449 lines
45 KiB
XML
449 lines
45 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1048.67564" ID-GBIF-Dataset="57d9ade8-67e7-4d65-9581-b85a80e88893" ID-PMC="PMC8292283" ID-Pensoft-Pub="1313-2970-1048-145" ID-Pensoft-UUID="93FE96A4127A5D8FAA3A39BCB3E4B729" ID-PubMed="34316267" ID-ZooBank="578795FD41BF4807B0016F7B5AC4D4B9" ModsDocID="1313-2970-1048-145" checkinTime="1626283267302" checkinUser="pensoft" docAuthor="Lantsov, Vladimir I. & Pilipenko, Valentin E." docDate="2021" docId="1D18358B839155CFB3A133D56252432A" docLanguage="en" docName="ZooKeys 1048: 145-175" docOrigin="ZooKeys 1048" docPubDate="2021-07-13" docSource="http://dx.doi.org/10.3897/zookeys.1048.67564" docTitle="Tipula (Lunatipula) talyshensis Savchenko 1964" docType="treatment" docVersion="4" id="93FE96A4127A5D8FAA3A39BCB3E4B729" lastPageNumber="145" masterDocId="93FE96A4127A5D8FAA3A39BCB3E4B729" masterDocTitle="A new Palaearctic species of the subgenus Lunatipula (Diptera, Tipulidae) from the West Caucasus with a survey of the caucasica species group" masterLastPageNumber="175" masterPageNumber="145" pageNumber="145" updateTime="1668150580932" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A new Palaearctic species of the subgenus Lunatipula (Diptera, Tipulidae) from the West Caucasus with a survey of the caucasica species group</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Lantsov, Vladimir I.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-8275-496X</mods:nameIdentifier>
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<mods:affiliation>Tembotov Institute of Ecology of Mountain Territories of Russian Academy of Sciences, I. Armand str., 37 a, Nalchik 360051, Russia</mods:affiliation>
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<mods:nameIdentifier type="email">lantsov@megalog.ru</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Pilipenko, Valentin E.</mods:namePart>
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<mods:affiliation>Lomonosov Moscow State University, GSP- 1, Leninskie Gory, Moscow 119991, Russia</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-07-13</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1048</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>145</mods:start>
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<mods:end>175</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1048.67564</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1048.67564</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1048-145</mods:identifier>
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<mods:identifier type="ZooBank">578795FD41BF4807B0016F7B5AC4D4B9</mods:identifier>
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<mods:identifier type="Pensoft-UUID">93FE96A4127A5D8FAA3A39BCB3E4B729</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="183543017" LSID="urn:lsid:plazi:treatment:1D18358B839155CFB3A133D56252432A" httpUri="http://treatment.plazi.org/id/1D18358B839155CFB3A133D56252432A" lastPageNumber="145" pageId="0" pageNumber="145">
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<subSubSection pageId="0" pageNumber="145" type="nomenclature">
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<paragraph pageId="0" pageNumber="145">
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<taxonomicName LSID="1D18358B-8391-55CF-B3A1-33D56252432A" authority="Savchenko, 1964" authorityName="Savchenko" authorityYear="1964" class="Insecta" family="Tipulidae" genus="Tipula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tipula (Lunatipula) talyshensis" order="Diptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="talyshensis" subGenus="Lunatipula">Tipula (Lunatipula) talyshensis Savchenko, 1964</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="145" type="description">
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<paragraph pageId="0" pageNumber="145">
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. T. (Lunatipula) talyshensis, male habitus A general lateral view B head and thorax, dorsal view. Scale bars: 10 mm (A), 1 mm (B)." figureDoi="10.3897/zookeys.1048.67564.figure10" httpUri="https://binary.pensoft.net/fig/566419" pageId="0" pageNumber="145">Figs 10</figureCitation>
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. T. (Lunatipula) talyshensis A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure11" httpUri="https://binary.pensoft.net/fig/566420" pageId="0" pageNumber="145">, 11</figureCitation>
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Female terminalia of Tipula caucasica species group A-C T. (Lunatipula) caucasica D-F T. (Lunatipula) quadridentata G-I T. (Lunatipula) talyshensis A, D, G ovipositor, lateral view B, E, H ovipositor ventral view C, F, I sternite 9 and furca, lateral view. Abbreviations: c-cerci; f-furca; hv-hypogynial valve; s-sternite; t-tergite. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure12" httpUri="https://binary.pensoft.net/fig/566421" pageId="0" pageNumber="145">, 12G-I</figureCitation>
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<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Heads of Tipula caucasica species group, dorsal view A T. (Lunatipula) caucasica B T. (Lunatipula) eleniya sp. nov. C T. (Lunatipula) paupera D T. (Lunatipula) quadridentata E T. (Lunatipula) Tipula talyshensis. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure13" httpUri="https://binary.pensoft.net/fig/566422" pageId="0" pageNumber="145">; 13E</figureCitation>
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<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Comparisons of some components of male terminalia of the Tipula caucasica species group A, F T. (Lunatipula) caucasica B, G T. (Lunatipula) eleniya sp. nov. C, I T. (Lunatipula) paupera D, J T. (Lunatipula) quadridentata E, K T. (Lunatipula) talyshensis A-E tergite 9, dorsal view F-K inner and outer gonostylus, lateral view. Arrows point to central notch at apex of tergite 9 (C, D), to the wedge-shaped projection posteriorly on inner gonostylus (F), to the rod-like outgrowth behind anterior beak of inner gonostylus (G), to the anterior (dorsal) margin of gonocoxite (I, J, K). Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure14" httpUri="https://binary.pensoft.net/fig/566423" pageId="0" pageNumber="145">, 14E, K</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="145" type="reference_group">
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<paragraph pageId="0" pageNumber="145">
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<taxonomicName authorityName="Savchenko" authorityYear="1964" class="Insecta" family="Tipulidae" genus="Tipula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tipula (Lunatipula) talyshensis" order="Diptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="talyshensis" subGenus="Lunatipula">Tipula (Lunatipula) talyshensis</taxonomicName>
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<bibRefCitation author="Savchenko, EN" journalOrPublisher="Zoologicheskiy Zhurnal" pageId="0" pageNumber="145" refId="B12" refString="Savchenko, EN, 1964. Crane-flies (Diptera, Tipulidae), Subfam. Tipulinae, Genus TipulaL., 2. Fauna USSR, N.S. 89, Nasekomye Dvukrylye [Diptera] 2(4): 1-503. [in Russian]" title="Crane-flies (Diptera, Tipulidae), Subfam. Tipulinae, Genus TipulaL., 2. Fauna USSR, N. S. 89, Nasekomye Dvukrylye [Diptera] 2 (4): 1 - 503. [in Russian]" year="1964">Savchenko 1964</bibRefCitation>
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: 391 (type locality: Azerbaijan, Lerik region); Oosterbroek, Theowald 1992: 121;
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<bibRefCitation author="Oosterbroek, P" journalOrPublisher="Fauna of the Stavropol region" pageId="0" pageNumber="145" publicationUrl="http://ccw.naturalis.nl/detail.php" refId="B8" refString="Oosterbroek, P, 2021. Catalogue of the Craneflies of the World. http://ccw.naturalis.nl/detail.php" title="Catalogue of the Craneflies of the World." url="http://ccw.naturalis.nl/detail.php" year="2021">Oosterbroek 2021</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="145" type="materials_examined">
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<paragraph pageId="0" pageNumber="145">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="145">
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<materialsCitation collectingDate="1954-06-26" collectorName="Jafarov, Original" country="Azerbaijan" determinerName="Savchenko" elevation="1115" latitude="38.775276" location="Azerbaijanskaya Soviet Socialist Republic" longLatPrecision="19" longitude="48.41528" specimenCount="1" specimenCount-male="1" stateProvince="Lerik" typeStatus="Holotype">
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<emphasis bold="true" pageId="0" pageNumber="145">
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<emphasis italics="true" pageId="0" pageNumber="145">
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<typeStatus>Holotype</typeStatus>
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</emphasis>
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:
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</emphasis>
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<collectingCountry name="Azerbaijan">Azerbaijan</collectingCountry>
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•
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<specimenCount type="male">1 male</specimenCount>
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,
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<normalizedToken originalValue="“р-н">"r-n</normalizedToken>
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<normalizedToken originalValue="Лерик">
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<collectingRegion country="Azerbaijan" name="Lerik">Lerik</collectingRegion>
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</normalizedToken>
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,
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<normalizedToken originalValue="Азербайдж">Azerbaidzh</normalizedToken>
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.
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<normalizedToken originalValue="ССР">SSR</normalizedToken>
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26. VI. [1]954
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<normalizedToken originalValue="Джафаров”">Dzhafarov"</normalizedToken>
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[
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<collectingRegion country="Azerbaijan" name="Lerik">Lerik region</collectingRegion>
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,
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<collectingCountry name="Azerbaijan">Azerbaijan</collectingCountry>
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. SSR. [
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<location LSID="urn:lsid:plazi:treatment:1D18358B839155CFB3A133D56252432A:EFB2ADBB14D953BAFACAC004ECB3936D" country="Azerbaijan" latitude="38.775276" longLatPrecision="19" longitude="48.41528" name="Azerbaijanskaya Soviet Socialist Republic" stateProvince="Lerik">Azerbaijanskaya Soviet Socialist Republic</location>
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] [alt. ~
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<quantity metricMagnitude="3" metricUnit="m" metricValue="1.115" unit="m" value="1115.0">
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<elevation metricMagnitude="3" metricUnit="m" metricValue="1.115" unit="m" value="1115.0">1115 m</elevation>
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</quantity>
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,
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<geoCoordinate degrees="38" direction="north" minutes="46" orientation="latitude" precision="15" seconds="31" value="38.775276">38°46'31"N</geoCoordinate>
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,
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<geoCoordinate degrees="48" direction="east" minutes="24" orientation="longitude" precision="15" seconds="55" value="48.41528">48°24'55"E</geoCoordinate>
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];
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<collectingDate value="1954-06-26">26 Jun. 1954</collectingDate>
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;
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<collectorName>Jafarov</collectorName>
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; / "
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<taxonomicName authorityName="Savchenko" authorityYear="1964" class="Insecta" family="Tipulidae" genus="Tipula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tipula talyshensis" order="Diptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="talyshensis">
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<emphasis italics="true" pageId="0" pageNumber="145">Tipula talyshensis</emphasis>
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</taxonomicName>
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det.
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<determinerName>Savchenko</determinerName>
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sp. n." [white label] / [
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<collectorName>Original</collectorName>
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red label without text] /
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<normalizedToken originalValue="“Holotypus”">
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"
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<typeStatus>Holotypus</typeStatus>
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"
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</normalizedToken>
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</materialsCitation>
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[red label]; ZISP.
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<typeStatus>Holotype</typeStatus>
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in good condition (Fig.
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. T. (Lunatipula) talyshensis, male habitus A general lateral view B head and thorax, dorsal view. Scale bars: 10 mm (A), 1 mm (B)." figureDoi="10.3897/zookeys.1048.67564.figure10" httpUri="https://binary.pensoft.net/fig/566419" pageId="0" pageNumber="145">10A, B</figureCitation>
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), however, some legs missing: fore legs - left missing, right up to femur; mid legs - left present, right missing; hind legs - left and right up to femur.
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<materialsCitation collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="Jafarov & Original" country="Azerbaijan" determinerName="Savchenko" elevation="1115" latitude="38.775276" location="Azerbaijan" longLatPrecision="19" longitude="48.41528" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Lerik" typeStatus="Paratypes">
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<emphasis bold="true" pageId="0" pageNumber="145">
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<emphasis italics="true" pageId="0" pageNumber="145">
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<typeStatus>Paratypes</typeStatus>
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</emphasis>
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.
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</emphasis>
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<collectingCountry name="Azerbaijan">Azerbaijan</collectingCountry>
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•
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<specimenCount type="male">2 males</specimenCount>
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,
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<specimenCount type="female">2 females</specimenCount>
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; same data as for holotype [printed on white paper] /
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<normalizedToken originalValue="“Paratypus”">
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"
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<typeStatus>Paratypus</typeStatus>
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"
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</normalizedToken>
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[
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<typeStatus>Paratype</typeStatus>
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not initially marked; red label]; ZISP
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</materialsCitation>
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.
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</paragraph>
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<caption doi="10.3897/zookeys.1048.67564.figure10" httpUri="https://binary.pensoft.net/fig/566419" pageId="0" pageNumber="145" start="Figure 10" startId="F10">
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<paragraph pageId="0" pageNumber="145">
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<emphasis bold="true" pageId="0" pageNumber="145">Figure 10.</emphasis>
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<taxonomicName lsidName="T. (Lunatipula) talyshensis" pageId="0" pageNumber="145" rank="species" species="talyshensis" subGenus="Lunatipula">T. (Lunatipula) talyshensis</taxonomicName>
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, male habitus
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<emphasis bold="true" pageId="0" pageNumber="145">A</emphasis>
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general lateral view
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<emphasis bold="true" pageId="0" pageNumber="145">B</emphasis>
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head and thorax, dorsal view. Scale bars:
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<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.0" unit="mm" value="10.0">10 mm</quantity>
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(
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<emphasis bold="true" pageId="0" pageNumber="145">A</emphasis>
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),
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<quantity metricMagnitude="-3" metricUnit="m" metricValue="1.0" unit="mm" value="1.0">1 mm</quantity>
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(
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<emphasis bold="true" pageId="0" pageNumber="145">B</emphasis>
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).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="145" type="diagnosis">
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<paragraph pageId="0" pageNumber="145">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="145">Male. Tergite 9 at apex with a denticle on either side of wide, shallow, flat median notch. Paired apical appendages of sternite 9 narrower distally, bearing tassel of long golden setae. Female. Hypogynial valve several times longer than width at base.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="145" type="redescription">
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<paragraph pageId="0" pageNumber="145">Redescription.</paragraph>
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<paragraph pageId="0" pageNumber="145">
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Adult male (Fig.
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. T. (Lunatipula) talyshensis, male habitus A general lateral view B head and thorax, dorsal view. Scale bars: 10 mm (A), 1 mm (B)." figureDoi="10.3897/zookeys.1048.67564.figure10" httpUri="https://binary.pensoft.net/fig/566419" pageId="0" pageNumber="145">10A, B</figureCitation>
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). General body coloration grey. Body length 15-20 mm, wings 18-19.5 mm.
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</paragraph>
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<paragraph pageId="0" pageNumber="145">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Head</emphasis>
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(Fig.
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<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Heads of Tipula caucasica species group, dorsal view A T. (Lunatipula) caucasica B T. (Lunatipula) eleniya sp. nov. C T. (Lunatipula) paupera D T. (Lunatipula) quadridentata E T. (Lunatipula) Tipula talyshensis. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure13" httpUri="https://binary.pensoft.net/fig/566422" pageId="0" pageNumber="145">13E</figureCitation>
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). Vertex light grey with indistinct median grey line, glabrous in middle and around eye with rows of setae between. Rostrum dorsally dark grey, whitish at base, ventrally light brown with grey median stripe. Nasus with long, whitish procumbent setae. Tempora and genae light grey to whitish (color depending on angle of inclination) with faint silvery pruinescence.
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</paragraph>
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<paragraph pageId="0" pageNumber="145">
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<emphasis bold="true" pageId="0" pageNumber="145">
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<emphasis italics="true" pageId="0" pageNumber="145">Antennae</emphasis>
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.
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</emphasis>
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Scape dark grey with rusty bristles, pedicel yellowish with indistinct brown line in middle, flagellum brown.
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</paragraph>
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<paragraph pageId="0" pageNumber="145">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Thorax</emphasis>
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(Fig.
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. T. (Lunatipula) talyshensis, male habitus A general lateral view B head and thorax, dorsal view. Scale bars: 10 mm (A), 1 mm (B)." figureDoi="10.3897/zookeys.1048.67564.figure10" httpUri="https://binary.pensoft.net/fig/566419" pageId="0" pageNumber="145">10A, B</figureCitation>
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). Pronotum grey with whitish setae. Scutum (Fig.
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. T. (Lunatipula) talyshensis, male habitus A general lateral view B head and thorax, dorsal view. Scale bars: 10 mm (A), 1 mm (B)." figureDoi="10.3897/zookeys.1048.67564.figure10" httpUri="https://binary.pensoft.net/fig/566419" pageId="0" pageNumber="145">10B</figureCitation>
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) with four brownish stripes on grey, short whitish setae between central and side stripes. Scutal lobe grey with faint silvery pruinescence, anterolaterally with group of whitish setae. Pleura and coxae grey with silvery pruinescence; katepisternum dorsally with sparse whitish setae, anepisternum and meron glabrous. Scutellum and mediotergite grey with whitish setae, scutellum with dark grey median line visible as specimen is rotated.
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</paragraph>
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<paragraph pageId="0" pageNumber="145">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Wings</emphasis>
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(Fig.
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. T. (Lunatipula) talyshensis, male habitus A general lateral view B head and thorax, dorsal view. Scale bars: 10 mm (A), 1 mm (B)." figureDoi="10.3897/zookeys.1048.67564.figure10" httpUri="https://binary.pensoft.net/fig/566419" pageId="0" pageNumber="145">10A</figureCitation>
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). Translucent, grey, with light brown pterostigma. Longitudinal veins
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<emphasis italics="true" pageId="0" pageNumber="145">C</emphasis>
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,
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<emphasis italics="true" pageId="0" pageNumber="145">Sc</emphasis>
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, and
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<emphasis italics="true" pageId="0" pageNumber="145">R</emphasis>
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and bases of
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<emphasis italics="true" pageId="0" pageNumber="145">A1</emphasis>
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and
|
||
<emphasis italics="true" pageId="0" pageNumber="145">A2</emphasis>
|
||
with macrotrichia.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<emphasis bold="true" pageId="0" pageNumber="145">
|
||
<taxonomicName genus="Halteres" lsidName="Halteres" pageId="0" pageNumber="145" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Halteres</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Stem yellowish with light setae, knob brown.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<emphasis bold="true" pageId="0" pageNumber="145">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Legs</emphasis>
|
||
.
|
||
</emphasis>
|
||
Coxae grey with silvery pruinescence and long, whitish setae. Femora light brown with procumbent dark brown setae. Claws of fifth tarsal segment without spine at base.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Abdomen</emphasis>
|
||
(Figs
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. T. (Lunatipula) talyshensis, male habitus A general lateral view B head and thorax, dorsal view. Scale bars: 10 mm (A), 1 mm (B)." figureDoi="10.3897/zookeys.1048.67564.figure10" httpUri="https://binary.pensoft.net/fig/566419" pageId="0" pageNumber="145">10A</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. T. (Lunatipula) talyshensis A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure11" httpUri="https://binary.pensoft.net/fig/566420" pageId="0" pageNumber="145">11A</figureCitation>
|
||
). Dark grey with short whitish setae. Posterior margins of tergites without noticeable light edging, lateral margins with wide whitish to yellowish edging.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Terminalia</emphasis>
|
||
(Figs
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. T. (Lunatipula) talyshensis A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure11" httpUri="https://binary.pensoft.net/fig/566420" pageId="0" pageNumber="145">11</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Comparisons of some components of male terminalia of the Tipula caucasica species group A, F T. (Lunatipula) caucasica B, G T. (Lunatipula) eleniya sp. nov. C, I T. (Lunatipula) paupera D, J T. (Lunatipula) quadridentata E, K T. (Lunatipula) talyshensis A-E tergite 9, dorsal view F-K inner and outer gonostylus, lateral view. Arrows point to central notch at apex of tergite 9 (C, D), to the wedge-shaped projection posteriorly on inner gonostylus (F), to the rod-like outgrowth behind anterior beak of inner gonostylus (G), to the anterior (dorsal) margin of gonocoxite (I, J, K). Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure14" httpUri="https://binary.pensoft.net/fig/566423" pageId="0" pageNumber="145">14E, K</figureCitation>
|
||
). Hypopygium (Fig.
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. T. (Lunatipula) talyshensis A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure11" httpUri="https://binary.pensoft.net/fig/566420" pageId="0" pageNumber="145">11A, B, C</figureCitation>
|
||
) not thickened. Tergite 9 (Figs
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. T. (Lunatipula) talyshensis A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure11" httpUri="https://binary.pensoft.net/fig/566420" pageId="0" pageNumber="145">11E, F</figureCitation>
|
||
;
|
||
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Comparisons of some components of male terminalia of the Tipula caucasica species group A, F T. (Lunatipula) caucasica B, G T. (Lunatipula) eleniya sp. nov. C, I T. (Lunatipula) paupera D, J T. (Lunatipula) quadridentata E, K T. (Lunatipula) talyshensis A-E tergite 9, dorsal view F-K inner and outer gonostylus, lateral view. Arrows point to central notch at apex of tergite 9 (C, D), to the wedge-shaped projection posteriorly on inner gonostylus (F), to the rod-like outgrowth behind anterior beak of inner gonostylus (G), to the anterior (dorsal) margin of gonocoxite (I, J, K). Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure14" httpUri="https://binary.pensoft.net/fig/566423" pageId="0" pageNumber="145">14E</figureCitation>
|
||
) trapezoidal, apex with rather wide, shallow median notch bounded by a denticles. Gonocoxites (Figs
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. T. (Lunatipula) talyshensis A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure11" httpUri="https://binary.pensoft.net/fig/566420" pageId="0" pageNumber="145">11G</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Comparisons of some components of male terminalia of the Tipula caucasica species group A, F T. (Lunatipula) caucasica B, G T. (Lunatipula) eleniya sp. nov. C, I T. (Lunatipula) paupera D, J T. (Lunatipula) quadridentata E, K T. (Lunatipula) talyshensis A-E tergite 9, dorsal view F-K inner and outer gonostylus, lateral view. Arrows point to central notch at apex of tergite 9 (C, D), to the wedge-shaped projection posteriorly on inner gonostylus (F), to the rod-like outgrowth behind anterior beak of inner gonostylus (G), to the anterior (dorsal) margin of gonocoxite (I, J, K). Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure14" httpUri="https://binary.pensoft.net/fig/566423" pageId="0" pageNumber="145">14K</figureCitation>
|
||
) with large ventral tooth, dorsal tooth truncated at apex. Outer gonostylus without significant features (Fig.
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. T. (Lunatipula) talyshensis A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure11" httpUri="https://binary.pensoft.net/fig/566420" pageId="0" pageNumber="145">11G</figureCitation>
|
||
). Inner gonostylus as in Figures
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. T. (Lunatipula) talyshensis A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure11" httpUri="https://binary.pensoft.net/fig/566420" pageId="0" pageNumber="145">11G, H</figureCitation>
|
||
;
|
||
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Comparisons of some components of male terminalia of the Tipula caucasica species group A, F T. (Lunatipula) caucasica B, G T. (Lunatipula) eleniya sp. nov. C, I T. (Lunatipula) paupera D, J T. (Lunatipula) quadridentata E, K T. (Lunatipula) talyshensis A-E tergite 9, dorsal view F-K inner and outer gonostylus, lateral view. Arrows point to central notch at apex of tergite 9 (C, D), to the wedge-shaped projection posteriorly on inner gonostylus (F), to the rod-like outgrowth behind anterior beak of inner gonostylus (G), to the anterior (dorsal) margin of gonocoxite (I, J, K). Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure14" httpUri="https://binary.pensoft.net/fig/566423" pageId="0" pageNumber="145">14K</figureCitation>
|
||
. Paired appendages of sternite 8 (Fig.
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. T. (Lunatipula) talyshensis A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure11" httpUri="https://binary.pensoft.net/fig/566420" pageId="0" pageNumber="145">11D</figureCitation>
|
||
) relatively small, with fringe of hairs along inner edge. Apical appendages of sternite 9 (Fig.
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. T. (Lunatipula) talyshensis A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure11" httpUri="https://binary.pensoft.net/fig/566420" pageId="0" pageNumber="145">11K, L</figureCitation>
|
||
) narrowed distally, bearing tassel of long golden setae. Gonocoxal fragment, semen pump and aedeagus as in Figure
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. T. (Lunatipula) talyshensis A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure11" httpUri="https://binary.pensoft.net/fig/566420" pageId="0" pageNumber="145">11I, J</figureCitation>
|
||
, respectively.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1048.67564.figure11" httpUri="https://binary.pensoft.net/fig/566420" pageId="0" pageNumber="145" start="Figure 11" startId="F11">
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<emphasis bold="true" pageId="0" pageNumber="145">Figure 11.</emphasis>
|
||
<taxonomicName lsidName="T. (Lunatipula) talyshensis" pageId="0" pageNumber="145" rank="species" species="talyshensis" subGenus="Lunatipula">T. (Lunatipula) talyshensis</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="145">A</emphasis>
|
||
terminal segments of male abdomen, lateral view (dry)
|
||
<emphasis bold="true" pageId="0" pageNumber="145">B-L</emphasis>
|
||
male terminalia (after KOH 10% treatment)
|
||
<emphasis bold="true" pageId="0" pageNumber="145">B</emphasis>
|
||
hypopygium, caudal view
|
||
<emphasis bold="true" pageId="0" pageNumber="145">C</emphasis>
|
||
hypopygium, lateral view
|
||
<emphasis bold="true" pageId="0" pageNumber="145">D</emphasis>
|
||
sternite 8, ventral view
|
||
<emphasis bold="true" pageId="0" pageNumber="145">E</emphasis>
|
||
tergite 9, caudal view
|
||
<emphasis bold="true" pageId="0" pageNumber="145">F</emphasis>
|
||
tergite 9, dorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="145">G</emphasis>
|
||
inner and outer gonostylus, lateral view
|
||
<emphasis bold="true" pageId="0" pageNumber="145">H</emphasis>
|
||
inner and outer gonostylus, caudal view
|
||
<emphasis bold="true" pageId="0" pageNumber="145">I</emphasis>
|
||
gonocoxal fragment, dorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="145">J</emphasis>
|
||
semen pump and aedeagus, lateral view
|
||
<emphasis bold="true" pageId="0" pageNumber="145">K</emphasis>
|
||
adminiculum and paired apical appendages of sternite 9, caudal view
|
||
<emphasis bold="true" pageId="0" pageNumber="145">L</emphasis>
|
||
adminiculum and paired apical appendages of sternite 9, lateral view. Scale bar: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<emphasis bold="true" pageId="0" pageNumber="145">Female.</emphasis>
|
||
Adult female body length 18-25 mm, wings 18-18.5 mm. Similar to male. Cercal hypogynial valve (Fig.
|
||
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Female terminalia of Tipula caucasica species group A-C T. (Lunatipula) caucasica D-F T. (Lunatipula) quadridentata G-I T. (Lunatipula) talyshensis A, D, G ovipositor, lateral view B, E, H ovipositor ventral view C, F, I sternite 9 and furca, lateral view. Abbreviations: c-cerci; f-furca; hv-hypogynial valve; s-sternite; t-tergite. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure12" httpUri="https://binary.pensoft.net/fig/566421" pageId="0" pageNumber="145">12G, H</figureCitation>
|
||
) several times longer than wide at base. Sternite 9 and furca as in Figure
|
||
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Female terminalia of Tipula caucasica species group A-C T. (Lunatipula) caucasica D-F T. (Lunatipula) quadridentata G-I T. (Lunatipula) talyshensis A, D, G ovipositor, lateral view B, E, H ovipositor ventral view C, F, I sternite 9 and furca, lateral view. Abbreviations: c-cerci; f-furca; hv-hypogynial valve; s-sternite; t-tergite. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure12" httpUri="https://binary.pensoft.net/fig/566421" pageId="0" pageNumber="145">12I</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1048.67564.figure12" httpUri="https://binary.pensoft.net/fig/566421" pageId="0" pageNumber="145" start="Figure 12" startId="F12">
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<emphasis bold="true" pageId="0" pageNumber="145">Figure 12.</emphasis>
|
||
Female terminalia of
|
||
<taxonomicName authorityName="Riedel" authorityYear="1920" class="Insecta" family="Tipulidae" genus="Tipula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tipula caucasica" order="Diptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="caucasica">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Tipula caucasica</emphasis>
|
||
</taxonomicName>
|
||
species group
|
||
<emphasis bold="true" pageId="0" pageNumber="145">A-C</emphasis>
|
||
<taxonomicName lsidName="T. (Lunatipula) caucasica" pageId="0" pageNumber="145" rank="species" species="caucasica" subGenus="Lunatipula">T. (Lunatipula) caucasica</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="145">D-F</emphasis>
|
||
<taxonomicName lsidName="T. (Lunatipula) quadridentata" pageId="0" pageNumber="145" rank="species" species="quadridentata" subGenus="Lunatipula">T. (Lunatipula) quadridentata</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="145">G-I</emphasis>
|
||
<taxonomicName lsidName="T. (Lunatipula) talyshensis" pageId="0" pageNumber="145" rank="species" species="talyshensis" subGenus="Lunatipula">T. (Lunatipula) talyshensis</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="145">A, D, G</emphasis>
|
||
ovipositor, lateral view
|
||
<emphasis bold="true" pageId="0" pageNumber="145">B, E, H</emphasis>
|
||
ovipositor ventral view
|
||
<emphasis bold="true" pageId="0" pageNumber="145">C, F, I</emphasis>
|
||
sternite 9 and furca, lateral view. Abbreviations: c-cerci; f-furca; hv-hypogynial valve; s-sternite; t-tergite. Scale bar: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="145" type="comparisons">
|
||
<paragraph pageId="0" pageNumber="145">Comparison with closely related species.</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
This species differs from other species of the
|
||
<taxonomicName authorityName="Riedel" authorityYear="1920" class="Insecta" family="Tipulidae" genus="Tipula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tipula caucasica" order="Diptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="caucasica">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Tipula caucasica</emphasis>
|
||
</taxonomicName>
|
||
group by the broad shallow notch at the apex of tergite 9, the tassel of golden setae at the tip of the apical appendage of sternite 9, and by the small size of the paired appendages of sternite 8. The unusually long hypogynial valve of the female distinguishes this species from that of
|
||
<taxonomicName lsidName="T. caucasica" pageId="0" pageNumber="145" rank="species" species="caucasica">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">T. caucasica</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="T. quadridentata" pageId="0" pageNumber="145" rank="species" species="quadridentata">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">T. quadridentata</emphasis>
|
||
</taxonomicName>
|
||
, the other known females of the
|
||
<taxonomicName authorityName="Riedel" authorityYear="1920" class="Insecta" family="Tipulidae" genus="Tipula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tipula caucasica" order="Diptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="caucasica">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Tipula caucasica</emphasis>
|
||
</taxonomicName>
|
||
group.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="145" type="elevation">
|
||
<paragraph pageId="0" pageNumber="145">Elevation.</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">Adults were collected at altitudes ~ 1115 m.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="145" type="flight period">
|
||
<paragraph pageId="0" pageNumber="145">Flight period.</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">Adults were collected on 26 June.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="145" type="habitat">
|
||
<paragraph pageId="0" pageNumber="145">Habitat.</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">No data.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="145" type="distribution">
|
||
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">Endemic to the Caucasus - currently only known from Talysh (Azerbaijan).</paragraph>
|
||
<caption doi="10.3897/zookeys.1048.67564.figure13" httpUri="https://binary.pensoft.net/fig/566422" pageId="0" pageNumber="145" start="Figure 13" startId="F13">
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<emphasis bold="true" pageId="0" pageNumber="145">Figure 13.</emphasis>
|
||
Heads of
|
||
<taxonomicName authorityName="Riedel" authorityYear="1920" class="Insecta" family="Tipulidae" genus="Tipula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tipula caucasica" order="Diptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="caucasica">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Tipula caucasica</emphasis>
|
||
</taxonomicName>
|
||
species group, dorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="145">A</emphasis>
|
||
<taxonomicName lsidName="T. (Lunatipula) caucasica" pageId="0" pageNumber="145" rank="species" species="caucasica" subGenus="Lunatipula">T. (Lunatipula) caucasica</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="145">B</emphasis>
|
||
<taxonomicName lsidName="T. (Lunatipula) eleniya" pageId="0" pageNumber="145" rank="species" species="eleniya" subGenus="Lunatipula">T. (Lunatipula) eleniya</taxonomicName>
|
||
sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="145">C</emphasis>
|
||
<taxonomicName lsidName="T. (Lunatipula) paupera" pageId="0" pageNumber="145" rank="species" species="paupera" subGenus="Lunatipula">T. (Lunatipula) paupera</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="145">D</emphasis>
|
||
<taxonomicName lsidName="T. (Lunatipula) quadridentata" pageId="0" pageNumber="145" rank="species" species="quadridentata" subGenus="Lunatipula">T. (Lunatipula) quadridentata</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="145">E</emphasis>
|
||
<taxonomicName genus="T." lsidName="T." pageId="0" pageNumber="145" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">T.</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName authorityName="Edwards" authorityYear="1931" class="Insecta" family="Tipulidae" genus="Lunatipula" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Lunatipula" order="Diptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Lunatipula</emphasis>
|
||
</taxonomicName>
|
||
)
|
||
<taxonomicName authorityName="Savchenko" authorityYear="1964" class="Insecta" family="Tipulidae" genus="Tipula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tipula talyshensis" order="Diptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="talyshensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Tipula talyshensis</emphasis>
|
||
</taxonomicName>
|
||
. Scale bar: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.1048.67564.figure14" httpUri="https://binary.pensoft.net/fig/566423" pageId="0" pageNumber="145" start="Figure 14" startId="F14">
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<emphasis bold="true" pageId="0" pageNumber="145">Figure 14.</emphasis>
|
||
Comparisons of some components of male terminalia of the
|
||
<taxonomicName authorityName="Riedel" authorityYear="1920" class="Insecta" family="Tipulidae" genus="Tipula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tipula caucasica" order="Diptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="caucasica">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Tipula caucasica</emphasis>
|
||
</taxonomicName>
|
||
species group
|
||
<emphasis bold="true" pageId="0" pageNumber="145">A, F</emphasis>
|
||
<taxonomicName lsidName="T. (Lunatipula) caucasica" pageId="0" pageNumber="145" rank="species" species="caucasica" subGenus="Lunatipula">T. (Lunatipula) caucasica</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="145">B, G</emphasis>
|
||
<taxonomicName lsidName="T. (Lunatipula) eleniya" pageId="0" pageNumber="145" rank="species" species="eleniya" subGenus="Lunatipula">T. (Lunatipula) eleniya</taxonomicName>
|
||
sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="145">C, I</emphasis>
|
||
<taxonomicName lsidName="T. (Lunatipula) paupera" pageId="0" pageNumber="145" rank="species" species="paupera" subGenus="Lunatipula">T. (Lunatipula) paupera</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="145">D, J</emphasis>
|
||
<taxonomicName lsidName="T. (Lunatipula) quadridentata" pageId="0" pageNumber="145" rank="species" species="quadridentata" subGenus="Lunatipula">T. (Lunatipula) quadridentata</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="145">E, K</emphasis>
|
||
<taxonomicName lsidName="T. (Lunatipula) talyshensis" pageId="0" pageNumber="145" rank="species" species="talyshensis" subGenus="Lunatipula">T. (Lunatipula) talyshensis</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="145">A-E</emphasis>
|
||
tergite 9, dorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="145">F-K</emphasis>
|
||
inner and outer gonostylus, lateral view. Arrows point to central notch at apex of tergite 9 (
|
||
<emphasis bold="true" pageId="0" pageNumber="145">C, D</emphasis>
|
||
), to the wedge-shaped projection posteriorly on inner gonostylus (
|
||
<emphasis bold="true" pageId="0" pageNumber="145">F</emphasis>
|
||
), to the rod-like outgrowth behind anterior beak of inner gonostylus (
|
||
<emphasis bold="true" pageId="0" pageNumber="145">G</emphasis>
|
||
), to the anterior (dorsal) margin of gonocoxite (
|
||
<emphasis bold="true" pageId="0" pageNumber="145">I, J, K</emphasis>
|
||
). Scale bar: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |