treatments-xml/data/E3/3C/87/E33C87BBFFE8FFF6368CFE5DFA90FCF4.xml
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<document id="C44792924038562D3A00A4743D567F3C" ID-CLB-Dataset="20427" ID-DOI="10.11646/phytotaxa.536.2.7" ID-GBIF-Dataset="c4d41e70-5b14-4264-a41a-ded2bf78026a" ID-ISSN="1179-3163" ID-Zenodo-Dep="6260329" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1645688326578" checkinUser="plazi" docAuthor="Adorador, Jiro T. &amp; Fernando, Edwino S." docDate="2022" docId="E33C87BBFFE8FFF6368CFE5DFA90FCF4" docLanguage="en" docName="phytotaxa.536.2.7.pdf" docOrigin="Phytotaxa 536 (2)" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Pinanga leonardcoi Adorador &amp; Fernando, Mt. Cagua 2022, sp. nov." docType="treatment" docVersion="8" lastPageNumber="188" masterDocId="1F05FFC3FFEAFFF33604FFBFFFCBFF90" masterDocTitle="Pinanga leonardcoi (Arecaceae), a new species from Mt. Cagua in northeast Luzon, Philippines" masterLastPageNumber="189" masterPageNumber="183" pageNumber="185" updateTime="1699316408570" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="58B40047EAF4D94FB8B5120BFF7A084B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E4BC59B8282BB6CB1B68A0A963AC51D4">
<mods:title id="3A1F4442E2EBD9FCD01CC41890F28103">Pinanga leonardcoi (Arecaceae), a new species from Mt. Cagua in northeast Luzon, Philippines</mods:title>
</mods:titleInfo>
<mods:name id="6A20A43ABE42CACCDFBDE27589B34A53" type="personal">
<mods:role id="109C207CCFAD82457C69FAA0BD440C49">
<mods:roleTerm id="42271BACB750C53DCD04A0A772A01E93">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1C3E8310C1C6D993A33693B36335C6A8">Adorador, Jiro T.</mods:namePart>
<mods:nameIdentifier id="14BF3413A79FCDA48B2B661D09039A23" type="ORCID">0000-0002-2860-7643</mods:nameIdentifier>
<mods:affiliation id="CC4302D52ED514D26EC14DB697731679">Institute of Biological Sciences, College of Arts and Sciences, University of the Philippines Los Baños, College, 4031 Laguna, Philippines &amp; jtadorador @ up. edu. ph; https: // orcid. org / 0000 - 0002 - 2860 - 7643</mods:affiliation>
<mods:nameIdentifier id="54FC9D566AD8EF6BD80C377352A4DCB1" type="email">jtadorador@up.edu.ph</mods:nameIdentifier>
</mods:name>
<mods:name id="9639A02197F5EF2BB939A6F9FD164F50" type="personal">
<mods:role id="B2B432879731ED358256BFC55FEB47A6">
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<mods:namePart id="5DB383F554D9DD2C2C5971851FC94FFB">Fernando, Edwino S.</mods:namePart>
<mods:nameIdentifier id="4193C3AFCA1F2BB6043111BDA54241B5" type="ORCID">0000-0001-9422-5451</mods:nameIdentifier>
<mods:affiliation id="CBA0591D32B5B34697B2C8664127A941">Department of Forest Biological Sciences, College of Forestry and Natural Resources, University of the Philippines Los Baños, College, 4031 Laguna, Philippines &amp; Museum of Natural History, University of the Philippines - Los Baños, College, 4031 Laguna, Philippines &amp; Institute of Biology, College of Science, University of the Philippines - Diliman, 1101 Quezon City, Philippines &amp; esfernando @ up. edu. ph; https: // orcid. org / 0000 - 0001 - 9422 - 5451</mods:affiliation>
<mods:nameIdentifier id="D3DE4B295B492A886C4CF7EC38F398DB" type="email">esfernando@up.edu.ph</mods:nameIdentifier>
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<mods:typeOfResource id="F178C76CD14E1E737AE019548AD57ED5">text</mods:typeOfResource>
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<mods:title id="84A3516744AFB2B81ABF11E57859060B">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="002D0D2010E8F2B659CCA5CAFF2E463A">
<mods:date id="A583623BE02982440E78736FE5D4C6B0">2022</mods:date>
<mods:detail id="3C508963B9C554DF6B922267B5B993BC" type="pubDate">
<mods:number id="2B4A3282A6DCE3E5EB19C38648B199FA">2022-02-24</mods:number>
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<mods:number id="F9F53A0B82AA2006EB4BB8974DCCBDAF">536</mods:number>
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<mods:classification id="084570E03FE57423988C64F3EF709F15">journal article</mods:classification>
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<mods:identifier id="359E29EA843FE395AEFD9FDA1B42B278" type="DOI">10.11646/phytotaxa.536.2.7</mods:identifier>
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<treatment id="E33C87BBFFE8FFF6368CFE5DFA90FCF4" ID-DOI="http://doi.org/10.5281/zenodo.6323881" ID-GBIF-Taxon="193852777" ID-Zenodo-Dep="6323881" LSID="urn:lsid:plazi:treatment:E33C87BBFFE8FFF6368CFE5DFA90FCF4" httpUri="http://treatment.plazi.org/id/E33C87BBFFE8FFF6368CFE5DFA90FCF4" lastPageId="5" lastPageNumber="188" pageId="2" pageNumber="185">
<subSubSection id="238F6526FFE8FFF1368CFE5DFD04FE6C" box="[136,719,482,509]" pageId="2" pageNumber="185" type="nomenclature">
<paragraph id="6B2A36ADFFE8FFF1368CFE5DFD04FE6C" blockId="2.[136,719,482,509]" box="[136,719,482,509]" pageId="2" pageNumber="185">
<heading id="306281C1FFE8FFF1368CFE5DFD04FE6C" box="[136,719,482,509]" fontSize="11" level="2" pageId="2" pageNumber="185" reason="3">
<taxonomicName id="AC954D2EFFE8FFF1368CFE5DFDA4FE6C" authority="Adorador &amp; Fernando" authorityName="Adorador &amp; Fernando, Mt. Cagua" authorityYear="2022" box="[136,623,482,509]" class="Liliopsida" family="Arecaceae" genus="Pinanga" kingdom="Plantae" order="Arecales" pageId="2" pageNumber="185" phylum="Tracheophyta" rank="species" species="leonardcoi" status="sp. nov.">
<emphasis id="59E1EABFFFE8FFF1368CFE5DFEA7FE6D" bold="true" box="[136,364,482,509]" italics="true" pageId="2" pageNumber="185">Pinanga leonardcoi</emphasis>
Adorador &amp; Fernando
</taxonomicName>
,
<emphasis id="59E1EABFFFE8FFF1347EFE5CFD04FE6C" box="[634,719,483,508]" italics="true" pageId="2" pageNumber="185">
<taxonomicNameLabel id="42D257C4FFE8FFF1347EFE5CFD04FE6C" box="[634,719,483,508]" pageId="2" pageNumber="185" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="238F6526FFE8FFF1368CFDB5FCD9FDD4" pageId="2" pageNumber="185" type="materials_examined">
<paragraph id="6B2A36ADFFE8FFF1368CFDB5FCD9FDD4" blockId="2.[136,1452,522,580]" pageId="2" pageNumber="185">
<materialsCitation id="DBFD3CF0FFE8FFF1368CFDB5FCD9FDD4" ID-GBIF-Occurrence="3503961302" collectingDate="2021-08-22" collectionCode="PNH, LBC" collectorName="J. T. Adorador" country="Philippines" county="Type" elevation="800" location="Gonzaga" municipality="Luzon Island" pageId="2" pageNumber="185" specimenCount="1" stateProvince="Cagayan" typeStatus="holotype">
<collectingCounty id="824B4E21FFE8FFF1368CFDB5FF72FD8F" box="[136,185,522,543]" pageId="2" pageNumber="185">Type</collectingCounty>
:—
<collectingCountry id="1382763DFFE8FFF136D1FDB5FEADFD8F" box="[213,358,522,543]" name="Philippines" pageId="2" pageNumber="185">PHILIPPINES</collectingCountry>
,
<collectingRegion id="A951F84FFFE8FFF13774FDB5FDE8FDB0" box="[368,547,522,544]" country="Philippines" name="Cagayan" pageId="2" pageNumber="185">Cagayan Province</collectingRegion>
:
<collectingMunicipality id="8B4EACD7FFE8FFF13429FDB5FD65FDB0" box="[557,686,522,544]" pageId="2" pageNumber="185">Luzon Island</collectingMunicipality>
,
<location id="6E4A6076FFE8FFF134BCFDB5FCDBFD8F" LSID="urn:lsid:plazi:treatment:E33C87BBFFE8FFF6368CFE5DFA90FCF4:6E4A6076FFE8FFF134BCFDB5FCDBFD8F" box="[696,784,522,543]" country="Philippines" county="Type" municipality="Luzon Island" name="Gonzaga" pageId="2" pageNumber="185" stateProvince="Cagayan">Gonzaga</location>
,
<location id="6E4A6076FFE8FFF1351DFDB5FC20FDB0" LSID="urn:lsid:plazi:treatment:E33C87BBFFE8FFF6368CFE5DFA90FCF4:6E4A6076FFE8FFF1351DFDB5FC20FDB0" box="[793,1003,522,544]" country="Philippines" county="Type" municipality="Luzon Island" name="Barangay Santa Clara" pageId="2" pageNumber="185" stateProvince="Cagayan">Barangay Santa Clara</location>
,
<location id="6E4A6076FFE8FFF135FAFDB5FBACFD8F" LSID="urn:lsid:plazi:treatment:E33C87BBFFE8FFF6368CFE5DFA90FCF4:6E4A6076FFE8FFF135FAFDB5FBACFD8F" box="[1022,1127,522,543]" country="Philippines" county="Type" municipality="Luzon Island" name="Mt. Cagua" pageId="2" pageNumber="185" stateProvince="Cagayan">Mt. Cagua</location>
, ±
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<elevation id="E0B8D19EFFE8FFF13280FDB5FB0AFD8F" box="[1156,1217,522,543]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="185" unit="m" value="800.0">800 m</elevation>
</quantity>
elev.,
<date id="1F2B106DFFE8FFF13301FDB5FA6CFD8F" box="[1285,1447,522,543]" pageId="2" pageNumber="185" value="2021-08-22">
<collectingDate id="0F6FE985FFE8FFF13301FDB5FA6CFD8F" box="[1285,1447,522,543]" pageId="2" pageNumber="185" value="2021-08-22">August 22, 2021</collectingDate>
</date>
,
<emphasis id="59E1EABFFFE8FFF136B9FD91FEA6FDD3" box="[189,365,558,579]" italics="true" pageId="2" pageNumber="185">
<collectorName id="C660537BFFE8FFF136B9FD91FE88FDD3" box="[189,323,558,579]" pageId="2" pageNumber="185">J.T. Adorador</collectorName>
201
</emphasis>
(
<typeStatus id="B42E880FFFE8FFF1377DFD91FE04FDD4" box="[377,463,558,580]" pageId="2" pageNumber="185" type="holotype">holotype</typeStatus>
<collectionCode id="0D84AE68FFE8FFF137D1FD91FDC6FDD3" box="[469,525,558,579]" country="Philippines" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15314" lsid="urn:lsid:biocol.org:col:15314" name="National Museum" pageId="2" pageNumber="185" type="Herbarium">PNH</collectionCode>
!; isotypes CAHUP!,
<collectionCode id="0D84AE68FFE8FFF134D0FD91FCC3FDD4" box="[724,776,558,580]" country="Philippines" lsid="urn:lsid:biocol.org:col:14633" name="University of the Philippines at Los Baños" pageId="2" pageNumber="185" type="Herbarium">LBC</collectionCode>
!)
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="238F6526FFE8FFF1368CFDCDFBDAFA84" pageId="2" pageNumber="185" type="diagnosis">
<paragraph id="6B2A36ADFFE8FFF1368CFDCDFA9EFD44" blockId="2.[136,1452,626,2093]" pageId="2" pageNumber="185">
<emphasis id="59E1EABFFFE8FFF1368CFDCDFECEFD1C" bold="true" box="[136,261,626,652]" pageId="2" pageNumber="185">Diagnosis:</emphasis>
<emphasis id="59E1EABFFFE8FFF13701FDCCFE7BFD1C" box="[261,432,626,652]" italics="true" pageId="2" pageNumber="185">
<taxonomicName id="AC954D2EFFE8FFF1371BFDCCFE7BFD1C" authorityName="Adorador &amp; Fernando, Mt. Cagua" authorityYear="2022" box="[287,432,626,652]" class="Liliopsida" family="Arecaceae" genus="Pinanga" kingdom="Plantae" order="Arecales" pageId="2" pageNumber="185" phylum="Tracheophyta" rank="species" species="leonardcoi">P. leonardcoi</taxonomicName>
</emphasis>
is closely related to members of the Urosperma group, namely,
<taxonomicName id="AC954D2EFFE8FFF13271FDCCFAC4FD1C" authorityName="Fernando" authorityYear="1994" box="[1141,1295,626,653]" class="Liliopsida" family="Arecaceae" genus="Pinanga" kingdom="Plantae" order="Arecales" pageId="2" pageNumber="185" phylum="Tracheophyta" rank="species" species="glaucifolia">
<emphasis id="59E1EABFFFE8FFF13271FDCCFB4DFD1C" box="[1141,1158,627,652]" italics="true" pageId="2" pageNumber="185">P</emphasis>
.
<emphasis id="59E1EABFFFE8FFF13297FDCDFAC4FD1C" box="[1171,1295,626,652]" italics="true" pageId="2" pageNumber="185">glaucifolia</emphasis>
</taxonomicName>
<emphasis id="59E1EABFFFE8FFF1330BFDCDFA60FD1C" box="[1295,1451,626,652]" italics="true" pageId="2" pageNumber="185">
,
<taxonomicName id="AC954D2EFFE8FFF1331EFDCCFA6CFD1C" authorityName="Beccari" authorityYear="1919" box="[1306,1447,627,652]" class="Liliopsida" family="Arecaceae" genus="Pinanga" kingdom="Plantae" order="Arecales" pageId="2" pageNumber="185" phylum="Tracheophyta" rank="species" species="samarana">P. samarana</taxonomicName>
,
</emphasis>
and
<taxonomicName id="AC954D2EFFE8FFF136BDFD28FE85FD20" authorityName="Beccari" authorityYear="1909" box="[185,334,663,688]" class="Liliopsida" family="Arecaceae" genus="Pinanga" kingdom="Plantae" order="Arecales" pageId="2" pageNumber="185" phylum="Tracheophyta" rank="species" species="urosperma">
<emphasis id="59E1EABFFFE8FFF136BDFD28FE85FD20" box="[185,334,663,688]" italics="true" pageId="2" pageNumber="185">P. urosperma</emphasis>
</taxonomicName>
but differs by the combination of the following characters: clustered stems with stoloniferous offshoots, very few rachillae (23) with spirally arranged fruits, and fruiting perianth with imbricate calyx lobes.
</paragraph>
<paragraph id="6B2A36ADFFE8FFF136B9FD61FBDAFA84" blockId="2.[136,1452,626,2093]" pageId="2" pageNumber="185">
Clustering moderately slender undergrowth palm, to
<quantity id="AC6D9B48FFE8FFF1351FFD61FC83FD69" box="[795,840,734,761]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="185" unit="m" value="4.0">4 m</quantity>
tall.
<emphasis id="59E1EABFFFE8FFF13581FD61FC09FD68" bold="true" box="[901,962,734,760]" pageId="2" pageNumber="185">Stem</emphasis>
to
<quantity id="AC6D9B48FFE8FFF135EEFD61FB9FFD69" box="[1002,1108,734,761]" metricMagnitude="-2" metricUnit="m" metricValue="3.75" metricValueMax="4.0" metricValueMin="3.5" pageId="2" pageNumber="185" unit="cm" value="3.75" valueMax="4.0" valueMin="3.5">3.54 cm</quantity>
diam., internodes about
<quantity id="AC6D9B48FFE8FFF13376FD60FA60FD69" box="[1394,1451,735,761]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="185" unit="cm" value="7.0">7 cm</quantity>
apart.
<emphasis id="59E1EABFFFE8FFF136CBFCBDFE97FC8C" bold="true" box="[207,348,770,796]" pageId="2" pageNumber="185">Crownshaft</emphasis>
elongate c.
<quantity id="AC6D9B48FFE8FFF137E1FCBDFDE1FC8D" box="[485,554,770,797]" metricMagnitude="-1" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="185" unit="cm" value="40.0">40 cm</quantity>
long, cylindrical, slightly swollen.
<emphasis id="59E1EABFFFE8FFF135BEFCBDFBC7FC8C" bold="true" box="[954,1036,770,796]" pageId="2" pageNumber="185">Leaves</emphasis>
to
<quantity id="AC6D9B48FFE8FFF13235FCBDFB96FC8C" box="[1073,1117,770,796]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="2" pageNumber="185" unit="in" value="6.0">6 in</quantity>
crown, sheath 2535 ×
<quantity id="AC6D9B48FFE8FFF13368FCBDFF66FCD1" metricMagnitude="-2" metricUnit="m" metricValue="4.25" metricValueMax="4.5" metricValueMin="4.0" pageId="2" pageNumber="185" unit="cm" value="4.25" valueMax="4.5" valueMin="4.0">44.5 cm</quantity>
, yellowish green, glaucescent, ligules narrowly triangular, inconspicuous, leaf (including petiole) without sheath
<quantity id="AC6D9B48FFE8FFF1368CFCF5FED9FCF5" box="[136,274,842,869]" metricMagnitude="0" metricUnit="m" metricValue="1.5699999999999998" metricValueMax="1.91" metricValueMin="1.23" pageId="2" pageNumber="185" unit="cm" value="157.0" valueMax="191.0" valueMin="123.0">123191 cm</quantity>
long, petiole 2544 ×
<quantity id="AC6D9B48FFE8FFF13409FCF5FD46FCF5" box="[525,653,842,869]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="2" pageNumber="185" unit="cm" value="1.35" valueMax="1.5" valueMin="1.2">1.21.5 cm</quantity>
, shallowly channeled adaxially, convex abaxially sparsely covered with brown scales, rachis angular, 69153 ×
<quantity id="AC6D9B48FFE8FFF1344EFCD1FD4EFC19" box="[586,645,878,905]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="185" unit="cm" value="1.0">1 cm</quantity>
, bifacial adaxially and rounded abaxially with indumentums as petiole. Leaflets 813 per side of the rachis, coriaceous, ± straight, generally multi-costulate except occasionally for basal leaflets, ± regularly arranged,
<quantity id="AC6D9B48FFE8FFF137D8FC09FD92FC41" box="[476,601,950,977]" metricMagnitude="-2" metricUnit="m" metricValue="5.4" metricValueMax="6.5" metricValueMin="4.3" pageId="2" pageNumber="185" unit="cm" value="5.4" valueMax="6.5" valueMin="4.3">4.36.5 cm</quantity>
apart, the apex long acuminate to subfalcate, glossy green adaxially drying grayish, much paler green and glabrescent abaxially drying brownish, with midfixed ramenta along the midrib to
<quantity id="AC6D9B48FFE8FFF13368FC65FA67FC65" box="[1388,1452,986,1013]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="185" unit="mm" value="4.0">4 mm</quantity>
long; basal leaflets 14 costulate, 3859 × 21.5
<quantity id="AC6D9B48FFE8FFF134A0FC41FD39FB89" box="[676,754,1022,1049]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" pageId="2" pageNumber="185" unit="cm" value="5.5">5.5 cm</quantity>
, middle leaflets 23-costulate, 52.579 × 3.56, apical leaflets 411 costulate, 2641 ×
<quantity id="AC6D9B48FFE8FFF137A3FB9DFDE9FBAD" box="[423,546,1058,1085]" metricMagnitude="-2" metricUnit="m" metricValue="6.75" metricValueMax="11.5" metricValueMin="2.0" pageId="2" pageNumber="185" unit="cm" value="6.75" valueMax="11.5" valueMin="2.0">211.5 cm</quantity>
, joined to
<quantity id="AC6D9B48FFE8FFF1349BFB9DFD2CFBAD" box="[671,743,1058,1085]" metricMagnitude="-1" metricUnit="m" metricValue="1.6" pageId="2" pageNumber="185" unit="cm" value="16.0">16 cm</quantity>
at the base along the rachis.
<emphasis id="59E1EABFFFE8FFF1323BFB9DFB26FBAC" bold="true" box="[1087,1261,1058,1084]" pageId="2" pageNumber="185">Infructescence</emphasis>
<quantity id="AC6D9B48FFE8FFF132F3FB9DFAA3FBAD" box="[1271,1384,1058,1085]" metricMagnitude="-1" metricUnit="m" metricValue="2.25" metricValueMax="2.9" metricValueMin="1.6" pageId="2" pageNumber="185" unit="cm" value="22.5" valueMax="29.0" valueMin="16.0">1629 cm</quantity>
long, the axes greenish yellow, prophyll unknown, peduncle 3.74 ×
<quantity id="AC6D9B48FFE8FFF13553FBF9FC6CFBF1" box="[855,935,1094,1121]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="2" pageNumber="185" unit="cm" value="1.4">1.4 cm</quantity>
, with just 23 ultimately pendulous rachillae, 1319.5 ×
<quantity id="AC6D9B48FFE8FFF136FBFBD5FE85FB15" box="[255,334,1130,1157]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="185" unit="cm" value="0.7">0.7 cm</quantity>
, rachillae bracts very shallow gibbous; flowers spirally arranged throughout the rachillae.
<emphasis id="59E1EABFFFE8FFF13336FBD5FA60FB14" bold="true" box="[1330,1451,1130,1156]" pageId="2" pageNumber="185">Staminate</emphasis>
and
<emphasis id="59E1EABFFFE8FFF136BEFB31FE4AFB38" bold="true" box="[186,385,1166,1192]" pageId="2" pageNumber="185">pistillate flowers</emphasis>
not known.
<emphasis id="59E1EABFFFE8FFF13410FB31FD99FB38" bold="true" box="[532,594,1166,1192]" pageId="2" pageNumber="185">Fruit</emphasis>
broad ovoid at maturity, up to 23 ×
<quantity id="AC6D9B48FFE8FFF1320DFB31FB40FB39" box="[1033,1163,1166,1193]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" metricValueMax="2.6" metricValueMin="1.5" pageId="2" pageNumber="185" unit="cm" value="2.05" valueMax="2.6" valueMin="1.5">1.52.6 cm</quantity>
, with short conical apex, spirally arranged, ripening greenish red, up to 37 fruits along rachis, fruiting perianth 2.53 ×
<quantity id="AC6D9B48FFE8FFF132C6FB0DFAE8FB5D" box="[1218,1315,1202,1229]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="2" pageNumber="185" unit="mm" value="6.5" valueMax="7.0" valueMin="6.0">67 mm</quantity>
, with broad imbricate calyx lobes, not contracted at the mouth.
<emphasis id="59E1EABFFFE8FFF134C8FB69FCCFFB60" bold="true" box="[716,772,1238,1264]" pageId="2" pageNumber="185">Seed</emphasis>
ovoid, to 2.1 ×
<quantity id="AC6D9B48FFE8FFF135BEFB69FBC2FB61" box="[954,1033,1238,1265]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="2" pageNumber="185" unit="cm" value="1.7">1.7 cm</quantity>
, rounded apically, obliquely concave below, endosperm ruminate, embryo short conical to
<quantity id="AC6D9B48FFE8FFF134D8FB45FCD6FA85" box="[732,797,1274,1301]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="185" unit="mm" value="3.0">3 mm</quantity>
long. (
<figureCitation id="F3AE2A28FFE8FFF1356FFB45FC1DFA84" box="[875,982,1274,1300]" captionStart="FIGURE 2" captionStartId="3.[136,229,1849,1871]" captionTargetBox="[182,1398,213,1795]" captionTargetId="figure-17@3.[151,1436,183,1832]" captionTargetPageId="3" captionText="FIGURE 2. Pinanga leonardcoi Adorador &amp; Fernando, clustering habit (A), leaves with 23-costulate segments (B), stem with infructescences (C) in different phases of development (DF), Close-up of fruits showing imbricate fruiting perianth (G), longitudinally dissected fruits showing ruminate endosperm and basal embryo (H), basal off-shoot with a stoloniferous stem (I). All from the holotype, J.T. Adorador 201 (PNH) except F from J.P. Bautista 902 (PUH). Scale bar: B—40 cm, C—15 cm, D &amp; E—15 cm, G—40 cm, I—8 cm. Photos by J.T. Adorador." figureDoi="http://doi.org/10.5281/zenodo.6260339" httpUri="https://zenodo.org/record/6260339/files/figure.png" pageId="2" pageNumber="185">Figures 2</figureCitation>
&amp;
<figureCitation id="F3AE2A28FFE8FFF135FEFB45FBC3FA84" box="[1018,1032,1274,1300]" captionStart="FIGURE 3" captionStartId="4.[136,229,1935,1957]" captionTargetBox="[178,1401,195,1905]" captionTargetId="figure-23@4.[172,1416,180,1919]" captionTargetPageId="4" captionText="FIGURE 3. Pinanga leonardcoi Adorador &amp; Fernando, clustering habit (A), middle leaflets with 3-costulate segments (B), apical leaflets (C), immature fruit showing imbricate fruiting perianth (D), infructescence showing spirally arranged fruits (E), longitudinally dissected fruits showing ruminate endosperm and basal embryo (F), seed with adherent portions of the pericarp (G), and basal of-shoot with a stoloniferous stem (H). BC from J.T. Adorador 202, DH from the holotype, J.T. Adorador 201.Scale bar: B &amp; C—6 cm, D—1 cm, E—7.5 cm, F—2 cm, G—1.8 cm, H—8 cm. Drawn by J.T. Adorador" figureDoi="http://doi.org/10.5281/zenodo.6260343" httpUri="https://zenodo.org/record/6260343/files/figure.png" pageId="2" pageNumber="185">3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="238F6526FFE8FFF136B9FAA1FEFEF89C" pageId="2" pageNumber="185" type="distribution">
<paragraph id="6B2A36ADFFE8FFF136B9FAA1FE3AFA10" blockId="2.[136,1452,626,2093]" pageId="2" pageNumber="185">
<emphasis id="59E1EABFFFE8FFF136B9FAA1FD90FAA8" bold="true" box="[189,603,1310,1336]" pageId="2" pageNumber="185">DISTRIBUTION AND HABITAT:</emphasis>
—There are just about eight isolated individuals encountered in the general area of occurrence. These accounted clusters are all situated inside the caldera of Mt. Cagua where it prefers welldrained gentle slopes or valleys.
</paragraph>
<paragraph id="6B2A36ADFFE8FFF136B9FA35FBFDFA59" blockId="2.[136,1452,626,2093]" pageId="2" pageNumber="185">
<emphasis id="59E1EABFFFE8FFF136B9FA35FE6BFA34" bold="true" box="[189,416,1418,1444]" pageId="2" pageNumber="185">LOCAL NAME:—</emphasis>
None recorded. With most common names of Philippine
<taxonomicName id="AC954D2EFFE8FFF13212FA34FBBEFA34" box="[1046,1141,1419,1444]" class="Liliopsida" family="Arecaceae" genus="Pinanga" kingdom="Plantae" order="Arecales" pageId="2" pageNumber="185" phylum="Tracheophyta" rank="genus">
<emphasis id="59E1EABFFFE8FFF13212FA34FBBEFA34" box="[1046,1141,1419,1444]" italics="true" pageId="2" pageNumber="185">Pinanga</emphasis>
</taxonomicName>
being referred as or derived from
<emphasis id="59E1EABFFFE8FFF136CBFA11FECCFA58" box="[207,263,1454,1480]" italics="true" pageId="2" pageNumber="185">abiki</emphasis>
, we hereby formalize this new species to be called as
<emphasis id="59E1EABFFFE8FFF1357CFA11FBE5FA58" box="[888,1070,1454,1480]" italics="true" pageId="2" pageNumber="185">Leonards abiki</emphasis>
.
</paragraph>
<paragraph id="6B2A36ADFFE8FFF136B9FA6DFAC4F9A4" blockId="2.[136,1452,626,2093]" pageId="2" pageNumber="185">
<emphasis id="59E1EABFFFE8FFF136B9FA6DFD53FA7C" bold="true" box="[189,664,1490,1516]" pageId="2" pageNumber="185">OTHER SPECIMENS EXAMINED:—</emphasis>
<materialsCitation id="DBFD3CF0FFE8FFF13493FA6DFB34F9A4" ID-GBIF-Occurrence="3503961301" collectingDate="2021-08-22" collectionCode="PNH, PUH" collectorName="J. T. Adorador &amp; J. P. Bautista" country="Philippines" county="Luzon Island" elevation="800" location="Barangay Santa Clara" municipality="Gonzaga" pageId="2" pageNumber="185" specimenCount="1" stateProvince="Cagayan">
<collectingCountry id="1382763DFFE8FFF13493FA6DFCF4FA7C" box="[663,831,1490,1516]" name="Philippines" pageId="2" pageNumber="185">PHILIPPINES</collectingCountry>
,
<collectingRegion id="A951F84FFFE8FFF13548FA6DFBD5FA7C" box="[844,1054,1490,1516]" country="Philippines" name="Cagayan" pageId="2" pageNumber="185">Cagayan Province</collectingRegion>
:
<collectingCounty id="824B4E21FFE8FFF1322EFA6CFB0BFA7C" box="[1066,1216,1490,1517]" pageId="2" pageNumber="185">Luzon Island</collectingCounty>
,
<collectingMunicipality id="8B4EACD7FFE8FFF132C9FA6DFAF8FA7C" box="[1229,1331,1490,1516]" pageId="2" pageNumber="185">Gonzaga</collectingMunicipality>
,
<location id="6E4A6076FFE8FFF13344FA6CFEC5F980" LSID="urn:lsid:plazi:treatment:E33C87BBFFE8FFF6368CFE5DFA90FCF4:6E4A6076FFE8FFF13344FA6CFEC5F980" country="Philippines" county="Luzon Island" municipality="Gonzaga" name="Barangay Santa Clara" pageId="2" pageNumber="185" stateProvince="Cagayan">Barangay Santa Clara</location>
,
<location id="6E4A6076FFE8FFF13718FA48FE5CF980" LSID="urn:lsid:plazi:treatment:E33C87BBFFE8FFF6368CFE5DFA90FCF4:6E4A6076FFE8FFF13718FA48FE5CF980" box="[284,407,1526,1553]" country="Philippines" county="Luzon Island" municipality="Gonzaga" name="Mt. Cagua" pageId="2" pageNumber="185" stateProvince="Cagayan">Mt. Cagua</location>
, ±
<quantity id="AC6D9B48FFE8FFF137B9FA49FDCDF981" box="[445,518,1526,1553]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="185" unit="m" value="800.0">
<elevation id="E0B8D19EFFE8FFF137B9FA49FDCDF981" box="[445,518,1526,1553]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="185" unit="m" value="800.0">800 m</elevation>
</quantity>
elev.,
<date id="1F2B106DFFE8FFF13455FA49FCD9F980" box="[593,786,1526,1552]" pageId="2" pageNumber="185" value="2021-08-22">
<collectingDate id="0F6FE985FFE8FFF13455FA49FCD9F980" box="[593,786,1526,1552]" pageId="2" pageNumber="185" value="2021-08-22">August 22, 2021</collectingDate>
</date>
,
<emphasis id="59E1EABFFFE8FFF1351BFA48FC24F980" box="[799,1007,1526,1552]" italics="true" pageId="2" pageNumber="185">
<collectorName id="C660537BFFE8FFF1351BFA48FC77F980" box="[799,956,1526,1552]" pageId="2" pageNumber="185">J.T. Adorador</collectorName>
202
</emphasis>
(LBC!; K!,
<collectionCode id="0D84AE68FFE8FFF13284FA49FB0AF980" box="[1152,1217,1526,1552]" country="Philippines" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15314" lsid="urn:lsid:biocol.org:col:15314" name="National Museum" pageId="2" pageNumber="185" type="Herbarium">PNH</collectionCode>
!),
<emphasis id="59E1EABFFFE8FFF132DFFA48FA60F980" box="[1243,1451,1526,1552]" italics="true" pageId="2" pageNumber="185">
<collectorName id="C660537BFFE8FFF132DFFA48FAB3F980" box="[1243,1400,1526,1552]" pageId="2" pageNumber="185">J.T. Adorador</collectorName>
203
</emphasis>
(LBC!:
<collectionCode id="0D84AE68FFE8FFF136E4F9A4FED6F9A5" box="[224,285,1563,1589]" country="Philippines" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15314" lsid="urn:lsid:biocol.org:col:15314" name="National Museum" pageId="2" pageNumber="185" type="Herbarium">PNH</collectionCode>
); [possibly from Barangay Magrafil], 19901995,
<emphasis id="59E1EABFFFE8FFF13557F9A4FBDBF9A4" box="[851,1040,1562,1588]" italics="true" pageId="2" pageNumber="185">
<collectorName id="C660537BFFE8FFF13557F9A4FC14F9A4" box="[851,991,1563,1588]" pageId="2" pageNumber="185">J.P. Bautista</collectorName>
902
</emphasis>
[PUH017387]
<collectionCode id="0D84AE68FFE8FFF132B9F9A5FB34F9A4" box="[1213,1279,1562,1588]" country="Philippines" lsid="urn:lsid:biocol.org:col:14636" name="University of the Philippines" pageId="2" pageNumber="185" type="Herbarium">PUH</collectionCode>
</materialsCitation>
!).
</paragraph>
<paragraph id="6B2A36ADFFE8FFF136B9F981FEFEF89C" blockId="2.[136,1452,626,2093]" pageId="2" pageNumber="185">
<emphasis id="59E1EABFFFE8FFF136B9F981FDF9F9C8" bold="true" box="[189,562,1598,1624]" pageId="2" pageNumber="185">CONSERVATION STATUS:—</emphasis>
Using the online GeoCAT conservation assessment tool (
<bibRefCitation id="0F044B5CFFE8FFF132BCF981FA54F9C8" author="Bachman, S. &amp; Moat, J. &amp; Hill, A. &amp; de la Torre, J. &amp; Scott, B." box="[1208,1439,1598,1625]" pageId="2" pageNumber="185" pagination="117 - 126" refId="ref3335" refString="Bachman, S., Moat, J., Hill, A., de la Torre, J. &amp; Scott, B. (2011) Supporting Red List threat assessment with GeoCAT: Geospatial conservation assessment tool. Zookeys 150: 117 - 126. https: // doi. org / 10.3897 / zookeys. 150.2109" type="journal article" year="2011">Bachman et al 2011</bibRefCitation>
), the new species AOO (area of occupancy) is calculated at
<quantity id="AC6D9B48FFE8FFF13544F9DDFCB6F9EC" box="[832,893,1634,1660]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="185" unit="km" value="4.0">4 km</quantity>
<superScript id="9CE09BE5FFE8FFF13578F9DDFC4FF9E0" attach="left" box="[892,900,1634,1648]" fontSize="6" pageId="2" pageNumber="185">2</superScript>
which is within the threshold for the Critically Endangered category (
<bibRefCitation id="0F044B5CFFE8FFF13785F939FDC1F930" author="IUCN" box="[385,522,1670,1696]" pageId="2" pageNumber="185" refId="ref3930" refString="IUCN (2012) IUCN Red List Categories and Criteria. Version 3.1, (2 nd edn). Gland and Cambridge, 32 pp." type="book" year="2012">IUCN 2012</bibRefCitation>
). Following
<bibRefCitation id="0F044B5CFFE8FFF13490F939FCE1F930" author="IUCN" box="[660,810,1670,1696]" pageId="2" pageNumber="185" refId="ref3930" refString="IUCN (2012) IUCN Red List Categories and Criteria. Version 3.1, (2 nd edn). Gland and Cambridge, 32 pp." type="book" year="2012">IUCN (2012)</bibRefCitation>
and
<bibRefCitation id="0F044B5CFFE8FFF13559F939FA6CF930" author="IUCN Standards &amp; Petitions Subcommittee" box="[861,1447,1670,1696]" pageId="2" pageNumber="185" refId="ref3957" refString="IUCN Standards and Petitions Subcommittee (2019) Guidelines for Using the IUCN Red List Categories and Criteria, Version 14. Prepared by the Standards and Petitions Subcommittee. Available from: http: // www. iucnredlist. org / documents / RedListGuidelines. pdf (accessed 6 October 2021)" type="url" year="2019">IUCN Standards and Petitions Subcommittee (2019)</bibRefCitation>
, we regard this species as Critically Endangered [CR B2 a b(v), D]. The new species is only recorded within the forested crater of Mt. Cagua with just eight surveyed mature individuals whose extinction is highly probable in the advent of a major eruption.
</paragraph>
</subSubSection>
<subSubSection id="238F6526FFE8FFF636B9F8A9FA90FCF4" lastPageId="5" lastPageNumber="188" pageId="2" pageNumber="185" type="etymology">
<paragraph id="6B2A36ADFFE8FFF136B9F8A9FB3DF80C" blockId="2.[136,1452,626,2093]" pageId="2" pageNumber="185">
<emphasis id="59E1EABFFFE8FFF136B9F8A9FE56F8A0" bold="true" box="[189,413,1814,1840]" pageId="2" pageNumber="185">ETYMOLOGY:—</emphasis>
This slender palm is named after the late Leonardo L. Co (also fondly called as just Leonard) of the Institute of Biology, College of Sciences, University of the
<collectingCountry id="1382763DFFE8FFF13565F885FC14F8C4" box="[865,991,1850,1876]" name="Philippines" pageId="2" pageNumber="185">Philippines</collectingCountry>
Diliman in honor of his selfless devotion to the advancement of understanding, exploration, and conservation of Philippine native plants. Leonard Co was lead botanist of the team that collected the first herbarium specimen of this new
<taxonomicName id="AC954D2EFFE8FFF135D1F83CFBFFF80C" box="[981,1076,1923,1948]" class="Liliopsida" family="Arecaceae" genus="Pinanga" kingdom="Plantae" order="Arecales" pageId="2" pageNumber="185" phylum="Tracheophyta" rank="genus">
<emphasis id="59E1EABFFFE8FFF135D1F83CFBFFF80C" box="[981,1076,1923,1948]" italics="true" pageId="2" pageNumber="185">Pinanga</emphasis>
</taxonomicName>
from Mt. Cagua.
</paragraph>
<paragraph id="6B2A36ADFFE8FFF636B9F819FD47FEFC" blockId="2.[136,1452,626,2093]" lastBlockId="5.[136,1452,158,869]" lastPageId="5" lastPageNumber="188" pageId="2" pageNumber="185">
<emphasis id="59E1EABFFFE8FFF136B9F819FE8AF850" bold="true" box="[189,321,1958,1984]" pageId="2" pageNumber="185">NOTES:—</emphasis>
<taxonomicName id="AC954D2EFFE8FFF13746F818FDEFF850" authorityName="Adorador &amp; Fernando, Mt. Cagua" authorityYear="2022" box="[322,548,1958,1984]" class="Liliopsida" family="Arecaceae" genus="Pinanga" kingdom="Plantae" order="Arecales" pageId="2" pageNumber="185" phylum="Tracheophyta" rank="species" species="leonardcoi">
<emphasis id="59E1EABFFFE8FFF13746F818FDEFF850" box="[322,548,1958,1984]" italics="true" pageId="2" pageNumber="185">Pinanga leonardcoi</emphasis>
</taxonomicName>
shares the fruiting perianth of imbricate calyx lobes with the more southern representatives,
<taxonomicName id="AC954D2EFFE8FFF13745F874FE13F874" authorityName="Fernando" authorityYear="1994" box="[321,472,1994,2020]" class="Liliopsida" family="Arecaceae" genus="Pinanga" kingdom="Plantae" order="Arecales" pageId="2" pageNumber="185" phylum="Tracheophyta" rank="species" species="glaucifolia">
<emphasis id="59E1EABFFFE8FFF13745F874FE13F874" box="[321,472,1994,2020]" italics="true" pageId="2" pageNumber="185">P. glaucifolia</emphasis>
</taxonomicName>
(of Bicol peninsula) and
<taxonomicName id="AC954D2EFFE8FFF13500F874FC58F874" authorityName="Beccari" authorityYear="1919" box="[772,915,1995,2020]" class="Liliopsida" family="Arecaceae" genus="Pinanga" kingdom="Plantae" order="Arecales" pageId="2" pageNumber="185" phylum="Tracheophyta" rank="species" species="samarana">
<emphasis id="59E1EABFFFE8FFF13500F874FC58F874" box="[772,915,1995,2020]" italics="true" pageId="2" pageNumber="185">P. samarana</emphasis>
</taxonomicName>
(of
<collectingRegion id="A951F84FFFE8FFF135C3F875FBC4F874" box="[967,1039,1994,2020]" country="Philippines" name="Samar" pageId="2" pageNumber="185">Samar</collectingRegion>
and
<collectingRegion id="A951F84FFFE8FFF1324FF875FB34F875" box="[1099,1279,1994,2021]" country="Philippines" name="Dinagat Islands" pageId="2" pageNumber="185">Dinagat Islands</collectingRegion>
) but these two species significantly differ in their solitary habit, having different abaxial leaf surface (glaucescent in
<taxonomicName id="AC954D2EFFE8FFF132EDF850FAB6F798" authorityName="Fernando" authorityYear="1994" box="[1257,1405,2030,2056]" class="Liliopsida" family="Arecaceae" genus="Pinanga" kingdom="Plantae" order="Arecales" pageId="2" pageNumber="185" phylum="Tracheophyta" rank="species" species="glaucifolia">
<emphasis id="59E1EABFFFE8FFF132EDF850FAB6F798" box="[1257,1405,2030,2056]" italics="true" pageId="2" pageNumber="185">P. glaucifolia</emphasis>
</taxonomicName>
and microlepidote in
<taxonomicName id="AC954D2EFFE8FFF13743F7ACFE19F7BC" authorityName="Beccari" authorityYear="1919" box="[327,466,2067,2092]" class="Liliopsida" family="Arecaceae" genus="Pinanga" kingdom="Plantae" order="Arecales" pageId="2" pageNumber="185" phylum="Tracheophyta" rank="species" species="samarana">
<emphasis id="59E1EABFFFE8FFF13743F7ACFE19F7BC" box="[327,466,2067,2092]" italics="true" pageId="2" pageNumber="185">P. samarana</emphasis>
</taxonomicName>
vs. glabrescent in
<taxonomicName id="AC954D2EFFE8FFF134A6F7ACFCFDF7BC" authorityName="Adorador &amp; Fernando, Mt. Cagua" authorityYear="2022" box="[674,822,2066,2092]" class="Liliopsida" family="Arecaceae" genus="Pinanga" kingdom="Plantae" order="Arecales" pageId="2" pageNumber="185" phylum="Tracheophyta" rank="species" species="leonardcoi">
<emphasis id="59E1EABFFFE8FFF134A6F7ACFCFDF7BC" box="[674,822,2066,2092]" italics="true" pageId="2" pageNumber="185">P. leonardcoi</emphasis>
</taxonomicName>
), greater number of rachillae (
<quantity id="AC6D9B48FFE8FFF1328FF7ADFB2AF7BC" box="[1163,1249,2066,2092]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" metricValueMax="2.54" metricValueMin="1.524" pageId="2" pageNumber="185" unit="in" value="8.0" valueMax="10.0" valueMin="6.0">610 in</quantity>
<taxonomicName id="AC954D2EFFE8FFF132ECF7ACFAB7F7BC" authorityName="Fernando" authorityYear="1994" box="[1256,1404,2066,2092]" class="Liliopsida" family="Arecaceae" genus="Pinanga" kingdom="Plantae" order="Arecales" pageId="2" pageNumber="185" phylum="Tracheophyta" rank="species" species="glaucifolia">
<emphasis id="59E1EABFFFE8FFF132ECF7ACFAB7F7BC" box="[1256,1404,2066,2092]" italics="true" pageId="2" pageNumber="185">P. glaucifolia</emphasis>
</taxonomicName>
and
<quantity id="AC6D9B48FFEFFFF6368CFF21FF20FF28" box="[136,235,158,184]" metricMagnitude="-1" metricUnit="m" metricValue="3.937" metricValueMax="5.334" metricValueMin="2.54" pageId="5" pageNumber="188" unit="in" value="15.5" valueMax="21.0" valueMin="10.0">1021 in</quantity>
<taxonomicName id="AC954D2EFFEFFFF636F6FF20FEB5FF28" authorityName="Beccari" authorityYear="1919" box="[242,382,159,184]" class="Liliopsida" family="Arecaceae" genus="Pinanga" kingdom="Plantae" order="Arecales" pageId="5" pageNumber="188" phylum="Tracheophyta" rank="species" species="samarana">
<emphasis id="59E1EABFFFEFFFF636F6FF20FEB5FF28" box="[242,382,159,184]" italics="true" pageId="5" pageNumber="188">P. samarana</emphasis>
</taxonomicName>
vs just
<quantity id="AC6D9B48FFEFFFF637D1FF21FDD7FF28" box="[469,540,158,184]" metricMagnitude="-2" metricUnit="m" metricValue="6.35" metricValueMax="7.62" metricValueMin="5.08" pageId="5" pageNumber="188" unit="in" value="2.5" valueMax="3.0" valueMin="2.0">23 in</quantity>
<taxonomicName id="AC954D2EFFEFFFF63427FF20FD7DFF28" authorityName="Adorador &amp; Fernando, Mt. Cagua" authorityYear="2022" box="[547,694,158,184]" class="Liliopsida" family="Arecaceae" genus="Pinanga" kingdom="Plantae" order="Arecales" pageId="5" pageNumber="188" phylum="Tracheophyta" rank="species" species="leonardcoi">
<emphasis id="59E1EABFFFEFFFF63427FF20FD7DFF28" box="[547,694,158,184]" italics="true" pageId="5" pageNumber="188">P. leonardcoi</emphasis>
</taxonomicName>
), and distichously arranged [sometimes seemingly becoming spiral in distal parts of rachillae due to mutual pressure upon fruit maturation] relatively smaller fruits, up to
<quantity id="AC6D9B48FFEFFFF6332AFF7DFAADFF4D" box="[1326,1382,194,221]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="188" unit="cm" value="2.0">2 cm</quantity>
×
<quantity id="AC6D9B48FFEFFFF6338CFF7DFF61FE91" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="5" pageNumber="188" unit="cm" value="1.8">1.8 cm</quantity>
(vs spirally arranged larger fruits, up to 3 ×
<quantity id="AC6D9B48FFEFFFF634B3FF59FD32FE90" box="[695,761,230,256]" metricMagnitude="-2" metricUnit="m" metricValue="6.604" pageId="5" pageNumber="188" unit="in" value="2.6">2.6 in</quantity>
<taxonomicName id="AC954D2EFFEFFFF63507FF58FC52FE90" authorityName="Adorador &amp; Fernando, Mt. Cagua" authorityYear="2022" box="[771,921,230,256]" class="Liliopsida" family="Arecaceae" genus="Pinanga" kingdom="Plantae" order="Arecales" pageId="5" pageNumber="188" phylum="Tracheophyta" rank="species" species="leonardcoi">
<emphasis id="59E1EABFFFEFFFF63507FF58FC52FE90" box="[771,921,230,256]" italics="true" pageId="5" pageNumber="188">P. leonardcoi</emphasis>
</taxonomicName>
). Meanwhile, the geographically sympatric
<taxonomicName id="AC954D2EFFEFFFF63393FF58FECBFEB4" authorityName="Beccari" authorityYear="1909" class="Liliopsida" family="Arecaceae" genus="Pinanga" kingdom="Plantae" order="Arecales" pageId="5" pageNumber="188" phylum="Tracheophyta" rank="species" species="urosperma">
<emphasis id="59E1EABFFFEFFFF63393FF58FECBFEB4" italics="true" pageId="5" pageNumber="188">P. urosperma</emphasis>
</taxonomicName>
(of northern to northeastern Luzon and immediate islands) differs in its solitary habit, the infructescence with 45 rachillae with distichously arranged fruits, fruit apex being narrow conical (vs. short conical in
<taxonomicName id="AC954D2EFFEFFFF6330BFE90FA69FED8" authorityName="Adorador &amp; Fernando, Mt. Cagua" authorityYear="2022" box="[1295,1442,302,328]" class="Liliopsida" family="Arecaceae" genus="Pinanga" kingdom="Plantae" order="Arecales" pageId="5" pageNumber="188" phylum="Tracheophyta" rank="species" species="leonardcoi">
<emphasis id="59E1EABFFFEFFFF6330BFE90FA69FED8" box="[1295,1442,302,328]" italics="true" pageId="5" pageNumber="188">P. leonardcoi</emphasis>
</taxonomicName>
) and fruiting perianth with valvate calyx lobes.
</paragraph>
<caption id="3FEA6625FFE9FFF0368CF886FEBDF855" ID-DOI="http://doi.org/10.5281/zenodo.6260339" ID-Zenodo-Dep="6260339" httpUri="https://zenodo.org/record/6260339/files/figure.png" pageId="3" pageNumber="186" startId="3.[136,229,1849,1871]" targetBox="[182,1398,213,1795]" targetPageId="3">
<paragraph id="6B2A36ADFFE9FFF0368CF886FEBDF855" blockId="3.[136,1452,1849,1989]" pageId="3" pageNumber="186">
<emphasis id="59E1EABFFFE9FFF0368CF886FEC8F8DF" bold="true" box="[136,259,1849,1871]" pageId="3" pageNumber="186">FIGURE 2.</emphasis>
<taxonomicName id="AC954D2EFFE9FFF0370AF885FD0BF8C0" authority="Adorador &amp; Fernando" authorityName="Adorador &amp; Fernando, Mt. Cagua" authorityYear="2022" box="[270,704,1850,1872]" class="Liliopsida" family="Arecaceae" genus="Pinanga" kingdom="Plantae" order="Arecales" pageId="3" pageNumber="186" phylum="Tracheophyta" rank="species" species="leonardcoi">
<emphasis id="59E1EABFFFE9FFF0370AF885FE19F8DF" box="[270,466,1850,1871]" italics="true" pageId="3" pageNumber="186">Pinanga leonardcoi</emphasis>
Adorador &amp; Fernando
</taxonomicName>
, clustering habit (A), leaves with 23-costulate segments (B), stem with infructescences (C) in different phases of development (DF), Close-up of fruits showing imbricate fruiting perianth (G), longitudinally dissected fruits showing ruminate endosperm and basal embryo (H), basal off-shoot with a stoloniferous stem (I). All from the holotype,
<emphasis id="59E1EABFFFE9FFF0368CF82DFEFCF837" box="[136,311,1938,1959]" italics="true" pageId="3" pageNumber="186">J.T. Adorador 201</emphasis>
(PNH) except F from
<emphasis id="59E1EABFFFE9FFF03414F82DFD79F837" box="[528,690,1938,1959]" italics="true" pageId="3" pageNumber="186">J.P. Bautista 902</emphasis>
(PUH). Scale bar: B—40 cm, C—15 cm, D &amp; E—15 cm, G—40 cm, I—8 cm. Photos by J.T. Adorador.
</paragraph>
</caption>
<caption id="3FEA6625FFEEFFF7368CF830FCEBF7B1" ID-DOI="http://doi.org/10.5281/zenodo.6260343" ID-Zenodo-Dep="6260343" httpUri="https://zenodo.org/record/6260343/files/figure.png" pageId="4" pageNumber="187" startId="4.[136,229,1935,1957]" targetBox="[178,1401,195,1905]" targetPageId="4">
<paragraph id="6B2A36ADFFEEFFF7368CF830FCEBF7B1" blockId="4.[136,1452,1935,2081]" pageId="4" pageNumber="187">
<emphasis id="59E1EABFFFEEFFF7368CF830FF36F834" bold="true" box="[136,253,1935,1957]" pageId="4" pageNumber="187">FIGURE 3.</emphasis>
<taxonomicName id="AC954D2EFFEEFFF73706F82FFD57F835" authority="Adorador &amp; Fernando" authorityName="Adorador &amp; Fernando, Mt. Cagua" authorityYear="2022" box="[258,668,1935,1957]" class="Liliopsida" family="Arecaceae" genus="Pinanga" kingdom="Plantae" order="Arecales" pageId="4" pageNumber="187" phylum="Tracheophyta" rank="species" species="leonardcoi">
<emphasis id="59E1EABFFFEEFFF73706F82FFE0BF834" box="[258,448,1935,1957]" italics="true" pageId="4" pageNumber="187">Pinanga leonardcoi</emphasis>
Adorador &amp; Fernando
</taxonomicName>
, clustering habit (A), middle leaflets with 3-costulate segments (B), apical leaflets (C), immature fruit showing imbricate fruiting perianth (D), infructescence showing spirally arranged fruits (E), longitudinally dissected fruits showing ruminate endosperm and basal embryo (F), seed with adherent portions of the pericarp (G), and basal of-shoot with a stoloniferous stem (H). BC from
<emphasis id="59E1EABFFFEEFFF737D8F857FD45F86C" box="[476,654,2023,2045]" italics="true" pageId="4" pageNumber="187">J.T. Adorador 202</emphasis>
, DH from the holotype,
<emphasis id="59E1EABFFFEEFFF73594F857FB88F86C" box="[912,1091,2023,2045]" italics="true" pageId="4" pageNumber="187">J.T. Adorador 201</emphasis>
.Scale bar: B &amp; C—6 cm, D—1 cm, E—7.5 cm, F—2 cm, G—1.8 cm, H—8 cm. Drawn by J.T. Adorador
</paragraph>
</caption>
<paragraph id="6B2A36ADFFEFFFF636B9FEC9FDA7FDF8" blockId="5.[136,1452,158,869]" pageId="5" pageNumber="188">
Among the Philippine congeners, only
<taxonomicName id="AC954D2EFFEFFFF6347DFEC8FBCDFE00" authority="Fernando (1994: 782)" authorityName="Fernando" authorityPageNumber="782" authorityYear="1994" box="[633,1030,374,400]" class="Liliopsida" family="Arecaceae" genus="Pinanga" kingdom="Plantae" order="Arecales" pageId="5" pageNumber="188" phylum="Tracheophyta" rank="species" species="sobolifera">
<emphasis id="59E1EABFFFEFFFF6347DFEC8FCC3FE00" box="[633,776,374,400]" italics="true" pageId="5" pageNumber="188">P. sobolifera</emphasis>
<bibRefCitation id="0F044B5CFFEFFFF6350BFEC9FBCDFE00" author="Fernando, E. S." box="[783,1030,374,400]" pageId="5" pageNumber="188" pagination="775 - 784" refId="ref3757" refString="Fernando, E. S. (1994) New species of Pinanga (Palmae: Arecoideae) from Luzon Island, Philippines. Kew Bulletin 49 (4): 775 - 784. https: // doi. org / 10.2307 / 4118071" type="journal article" year="1994">Fernando (1994: 782)</bibRefCitation>
</taxonomicName>
of the Philippinensis group, likewise possess the combination of stoloniferous basal shoots and spirally-arranged fruits but
<taxonomicName id="AC954D2EFFEFFFF6324DFE24FB16FE24" authorityName="Adorador &amp; Fernando, Mt. Cagua" authorityYear="2022" box="[1097,1245,410,436]" class="Liliopsida" family="Arecaceae" genus="Pinanga" kingdom="Plantae" order="Arecales" pageId="5" pageNumber="188" phylum="Tracheophyta" rank="species" species="leonardcoi">
<emphasis id="59E1EABFFFEFFFF6324DFE24FB16FE24" box="[1097,1245,410,436]" italics="true" pageId="5" pageNumber="188">P. leonardcoi</emphasis>
</taxonomicName>
differs in its more robust stems (
<quantity id="AC6D9B48FFEFFFF63726FE01FE41FE49" box="[290,394,446,473]" metricMagnitude="-2" metricUnit="m" metricValue="3.75" metricValueMax="4.0" metricValueMin="3.5" pageId="5" pageNumber="188" unit="cm" value="3.75" valueMax="4.0" valueMin="3.5">3.54 cm</quantity>
vs to
<quantity id="AC6D9B48FFEFFFF637CFFE01FDDDFE49" box="[459,534,446,473]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="5" pageNumber="188" unit="cm" value="2.5">2.5 cm</quantity>
in
<taxonomicName id="AC954D2EFFEFFFF6343CFE00FD0FFE48" authorityName="Fernando" authorityYear="1994" box="[568,708,446,472]" class="Liliopsida" family="Arecaceae" genus="Pinanga" kingdom="Plantae" order="Arecales" pageId="5" pageNumber="188" phylum="Tracheophyta" rank="species" species="sobolifera">
<emphasis id="59E1EABFFFEFFFF6343CFE00FD0FFE48" box="[568,708,446,472]" italics="true" pageId="5" pageNumber="188">P. sobolifera</emphasis>
</taxonomicName>
), fewer rachillae (vs.
<quantity id="AC6D9B48FFEFFFF635B1FE01FBC3FE48" box="[949,1032,446,472]" metricMagnitude="-1" metricUnit="m" metricValue="2.413" metricValueMax="2.794" metricValueMin="2.032" pageId="5" pageNumber="188" unit="in" value="9.5" valueMax="11.0" valueMin="8.0">811 in</quantity>
<taxonomicName id="AC954D2EFFEFFFF6320AFE00FB51FE48" authorityName="Fernando" authorityYear="1994" box="[1038,1178,446,472]" class="Liliopsida" family="Arecaceae" genus="Pinanga" kingdom="Plantae" order="Arecales" pageId="5" pageNumber="188" phylum="Tracheophyta" rank="species" species="sobolifera">
<emphasis id="59E1EABFFFEFFFF6320AFE00FB51FE48" box="[1038,1178,446,472]" italics="true" pageId="5" pageNumber="188">P. sobolifera</emphasis>
</taxonomicName>
), and much larger fruits, averaging
<quantity id="AC6D9B48FFEFFFF636F9FE5DFE82FE6D" box="[253,329,482,509]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="5" pageNumber="188" unit="cm" value="2.4">2.4 cm</quantity>
×
<quantity id="AC6D9B48FFEFFFF63763FE5DFE78FE6D" box="[359,435,482,509]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="5" pageNumber="188" unit="cm" value="1.8">1.8 cm</quantity>
(vs 1.2 ×
<quantity id="AC6D9B48FFEFFFF63427FE5DFDA4FE6D" box="[547,623,482,509]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="188" unit="cm" value="0.8">0.8 cm</quantity>
in
<taxonomicName id="AC954D2EFFEFFFF63497FE5CFCEBFE6C" authorityName="Fernando" authorityYear="1994" box="[659,800,482,508]" class="Liliopsida" family="Arecaceae" genus="Pinanga" kingdom="Plantae" order="Arecales" pageId="5" pageNumber="188" phylum="Tracheophyta" rank="species" species="sobolifera">
<emphasis id="59E1EABFFFEFFFF63497FE5CFCEBFE6C" box="[659,800,482,508]" italics="true" pageId="5" pageNumber="188">P. sobolifera</emphasis>
</taxonomicName>
). Meanwhile, the other locally sympatric but more widely occurring congeners,
<emphasis id="59E1EABFFFEFFFF63778FDB8FE69FDB0" box="[380,418,519,544]" italics="true" pageId="5" pageNumber="188">viz.</emphasis>
<taxonomicName id="AC954D2EFFEFFFF637AEFDB8FC9BFDB0" authority="Beccari (1889: 180)" authorityName="Beccari" authorityPageNumber="180" authorityYear="1889" box="[426,848,518,544]" class="Liliopsida" family="Arecaceae" genus="Pinanga" kingdom="Plantae" order="Arecales" pageId="5" pageNumber="188" phylum="Tracheophyta" rank="species" species="philippinensis">
<emphasis id="59E1EABFFFEFFFF637AEFDB8FDAFFDB0" box="[426,612,518,544]" italics="true" pageId="5" pageNumber="188">P. philippinensis</emphasis>
<bibRefCitation id="0F044B5CFFEFFFF63468FDB9FC9BFDB0" author="Beccari, O." box="[620,848,518,544]" pageId="5" pageNumber="188" pagination="169 - 200" refId="ref3396" refString="Beccari, O. (1889) Nuove palme Asiatiche. Malesia 3: 169 - 200." type="journal article" year="1889">Beccari (1889: 180)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="AC954D2EFFEFFFF63559FDB8FB31FDB0" authority="Porte ex Lemaire (1863" authorityName="Lemaire" authorityYear="1863" box="[861,1274,518,545]" class="Liliopsida" family="Arecaceae" genus="Pinanga" kingdom="Plantae" order="Arecales" pageId="5" pageNumber="188" phylum="Tracheophyta" rank="species" species="maculata">
<emphasis id="59E1EABFFFEFFFF63559FDB8FC29FDB0" box="[861,994,518,544]" italics="true" pageId="5" pageNumber="188">P. maculata</emphasis>
Porte ex
<bibRefCitation id="0F044B5CFFEFFFF63254FDB9FB31FDB0" author="Lemaire, C." box="[1104,1274,518,544]" pageId="5" pageNumber="188" refId="ref4010" refString="Lemaire, C. (1863) Pinanga maculata. L'Illustration Horticole 10: pl. 361." type="journal volume" year="1863">Lemaire (1863</bibRefCitation>
</taxonomicName>
: pl. 361) and
<taxonomicName id="AC954D2EFFEFFFF63393FDB8FE07FDD4" authority="Beccari (1907: 223)" authorityName="Beccari" authorityPageNumber="223" authorityYear="1907" class="Liliopsida" family="Arecaceae" genus="Pinanga" kingdom="Plantae" order="Arecales" pageId="5" pageNumber="188" phylum="Tracheophyta" rank="species" species="insignis">
<emphasis id="59E1EABFFFEFFFF63393FDB8FF14FDD4" italics="true" pageId="5" pageNumber="188">P. insignis</emphasis>
<bibRefCitation id="0F044B5CFFEFFFF636E3FD95FE07FDD4" author="Beccari, O." box="[231,460,554,580]" pageId="5" pageNumber="188" pagination="219 - 240" refId="ref3414" refString="Beccari, O. (1907) Notes on Philippine palms, I. Philippine Journal of Science (Botany) 2: 219 - 240." type="journal article" year="1907">Beccari (1907: 223)</bibRefCitation>
</taxonomicName>
, cannot be confused with
<taxonomicName id="AC954D2EFFEFFFF634F1FD94FC42FDD4" authorityName="Adorador &amp; Fernando, Mt. Cagua" authorityYear="2022" box="[757,905,554,580]" class="Liliopsida" family="Arecaceae" genus="Pinanga" kingdom="Plantae" order="Arecales" pageId="5" pageNumber="188" phylum="Tracheophyta" rank="species" species="leonardcoi">
<emphasis id="59E1EABFFFEFFFF634F1FD94FC42FDD4" box="[757,905,554,580]" italics="true" pageId="5" pageNumber="188">P. leonardcoi</emphasis>
</taxonomicName>
as these three taxa belong to separate groups of Philippine
<taxonomicName id="AC954D2EFFEFFFF63706FDF0FDA3FDF8" authority="(Adorador et al. 2020)" baseAuthorityName="Adorador" baseAuthorityYear="2020" box="[258,616,590,616]" class="Liliopsida" family="Arecaceae" genus="Pinanga" kingdom="Plantae" order="Arecales" pageId="5" pageNumber="188" phylum="Tracheophyta" rank="genus">
<emphasis id="59E1EABFFFEFFFF63706FDF0FEAAFDF8" box="[258,353,591,616]" italics="true" pageId="5" pageNumber="188">Pinanga</emphasis>
(
<bibRefCitation id="0F044B5CFFEFFFF6376BFDF1FDABFDF8" author="Adorador, J. T. &amp; Meneses, Z. D. &amp; Fernando, E. S." box="[367,608,590,616]" pageId="5" pageNumber="188" pagination="120 - 134" refId="ref3265" refString="Adorador, J. T., Meneses, Z. D. &amp; Fernando, E. S. (2020) Pinanga gruezoi (Arecaceae), a new slender clustering palm from the Philippines with notes on an amended description of P. samarana. Phytotaxa 429 (2): 120 - 134. https: // doi. org / 10.11646 / phytotaxa. 429.2.3" type="journal article" year="2020">
Adorador
<emphasis id="59E1EABFFFEFFFF637E0FDF0FDD7FDF8" box="[484,540,590,616]" italics="true" pageId="5" pageNumber="188">et al.</emphasis>
2020
</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="6B2A36ADFFEFFFF636B9FDCCFA90FCF4" blockId="5.[136,1452,158,869]" pageId="5" pageNumber="188">
It is curious that
<taxonomicName id="AC954D2EFFEFFFF6377FFDCCFDC4FD1C" authorityName="Adorador &amp; Fernando, Mt. Cagua" authorityYear="2022" box="[379,527,626,652]" class="Liliopsida" family="Arecaceae" genus="Pinanga" kingdom="Plantae" order="Arecales" pageId="5" pageNumber="188" phylum="Tracheophyta" rank="species" species="leonardcoi">
<emphasis id="59E1EABFFFEFFFF6377FFDCCFDC4FD1C" box="[379,527,626,652]" italics="true" pageId="5" pageNumber="188">P. leonardcoi</emphasis>
</taxonomicName>
has only been collected once before as compared to the other generally sympatric
<taxonomicName id="AC954D2EFFEFFFF6368CFD28FF2CFD20" box="[136,231,663,688]" class="Liliopsida" family="Arecaceae" genus="Pinanga" kingdom="Plantae" order="Arecales" pageId="5" pageNumber="188" phylum="Tracheophyta" rank="genus">
<emphasis id="59E1EABFFFEFFFF6368CFD28FF2CFD20" box="[136,231,663,688]" italics="true" pageId="5" pageNumber="188">Pinanga</emphasis>
</taxonomicName>
which rather indicates its rarity in this sub-biogeographic region. Apparently, this palm has successfully established within Mt. Caguas caldera—a geologically recent habitat with the last known eruption in 1860 and strong solfataric activity in 1907 (
<bibRefCitation id="0F044B5CFFEFFFF6340CFD61FC9CFD68" author="Smithsonian Institution" box="[520,855,734,760]" pageId="5" pageNumber="188" refId="ref4307" refString="Smithsonian Institution. (2013) Global Volcanism Program. Available from: https: // volcano. si. edu / (accessed 13 October 2021)." type="url" year="2013">Smithsonian Institution 2013</bibRefCitation>
). Likewise, the calderas average elevation of
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approximates the general elevational species turn-over for palms at around 1,000 (± 200) m elev. (authors personal observations). Thus, its preference to this unique environment and the paucity of additional distribution records altogether indicates that
<taxonomicName id="AC954D2EFFEFFFF6379FFCF4FDE5FCF4" authorityName="Adorador &amp; Fernando, Mt. Cagua" authorityYear="2022" box="[411,558,842,868]" class="Liliopsida" family="Arecaceae" genus="Pinanga" kingdom="Plantae" order="Arecales" pageId="5" pageNumber="188" phylum="Tracheophyta" rank="species" species="leonardcoi">
<emphasis id="59E1EABFFFEFFFF6379FFCF4FDE5FCF4" box="[411,558,842,868]" italics="true" pageId="5" pageNumber="188">P. leonardcoi</emphasis>
</taxonomicName>
is a very narrow endemic palm or could even be a neo-endemic species.
</paragraph>
</subSubSection>
</treatment>
</document>