199,345,655,681]" pageId="4" pageNumber="393">Description.</emphasis>
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CMM-I-4600, preserved inside gastropod shell (
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<taxonomicName id="9BCBF2BF2133FF92FC15DBCB0947FD14" box="[905,1042,655,681]" class="Gastropoda" family="Buccinidae" genus="Busycon" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="393" phylum="Mollusca" rank="species" species="undetermined">
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<emphasis id="6EBF552E2133FF92FC15DBCB0EBDFD14" box="[905,1000,656,681]" italics="true" pageId="4" pageNumber="393">Busycon</emphasis>
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sp.
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</taxonomicName>
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), length
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<quantity id="9B3324D92133FF92FBE4DBD4099EFD17" box="[1144,1227,655,682]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="393" unit="mm" value="25.0">25 mm</quantity>
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, greatest width
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<quantity id="9B3324D92133FF92FA1CDBD40D93FD73" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="4" pageNumber="393" unit="mm" value="17.0">17 mm</quantity>
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. Manus missing from major (left) cheliped, carpus and merus preserved. Keeled merus with row of small teeth on distal margin. Minor (right) cheliped comprising complete propodus, lodged in gastropod aperture, measuring
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<quantity id="9B3324D92133FF92FA12DB830D96FCAB" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="393" unit="mm" value="7.0">7 mm</quantity>
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length,
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<quantity id="9B3324D92133FF92FE87DBA00C0EFCAB" box="[283,347,763,790]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="393" unit="mm" value="2.0">2 mm</quantity>
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maximum width. Moveable finger corneous, fixed finger covered with rows of alveolate tubercles decreasing in size on distal end; most tubercles with alveoli, indicative of setal insertions; upper margin covered with tubercles; cutting edge with several teeth proximally; distal side not visible.
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</paragraph>
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<caption id="08B4D9B42133FF92FF0BDFC709E8F96A" ID-DOI="http://doi.org/10.5281/zenodo.7518852" ID-Zenodo-Dep="7518852" httpUri="https://zenodo.org/record/7518852/files/figure.png" pageId="4" pageNumber="393" startId="4.[151,250,1692,1716]" targetBox="[151,1436,885,1668]" targetPageId="4">
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<paragraph id="5C74893C2133FF92FF0BDFC709E8F96A" blockId="4.[151,1436,1692,1752]" pageId="4" pageNumber="393">
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<emphasis id="6EBF552E2133FF92FF0BDFC70C47F909" bold="true" box="[151,274,1692,1716]" pageId="4" pageNumber="393">FIGURE 2.</emphasis>
|
||
<taxonomicName id="9BCBF2BF2133FF92FE84DFC70C9EF90E" authority="Wallaard & Fraaije & Van Bakel & Nance & Lindholm & Jagt, 2023" authorityName="Wallaard & Fraaije & Van Bakel & Nance & Lindholm & Jagt" authorityYear="2023" box="[280,459,1692,1715]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="393" phylum="Arthropoda" rank="species" species="timoni" status="sp. nov.">
|
||
<emphasis id="6EBF552E2133FF92FE84DFC70C9EF90E" box="[280,459,1692,1715]" italics="true" pageId="4" pageNumber="393">Paguristes timoni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="6EBF552E2133FF92FE4CDFC70F5DF909" bold="true" box="[464,520,1692,1716]" pageId="4" pageNumber="393">
|
||
<taxonomicNameLabel id="758CE8552133FF92FE4CDFC70F5DF909" box="[464,520,1692,1716]" pageId="4" pageNumber="393" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, CMM-I-4600, from the upper Miocene St. Marys Formation in the cliff south of Little Cove Point, Calvert County (Maryland), in its host shell, and in ventral (A) and dorsal (B) aspects.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="5C74893C2133FF92FF5BDE710998F8F9" blockId="4.[151,1436,1834,2041]" box="[199,1229,1834,1861]" pageId="4" pageNumber="393">Carapace fragment preserved but covered; small portion visible lacking any specific details;</paragraph>
|
||
<paragraph id="5C74893C2133FF92FF5BDE150E6AF831" blockId="4.[151,1436,1834,2041]" pageId="4" pageNumber="393">Single fragment of walking leg preserved showing keeled edge with row of tubercles. Several other fragments preserved, but unidentifiable due to weathering of specimen.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="14D1DAB72133FF93FF5BDECD0F66FED8" lastPageId="5" lastPageNumber="394" pageId="4" pageNumber="393" type="discussion">
|
||
<paragraph id="5C74893C2133FF92FF5BDECD09EEF845" blockId="4.[151,1436,1834,2041]" pageId="4" pageNumber="393">
|
||
<emphasis id="6EBF552E2133FF92FF5BDECD0C6EF80D" bold="true" box="[199,315,1942,1968]" pageId="4" pageNumber="393">Remarks.</emphasis>
|
||
In this novel form, the left cheliped is larger than the right one, which is diagnostic feature of diogenid and annuntidiogenid hermit crabs (
|
||
<bibRefCitation id="385AF4CD2133FF92FD80DEE10FBFF869" author="McLaughlin, P. A." box="[540,746,1978,2004]" pageId="4" pageNumber="393" pagination="111 - 144" refId="ref6685" refString="McLaughlin, P. A. (2003) Illustrated keys to families and genera of the superfamily Paguroidea (Crustacea: Decapoda: Anomura), with diagnoses of genera of Paguridae. Memoirsof Museum Victoria, 60 (1), 111 - 144." type="journal article" year="2003">McLaughlin 2003</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="385AF4CD2133FF92FD6ADEE10ED2F869" author="Fraaije, R. H. B." box="[758,903,1978,2004]" pageId="4" pageNumber="393" pagination="121 - 145" refId="ref4724" refString="Fraaije, R. H. B. (2014) Diverse Late Jurassic anomuran assemblages from the Swabian Alb and evolutionary history of paguroids based on carapace morphology. Neues Jahrbuch fur Geologie und Palaontologie Abhandlungen, 273 (2), 121 - 145." type="journal article" year="2014">Fraaije 2014</bibRefCitation>
|
||
). The Maryland specimen is remarkably similar to the extant
|
||
<taxonomicName id="9BCBF2BF2133FF92FEB5DE840E32F844" authority="De Matos-Pita & Ramil, 2015" authorityName="De Matos-Pita & Ramil" authorityYear="2015" box="[297,871,2014,2041]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="393" phylum="Arthropoda" rank="species" species="candelae">
|
||
<emphasis id="6EBF552E2133FF92FEB5DE840F59F845" box="[297,524,2014,2040]" italics="true" pageId="4" pageNumber="393">Paguristes candelae</emphasis>
|
||
<bibRefCitation id="385AF4CD2133FF92FD8FDE840E32F844" author="De Matos-Pita, S. S. & Ramil, F." box="[531,871,2014,2041]" pageId="4" pageNumber="393" pagination="151 - 190" refId="ref4199" refString="De Matos-Pita, S. S. & Ramil, F. (2015) Hermit crabs (Decapoda: Crustacea) from deep Mauritanian waters (NW Africa) with the description of a new species. Zootaxa, 3926 (2), 151 - 190." type="journal article" year="2015">De Matos-Pita & Ramil, 2015</bibRefCitation>
|
||
</taxonomicName>
|
||
, from
|
||
<collectingCountry id="24DCC9AC2133FF92FC2CDE85097EF845" box="[944,1067,2014,2040]" name="Mauritania" pageId="4" pageNumber="393">Mauritania</collectingCountry>
|
||
(see below).
|
||
</paragraph>
|
||
<paragraph id="5C74893C2132FF93FF5BD9CD0F66FED8" blockId="5.[151,1437,150,357]" pageId="5" pageNumber="394">
|
||
<emphasis id="6EBF552E2132FF93FF5BD9CD0C19FF0D" bold="true" box="[199,332,150,176]" pageId="5" pageNumber="394">Discussion.</emphasis>
|
||
Cheliped shape and ornament in
|
||
<taxonomicName id="9BCBF2BF2132FF93FD5ED9CC0E70FF0D" authority="Wallaard & Fraaije & Van Bakel & Nance & Lindholm & Jagt, 2023" authorityName="Wallaard & Fraaije & Van Bakel & Nance & Lindholm & Jagt" authorityYear="2023" box="[706,805,151,176]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="394" phylum="Arthropoda" rank="species" species="timoni" status="sp. nov.">
|
||
<emphasis id="6EBF552E2132FF93FD5ED9CC0E70FF0D" box="[706,805,151,176]" italics="true" pageId="5" pageNumber="394">P. timoni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="6EBF552E2132FF93FCB0D9CC0E3FFF0C" bold="true" box="[812,874,151,177]" pageId="5" pageNumber="394">
|
||
<taxonomicNameLabel id="758CE8552132FF93FCB0D9CC0E3FFF0C" box="[812,874,151,177]" pageId="5" pageNumber="394" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
compare closely with those of
|
||
<taxonomicName id="9BCBF2BF2132FF93FB5BD9CC081DFF0C" authorityName="De Matos-Pita & Ramil" authorityYear="2015" box="[1223,1352,151,177]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="394" phylum="Arthropoda" rank="species" species="candelae">
|
||
<emphasis id="6EBF552E2132FF93FB5BD9CC081DFF0C" box="[1223,1352,151,177]" italics="true" pageId="5" pageNumber="394">P. candelae</emphasis>
|
||
</taxonomicName>
|
||
, in that both have rows of tubercles and setae and tubercles decrease in size toward the distal end. The dorsomesial margin in
|
||
<taxonomicName id="9BCBF2BF2132FF93FF2FD9840C67FF44" authorityName="De Matos-Pita & Ramil" authorityYear="2015" box="[179,306,223,249]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="394" phylum="Arthropoda" rank="species" species="candelae">
|
||
<emphasis id="6EBF552E2132FF93FF2FD9840C67FF44" box="[179,306,223,249]" italics="true" pageId="5" pageNumber="394">P. candelae</emphasis>
|
||
</taxonomicName>
|
||
has three large tubercles; these are absent from
|
||
<taxonomicName id="9BCBF2BF2132FF93FCA3D9840EF7FF45" authority="Wallaard & Fraaije & Van Bakel & Nance & Lindholm & Jagt, 2023" authorityName="Wallaard & Fraaije & Van Bakel & Nance & Lindholm & Jagt" authorityYear="2023" box="[831,930,223,248]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="394" phylum="Arthropoda" rank="species" species="timoni" status="sp. nov.">
|
||
<emphasis id="6EBF552E2132FF93FCA3D9840EF7FF45" box="[831,930,223,248]" italics="true" pageId="5" pageNumber="394">P. timoni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="6EBF552E2132FF93FC3BD9840EB1FF44" bold="true" box="[935,996,223,249]" pageId="5" pageNumber="394">
|
||
<taxonomicNameLabel id="758CE8552132FF93FC3BD9840EB1FF44" box="[935,996,223,249]" pageId="5" pageNumber="394" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
The dorsal cheliped surface in the latter appears to be less convex in comparison with that of
|
||
<taxonomicName id="9BCBF2BF2132FF93FD7ED8580E37FEA0" authorityName="De Matos-Pita & Ramil" authorityYear="2015" box="[738,866,259,285]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="394" phylum="Arthropoda" rank="species" species="candelae">
|
||
<emphasis id="6EBF552E2132FF93FD7ED8580E37FEA0" box="[738,866,259,285]" italics="true" pageId="5" pageNumber="394">P. candelae</emphasis>
|
||
</taxonomicName>
|
||
, while the teeth along the cutting edge appear to be larger in the extinct form. The walking legs in both taxa show a keeled ridge with a row of spines, although the one in
|
||
<taxonomicName id="9BCBF2BF2132FF93FF28D8100C42FED9" authority="Wallaard & Fraaije & Van Bakel & Nance & Lindholm & Jagt, 2023" authorityName="Wallaard & Fraaije & Van Bakel & Nance & Lindholm & Jagt" authorityYear="2023" box="[180,279,331,356]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="394" phylum="Arthropoda" rank="species" species="timoni" status="sp. nov.">
|
||
<emphasis id="6EBF552E2132FF93FF28D8100C42FED9" box="[180,279,331,356]" italics="true" pageId="5" pageNumber="394">P. timoni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="6EBF552E2132FF93FE82D8100C08FED8" bold="true" box="[286,349,331,357]" pageId="5" pageNumber="394">
|
||
<taxonomicNameLabel id="758CE8552132FF93FE82D8100C08FED8" box="[286,349,331,357]" pageId="5" pageNumber="394" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
seems more acute.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |