339 lines
30 KiB
XML
339 lines
30 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.816.29738" ID-GBIF-Dataset="31bce27d-7094-467b-95ae-009a27e00cf5" ID-PMC="PMC6341056" ID-Pensoft-Pub="1313-2970-816-1" ID-PubMed="30686917" ID-ZBK="51CEEF2E1E1040A8A6731140426ED5A7" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1313-2970-816-1" ModsDocOrigin="ZooKeys 816" ModsDocTitle="A taxonomic review of the pericaline ground-beetles in Taiwan, with descriptions of new species (Coleoptera, Carabidae, Lebiini)" checkinTime="1547561011487" checkinUser="pensoft" docAuthor="Hunting, Wesley & Yang, Man-Miao" docDate="2019" docId="A7CF425B4C3DE9D2255D47D7FC4376E6" docLanguage="en" docName="ZooKeys 816: 1-164" docOrigin="ZooKeys 816" docSource="http://dx.doi.org/10.3897/zookeys.816.29738" docTitle="Catascopus (Catascopoides) horni Jedlicka" docType="treatment" docVersion="4" lastPageNumber="50" masterDocId="AA26FF84157BFFAE490E7F133744FF98" masterDocTitle="A taxonomic review of the pericaline ground-beetles in Taiwan, with descriptions of new species (Coleoptera, Carabidae, Lebiini)" masterLastPageNumber="164" masterPageNumber="1" pageNumber="46" updateTime="1668166739332" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A taxonomic review of the pericaline ground-beetles in Taiwan, with descriptions of new species (Coleoptera, Carabidae, Lebiini)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Hunting, Wesley</mods:namePart>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Yang, Man-Miao</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="volume">
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<mods:number>816</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>164</mods:end>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.816.29738</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.816.29738</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-816-1</mods:identifier>
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<mods:identifier type="ZBK">51CEEF2E1E1040A8A6731140426ED5A7</mods:identifier>
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<mods:identifier type="ZooBank">51CEEF2E1E1040A8A6731140426ED5A7</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="154126005" LSID="urn:lsid:plazi:treatment:A7CF425B4C3DE9D2255D47D7FC4376E6" httpUri="http://treatment.plazi.org/id/A7CF425B4C3DE9D2255D47D7FC4376E6" lastPageId="49" lastPageNumber="50" pageId="45" pageNumber="46">
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<subSubSection pageId="45" pageNumber="46" type="nomenclature">
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<paragraph pageId="45" pageNumber="46">
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<taxonomicName authority="Jedlicka" class="Insecta" family="Carabidae" genus="Catascopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Catascopus (Catascopoides) horni" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="horni" subGenus="Catascopoides">
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Catascopus (Catascopoides) horni
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<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
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</taxonomicName>
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Figs 23B, 30, 31, 32
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<normalizedToken originalValue="A–D">A-D</normalizedToken>
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, 33A, 34D, 35
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</paragraph>
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</subSubSection>
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<subSubSection pageId="45" pageNumber="46" type="reference_group">
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<paragraph pageId="45" pageNumber="46">
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<taxonomicName class="Insecta" family="Carabidae" genus="Catascopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Catascopus (Catascopoides) horni" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="horni" subGenus="Catascopoides">Catascopus (Catascopoides) horni</taxonomicName>
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<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
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, 1932: 82;
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<bibRefCitation pageId="45" pageNumber="46">
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<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
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1963
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</bibRefCitation>
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: 383;
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<bibRefCitation author="Straneo, SL" journalOrPublisher="Elytron" pageId="154" pageNumber="155" pagination="141 - 172" title="Sulle species Orientali del genere Catascopus Kirby, 1825 (Coleoptera: Carabidae)." volume="8" year="1994">Straneo 1994</bibRefCitation>
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: 143;
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<bibRefCitation author="Lorenz, W" journalOrPublisher="Paussinae, Cicindelinae, Rhysodinae). Second ed. Germany" pageId="152" pageNumber="153" title="Systematic List of Extant Ground Beetles of the World. (Insecta, Coleoptera, Adephaga: Trachypachidae & Carabidae incl." year="2005">Lorenz 2005</bibRefCitation>
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: 454.
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</paragraph>
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<paragraph pageId="45" pageNumber="46">
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<taxonomicName class="Insecta" family="Carabidae" genus="Catascopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Catascopus (Dentiscopus) horni" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="horni" subGenus="Dentiscopus">Catascopus (Dentiscopus) horni</taxonomicName>
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<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
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: Straneo, 1994: 148.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="45" pageNumber="46" type="type material">
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<paragraph pageId="45" pageNumber="46">Types and other material examined.</paragraph>
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<paragraph pageId="45" pageNumber="46">
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Holotype (female) labeled: "Banshoryo - Disfr./Sokutsu (Formosa)/H. Sauter VI. 1912";
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<normalizedToken originalValue="“Holotypus”">"Holotypus"</normalizedToken>
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[rectangular, red paper];
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<normalizedToken originalValue="“TYPE”">"TYPE"</normalizedToken>
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[rectangular, red paper, black border]; "
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<taxonomicName class="Insecta" family="Carabidae" genus="Catascopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Catascopus" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">Catascopus</taxonomicName>
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Horni sp. n./mihi/DET.ING.JEDLICKA"; "DEI
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<taxonomicName lsidName="" order="Coleoptera" pageId="45" pageNumber="46" rank="order">Coleoptera</taxonomicName>
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/# 200416". 23 specimens of
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<taxonomicName lsidName="C. horni" pageId="45" pageNumber="46" rank="species" species="horni">C. horni</taxonomicName>
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: seven males and 16 females. For further details see EH Strickland Virtual Entomology Museum Database.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="45" pageNumber="46" type="type locality">
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<paragraph pageId="45" pageNumber="46">Type locality.</paragraph>
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<paragraph pageId="45" pageNumber="46">"Formosa, Sokutsu". Formosa is the old name for Taiwan and Sokutsu refers to Hsiaolin, Kaohsiung county.</paragraph>
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</subSubSection>
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<subSubSection pageId="45" pageNumber="46" type="taxonomic notes">
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<paragraph pageId="45" pageNumber="46">Taxonomic notes.</paragraph>
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<paragraph pageId="45" pageNumber="46">
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Both
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<taxonomicName genus="C." lsidName="C." pageId="45" pageNumber="46" rank="genus">C.</taxonomicName>
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(
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<taxonomicName lsidName="(C.)" pageId="45" pageNumber="46" rank="subGenus" subGenus="C.">C.</taxonomicName>
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)
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<taxonomicName lsidName="mirabilis" pageId="45" pageNumber="46" rank="species" species="mirabilis">mirabilis</taxonomicName>
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and
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<taxonomicName genus="C." lsidName="C." pageId="45" pageNumber="46" rank="genus">C.</taxonomicName>
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(
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<taxonomicName lsidName="(C.)" pageId="45" pageNumber="46" rank="subGenus" subGenus="C.">C.</taxonomicName>
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)
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<taxonomicName lsidName="horni" pageId="45" pageNumber="46" rank="species" species="horni">horni</taxonomicName>
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were recorded from Taiwan (Taiwan being the type locality of
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<taxonomicName genus="C." lsidName="C." pageId="45" pageNumber="46" rank="genus">C.</taxonomicName>
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(
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<taxonomicName lsidName="(C.)" pageId="45" pageNumber="46" rank="subGenus" subGenus="C.">C.</taxonomicName>
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)
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<taxonomicName lsidName="horni" pageId="45" pageNumber="46" rank="species" species="horni">horni</taxonomicName>
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<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
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). From the images and illustrations, it appeared that two species were superficially very similar so it is possible that they might be conspecific. All of the major collections in Taiwan were examined, and most known Taiwanese material was borrowed. Fresh material was collected from the wild. The holotype of
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<taxonomicName genus="C." lsidName="C." pageId="45" pageNumber="46" rank="genus">C.</taxonomicName>
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(
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<taxonomicName lsidName="(C.)" pageId="45" pageNumber="46" rank="subGenus" subGenus="C.">C.</taxonomicName>
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)
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<taxonomicName lsidName="horni" pageId="45" pageNumber="46" rank="species" species="horni">horni</taxonomicName>
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and three paratypes of
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<taxonomicName genus="C." lsidName="C." pageId="45" pageNumber="46" rank="genus">C.</taxonomicName>
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(
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<taxonomicName lsidName="(C.)" pageId="45" pageNumber="46" rank="subGenus" subGenus="C.">C.</taxonomicName>
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)
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<taxonomicName lsidName="mirabilis" pageId="45" pageNumber="46" rank="species" species="mirabilis">mirabilis</taxonomicName>
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are deposited SDEI, Germany where, upon examination, it was clear that the species could be distinguished from one another but that it was more of a total collection of subtleties than any one obvious external character that set them apart.
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<taxonomicName class="Insecta" family="Carabidae" genus="Catascopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Catascopus" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">Catascopus</taxonomicName>
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(
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<taxonomicName lsidName="(C.)" pageId="45" pageNumber="46" rank="subGenus" subGenus="C.">C.</taxonomicName>
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)
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<taxonomicName lsidName="mirabilis" pageId="45" pageNumber="46" rank="species" species="mirabilis">mirabilis</taxonomicName>
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differs from
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<taxonomicName genus="C." lsidName="C." pageId="45" pageNumber="46" rank="genus">C.</taxonomicName>
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(
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<taxonomicName lsidName="(C.)" pageId="45" pageNumber="46" rank="subGenus" subGenus="C.">C.</taxonomicName>
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)
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<taxonomicName lsidName="horni" pageId="45" pageNumber="46" rank="species" species="horni">horni</taxonomicName>
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in that it has slightly larger eyes, a slightly more cupreous sheen, apical and basal angles of pronotum that are that are slightly more lobed and explanate, elytral microsculpture that is more granulate and raised, giving a duller appearance, and broken striae (primarily 3, 5, 7) that are somewhat more raised. All of these characters are slightly variable and difficult to define so it became clear that without a lot of material to compare, confusing these two species would be easy. After going through the material however, a few good characters were consistent and allowed for discrimination of species.
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<taxonomicName genus="C." lsidName="C." pageId="45" pageNumber="46" rank="genus">C.</taxonomicName>
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(
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<taxonomicName lsidName="(C.)" pageId="45" pageNumber="46" rank="subGenus" subGenus="C.">C.</taxonomicName>
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)
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<taxonomicName lsidName="mirabilis" pageId="45" pageNumber="46" rank="species" species="mirabilis">mirabilis</taxonomicName>
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has females with the ventral surface of the fore-femur with a dense field of yellow setae (more than 20), while
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<taxonomicName genus="C." lsidName="C." pageId="45" pageNumber="46" rank="genus">C.</taxonomicName>
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(
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<taxonomicName lsidName="(C.)" pageId="45" pageNumber="46" rank="subGenus" subGenus="C.">C.</taxonomicName>
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)
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<taxonomicName lsidName="horni" pageId="45" pageNumber="46" rank="species" species="horni">horni</taxonomicName>
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only has a field of ~twelve setae. Females of
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<taxonomicName genus="C." lsidName="C." pageId="45" pageNumber="46" rank="genus">C.</taxonomicName>
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(
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<taxonomicName lsidName="(C.)" pageId="45" pageNumber="46" rank="subGenus" subGenus="C.">C.</taxonomicName>
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)
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<taxonomicName lsidName="mirabilis" pageId="45" pageNumber="46" rank="species" species="mirabilis">mirabilis</taxonomicName>
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also have a genitalic character, the bursa copulatrix sclerite (bsc, Fig. 33C), that makes them easily distinguishable from
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<taxonomicName genus="C." lsidName="C." pageId="45" pageNumber="46" rank="genus">C.</taxonomicName>
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(
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<taxonomicName lsidName="(C.)" pageId="45" pageNumber="46" rank="subGenus" subGenus="C.">C.</taxonomicName>
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)
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<taxonomicName lsidName="horni" pageId="45" pageNumber="46" rank="species" species="horni">horni</taxonomicName>
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. After dissecting the available material including the specimens that were examined and thought to be
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<taxonomicName genus="C." lsidName="C." pageId="45" pageNumber="46" rank="genus">C.</taxonomicName>
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(
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<taxonomicName lsidName="(C.)" pageId="45" pageNumber="46" rank="subGenus" subGenus="C.">C.</taxonomicName>
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)
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<taxonomicName lsidName="mirabilis" pageId="45" pageNumber="46" rank="species" species="mirabilis">mirabilis</taxonomicName>
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by both
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<bibRefCitation pageId="45" pageNumber="46">
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<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
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(1963)
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</bibRefCitation>
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and
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<bibRefCitation author="Habu, A" journalOrPublisher="Tokyo Electrical Engineering College Press, Tokyo" pageId="151" pageNumber="152" title="Carabidae. Truncatipennes group. (Insecta: Coleoptera). Fauna Japonica." year="1967">Habu (1967)</bibRefCitation>
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, it seems apparent that
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<taxonomicName genus="C." lsidName="C." pageId="45" pageNumber="46" rank="genus">C.</taxonomicName>
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(
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<taxonomicName lsidName="(C.)" pageId="45" pageNumber="46" rank="subGenus" subGenus="C.">C.</taxonomicName>
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)
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<taxonomicName lsidName="horni" pageId="45" pageNumber="46" rank="species" species="horni">horni</taxonomicName>
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is restricted to Taiwan while
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<taxonomicName genus="C." lsidName="C." pageId="45" pageNumber="46" rank="genus">C.</taxonomicName>
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(
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<taxonomicName lsidName="(C.)" pageId="45" pageNumber="46" rank="subGenus" subGenus="C.">C.</taxonomicName>
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)
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<taxonomicName lsidName="mirabilis" pageId="45" pageNumber="46" rank="species" species="mirabilis">mirabilis</taxonomicName>
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is known from Vietnam, Laos and China. See also "Geographical distribution".
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</paragraph>
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</subSubSection>
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<subSubSection pageId="45" pageNumber="46" type="diagnosis">
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<paragraph pageId="45" pageNumber="46">Diagnosis.</paragraph>
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<paragraph pageId="45" pageNumber="46">Specimens of this species are easily distinguished from other pericalines by the distinctly asymmetrical mandibles and metallic black dorsal coloration.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="49" lastPageNumber="50" pageId="45" pageNumber="46" type="redescription">
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<paragraph pageId="45" pageNumber="46">Redescription.</paragraph>
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<paragraph pageId="45" pageNumber="46">OBL 15.5 - 19 mm. Length (n = seven males, ten females): head 1.24 - 1.56, pronotum, 2.76 - 3.44, elytra 8.25 - 10.50, metepisternum 2.08 - 2.68 mm; width: head 3.00 - 3.72, pronotum 2.92 - 3.76, elytra 5.00 - 5.83, metepisternum 1.08 - 1.20 mm.</paragraph>
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<paragraph pageId="45" pageNumber="46">Body proportions. HW/HL 2.35 - 2.49; PWM/PL 1.07 - 1.16; EL/EW 1.65 - 1.83; ML/MW 1.93 - 2.31.</paragraph>
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<paragraph pageId="45" pageNumber="46">Color. Fig. 30. Dorsum of head brunneo-piceous to piceous, with faintly metallic violaceous sheen; antennae with segments 1-4 rufo-piceous to piceous, segments 5-11 brunneous to rufo-piceous; palpi rufo-brunneous to rufo-piceous; clypeus and mentum rufo-piceous; pronotum piceous, with faintly metallic violaceous sheen; elytra piceous, with faintly metallic violaceous sheen; elytral epipleura piceous; thoracic sclerites rufo-piceous to piceous; abdominal sterna rufo-piceous medially, rufo-piceous to piceous at lateral margins; legs with trochanter and femora rufo-piceous, tibia rufo-piceous to piceous.</paragraph>
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<caption pageId="45" pageNumber="46">
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<paragraph pageId="45" pageNumber="46">
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Figure 30. Dorsal habitus and color pattern of
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<taxonomicName lsidName="C. (Catascopoides) horni" pageId="45" pageNumber="46" rank="species" species="horni" subGenus="Catascopoides">C. (Catascopoides) horni</taxonomicName>
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<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
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(OBL 17.20 mm).
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</paragraph>
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</caption>
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<paragraph pageId="46" pageNumber="47">
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<pageBreakToken pageId="46" pageNumber="47" start="start">Microsculpture</pageBreakToken>
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. Dorsum of head with microsculpture not visible at 50
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<normalizedToken originalValue="×">x</normalizedToken>
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magnification; pronotum with transverse mesh pattern faintly visible at 50
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<normalizedToken originalValue="×">x</normalizedToken>
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magnification; elytra with shallow, transverse sculpticells on majority of disc, lower depressions of striae are nearly isodiametric, easily visible at 50
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<normalizedToken originalValue="×">x</normalizedToken>
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magnification; ventral surface of head, prosternum, proepipleuron, mesepisternum, and metepisternum with sculpticells forming a shallow transverse mesh.
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</paragraph>
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<paragraph pageId="46" pageNumber="47">Macrosculpture. Dorsum of head with scattered punctures laterally in front of eye to mid-way between basal supraorbital setae and pronotum apex, punctures near base of head are deep and confluent, more separated and shallow towards apex; pronotum rugulose; elytra with 3-4 diagonal impressions on disc, impressions evenly spaced with first impression near lateroapical angle of elytral base to suture, joining suture ~1/3 from base, first and third impressions deeper than others; intervals carinate, intervals 3,5,7, more strongly carinate in basal half than others, intervals convex to flat where interrupted by diagonal impressions; intervals 7 and 8 distinctly carinate and rounded at apical 1/4, disrupting normal contour of elytra; striae punctate along length; ventrally: prosternum, prosternal process, mesosternum, mesocoxa, and mesosternal intercoxal process, metasternum, hind coxa and base of abdominal sternite 3 with scattered and deep punctation, often confluent and rugulose in appearance; metasternum with several lateral striations on either side of suture (comb-like in appearance); abdominal sterna with scattered, shallow punctures; fore femur of males with more than twenty deep punctures in basal half of ventral surface, females with less (eight to fifteen), not all bearing setae.</paragraph>
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<paragraph pageId="46" pageNumber="47">Pilosity. Fig. 31. Dorsum of head, pronotum and disc of elytra with scattered micro-punctures; ventrally: prosternum, prosternal process, mesosternum, mesocoxa, and mesosternal intercoxal process and metasternum with moderate to long, blonde, setae associated with deep punctures; fore and mid femora of males with long, blonde setae associated with each puncture; fore and mid femora of females with several (fewer than males) blonde setae of various lengths in most punctures, some punctures without setae.</paragraph>
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<caption pageId="46" pageNumber="47">
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<paragraph pageId="46" pageNumber="47">
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Figure 31. Digital image of the upper ventral surface and associated seta pattern of
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<taxonomicName class="Insecta" family="Carabidae" genus="Catascopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Catascopus (Catascopoides) horni" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="horni" subGenus="Catascopoides">Catascopus (Catascopoides) horni</taxonomicName>
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<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
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.
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</paragraph>
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</caption>
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<paragraph pageId="46" pageNumber="47">Fixed setae. Two pairs of supraorbital setae; clypeus with two lateral setae; labrum with six setae along apical margin; one pair of suborbital setae; pronotum with one seta at base of lateral margin; elytra with one seta at basal quarter of interval 3, one seta in interval 3 at mid-length, one seta in apical quarter of interval 3; 16-17 lateral (umbilical) setae in interval 9; two setae on each of abdominal sterna III to VI, two setae along apical margin of sternum VII in males, four setae along apical margin of sternum VII.</paragraph>
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<paragraph pageId="46" pageNumber="47">Luster. Head capsule, pronotum and elytra moderately glossy to glossy; ventral thoracic sterna and abdominal sterna moderately glossy.</paragraph>
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<paragraph pageId="46" pageNumber="47">Head. Fig. 23B. Mandible long and almost straight, obtuse tooth near apex of inside margin of each mandible, tooth of left mandible larger and situated lower than right, left mandible with additional tooth on inside cutting surface above mid-length of mandible; labrum unevenly bilobed, right lobe always longer that left; mentum with single broad tooth; eyes convex; palpi cylindrical, elongate.</paragraph>
|
||
<paragraph pageId="47" pageNumber="48">
|
||
<pageBreakToken pageId="47" pageNumber="48" start="start">Pronotum</pageBreakToken>
|
||
. Anterior transverse impression moderately deep; posterior transverse impression and median longitudinal impression deep; apical margin narrowly curved forming short, acute latero-apical lobes; lateral margins sinuate toward base; posterio-lateral margins almost right-angled.
|
||
</paragraph>
|
||
<paragraph pageId="47" pageNumber="48">Elytra. Humeri broadly rounded; elytral margin shallowly impressed in basal 1/3; elytral apices each with one small lateral spine and two apical spines, outside apical spine always longer than inside apical spine.</paragraph>
|
||
<paragraph pageId="47" pageNumber="48">Hind wings. Macropterous.</paragraph>
|
||
<paragraph pageId="47" pageNumber="48">Legs. Tarsal claws smooth. Males with adhesive vestiture ventrally, two rows of squamo-setae on tarsomeres 1-3 of fore-leg.</paragraph>
|
||
<paragraph pageId="48" pageNumber="49">
|
||
<pageBreakToken pageId="48" pageNumber="49" start="start">Abdominal</pageBreakToken>
|
||
sterna. Abdominal sternum VII bilobed, with shallow notch apically.
|
||
</paragraph>
|
||
<paragraph pageId="48" pageNumber="49">
|
||
Male genitalia. Fig. 32
|
||
<normalizedToken originalValue="A–D">A-D</normalizedToken>
|
||
. Length 3.52 - 4.00 mm. Ostium left pleuropic. Phallus cylindrical, narrowest at base of shaft, expanded on left side from base towards median in ventral view, constricted again before apex; apical area distinctively long and narrow, apical curve visible in left and right view. Endophallus relatively long and distinctly curled left when viewed from left lateral aspect; sclerite near endophallus base, left side of ostium when viewed from left lateral aspect; one basal endophallic lobe near center of ostium opening; one meso-endophallic lobe on inside of the curled endophallus when viewed from left lateral aspect; two microtrichial fields present (see fig. 32A)
|
||
</paragraph>
|
||
<caption pageId="48" pageNumber="49">
|
||
<paragraph pageId="48" pageNumber="49">
|
||
Figure 32. Digital images of male genitalia of
|
||
<taxonomicName lsidName="C. (Catascopoides) horni" pageId="48" pageNumber="49" rank="species" species="horni" subGenus="Catascopoides">C. (Catascopoides) horni</taxonomicName>
|
||
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
|
||
A left lateral aspect endophallus everted B right lateral aspect C ventral aspect D left lateral aspect. Legend: el endophallic lobe; mtf microtrichial field.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="49" pageNumber="50">
|
||
<pageBreakToken pageId="49" pageNumber="50" start="start">Female</pageBreakToken>
|
||
genitalia. Figs 33A, 34D. Width 1.84 - 2.12 mm. Gonocoxite 2 (gc2) wide at base, narrowing sharply toward apex, highly curved; two lateral ensiform setae (les) and one dorsal ensiform seta (des) present (not visible from dorsal view). Sensory furrow, furrow pegs and associated nematiform setae not observed; Bursa copulatrix highly textured with many infoldings; large, distinctively shaped sclerite (bsc) (type 1) internally at base, between apex of lateral tergites (not visible from dorsal view), differing from the bursal sclerite of
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Horniulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horniulus andrewesi" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="andrewesi">Horniulus andrewesi</taxonomicName>
|
||
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
|
||
(Fig. 94B) in that the sclerite appears to be inside the tissue of the dorsal surface of the bursa and not open to the interior cavity; one spermatheca (sp1) long and narrow; One spermathecal accessory gland (sg) long and narrow; spermathecal gland duct (sgd) very long and narrow, attachment site at base of spermatheca.
|
||
</paragraph>
|
||
<caption pageId="49" pageNumber="50">
|
||
<paragraph pageId="49" pageNumber="50">
|
||
Figure 33. Line drawings comparing the bursal sclerite, ventral aspect of A
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Catascopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Catascopus (Catascopoides) horni" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="horni" subGenus="Catascopoides">Catascopus (Catascopoides) horni</taxonomicName>
|
||
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
|
||
B
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Catascopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Catascopus" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="genus">Catascopus</taxonomicName>
|
||
(s. str.)
|
||
<taxonomicName lsidName="sauteri" pageId="49" pageNumber="50" rank="species" species="sauteri">sauteri</taxonomicName>
|
||
Dupuis C
|
||
<taxonomicName genus="C." lsidName="C." pageId="49" pageNumber="50" rank="genus">C.</taxonomicName>
|
||
(
|
||
<taxonomicName lsidName="(Catascopoides)" pageId="49" pageNumber="50" rank="subGenus" subGenus="Catascopoides">Catascopoides</taxonomicName>
|
||
)
|
||
<taxonomicName lsidName="mirabilis" pageId="49" pageNumber="50" rank="species" species="mirabilis">mirabilis</taxonomicName>
|
||
Bates (not present in Taiwan) Legend: bc bursa copulatrix; bsc bursal sclerite (type 1); lt lateral tergites.
|
||
</paragraph>
|
||
</caption>
|
||
<caption pageId="49" pageNumber="50">
|
||
<paragraph pageId="49" pageNumber="50">
|
||
Figure 34. Line drawings of the female reproductive tract of species of the genus
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Catascopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Catascopus" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="genus">Catascopus</taxonomicName>
|
||
Kirby, known from Taiwan, ventral aspect. A
|
||
<taxonomicName genus="C." lsidName="C." pageId="49" pageNumber="50" rank="genus">C.</taxonomicName>
|
||
(s. str.)
|
||
<taxonomicName lsidName="ignicinctus" pageId="49" pageNumber="50" rank="species" species="ignicinctus">ignicinctus</taxonomicName>
|
||
Bates B
|
||
<taxonomicName genus="C." lsidName="C." pageId="49" pageNumber="50" rank="genus">C.</taxonomicName>
|
||
(s. str.)
|
||
<taxonomicName lsidName="sauteri" pageId="49" pageNumber="50" rank="species" species="sauteri">sauteri</taxonomicName>
|
||
Dupuis C
|
||
<taxonomicName genus="C." lsidName="C." pageId="49" pageNumber="50" rank="genus">C.</taxonomicName>
|
||
(s. str.)
|
||
<taxonomicName lsidName="viridiorchis" pageId="49" pageNumber="50" rank="species" species="viridiorchis">viridiorchis</taxonomicName>
|
||
sp. n. D
|
||
<taxonomicName lsidName="C. (Catascopoides) horni" pageId="49" pageNumber="50" rank="species" species="horni" subGenus="Catascopoides">C. (Catascopoides) horni</taxonomicName>
|
||
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
|
||
. Legend: bc bursa copulatrix; bsc bursal sclerite; co common oviduct; des dorsal ensiform setae; div diverticulum; gc1 gonocoxite 1; gc2 gonocoxite 2; les lateral ensiform setae; lt lateral tergite; sg spermathecal gland; sgd spermathecal gland duct; sp1 spermatheca 1. Scale bars: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="49" pageNumber="50" type="habitat">
|
||
<paragraph pageId="49" pageNumber="50">Habitat, habits, and seasonal occurrence.</paragraph>
|
||
<paragraph pageId="49" pageNumber="50">
|
||
The known elevational range of
|
||
<taxonomicName lsidName="C. horni" pageId="49" pageNumber="50" rank="species" species="horni">C. horni</taxonomicName>
|
||
is from 300 to 2000 meters. Only two specimens have been collected below 1000 meters altitude, with most specimens being collected from 1500 to 2000 meters. Adults of this species are found in mixed primary and secondary forest of montane areas. Adults are crepuscular or nocturnal with most activity observed on trunks of fallen or dying trees at night. Specimens have been collected from March to December but are most commonly collected from March to June. Methods of collecting include u.v. light, sugar baits painted on tree trunks (have not been observed at actual bait, only near), hand collecting and malaise trap. The only confirmed tree species from which
|
||
<taxonomicName genus="C." lsidName="C." pageId="49" pageNumber="50" rank="genus">C.</taxonomicName>
|
||
(
|
||
<taxonomicName lsidName="(C.)" pageId="49" pageNumber="50" rank="subGenus" subGenus="C.">C.</taxonomicName>
|
||
)
|
||
<taxonomicName lsidName="horni" pageId="49" pageNumber="50" rank="species" species="horni">horni</taxonomicName>
|
||
has been collected is
|
||
<taxonomicName class="Pinopsida" family="Pinaceae" genus="Pinus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pinus morrisonicola" order="Pinales" pageId="49" pageNumber="50" phylum="Tracheophyta" rank="species" species="morrisonicola">Pinus morrisonicola</taxonomicName>
|
||
Hayata. Adults are very fast runners and when they are lit at night, they will quickly run to the dark side of the tree.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="49" pageNumber="50" type="distribution">
|
||
<paragraph pageId="49" pageNumber="50">Geographical distribution.</paragraph>
|
||
<paragraph pageId="49" pageNumber="50">
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Catascopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Catascopus horni" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="horni">Catascopus horni</taxonomicName>
|
||
is known only from Taiwan. See Figure 35.
|
||
</paragraph>
|
||
<caption pageId="49" pageNumber="50">
|
||
<paragraph pageId="49" pageNumber="50">
|
||
Figure 35. Map showing known localities for species of the genus
|
||
<taxonomicName authority="Kirby" class="Insecta" family="Carabidae" genus="Catascopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Catascopus" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="authority">Catascopus Kirby</taxonomicName>
|
||
and subgenus
|
||
<taxonomicName lsidName="(Catascopoides)" pageId="49" pageNumber="50" rank="subGenus" subGenus="Catascopoides">Catascopoides</taxonomicName>
|
||
Habu, in Taiwan.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |