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<document id="2AA8F1768C8CF0F219F59D1E27509B52" ID-DOI="10.11646/zootaxa.4039.4.1" ID-GBIF-Dataset="ca91ab86-49b9-4d90-ac6f-2051ca9d2f85" ID-ISSN="1175-5326" ID-Zenodo-Dep="232034" ID-ZooBank="4B12AA52-6A3F-44B9-BBEC-EBABF0E22DF5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461332656152" checkinUser="plazi" docAuthor="Blake, James A. &amp; Göransson, Peter" docDate="2015" docId="EE2B2061FFC30270FF2F0BCEFC03FA62" docLanguage="en" docName="zt04039p515.pdf" docOrigin="Zootaxa 4039 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tharyx maryae Blake &amp; Göransson, 2015, new species" docType="treatment" docVersion="8" lastPageNumber="508" masterDocId="12125819FFC70277FFB8095EFFE8FF8E" masterDocTitle="Redescription of Tharyx killariensis (Southern) from Ireland and description of two new species of Tharyx from the Kattegat, Sweden (Polychaeta, Cirratulidae)" masterLastPageNumber="515" masterPageNumber="501" pageNumber="505" updateTime="1698618770507" updateUser="plazi">
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<mods:title id="1B58C10FE66C64E83748006B38030568">Redescription of Tharyx killariensis (Southern) from Ireland and description of two new species of Tharyx from the Kattegat, Sweden (Polychaeta, Cirratulidae)</mods:title>
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<mods:namePart id="495DF2F864412500BC5FCAC4D20173D7">Blake, James A.</mods:namePart>
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<mods:namePart id="9E27108ADAD1139EACC0662DA73463AC">Göransson, Peter</mods:namePart>
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<mods:date id="9B474128EF5A4D3FCB8C4B558789754B">2015</mods:date>
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<subSubSection id="2E98C2FCFFC30273FF2F0BCEFEB9FD45" pageId="4" pageNumber="505" type="nomenclature">
<paragraph id="663D9177FFC30273FF2F0BCEFE0FFD24" blockId="4.[151,487,656,715]" box="[151,487,656,682]" pageId="4" pageNumber="505">
<heading id="3D75261BFFC30273FF2F0BCEFE0FFD24" bold="true" box="[151,487,656,682]" fontSize="11" level="1" pageId="4" pageNumber="505" reason="1">
<emphasis id="54F64D65FFC30273FF2F0BCEFE0FFD24" bold="true" box="[151,487,656,682]" pageId="4" pageNumber="505">
<taxonomicName id="A182EAF4FFC30273FF2F0BCEFEA7FD24" ID-CoL="7C6Y6" box="[151,335,656,682]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="505" phylum="Annelida" rank="species" species="maryae" status="sp. nov.">
<emphasis id="54F64D65FFC30273FF2F0BCEFEA7FD24" bold="true" box="[151,335,656,682]" italics="true" pageId="4" pageNumber="505">Tharyx maryae</emphasis>
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<taxonomicNameLabel id="4FC5F01EFFC30273FEEF0BCEFE0FFD24" box="[343,487,656,682]" pageId="4" pageNumber="505" rank="species">new species</taxonomicNameLabel>
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</heading>
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<paragraph id="663D9177FFC30273FF2F0BECFEB9FD45" blockId="4.[151,487,656,715]" box="[151,337,690,715]" pageId="4" pageNumber="505">
<figureCitation id="FEB98DF2FFC30273FF2F0BECFEEAFD45" box="[151,258,690,715]" captionStart="FIGURE 2" captionStartId="5.[151,250,1885,1907]" captionTargetBox="[232,1354,201,1840]" captionTargetId="figure@5.[189,1383,193,1864]" captionTargetPageId="5" captionText="FIGURE 2. Tharyx maryae n. sp. Holotype (SMNH 8754): A, anterior end, dorsal view; B, posterior end, right lateral view; C, far posterior parapodium; D G, posterior neuroacicular spines: D, pointed transitional spine from fifth to last setiger on body; E, blunt-tipped spine from fourth to last setiger on body; F, short blunt-tipped spine from second to last setiger on body; G, knob-tipped spine and capillary from last setiger on body. Abbreviations: br, branchiae; per, peristomium; pr, prostomium; tn, tentacle." httpUri="https://zenodo.org/record/232036/files/figure.png" pageId="4" pageNumber="505">Figures 2</figureCitation>
,
<figureCitation id="FEB98DF2FFC30273FEB60BEDFEF2FD45" box="[270,282,691,715]" captionStart="FIGURE 3" captionStartId="6.[151,250,1802,1824]" captionTargetBox="[157,1428,199,1777]" captionTargetId="figure@6.[151,1436,193,1780]" captionTargetPageId="6" captionText="FIGURE 3. Tharyx maryae n. sp. Holotype (SMNH 8754): A, anterior end, right lateral view; B, posterior end, right lateral view. Paratype (SMNH 8755): C, anterior two thirds of body in ventrolateral view. Tharyx robustus n. sp. Holotype (SMNH 8756): D, entire animal in ventral view. A B stained with Shirlastain A; C, stained with methyl green. Abbreviations: MG, methyl green; per, peristomium; pr, prostomium." httpUri="https://zenodo.org/record/232037/files/figure.png" pageId="4" pageNumber="505">3</figureCitation>
AC
</paragraph>
</subSubSection>
<subSubSection id="2E98C2FCFFC30273FF2F0BA4FF37FCD5" pageId="4" pageNumber="505" type="materials_examined">
<paragraph id="663D9177FFC30273FF2F0BA4FF37FCD5" blockId="4.[151,1437,762,2011]" pageId="4" pageNumber="505">
<emphasis id="54F64D65FFC30273FF2F0BA4FE62FC9D" bold="true" box="[151,394,762,787]" pageId="4" pageNumber="505">Material examined.</emphasis>
<collectingCountry id="1E95D1E7FFC30273FE230BA5FDFBFC9D" box="[411,531,763,787]" name="Sweden" pageId="4" pageNumber="505">SWEDEN</collectingCountry>
, Kattegat, near Varberg, 0
<date id="123CB7B7FFC30273FCE70BA5FC11FC9D" box="[863,1017,763,787]" pageId="4" pageNumber="505" value="2012-08-03">3 Aug 2012</date>
, in sediments near eelgrass beds,
<geoCoordinate id="03B6F7B0FFC30273FF2F0A40FECAFCB9" box="[151,290,798,823]" direction="north" orientation="latitude" pageId="4" pageNumber="505" precision="1" value="57.11501">57.11501°N</geoCoordinate>
;
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,
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depth, coll. P. Göransson,
<typeStatus id="B9392FD5FFC30273FCCA0A40FC3CFCB9" box="[882,980,798,823]" pageId="4" pageNumber="505" type="holotype">holotype</typeStatus>
(
<collectionCode id="009309B2FFC30273FC5E0A41FBD1FCB9" box="[998,1081,799,823]" httpUri="http://grbio.org/cool/8z6v-f8is" name="Saskatchewan Museum of Natural History" pageId="4" pageNumber="505">SMNH</collectionCode>
8754), four
<typeStatus id="B9392FD5FFC30273FB750A41FAD0FCB9" box="[1229,1336,799,823]" pageId="4" pageNumber="505" type="paratype">paratypes</typeStatus>
(
<collectionCode id="009309B2FFC30273FAF10A41FA74FCB9" box="[1353,1436,799,823]" httpUri="http://grbio.org/cool/8z6v-f8is" name="Saskatchewan Museum of Natural History" pageId="4" pageNumber="505">SMNH</collectionCode>
8755).
</paragraph>
</subSubSection>
<subSubSection id="2E98C2FCFFC30273FF7F0A38FD53F8E1" pageId="4" pageNumber="505" type="description">
<paragraph id="663D9177FFC30273FF7F0A38FBAEFB69" blockId="4.[151,1437,762,2011]" pageId="4" pageNumber="505">
<emphasis id="54F64D65FFC30273FF7F0A38FEB1FCF1" bold="true" box="[199,345,870,895]" pageId="4" pageNumber="505">Description.</emphasis>
A moderate-sized species,
<typeStatus id="B9392FD5FFC30273FD360A38FD18FCF1" box="[654,752,870,895]" pageId="4" pageNumber="505" type="holotype">holotype</typeStatus>
complete,
<quantity id="A17A3C92FFC30273FCD50A39FC55FCF0" box="[877,957,871,895]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="505" unit="mm" value="10.0">10 mm</quantity>
long,
<quantity id="A17A3C92FFC30273FBBD0A38FBB4FCF0" box="[1029,1116,870,895]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="505" unit="mm" value="0.5">0.5 mm</quantity>
wide across thorax,
<quantity id="A17A3C92FFC30273FAFE0A39FA74FCF0" box="[1350,1436,871,895]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="505" unit="mm" value="0.8">0.8 mm</quantity>
wide across expanded posterior end for 115 setigerous segments;
<typeStatus id="B9392FD5FFC30273FC390AD5FC04FC2D" box="[897,1004,907,931]" pageId="4" pageNumber="505" type="paratype">paratypes</typeStatus>
all incomplete (lacking posteriormost segments), up to
<quantity id="A17A3C92FFC30273FEEE0AF1FE7FFC48" box="[342,407,943,967]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="4" pageNumber="505" unit="mm" value="9.0">9 mm</quantity>
long,
<quantity id="A17A3C92FFC30273FE660AF0FDDBFC48" box="[478,563,942,967]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="505" unit="mm" value="0.5">0.5 mm</quantity>
wide, with about 70 setigerous segments. Color in alcohol light tan to opaque white; lateral black pigment spots present on peristomium of some specimens (
<figureCitation id="FEB98DF2FFC30273FBAF0A8CFBB1FC65" box="[1047,1113,978,1003]" captionStart="FIGURE 3" captionStartId="6.[151,250,1802,1824]" captionTargetBox="[157,1428,199,1777]" captionTargetId="figure@6.[151,1436,193,1780]" captionTargetPageId="6" captionText="FIGURE 3. Tharyx maryae n. sp. Holotype (SMNH 8754): A, anterior end, right lateral view; B, posterior end, right lateral view. Paratype (SMNH 8755): C, anterior two thirds of body in ventrolateral view. Tharyx robustus n. sp. Holotype (SMNH 8756): D, entire animal in ventral view. A B stained with Shirlastain A; C, stained with methyl green. Abbreviations: MG, methyl green; per, peristomium; pr, prostomium." httpUri="https://zenodo.org/record/232037/files/figure.png" pageId="4" pageNumber="505">Fig. 3</figureCitation>
A), faded or not apparent on others; some specimens with brown pigment spots in intersegmental grooves along body. Body elongate, laterally expanded through first 2025 thoracic setigers with narrow crowded segments (
<figureCitation id="FEB98DF2FFC30273FBA80D44FBB5FBBD" box="[1040,1117,1050,1075]" captionStart="FIGURE 2" captionStartId="5.[151,250,1885,1907]" captionTargetBox="[232,1354,201,1840]" captionTargetId="figure@5.[189,1383,193,1864]" captionTargetPageId="5" captionText="FIGURE 2. Tharyx maryae n. sp. Holotype (SMNH 8754): A, anterior end, dorsal view; B, posterior end, right lateral view; C, far posterior parapodium; D G, posterior neuroacicular spines: D, pointed transitional spine from fifth to last setiger on body; E, blunt-tipped spine from fourth to last setiger on body; F, short blunt-tipped spine from second to last setiger on body; G, knob-tipped spine and capillary from last setiger on body. Abbreviations: br, branchiae; per, peristomium; pr, prostomium; tn, tentacle." httpUri="https://zenodo.org/record/232036/files/figure.png" pageId="4" pageNumber="505">Figs. 2</figureCitation>
A, 3C); abdominal segments narrower, longer and rounded, but not moniliform; posterior 2025 segments becoming enlarged, forming expanded posterior end with narrow segments, tapering to pygidium with dorsal anus and bulbous ventral lobe (
<figureCitation id="FEB98DF2FFC30273FF270DD8FF07FB11" box="[159,239,1158,1183]" captionStart="FIGURE 2" captionStartId="5.[151,250,1885,1907]" captionTargetBox="[232,1354,201,1840]" captionTargetId="figure@5.[189,1383,193,1864]" captionTargetPageId="5" captionText="FIGURE 2. Tharyx maryae n. sp. Holotype (SMNH 8754): A, anterior end, dorsal view; B, posterior end, right lateral view; C, far posterior parapodium; D G, posterior neuroacicular spines: D, pointed transitional spine from fifth to last setiger on body; E, blunt-tipped spine from fourth to last setiger on body; F, short blunt-tipped spine from second to last setiger on body; G, knob-tipped spine and capillary from last setiger on body. Abbreviations: br, branchiae; per, peristomium; pr, prostomium; tn, tentacle." httpUri="https://zenodo.org/record/232036/files/figure.png" pageId="4" pageNumber="505">Figs. 2</figureCitation>
B, 3B). Dorsum of thoracic region broad between notopodia, with segmentation evident (
<figureCitation id="FEB98DF2FFC30273FB5E0DD8FAC3FB11" box="[1254,1323,1158,1183]" captionStart="FIGURE 2" captionStartId="5.[151,250,1885,1907]" captionTargetBox="[232,1354,201,1840]" captionTargetId="figure@5.[189,1383,193,1864]" captionTargetPageId="5" captionText="FIGURE 2. Tharyx maryae n. sp. Holotype (SMNH 8754): A, anterior end, dorsal view; B, posterior end, right lateral view; C, far posterior parapodium; D G, posterior neuroacicular spines: D, pointed transitional spine from fifth to last setiger on body; E, blunt-tipped spine from fourth to last setiger on body; F, short blunt-tipped spine from second to last setiger on body; G, knob-tipped spine and capillary from last setiger on body. Abbreviations: br, branchiae; per, peristomium; pr, prostomium; tn, tentacle." httpUri="https://zenodo.org/record/232036/files/figure.png" pageId="4" pageNumber="505">Fig. 2</figureCitation>
A); dorsal groove absent. Venter generally flattened with mid-ventral line of ridges along entire body from setigers 23 (
<figureCitation id="FEB98DF2FFC30273FAD70DF4FF4DFB69" captionStart="FIGURE 3" captionStartId="6.[151,250,1802,1824]" captionTargetBox="[157,1428,199,1777]" captionTargetId="figure@6.[151,1436,193,1780]" captionTargetPageId="6" captionText="FIGURE 3. Tharyx maryae n. sp. Holotype (SMNH 8754): A, anterior end, right lateral view; B, posterior end, right lateral view. Paratype (SMNH 8755): C, anterior two thirds of body in ventrolateral view. Tharyx robustus n. sp. Holotype (SMNH 8756): D, entire animal in ventral view. A B stained with Shirlastain A; C, stained with methyl green. Abbreviations: MG, methyl green; per, peristomium; pr, prostomium." httpUri="https://zenodo.org/record/232037/files/figure.png" pageId="4" pageNumber="505">Fig. 3</figureCitation>
C); ridges formed from ventromedial bulges or pads associated with each segment.
</paragraph>
<paragraph id="663D9177FFC30273FF7F0DACFD26FA31" blockId="4.[151,1437,762,2011]" pageId="4" pageNumber="505">
Prostomium triangular, tapering to pointed tip on anterior margin (
<figureCitation id="FEB98DF2FFC30273FC640DACFBC6FA85" box="[988,1070,1266,1291]" captionStart="FIGURE 2" captionStartId="5.[151,250,1885,1907]" captionTargetBox="[232,1354,201,1840]" captionTargetId="figure@5.[189,1383,193,1864]" captionTargetPageId="5" captionText="FIGURE 2. Tharyx maryae n. sp. Holotype (SMNH 8754): A, anterior end, dorsal view; B, posterior end, right lateral view; C, far posterior parapodium; D G, posterior neuroacicular spines: D, pointed transitional spine from fifth to last setiger on body; E, blunt-tipped spine from fourth to last setiger on body; F, short blunt-tipped spine from second to last setiger on body; G, knob-tipped spine and capillary from last setiger on body. Abbreviations: br, branchiae; per, peristomium; pr, prostomium; tn, tentacle." httpUri="https://zenodo.org/record/232036/files/figure.png" pageId="4" pageNumber="505">Figs. 2</figureCitation>
A, 3A), broadening posteriorly, nuchal organs narrow slits on posterior margin of prostomium, inconspicuous, not pigmented; eyes absent. Peristomium as long as wide, with two annuli surmounted by longitudinal dorsal crest (
<figureCitation id="FEB98DF2FFC30273FB2A0C64FB31FADD" box="[1170,1241,1338,1363]" captionStart="FIGURE 2" captionStartId="5.[151,250,1885,1907]" captionTargetBox="[232,1354,201,1840]" captionTargetId="figure@5.[189,1383,193,1864]" captionTargetPageId="5" captionText="FIGURE 2. Tharyx maryae n. sp. Holotype (SMNH 8754): A, anterior end, dorsal view; B, posterior end, right lateral view; C, far posterior parapodium; D G, posterior neuroacicular spines: D, pointed transitional spine from fifth to last setiger on body; E, blunt-tipped spine from fourth to last setiger on body; F, short blunt-tipped spine from second to last setiger on body; G, knob-tipped spine and capillary from last setiger on body. Abbreviations: br, branchiae; per, peristomium; pr, prostomium; tn, tentacle." httpUri="https://zenodo.org/record/232036/files/figure.png" pageId="4" pageNumber="505">Fig. 2</figureCitation>
A). Paired dorsal tentacles on posterior margin of peristomium, with first pair of branchiae located immediately posterior to tentacles on peristomium (
<figureCitation id="FEB98DF2FFC30273FEEE0CDCFE71FA15" box="[342,409,1410,1435]" captionStart="FIGURE 2" captionStartId="5.[151,250,1885,1907]" captionTargetBox="[232,1354,201,1840]" captionTargetId="figure@5.[189,1383,193,1864]" captionTargetPageId="5" captionText="FIGURE 2. Tharyx maryae n. sp. Holotype (SMNH 8754): A, anterior end, dorsal view; B, posterior end, right lateral view; C, far posterior parapodium; D G, posterior neuroacicular spines: D, pointed transitional spine from fifth to last setiger on body; E, blunt-tipped spine from fourth to last setiger on body; F, short blunt-tipped spine from second to last setiger on body; G, knob-tipped spine and capillary from last setiger on body. Abbreviations: br, branchiae; per, peristomium; pr, prostomium; tn, tentacle." httpUri="https://zenodo.org/record/232036/files/figure.png" pageId="4" pageNumber="505">Fig. 2</figureCitation>
A); second pair of branchiae on posterior margin of setiger 1, dorsal to notosetae. Branchiae present on most segments through middle of body.
</paragraph>
<paragraph id="663D9177FFC30273FF7F0C95FE77F935" blockId="4.[151,1437,762,2011]" pageId="4" pageNumber="505">
Noto- and neuropodia with setal fascicles arising close together; anterior parapodia with spreading fascicles of about 56 capillary setae in notopodia and 67 capillaries in neuropodia; middle body segments with about 910 capillaries in notopodia and
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neuropodia; far posterior notopodia with 45 narrow capillaries; neuropodia with 34 capillaries and 13 short, curved, spines in posteriormost 45 setigers (
<figureCitation id="FEB98DF2FFC30273FB840F68FB6AF9C1" box="[1084,1154,1590,1615]" captionStart="FIGURE 2" captionStartId="5.[151,250,1885,1907]" captionTargetBox="[232,1354,201,1840]" captionTargetId="figure@5.[189,1383,193,1864]" captionTargetPageId="5" captionText="FIGURE 2. Tharyx maryae n. sp. Holotype (SMNH 8754): A, anterior end, dorsal view; B, posterior end, right lateral view; C, far posterior parapodium; D G, posterior neuroacicular spines: D, pointed transitional spine from fifth to last setiger on body; E, blunt-tipped spine from fourth to last setiger on body; F, short blunt-tipped spine from second to last setiger on body; G, knob-tipped spine and capillary from last setiger on body. Abbreviations: br, branchiae; per, peristomium; pr, prostomium; tn, tentacle." httpUri="https://zenodo.org/record/232036/files/figure.png" pageId="4" pageNumber="505">Fig. 2</figureCitation>
C). Neuroacicular spines initially as straight pointed spine (
<figureCitation id="FEB98DF2FFC30273FDAA0F04FDBCF9FD" box="[530,596,1626,1651]" captionStart="FIGURE 2" captionStartId="5.[151,250,1885,1907]" captionTargetBox="[232,1354,201,1840]" captionTargetId="figure@5.[189,1383,193,1864]" captionTargetPageId="5" captionText="FIGURE 2. Tharyx maryae n. sp. Holotype (SMNH 8754): A, anterior end, dorsal view; B, posterior end, right lateral view; C, far posterior parapodium; D G, posterior neuroacicular spines: D, pointed transitional spine from fifth to last setiger on body; E, blunt-tipped spine from fourth to last setiger on body; F, short blunt-tipped spine from second to last setiger on body; G, knob-tipped spine and capillary from last setiger on body. Abbreviations: br, branchiae; per, peristomium; pr, prostomium; tn, tentacle." httpUri="https://zenodo.org/record/232036/files/figure.png" pageId="4" pageNumber="505">Fig. 2</figureCitation>
D) transitioning over next four setigers to blunt-tipped spine (
<figureCitation id="FEB98DF2FFC30273FABA0F04FAADF9FD" box="[1282,1349,1626,1651]" captionStart="FIGURE 2" captionStartId="5.[151,250,1885,1907]" captionTargetBox="[232,1354,201,1840]" captionTargetId="figure@5.[189,1383,193,1864]" captionTargetPageId="5" captionText="FIGURE 2. Tharyx maryae n. sp. Holotype (SMNH 8754): A, anterior end, dorsal view; B, posterior end, right lateral view; C, far posterior parapodium; D G, posterior neuroacicular spines: D, pointed transitional spine from fifth to last setiger on body; E, blunt-tipped spine from fourth to last setiger on body; F, short blunt-tipped spine from second to last setiger on body; G, knob-tipped spine and capillary from last setiger on body. Abbreviations: br, branchiae; per, peristomium; pr, prostomium; tn, tentacle." httpUri="https://zenodo.org/record/232036/files/figure.png" pageId="4" pageNumber="505">Fig. 2</figureCitation>
E), then spines with knob-like tips (
<figureCitation id="FEB98DF2FFC30273FE7F0F20FDFDF919" box="[455,533,1662,1687]" captionStart="FIGURE 2" captionStartId="5.[151,250,1885,1907]" captionTargetBox="[232,1354,201,1840]" captionTargetId="figure@5.[189,1383,193,1864]" captionTargetPageId="5" captionText="FIGURE 2. Tharyx maryae n. sp. Holotype (SMNH 8754): A, anterior end, dorsal view; B, posterior end, right lateral view; C, far posterior parapodium; D G, posterior neuroacicular spines: D, pointed transitional spine from fifth to last setiger on body; E, blunt-tipped spine from fourth to last setiger on body; F, short blunt-tipped spine from second to last setiger on body; G, knob-tipped spine and capillary from last setiger on body. Abbreviations: br, branchiae; per, peristomium; pr, prostomium; tn, tentacle." httpUri="https://zenodo.org/record/232036/files/figure.png" pageId="4" pageNumber="505">Figs. 2</figureCitation>
FG). On
<typeStatus id="B9392FD5FFC30273FD3F0F20FD01F919" box="[647,745,1662,1687]" pageId="4" pageNumber="505" type="holotype">holotype</typeStatus>
of 115 setigers, neuroacicular spines limited to posteriormost five setigers (111115).
</paragraph>
<paragraph id="663D9177FFC30273FF7F0F98FD53F8E1" blockId="4.[151,1437,762,2011]" pageId="4" pageNumber="505">
<emphasis id="54F64D65FFC30273FF7F0F98FE57F951" bold="true" box="[199,447,1734,1759]" pageId="4" pageNumber="505">Methyl Green Stain.</emphasis>
Most of body staining initially, tip of prostomium and dorsal peristomial crest clear; stain retained laterally in irregular patches on sides of peristomium and intersegmentally on dorsum, between parapodia, and mid-ventrally on thoracic region where ventral ridges are located sometimes producing a line of paired spots along the venter. The most conspicuous stain is present intersegmentally between the parapodia; no distinct pattern on prostomium and peristomium.
</paragraph>
</subSubSection>
<subSubSection id="2E98C2FCFFC30270FF7F0E24FD7FFD7E" lastPageId="7" lastPageNumber="508" pageId="4" pageNumber="505" type="discussion">
<paragraph id="663D9177FFC30273FF7F0E24FA65F855" blockId="4.[151,1437,762,2011]" pageId="4" pageNumber="505">
<emphasis id="54F64D65FFC30273FF7F0E24FED3F81D" bold="true" box="[199,315,1914,1939]" pageId="4" pageNumber="505">Remarks.</emphasis>
Because of the expanded posterior end on the
<typeStatus id="B9392FD5FFC30273FCD30E24FC25F81D" box="[875,973,1914,1939]" pageId="4" pageNumber="505" type="holotype">holotype</typeStatus>
of
<taxonomicName id="A182EAF4FFC30273FC420E22FB84F81D" box="[1018,1132,1916,1939]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="505" phylum="Annelida" rank="species" species="maryae" status="sp. nov.">
<emphasis id="54F64D65FFC30273FC420E22FB84F81D" box="[1018,1132,1916,1939]" italics="true" pageId="4" pageNumber="505">T. maryae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FC5F01EFFC30273FBC10E24FB53F81C" box="[1145,1211,1914,1939]" pageId="4" pageNumber="505" rank="species">
<emphasis id="54F64D65FFC30273FBC10E24FB5EF81D" bold="true" box="[1145,1206,1914,1939]" pageId="4" pageNumber="505">n. sp</emphasis>
.
</taxonomicNameLabel>
, the specimen was initially thought to be a species of
<taxonomicName id="A182EAF4FFC30273FD9A0EC0FD54F839" box="[546,700,1950,1975]" class="Polychaeta" family="Cirratulidae" genus="Aphelochaeta" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="505" phylum="Annelida" rank="genus">
<emphasis id="54F64D65FFC30273FD9A0EC0FD54F839" box="[546,700,1950,1975]" italics="true" pageId="4" pageNumber="505">Aphelochaeta</emphasis>
</taxonomicName>
. However, careful inspection of the posterior parapodia revealed the presence of short, curved, blunt-tipped spines ventral to the capillaries in the posteriormost five neuropodia.
</paragraph>
<caption id="32FDC1FFFFC20272FF2F0E03FF05F860" httpUri="https://zenodo.org/record/232036/files/figure.png" pageId="5" pageNumber="506" targetBox="[232,1354,201,1840]" targetPageId="5">
<paragraph id="663D9177FFC20272FF2F0E03FF05F860" blockId="5.[151,1436,1885,2030]" pageId="5" pageNumber="506">
<emphasis id="54F64D65FFC20272FF2F0E03FEFAF8FD" bold="true" box="[151,274,1885,1907]" pageId="5" pageNumber="506">FIGURE 2.</emphasis>
<taxonomicName id="A182EAF4FFC20272FEA00E03FE59F8FA" box="[280,433,1885,1908]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="5" pageNumber="506" phylum="Annelida" rank="species" species="maryae" status="sp. nov.">
<emphasis id="54F64D65FFC20272FEA00E03FE59F8FA" box="[280,433,1885,1908]" italics="true" pageId="5" pageNumber="506">Tharyx maryae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FC5F01EFFC20272FE0F0E00FE18F8FD" box="[439,496,1886,1907]" pageId="5" pageNumber="506" rank="species">
<emphasis id="54F64D65FFC20272FE0F0E00FE18F8FD" bold="true" box="[439,496,1886,1907]" pageId="5" pageNumber="506">n. sp.</emphasis>
</taxonomicNameLabel>
Holotype (SMNH 8754): A, anterior end, dorsal view; B, posterior end, right lateral view; C, far posterior parapodium; DG, posterior neuroacicular spines: D, pointed transitional spine from fifth to last setiger on body; E, blunt-tipped spine from fourth to last setiger on body; F, short blunt-tipped spine from second to last setiger on body; G, knob-tipped spine and capillary from last setiger on body. Abbreviations: br, branchiae; per, peristomium; pr, prostomium; tn, tentacle.
</paragraph>
</caption>
<caption id="32FDC1FFFFC10271FF2F0E54FD6DF8F5" httpUri="https://zenodo.org/record/232037/files/figure.png" pageId="6" pageNumber="507" targetBox="[157,1428,199,1777]" targetPageId="6">
<paragraph id="663D9177FFC10271FF2F0E54FD6DF8F5" blockId="6.[151,1436,1802,1915]" pageId="6" pageNumber="507">
<emphasis id="54F64D65FFC10271FF2F0E54FEE6F891" bold="true" box="[151,270,1802,1824]" pageId="6" pageNumber="507">FIGURE 3</emphasis>
.
<taxonomicName id="A182EAF4FFC10271FEA40E54FE5FF8AE" box="[284,439,1802,1824]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="6" pageNumber="507" phylum="Annelida" rank="species" species="maryae" status="sp. nov.">
<emphasis id="54F64D65FFC10271FEA40E54FE5FF8AE" box="[284,439,1802,1824]" italics="true" pageId="6" pageNumber="507">Tharyx maryae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FC5F01EFFC10271FE070E54FE11F891" box="[447,505,1802,1823]" pageId="6" pageNumber="507" rank="species">
<emphasis id="54F64D65FFC10271FE070E54FE11F891" bold="true" box="[447,505,1802,1823]" pageId="6" pageNumber="507">n. sp.</emphasis>
</taxonomicNameLabel>
Holotype (SMNH 8754): A, anterior end, right lateral view; B, posterior end, right lateral view. Paratype (SMNH 8755): C, anterior two thirds of body in ventrolateral view.
<taxonomicName id="A182EAF4FFC10271FC4C0E76FB72F8B0" box="[1012,1178,1832,1854]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="6" pageNumber="507" phylum="Annelida" rank="species" species="robustus" status="sp. nov.">
<emphasis id="54F64D65FFC10271FC4C0E76FB72F8B0" box="[1012,1178,1832,1854]" italics="true" pageId="6" pageNumber="507">Tharyx robustus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FC5F01EFFC10271FB1A0E77FB35F8B0" box="[1186,1245,1833,1854]" pageId="6" pageNumber="507" rank="species">
<emphasis id="54F64D65FFC10271FB1A0E77FB35F8B0" bold="true" box="[1186,1245,1833,1854]" pageId="6" pageNumber="507">n. sp.</emphasis>
</taxonomicNameLabel>
Holotype (SMNH 8756): D, entire animal in ventral view. AB stained with Shirlastain A; C, stained with methyl green. Abbreviations: MG, methyl green; per, peristomium; pr, prostomium.
</paragraph>
</caption>
<paragraph id="663D9177FFC00270FF2F09C9FD30FE5E" blockId="7.[151,1437,151,1517]" pageId="7" pageNumber="508">
These spines transition from curved pointed setae among capillaries to short curved blunt-tipped spines with knoblike tips in the very last setigers. No spines were observed in the notopodia. Because the neuropodial spines are limited to the very posteriormost setigers, they will not be observed in fragmented specimens. Therefore other characters will be required in order to identify the species. Other features of the body are similar to related species of
<taxonomicName id="A182EAF4FFC00270FF0E0879FEEDFECE" box="[182,261,295,320]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="7" pageNumber="508" phylum="Annelida" rank="genus">
<emphasis id="54F64D65FFC00270FF0E0879FEEDFECE" box="[182,261,295,320]" italics="true" pageId="7" pageNumber="508">Tharyx</emphasis>
</taxonomicName>
. These include the raised crest on the peristomium, the occurrence of lateral black pigment spots on the peristomium; the position of the paired dorsal tentacles on the posterior margin of the peristomium followed by the first pair of branchiae also on the peristomium and the generally elongated form of the body. In addition, the methyl green staining pattern in conjunction with the paired peristomial pigment spots readily differentiates this species from other cirratulids observed in these collections.
</paragraph>
<paragraph id="663D9177FFC00270FF7F0882FD7FFD7E" blockId="7.[151,1437,151,1517]" pageId="7" pageNumber="508">
<taxonomicName id="A182EAF4FFC00270FF7F0882FE99FE7A" box="[199,369,476,501]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="7" pageNumber="508" phylum="Annelida" rank="species" species="maryae" status="sp. nov.">
<emphasis id="54F64D65FFC00270FF7F0882FE99FE7A" box="[199,369,476,501]" italics="true" pageId="7" pageNumber="508">Tharyx maryae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FC5F01EFFC00270FEC10882FE50FE7B" box="[377,440,476,501]" pageId="7" pageNumber="508" rank="species">
<emphasis id="54F64D65FFC00270FEC10882FE50FE7B" bold="true" box="[377,440,476,501]" pageId="7" pageNumber="508">n. sp.</emphasis>
</taxonomicNameLabel>
differs from all other species of the genus in having an expanded posterior end and with the neuropodial spines limited to the five posteriormost setigers. The expanded posterior end suggests a close relationship of
<taxonomicName id="A182EAF4FFC00270FEFC0B7BFE5CFDB2" box="[324,436,549,572]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="7" pageNumber="508" phylum="Annelida" rank="species" species="maryae" status="sp. nov.">
<emphasis id="54F64D65FFC00270FEFC0B7BFE5CFDB2" box="[324,436,549,572]" italics="true" pageId="7" pageNumber="508">T. maryae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FC5F01EFFC00270FE050B7AFE16FDB3" box="[445,510,548,573]" pageId="7" pageNumber="508" rank="species">
<emphasis id="54F64D65FFC00270FE050B7AFE16FDB3" bold="true" box="[445,510,548,573]" pageId="7" pageNumber="508">n. sp.</emphasis>
</taxonomicNameLabel>
with species of
<taxonomicName id="A182EAF4FFC00270FD030B7AFCBDFDB3" box="[699,853,548,573]" class="Polychaeta" family="Cirratulidae" genus="Aphelochaeta" kingdom="Animalia" order="Terebellida" pageId="7" pageNumber="508" phylum="Annelida" rank="genus">
<emphasis id="54F64D65FFC00270FD030B7AFCBDFDB3" box="[699,853,548,573]" italics="true" pageId="7" pageNumber="508">Aphelochaeta</emphasis>
</taxonomicName>
also having this character. Among eight species of
<taxonomicName id="A182EAF4FFC00270FF2F0B19FEDAFDEE" box="[151,306,583,608]" class="Polychaeta" family="Cirratulidae" genus="Aphelochaeta" kingdom="Animalia" order="Terebellida" pageId="7" pageNumber="508" phylum="Annelida" rank="genus">
<emphasis id="54F64D65FFC00270FF2F0B19FEDAFDEE" box="[151,306,583,608]" italics="true" pageId="7" pageNumber="508">Aphelochaeta</emphasis>
</taxonomicName>
described by
<bibRefCitation id="0213EC86FFC00270FE6C0B19FD81FDEE" author="Blake" box="[468,617,583,608]" pageId="7" pageNumber="508" refString="Blake, J. A. (1996) Chapter 8. Family Cirratulidae. In: Blake, J. A., Hilbig, B. &amp; Scott, P. H. (Eds.), Taxonomic Atlas of the Santa Maria Basin and Western Santa Barbara Channel. Vol. 6. Annelida Part 3. Polychaeta: Orbiniidae to Cossuridae. Santa Barbara Museum of Natural History, California, pp. 263 - 384." type="journal article" year="1996">Blake (1996)</bibRefCitation>
from the eastern Pacific, six species had the posterior end expanded in some form. Of these, none had the first pair of branchiae positioned immediately posterior to the dorsal tentacle on the peristomium as is typical for species of
<taxonomicName id="A182EAF4FFC00270FD310BD1FD30FD26" box="[649,728,655,680]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="7" pageNumber="508" phylum="Annelida" rank="genus">
<emphasis id="54F64D65FFC00270FD310BD1FD30FD26" box="[649,728,655,680]" italics="true" pageId="7" pageNumber="508">Tharyx</emphasis>
</taxonomicName>
. When branchiae were present on the peristomium, they were positioned lateral to the tentacles. There are no described species of
<taxonomicName id="A182EAF4FFC00270FC2C0BEAFBC6FD43" box="[916,1070,692,717]" class="Polychaeta" family="Cirratulidae" genus="Aphelochaeta" kingdom="Animalia" order="Terebellida" pageId="7" pageNumber="508" phylum="Annelida" rank="genus">
<emphasis id="54F64D65FFC00270FC2C0BEAFBC6FD43" box="[916,1070,692,717]" italics="true" pageId="7" pageNumber="508">Aphelochaeta</emphasis>
</taxonomicName>
from European waters that have characters similar to those of
<taxonomicName id="A182EAF4FFC00270FE590B87FDA7FD7E" box="[481,591,729,752]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="7" pageNumber="508" phylum="Annelida" rank="species" species="maryae" status="sp. nov.">
<emphasis id="54F64D65FFC00270FE590B87FDA7FD7E" box="[481,591,729,752]" italics="true" pageId="7" pageNumber="508">T. maryae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FC5F01EFFC00270FDE00B89FD7FFD7E" box="[600,663,727,752]" pageId="7" pageNumber="508" rank="species">
<emphasis id="54F64D65FFC00270FDE00B89FD7FFD7E" bold="true" box="[600,663,727,752]" pageId="7" pageNumber="508">n. sp.</emphasis>
</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="2E98C2FCFFC00270FF7F0BA2FC6FFCB6" pageId="7" pageNumber="508" type="etymology">
<paragraph id="663D9177FFC00270FF7F0BA2FC6FFCB6" blockId="7.[151,1437,151,1517]" pageId="7" pageNumber="508">
<emphasis id="54F64D65FFC00270FF7F0BA2FEA6FC9B" bold="true" box="[199,334,764,789]" pageId="7" pageNumber="508">Etymology.</emphasis>
This species is named for the late Dr. Mary Elizabeth Petersen, in recognition of her numerous contributions to polychaete systematics and research on cirratulids.
</paragraph>
</subSubSection>
<subSubSection id="2E98C2FCFFC00270FF7F0A1AFD3EFA47" pageId="7" pageNumber="508" type="biology_ecology">
<paragraph id="663D9177FFC00270FF7F0A1AFDD8FB43" blockId="7.[151,1437,151,1517]" pageId="7" pageNumber="508">
<emphasis id="54F64D65FFC00270FF7F0A1AFECFFCD3" bold="true" box="[199,295,836,861]" pageId="7" pageNumber="508">Biology.</emphasis>
The specimens of
<taxonomicName id="A182EAF4FFC00270FE460A1AFD42FCD2" box="[510,682,836,861]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="7" pageNumber="508" phylum="Annelida" rank="species" species="maryae" status="sp. nov.">
<emphasis id="54F64D65FFC00270FE460A1AFD42FCD2" box="[510,682,836,861]" italics="true" pageId="7" pageNumber="508">Tharyx maryae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FC5F01EFFC00270FD0A0A1AFD1AFCD3" box="[690,754,836,861]" pageId="7" pageNumber="508" rank="species">
<emphasis id="54F64D65FFC00270FD0A0A1AFD1AFCD3" bold="true" box="[690,754,836,861]" pageId="7" pageNumber="508">n. sp.</emphasis>
</taxonomicNameLabel>
were collected by one us (PG) in connection with a benthic macrofauna mapping project in sediments in and outside eelgrass (
<taxonomicName id="A182EAF4FFC00270FC310A37FBDFFC0E" box="[905,1079,873,896]" class="Liliopsida" family="Zosteraceae" genus="Zostera" kingdom="Plantae" order="Alismatales" pageId="7" pageNumber="508" phylum="Tracheophyta" rank="species" species="marina">
<emphasis id="54F64D65FFC00270FC310A37FBDFFC0E" box="[905,1079,873,896]" italics="true" pageId="7" pageNumber="508">Zostera marina</emphasis>
</taxonomicName>
) beds near Varberg on the west coast of
<collectingCountry id="1E95D1E7FFC00270FF4E0AD2FEBAFC2B" box="[246,338,908,933]" name="Sweden" pageId="7" pageNumber="508">Sweden</collectingCountry>
. Salinity and temperature variation in this shallow area is high (2030 PSU; &lt;0°C to&gt;25°C). The worms occurred in four of 30 quantitative haps-corer samples, calculated to each contain 160400 ind/m2 with a biomass of &lt;
<quantity id="A17A3C92FFC00270FE920A84FE4AFC7C" box="[298,418,986,1010]" metricMagnitude="-4" metricUnit="kg" metricValue="4.05" metricValueMax="8.0" metricValueMin="0.1" pageId="7" pageNumber="508" unit="g" value="0.405" valueMax="0.8" valueMin="0.01">0.010.8 g</quantity>
/m2 (variation in specimen length was
<quantity id="A17A3C92FFC00270FCEF0A87FC21FC7C" box="[855,969,985,1010]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="12.0" metricValueMin="6.0" pageId="7" pageNumber="508" unit="mm" value="9.0" valueMax="12.0" valueMin="6.0">612 mm</quantity>
). These samples were taken at
<quantity id="A17A3C92FFC00270FA920A84FA74FC7F" box="[1322,1436,986,1010]" metricMagnitude="0" metricUnit="m" metricValue="1.0" metricValueMax="1.7" metricValueMin="0.3" pageId="7" pageNumber="508" unit="m" value="1.0" valueMax="1.7" valueMin="0.3">0.31.7 m</quantity>
depth in coarse to fine sand with a low organic content (about 12 % loss on ignition) and almost no vegetation, outside or in the margin of the eelgrass bed. The associated fauna were relatively scarce in most samples (612 species and 3,093 ind/m2) and were dominated by
<taxonomicName id="A182EAF4FFC00270FD450D16FB34FBEF" authority="O.F. Muller, 1776" authorityName="O.F. Muller" authorityYear="1776" box="[765,1244,1096,1121]" class="Polychaeta" family="Nereididae" genus="Hediste" kingdom="Animalia" order="Phyllodocida" pageId="7" pageNumber="508" phylum="Annelida" rank="species" species="diversicolor">
<emphasis id="54F64D65FFC00270FD450D16FC0FFBEF" box="[765,999,1096,1121]" italics="true" pageId="7" pageNumber="508">Hediste diversicolor</emphasis>
(
<collectingCountry id="1E95D1E7FFC00270FC460D17FBFCFBEF" box="[1022,1044,1097,1121]" name="Iceland" pageId="7" pageNumber="508">O</collectingCountry>
.F. Müller, 1776)
</taxonomicName>
,
<taxonomicName id="A182EAF4FFC00270FB480D17FED1FB0B" authority="Pennant, 1777" authorityName="Pennant" authorityYear="1777" class="Gastropoda" family="Hydrobiidae" genus="Peringia" kingdom="Animalia" order="Littorinimorpha" pageId="7" pageNumber="508" phylum="Mollusca" rank="species" species="ulvae">
<emphasis id="54F64D65FFC00270FB480D17FA74FBEF" box="[1264,1436,1096,1121]" italics="true" pageId="7" pageNumber="508">Peringia ulvae</emphasis>
Pennant, 1777
</taxonomicName>
,
<taxonomicName id="A182EAF4FFC00270FEF00D32FD36FB0B" authority="Udekem, 1855" authorityName="Udekem" authorityYear="1855" box="[328,734,1132,1157]" class="Clitellata" family="Tubificidae" genus="Tubificoides" kingdom="Animalia" order="Haplotaxida" pageId="7" pageNumber="508" phylum="Annelida" rank="species" species="benedii">
<emphasis id="54F64D65FFC00270FEF00D32FDC3FB0B" box="[328,555,1132,1157]" italics="true" pageId="7" pageNumber="508">Tubificoides benedii</emphasis>
Udekem, 1855
</taxonomicName>
, and
<taxonomicName id="A182EAF4FFC00270FCA40D32FB3CFB0B" authority="O.F. Muller, 1776" authorityName="O.F. Muller" authorityYear="1776" box="[796,1236,1132,1157]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="7" pageNumber="508" phylum="Annelida" rank="species" species="armiger">
<emphasis id="54F64D65FFC00270FCA40D32FC03FB0A" box="[796,1003,1132,1157]" italics="true" pageId="7" pageNumber="508">Scoloplos armiger</emphasis>
(
<collectingCountry id="1E95D1E7FFC00270FC440D33FBFAFB0B" box="[1020,1042,1133,1157]" name="Iceland" pageId="7" pageNumber="508">O</collectingCountry>
.F. Müller, 1776)
</taxonomicName>
. The fauna in the eelgrass beds was richer and more diverse in most samples, with a representation of polychaetes and several species of crustaceans and molluscs.
</paragraph>
<paragraph id="663D9177FFC00270FF7F0D86FD3EFA47" blockId="7.[151,1437,151,1517]" pageId="7" pageNumber="508">
Both
<taxonomicName id="A182EAF4FFC00270FEBE0D86FE5AFB7E" box="[262,434,1240,1265]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="7" pageNumber="508" phylum="Annelida" rank="species" species="maryae">
<emphasis id="54F64D65FFC00270FEBE0D86FE5AFB7E" box="[262,434,1240,1265]" italics="true" pageId="7" pageNumber="508">Tharyx maryae</emphasis>
</taxonomicName>
<emphasis id="54F64D65FFC00270FE030D86FE1CFB7F" bold="true" box="[443,500,1240,1265]" pageId="7" pageNumber="508">n. sp</emphasis>
and
<taxonomicName id="A182EAF4FFC00270FD970D87FD43FB7F" box="[559,683,1240,1265]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="7" pageNumber="508" phylum="Annelida" rank="species" species="robustus" status="sp. nov.">
<emphasis id="54F64D65FFC00270FD970D87FD43FB7F" box="[559,683,1240,1265]" italics="true" pageId="7" pageNumber="508">T. robustus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FC5F01EFFC00270FD0C0D86FD1DFB7F" box="[692,757,1240,1265]" pageId="7" pageNumber="508" rank="species">
<emphasis id="54F64D65FFC00270FD0C0D86FD1DFB7F" bold="true" box="[692,757,1240,1265]" pageId="7" pageNumber="508">n. sp.</emphasis>
</taxonomicNameLabel>
(described below) were found in the same sample together with an anterior fragment of
<taxonomicName id="A182EAF4FFC00270FE6F0DA2FD99FA9B" box="[471,625,1276,1301]" class="Polychaeta" family="Cirratulidae" genus="Aphelochaeta" kingdom="Animalia" order="Terebellida" pageId="7" pageNumber="508" phylum="Annelida" rank="genus">
<emphasis id="54F64D65FFC00270FE6F0DA2FD99FA9B" box="[471,625,1276,1301]" italics="true" pageId="7" pageNumber="508">Aphelochaeta</emphasis>
</taxonomicName>
sp., also likely new to science. These specimens were taken in relatively fine sediment with higher organic content outside the eel grass beds where small patches of the algae
<taxonomicName id="A182EAF4FFC00270FAAD0C7FFA8EFAB6" box="[1301,1382,1313,1336]" class="Liliopsida" family="Ruppiaceae" genus="Ruppia" kingdom="Plantae" order="Alismatales" pageId="7" pageNumber="508" phylum="Tracheophyta" rank="genus">
<emphasis id="54F64D65FFC00270FAAD0C7FFA8EFAB6" box="[1301,1382,1313,1336]" italics="true" pageId="7" pageNumber="508">Ruppia</emphasis>
</taxonomicName>
spp. and
<taxonomicName id="A182EAF4FFC00270FF740C1BFD85FAD3" authority="Linnaeus, 1753" authorityName="Linnaeus" authorityYear="1753" box="[204,621,1348,1373]" class="Phaeophyceae" family="Fucaceae" genus="Fucus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fucales" pageId="7" pageNumber="508" phylum="Heterokontophyta" rank="species" species="vesiculosus">
<emphasis id="54F64D65FFC00270FF740C1BFE74FAD3" box="[204,412,1348,1373]" italics="true" pageId="7" pageNumber="508">Fucus vesiculosus</emphasis>
(Linnaeus, 1753)
</taxonomicName>
were found; however, cirratulids were recorded in sediments almost entirely free of vegetation in three of the four samples. These results suggest that
<taxonomicName id="A182EAF4FFC00270FB890C37FB49FA0E" box="[1073,1185,1385,1408]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="7" pageNumber="508" phylum="Annelida" rank="species" species="maryae" status="sp. nov.">
<emphasis id="54F64D65FFC00270FB890C37FB49FA0E" box="[1073,1185,1385,1408]" italics="true" pageId="7" pageNumber="508">T. maryae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FC5F01EFFC00270FB110C36FB00FA0F" box="[1193,1256,1384,1409]" pageId="7" pageNumber="508" rank="species">
<emphasis id="54F64D65FFC00270FB110C36FB00FA0F" bold="true" box="[1193,1256,1384,1409]" pageId="7" pageNumber="508">n. sp.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="A182EAF4FFC00270FA980C37FA73FA0F" box="[1312,1435,1384,1409]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="7" pageNumber="508" phylum="Annelida" rank="species" species="robustus" status="sp. nov.">
<emphasis id="54F64D65FFC00270FA980C37FA73FA0F" box="[1312,1435,1384,1409]" italics="true" pageId="7" pageNumber="508">T. robustus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FC5F01EFFC00270FF2F0CD2FF30FA2B" box="[151,216,1420,1445]" pageId="7" pageNumber="508" rank="species">
<emphasis id="54F64D65FFC00270FF2F0CD2FF30FA2B" bold="true" box="[151,216,1420,1445]" pageId="7" pageNumber="508">n. sp.</emphasis>
</taxonomicNameLabel>
are very tolerant of variations in salinity and temperature and prefer relatively coarse sand with almost no vegetation and low competition from other species.
</paragraph>
</subSubSection>
<subSubSection id="2E98C2FCFFC00270FF7F0C8AFC03FA62" box="[199,1003,1492,1517]" pageId="7" pageNumber="508" type="distribution">
<paragraph id="663D9177FFC00270FF7F0C8AFC03FA62" blockId="7.[151,1437,151,1517]" box="[199,1003,1492,1517]" pageId="7" pageNumber="508">
<emphasis id="54F64D65FFC00270FF7F0C8AFE88FA63" bold="true" box="[199,352,1492,1517]" pageId="7" pageNumber="508">Distribution.</emphasis>
Known only from the west coast of
<collectingCountry id="1E95D1E7FFC00270FD410C8AFCBAFA63" box="[761,850,1492,1517]" name="Sweden" pageId="7" pageNumber="508">Sweden</collectingCountry>
in low water.
</paragraph>
</subSubSection>
</treatment>
</document>