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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.185.3082" ID-GBIF-Dataset="f54227ef-6fd3-41eb-a6fc-6203e8ed2ec5" ID-PMC="PMC3345791" ID-Pensoft-Pub="1313-2970-185-1" ID-PubMed="22577310" ModsDocAuthor="" ModsDocDate="2012" ModsDocID="1313-2970-185-1" ModsDocOrigin="ZooKeys 185" ModsDocTitle="Three new cavernicolous species of dragon millipedes, genus Desmoxytes Chamberlin, 1923, from southern China, with notes on a formal congener from the Philippines (Diplopoda, Polydesmida, Paradoxosomatidae)" checkinTime="1451249126133" checkinUser="pensoft" docAuthor="Golovatch, Sergei I., Li, Youbang, Liu, Weixin &amp; Geoffroy, Jean-Jacques" docDate="2012" docId="CEA4EBA5B3265B0BD1EA920A79FA0389" docLanguage="en" docName="ZooKeys 185: 1-17" docOrigin="ZooKeys 185" docSource="http://dx.doi.org/10.3897/zookeys.185.3082" docTitle="Desmoxytes spinissima Golovatch, Li, Liu &amp; Geoffroy, 2012, sp. n." docType="treatment" docVersion="4" lastPageNumber="9" masterDocId="B12DC7150217FFBB535C5A786D46FFD9" masterDocTitle="Three new cavernicolous species of dragon millipedes, genus Desmoxytes Chamberlin, 1923, from southern China, with notes on a formal congener from the Philippines (Diplopoda, Polydesmida, Paradoxosomatidae)" masterLastPageNumber="17" masterPageNumber="1" pageNumber="6" updateTime="1668153676840" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Three new cavernicolous species of dragon millipedes, genus Desmoxytes Chamberlin, 1923, from southern China, with notes on a formal congener from the Philippines (Diplopoda, Polydesmida, Paradoxosomatidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Golovatch, Sergei I.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Li, Youbang</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Liu, Weixin</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Geoffroy, Jean-Jacques</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2012</mods:date>
<mods:detail type="volume">
<mods:number>185</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>17</mods:end>
</mods:extent>
</mods:part>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.185.3082</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.185.3082</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-185-1</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152034746" LSID="urn:lsid:zoobank.org:act:805A9FDB-02DC-4449-93AE-5C514FD59D0E" httpUri="http://treatment.plazi.org/id/CEA4EBA5B3265B0BD1EA920A79FA0389" lastPageId="8" lastPageNumber="9" pageId="5" pageNumber="6">
<subSubSection pageId="5" pageNumber="6" type="nomenclature">
<paragraph pageId="5" pageNumber="6">
<taxonomicName LSID="urn:lsid:zoobank.org:act:805A9FDB-02DC-4449-93AE-5C514FD59D0E" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmoxytes spinissima" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="spinissima">Desmoxytes spinissima</taxonomicName>
<taxonomicNameLabel pageId="5" pageNumber="6">sp. n.</taxonomicNameLabel>
Figs 34
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="6" type="holotype">
<paragraph pageId="5" pageNumber="6">Holotype.</paragraph>
<paragraph pageId="5" pageNumber="6">♂(IZAS), China, Guangxi Prov., Fuchuan County, Guanyuan, Cave Guanyuan Dong, 25.I.2012, leg. Li Youbang, Li Youting and Tang Kewen (CHIgx12-LYB04).</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="6" type="paratype">
<paragraph pageId="5" pageNumber="6">Paratype.</paragraph>
<paragraph pageId="5" pageNumber="6">1 ♀ (GNUG), same locality, together with holotype.</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="6" type="name">
<paragraph pageId="5" pageNumber="6">Name.</paragraph>
<paragraph pageId="5" pageNumber="6">To emphasize the extremely long and spiniform paraterga.</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="6" type="diagnosis">
<paragraph pageId="5" pageNumber="6">Diagnosis.</paragraph>
<paragraph pageId="5" pageNumber="6">Differs in the paraterga being long and spiniform throughout, combined with a strongly condensed solenophore and both ♂ femora 6 and 7 being humped. See also Key below.</paragraph>
</subSubSection>
<subSubSection lastPageId="8" lastPageNumber="9" pageId="5" pageNumber="6" type="description">
<paragraph pageId="5" pageNumber="6">Description.</paragraph>
<paragraph pageId="5" pageNumber="6">Length ca 27 (♂) or 36 mm (♀); width of pro- and metaterga together with paraterga 1.6 and 4.0 (♂) or 2.0 and 4.0 mm (♀), respectively.</paragraph>
<paragraph pageId="5" pageNumber="6">Body strongly moniliform (Fig. 3A, B). Coloration of alcohol material rather uniformly light pink-brownish to nearly pallid, anterior body part a little darker, pinkish (Fig. 3). Antennomere 7 dark brown. In width, head &lt;&lt; collum = segment 2 = 3 &lt;4 &lt;5 = 16; thereafter body gradually tapering towards telson. Head densely setose throughout, epicranial suture distinct. Antennae very long and slender, reaching back to segment 8 (♂) or 6 (♀) dorsally, antennomeres 5 and 6 each with an apicodorsal compact group of bacilliform sensilla.</paragraph>
<paragraph pageId="5" pageNumber="6">Tegument rather smooth and poorly shining, prozona very delicately microalveolate, metaterga finely shagreened to microgranulate/microspiculate, surface below paraterga finely shagreened. Collum with 3+3 evident setigerous spines in a row at front margin (growing increasingly long laterad), behind it with 1+1 and 2+2 similar spinules in two transverse rows. Following metaterga with a pattern of similar 1+1 and 2+2 spinules in two transverse rows. Collum and all following metaterga with extremely long, straight, spiniform paraterga (Fig. 3A, B) directed more dorsally than laterally and ending up clearly above dorsum on collum and in segments 2-18; only paraterga 19 subhorizontal, about level with dorsum, directed clearly caudad and reaching behind until about midlength along telson. Paraterga ca 1.2-1.3 (♂) or 0.9 times (♀) as long as midbody height. Paraterga 1-18 with 2-3 evident indentations/spinules frontally (these growing increasingly inconspicuous towards telson) and a short, but evident tooth posteriorly at base. Stricture between pro- and metazona rather narrow and shallow, always smooth at bottom (Fig. 3B). Pore formula normal; ozopores inconspicuous, located just behind last indentation on ventral side of poriferous paraterga. Transverse sulcus missing. Pleurosternal carinae evident only in segments 2 and 3. Epiproct (Fig. 3C, D) rather simple, subapical and, especially, pre-apical papillae very distinct, finger-shaped. Hypoproct (Fig. 3D) subtrapeziform, caudal margin emarginate, setigerous cones at caudal edge small, but evident, widely separated. Axial line missing.</paragraph>
<paragraph lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
Sterna quite densely setose, cross-impressions faint. A paramedian pair of entirely separated, short, rounded, setose tubercles between ♂ coxae 4 (Fig. 4A). Legs (
<pageBreakToken pageId="6" pageNumber="7" start="start">Fig</pageBreakToken>
. 3
<normalizedToken originalValue="AC">A-C</normalizedToken>
) extremely long and slender,&gt; 4.0 (♂) or 3.2 (♀) times longer than midbody height. ♂femora 6 and 7 strongly inflated ventrally in distal quarter (Fig. 3E)
</paragraph>
<paragraph lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
Gonopods (Fig. 4B, C) simple. Coxite rather short, subcylindrical, poorly setose distodorsally, about 1/3 as long as telopodite. Prefemoral portion (= densely setose)
<pageBreakToken pageId="7" pageNumber="8" start="start">about</pageBreakToken>
half as long as acropodite. Femorite (fe) slender, elongate, only slightly enlarged distad, with seminal groove running entirely on mesal face, also with a distinct sulcus demarcating a short, strongly condensed solenophore (sph). The latter distinguished by a smaller, somewhat folded lamina medialis (lm) and a simpler and larger lamina lateralis (ll). Solenomere (sl) similarly short, flagelliform, rather faintly separated at base from solenophore.
</paragraph>
<caption pageId="8" pageNumber="9">
<paragraph pageId="8" pageNumber="9">
<pageBreakToken pageId="8" pageNumber="9" start="start">Figure</pageBreakToken>
3.
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmoxytes spinissima" order="Polydesmida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="spinissima">Desmoxytes spinissima</taxonomicName>
sp. n., ♂ holotype. A anterior part of body, lateral view B midbody segments, lateral view C posterior part of body, dorsal view D telson, ventral view E femora 6 and 7, lateral view.Photographed not to scale.
</paragraph>
</caption>
<caption pageId="8" pageNumber="9">
<paragraph pageId="8" pageNumber="9">
Figure 4.
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmoxytes spinissima" order="Polydesmida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="spinissima">Desmoxytes spinissima</taxonomicName>
sp. n., ♂ holotype. A sternal cones between coxae 4, ventral view
<normalizedToken originalValue="BC">B-C</normalizedToken>
left gonopod, mesal and lateral views, respectively. Scale bar: 1.0 (A) and 0.5 mm (B, C). Designations: fe femorite sph solenophore sl solenomere ll lamina lateralis lm lamina medialis
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>