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<mods:title id="AD9F10C5FBB50241B71834C26D216B55">Eusyllinae and “ Incertae sedis ” syllids (Annelida: Syllidae) from South America, with a new species from Brazil and a new combination for a Peruvian species</mods:title>
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<emphasis id="1FBAEA93FFF5FFC1FF68FF6AD4EFA5DB" bold="true" box="[151,414,151,177]" italics="true" pageId="19" pageNumber="526">Opisthodonta morena</emphasis>
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<figureCitation id="B5F52A04FFF5FFC1FF68FF47D446A5B9" box="[151,311,186,211]" captionStart-0="FIGURE 11" captionStart-1="FIGURE 12" captionStart-2="FIGURE 13" captionStartId-0="20.[151,250,1951,1973]" captionStartId-1="21.[151,250,1594,1616]" captionStartId-2="22.[151,250,1934,1956]" captionTargetBox-0="[184,1403,205,1926]" captionTargetBox-1="[156,1428,198,1565]" captionTargetBox-2="[156,1429,686,1906]" captionTargetId-0="figure@20.[165,1421,193,1930]" captionTargetId-1="figure@21.[152,1435,193,1572]" captionTargetId-2="figure@22.[152,1435,681,1912]" captionTargetPageId-0="20" captionTargetPageId-1="21" captionTargetPageId-2="22" captionText-0="FIGURE 11. Opisthodonta morena. (A) anterior body, dorsal view; (B) falcigers, anterior body; (C) falcigers, mid- and posterior body; (D) dorsal simple chaeta; (E) ventral simple chaeta; (F) aciculae, anterior body; (G) acicula, posterior body. Scale bars: A = 200 µm; B – G = 10 µm." captionText-1="FIGURE 12. Opisthodonta morena, SEM. (A) Anterior body, dorsal view; (B) prostomium, dorsal view; (C) posterior body, ventral view; (D) anterior body parapodia, dorsal view. Scale bars: A, C = 100 µm; B = 50 µm; D = 20 µm." captionText-2="FIGURE 13. Opisthodonta morena, SEM. (A) Anterior body, fronto-ventral view; (B) midbody, dorsal view; (C) anterior body, ventro-lateral view; (D) tufts of cilia on dorsum, midboy. Scale bars: A – B = 100 µm; C = 50 µm; D = 20 µm." httpUri-0="https://zenodo.org/record/233503/files/figure.png" httpUri-1="https://zenodo.org/record/233504/files/figure.png" httpUri-2="https://zenodo.org/record/233505/files/figure.png" pageId="19" pageNumber="526">Figures 11–13</figureCitation>
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, fig. 12; 
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–57, figs 15–16; 
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–308, figs 40–42A-B; 
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–63, fig. 16.
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1007), 
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Project ' 
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'. 
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2396), 
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7 
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2405), 
<date id="59701041FFF5FFC1FF19FD79D41CA7F6" box="[230,365,644,669]" pageId="19" pageNumber="526" value="2009-07-21">21 Jul 2009</date>
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<quantity id="EA369B64FFF5FFC1FD6AFD79D7A2A7F6" box="[661,723,644,669]" metricMagnitude="1" metricUnit="m" metricValue="5.2" pageId="19" pageNumber="526" unit="m" value="52.0">52 m</quantity>
: 1 spec. (ZUEC-POL 16127), 
<date id="59701041FFF5FFC1FBD6FD79D1C1A7F6" box="[1065,1200,644,669]" pageId="19" pageNumber="526" value="2009-03-05">
5 
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</date>
, and 1 spec. (ZUEC- POL 16136), 
<date id="59701041FFF5FFC1FECBFD55D4CCA7AA" box="[308,445,679,704]" pageId="19" pageNumber="526" value="2009-07-21">21 Jul 2009</date>
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: 1 spec. (ZUEC-POL 16137), 
<date id="59701041FFF5FFC1FB79FD55D07EA7AA" box="[1158,1295,679,704]" pageId="19" pageNumber="526" value="2009-07-21">21 Jul 2009</date>
; 
<geoCoordinate id="48FA5046FFF5FFC1FAE2FD55D0EDA7AA" box="[1309,1436,680,704]" direction="south" orientation="latitude" pageId="19" pageNumber="526" precision="15" value="-21.411943">21°24'43&quot;S</geoCoordinate>
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<quantity id="EA369B64FFF5FFC1FED1FD31D41CA78E" box="[302,365,716,740]" metricMagnitude="1" metricUnit="m" metricValue="3.3" pageId="19" pageNumber="526" unit="m" value="33.0">33 m</quantity>
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2386), 
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2398; ZUEC-POL 16144), 
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9 
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<geoCoordinate id="48FA5046FFF5FFC1FD5EFD0DD65BA662" box="[673,810,752,776]" direction="west" orientation="longitude" pageId="19" pageNumber="526" precision="15" value="-40.19167">40°11'30&quot;W</geoCoordinate>
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<quantity id="EA369B64FFF5FFC1FCC9FD0DD605A66D" box="[822,884,752,776]" metricMagnitude="1" metricUnit="m" metricValue="7.1" pageId="19" pageNumber="526" unit="m" value="71.0">71 m</quantity>
: 1 spec. (
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2404), 
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, 
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2387), 
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; 
<geoCoordinate id="48FA5046FFF5FFC1FCC7FCE9D6D8A646" box="[824,937,788,812]" direction="south" orientation="latitude" pageId="19" pageNumber="526" precision="15" value="-22.060555">22°3'38&quot;S</geoCoordinate>
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, 
<quantity id="EA369B64FFF5FFC1FBC1FCE9D10FA646" box="[1086,1150,788,812]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="19" pageNumber="526" unit="m" value="90.0">90 m</quantity>
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2403; ZUEC-POL 16145), 
<date id="59701041FFF5FFC1FE7BFCC5D48DA63A" box="[388,508,823,848]" pageId="19" pageNumber="526" value="2009-07-07">7 Jul 2009</date>
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<quantity id="EA369B64FFF5FFC1FCF5FCCAD636A625" box="[778,839,823,848]" metricMagnitude="1" metricUnit="m" metricValue="7.5" pageId="19" pageNumber="526" unit="m" value="75.0">75 m</quantity>
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2394; ZUEC-POL 16125), 
<date id="59701041FFF5FFC1FAB0FCCAD5A3A61E" pageId="19" pageNumber="526" value="2009-02-25">25 Feb 2009</date>
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2402; ZUEC-POL 16133), 
<date id="59701041FFF5FFC1FCEDFCA1D6FCA61E" box="[786,909,860,885]" pageId="19" pageNumber="526" value="2009-07-06">6 Jul 2009</date>
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<geoCoordinate id="48FA5046FFF5FFC1FBEBFCA1D1F2A61F" box="[1044,1155,860,885]" direction="west" orientation="longitude" pageId="19" pageNumber="526" precision="15" value="-40.051388">40°3'5&quot;W</geoCoordinate>
, 
<quantity id="EA369B64FFF5FFC1FB70FCA1D1ADA61E" box="[1167,1244,860,885]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="19" pageNumber="526" unit="m" value="150.0">150 m</quantity>
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<date id="59701041FFF5FFC1FECAFC7DD4C2A6F2" box="[309,435,895,920]" pageId="19" pageNumber="526" value="2009-07-07">7 Jul 2009</date>
; 
<geoCoordinate id="48FA5046FFF5FFC1FE3EFC7DD731A6F2" box="[449,576,896,920]" direction="south" orientation="latitude" pageId="19" pageNumber="526" precision="15" value="-22.228334">22°13'42&quot;S</geoCoordinate>
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, 
<quantity id="EA369B64FFF5FFC1FD1DFC7DD653A6FD" box="[738,802,896,920]" metricMagnitude="1" metricUnit="m" metricValue="7.0" pageId="19" pageNumber="526" unit="m" value="70.0">70 m</quantity>
: 1 spec. (ZUEC-POL 16117), 
<date id="59701041FFF5FFC1FB7CFC7DD07FA6F2" box="[1155,1294,895,920]" pageId="19" pageNumber="526" value="2009-07-24">24 Jul 2009</date>
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<geoCoordinate id="48FA5046FFF5FFC1FAE2FC7DD0EDA6F2" box="[1309,1436,896,920]" direction="south" orientation="latitude" pageId="19" pageNumber="526" precision="15" value="-22.295">22°17'42&quot;S</geoCoordinate>
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<quantity id="EA369B64FFF5FFC1FED3FC59D406A6D6" box="[300,375,932,956]" metricMagnitude="2" metricUnit="m" metricValue="1.04" pageId="19" pageNumber="526" unit="m" value="104.0">104 m</quantity>
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2395), 
<date id="59701041FFF5FFC1FD6EFC59D66EA6D6" box="[657,799,932,957]" pageId="19" pageNumber="526" value="2009-02-23">23 Feb 2009</date>
, and 9 specs (
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2401; ZUEC-POL 16131, 16143), 
<date id="59701041FFF5FFC1FF68FC35D47EA68A" box="[151,271,967,992]" pageId="19" pageNumber="526" value="2009-07-04">4 Jul 2009</date>
; 
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<quantity id="EA369B64FFF5FFC1FDE3FC3AD716A6B5" box="[540,615,967,992]" metricMagnitude="2" metricUnit="m" metricValue="3.86" pageId="19" pageNumber="526" unit="m" value="386.0">386 m</quantity>
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2385), 
<date id="59701041FFF5FFC1FC7AFC35D161A68A" box="[901,1040,968,992]" pageId="19" pageNumber="526" value="2009-01-30">30 Jan 2009</date>
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<quantity id="EA369B64FFF5FFC1FAC7FC3AD004A6B5" box="[1336,1397,967,992]" metricMagnitude="1" metricUnit="m" metricValue="7.5" pageId="19" pageNumber="526" unit="m" value="75.0">75 m</quantity>
: 12 specs (
<collectionCode id="4BDFAE44FFF5FFC1FF1BFC11D432A16E" box="[228,323,1004,1028]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="19" pageNumber="526">MZUSP</collectionCode>
2393; ZUEC-POL 16126), 
<date id="59701041FFF5FFC1FD79FC11D66FA16E" box="[646,798,1004,1029]" pageId="19" pageNumber="526" value="2009-03-15">
15 
<bibRefCitation id="495F4B70FFF5FFC1FD54FC10D66FA16E" author="Mar" box="[683,798,1004,1028]" pageId="19" pageNumber="526" refString="San Martin, G., Lopez, E. &amp; Aguado, M. T. (2009) Revision of the genus Pionosyllis (Polychaeta: Syllidae: Eusyllinae), with a cladistic analysis, and the description of five new genera and two new species. Journal of the Marine Biological Association of the United Kingdom, 89, 1455 - 1498. http: // dx. doi. org / 10.1017 / S 0025315409003099" type="journal article" year="2009">Mar 2009</bibRefCitation>
</date>
, and 8 specs (
<collectionCode id="4BDFAE44FFF5FFC1FC3DFC11D150A16E" box="[962,1057,1004,1028]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="19" pageNumber="526">MZUSP</collectionCode>
2400; ZUEC-POL 16129), 
<date id="59701041FFF5FFC1FA9BFC11D5A2A142" pageId="19" pageNumber="526" value="2009-07-04">4 Jul 2009</date>
; 
<geoCoordinate id="48FA5046FFF5FFC1FF21FBEDD42CA142" box="[222,349,1040,1064]" direction="south" orientation="latitude" pageId="19" pageNumber="526" precision="15" value="-22.389166">22°23'21&quot;S</geoCoordinate>
<geoCoordinate id="48FA5046FFF5FFC1FE9AFBF2D49EA142" box="[357,495,1039,1064]" direction="west" orientation="longitude" pageId="19" pageNumber="526" precision="15" value="-40.5825">40°34'57&quot;W</geoCoordinate>
, 
<quantity id="EA369B64FFF5FFC1FE04FBEDD737A142" box="[507,582,1040,1064]" metricMagnitude="2" metricUnit="m" metricValue="1.1" pageId="19" pageNumber="526" unit="m" value="110.0">110 m</quantity>
: 1 spec. (ZUEC-POL 16135), 
<date id="59701041FFF5FFC1FC62FBF2D155A142" box="[925,1060,1039,1064]" pageId="19" pageNumber="526" value="2009-07-25">25 Jul 2009</date>
; 
<geoCoordinate id="48FA5046FFF5FFC1FBCFFBEDD1DEA142" box="[1072,1199,1040,1064]" direction="south" orientation="latitude" pageId="19" pageNumber="526" precision="15" value="-22.394167">22°23'39&quot;S</geoCoordinate>
<geoCoordinate id="48FA5046FFF5FFC1FB48FBEDD030A142" box="[1207,1345,1040,1064]" direction="west" orientation="longitude" pageId="19" pageNumber="526" precision="15" value="-40.344444">40°20'40&quot;W</geoCoordinate>
, 
<quantity id="EA369B64FFF5FFC1FAB2FBF2D0E8A142" box="[1357,1433,1039,1064]" metricMagnitude="2" metricUnit="m" metricValue="1.53" pageId="19" pageNumber="526" unit="m" value="153.0">153 m</quantity>
: 1 spec. (ZUEC-POL 16124), 
<date id="59701041FFF5FFC1FE18FBC9D708A126" box="[487,633,1076,1101]" pageId="19" pageNumber="526" value="2009-02-23">23 Feb 2009</date>
, and 2 specs (ZUEC-POL 16132), 
<date id="59701041FFF5FFC1FBF4FBC9D1F7A126" box="[1035,1158,1076,1101]" pageId="19" pageNumber="526" value="2009-07-04">4 Jul 2009</date>
; 
<geoCoordinate id="48FA5046FFF5FFC1FB6BFBC9D074A126" box="[1172,1285,1076,1100]" direction="south" orientation="latitude" pageId="19" pageNumber="526" precision="15" value="-22.51861">22°31'7&quot;S</geoCoordinate>
<geoCoordinate id="48FA5046FFF5FFC1FAF1FBC9D0E9A126" box="[1294,1432,1076,1100]" direction="west" orientation="longitude" pageId="19" pageNumber="526" precision="15" value="-40.525555">40°31'32&quot;W</geoCoordinate>
, 
<quantity id="EA369B64FFF5FFC1FF68FBA5D597A11A" box="[151,230,1112,1136]" metricMagnitude="2" metricUnit="m" metricValue="1.39" pageId="19" pageNumber="526" unit="m" value="139.0">139 m</quantity>
: 1 spec. (ZUEC-POL 16123), 
<date id="59701041FFF5FFC1FDAEFBA5D798A11A" box="[593,745,1111,1136]" pageId="19" pageNumber="526" value="2009-02-23">23 Feb 2009</date>
, and 2 specs (ZUEC-POL 16130), 
<date id="59701041FFF5FFC1FB73FBA5D07CA11A" box="[1164,1293,1111,1136]" pageId="19" pageNumber="526" value="2009-07-03">3 Jul 2009</date>
; 
<geoCoordinate id="48FA5046FFF5FFC1FAE2FBAAD0EDA11A" box="[1309,1436,1111,1136]" direction="south" orientation="latitude" pageId="19" pageNumber="526" precision="15" value="-22.781666">22°46'54&quot;S</geoCoordinate>
<geoCoordinate id="48FA5046FFF5FFC1FF68FB81D462A1FE" box="[151,275,1148,1172]" direction="west" orientation="longitude" pageId="19" pageNumber="526" precision="15" value="-41.059166">41°3'33&quot;W</geoCoordinate>
, 
<quantity id="EA369B64FFF5FFC1FEE0FB80D42FA1FE" box="[287,350,1149,1172]" metricMagnitude="1" metricUnit="m" metricValue="7.7" pageId="19" pageNumber="526" unit="m" value="77.0">77 m</quantity>
: 3 specs (
<collectionCode id="4BDFAE44FFF5FFC1FE33FB81D75AA1FE" box="[460,555,1148,1172]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="19" pageNumber="526">MZUSP</collectionCode>
2392; ZUEC-POL 16122), 
<date id="59701041FFF5FFC1FC96FB81D68AA1FE" box="[873,1019,1148,1173]" pageId="19" pageNumber="526" value="2009-02-22">22 Feb 2009</date>
, and 3 specs (
<collectionCode id="4BDFAE44FFF5FFC1FB65FB81D188A1FE" box="[1178,1273,1148,1172]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="19" pageNumber="526">MZUSP</collectionCode>
2399; ZUEC- POL 16128), 
<date id="59701041FFF5FFC1FECCFB5DD4DCA1D2" box="[307,429,1183,1208]" pageId="19" pageNumber="526" value="2009-07-02">2 Jul 2009</date>
; 
<geoCoordinate id="48FA5046FFF5FFC1FE45FB5DD75AA1D2" box="[442,555,1184,1208]" direction="south" orientation="latitude" pageId="19" pageNumber="526" precision="15" value="-23.167778">23°10'4&quot;S</geoCoordinate>
<geoCoordinate id="48FA5046FFF5FFC1FDCBFB62D7D0A1D2" box="[564,673,1183,1208]" direction="west" orientation="longitude" pageId="19" pageNumber="526" precision="15" value="-41.051666">41°3'6&quot;W</geoCoordinate>
, 
<quantity id="EA369B64FFF5FFC1FD51FB62D78AA1D2" box="[686,763,1183,1208]" metricMagnitude="2" metricUnit="m" metricValue="1.05" pageId="19" pageNumber="526" unit="m" value="105.0">105 m</quantity>
: 4 specs (
<collectionCode id="4BDFAE44FFF5FFC1FC96FB5DD6B9A1D2" box="[873,968,1184,1208]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="19" pageNumber="526">MZUSP</collectionCode>
2388; ZUEC-POL 16118), 
<date id="59701041FFF5FFC1FAF8FB5DD0E9A1D2" box="[1287,1432,1183,1208]" pageId="19" pageNumber="526" value="2009-02-21">21 Feb 2009</date>
, and 3 specs (
<collectionCode id="4BDFAE44FFF5FFC1FECEFB39D4E1A1B6" box="[305,400,1220,1244]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="19" pageNumber="526">MZUSP</collectionCode>
2389; ZUEC-POL 16119), 
<date id="59701041FFF5FFC1FD28FB39D627A1B6" box="[727,854,1220,1245]" pageId="19" pageNumber="526" value="2009-07-02">2 Jul 2009</date>
; 
<geoCoordinate id="48FA5046FFF5FFC1FC99FB39D695A1B6" box="[870,996,1220,1244]" direction="south" orientation="latitude" pageId="19" pageNumber="526" precision="15" value="-23.19111">23°11'28&quot;S</geoCoordinate>
<geoCoordinate id="48FA5046FFF5FFC1FC11FB39D11AA1B6" box="[1006,1131,1220,1244]" direction="west" orientation="longitude" pageId="19" pageNumber="526" precision="15" value="-41.01361">41°0'49&quot;W</geoCoordinate>
, 
<quantity id="EA369B64FFF5FFC1FB85FB39D1B6A1B6" box="[1146,1223,1220,1244]" metricMagnitude="2" metricUnit="m" metricValue="1.17" pageId="19" pageNumber="526" unit="m" value="117.0">117 m</quantity>
: 8 specs (
<collectionCode id="4BDFAE44FFF5FFC1FAC2FB39D0EDA1B6" box="[1341,1436,1220,1244]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="19" pageNumber="526">MZUSP</collectionCode>
2391; ZUEC-POL 16121), 
<date id="59701041FFF5FFC1FE28FB15D71EA06A" box="[471,623,1255,1280]" pageId="19" pageNumber="526" value="2009-02-21">21 Feb 2009</date>
; 
<geoCoordinate id="48FA5046FFF5FFC1FD80FB15D780A06A" box="[639,753,1256,1280]" direction="south" orientation="latitude" pageId="19" pageNumber="526" precision="15" value="-23.202223">23°12'8&quot;S</geoCoordinate>
<geoCoordinate id="48FA5046FFF5FFC1FD03FB1AD6F7A06A" box="[764,902,1255,1280]" direction="west" orientation="longitude" pageId="19" pageNumber="526" precision="15" value="-40.993057">40°59'35&quot;W</geoCoordinate>
, 
<quantity id="EA369B64FFF5FFC1FC68FB15D697A06A" box="[919,998,1256,1280]" metricMagnitude="2" metricUnit="m" metricValue="1.42" pageId="19" pageNumber="526" unit="m" value="142.0">142 m</quantity>
: 2 specs (
<collectionCode id="4BDFAE44FFF5FFC1FBA0FB15D1CFA06A" box="[1119,1214,1256,1280]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="19" pageNumber="526">MZUSP</collectionCode>
2390; ZUEC-POL 16120), 
<date id="59701041FFF5FFC1FF0BFAF1D41CA04E" box="[244,365,1292,1317]" pageId="19" pageNumber="526" value="2009-07-02">2 Jul 2009</date>
.
</paragraph>
<paragraph id="2D713681FFF5FFC1FF38FAD2D436A34A" blockId="19.[151,1437,356,2037]" pageId="19" pageNumber="526">
<emphasis id="1FBAEA93FFF5FFC1FF38FAD2D73FA022" bold="true" box="[199,590,1327,1352]" pageId="19" pageNumber="526">Additional material examined.</emphasis>
<taxonomicName id="EACE4D02FFF5FFC1FD9DFAD2D613A022" box="[610,866,1327,1352]" class="Polychaeta" family="Syllidae" genus="Opisthodonta" kingdom="Animalia" order="Phyllodocida" pageId="19" pageNumber="526" phylum="Annelida" rank="species" species="morena">
<emphasis id="1FBAEA93FFF5FFC1FD9DFAD2D613A022" box="[610,866,1327,1352]" italics="true" pageId="19" pageNumber="526">Opisthodonta morena</emphasis>
</taxonomicName>
—Atlantic Ocean, Josephine Bank (
<geoCoordinate id="48FA5046FFF5FFC1FAD6FAD2D0EDA022" box="[1321,1436,1327,1352]" direction="north" orientation="latitude" pageId="19" pageNumber="526" precision="92" value="36.678333">36°40.7'N</geoCoordinate>
<geoCoordinate id="48FA5046FFF5FFC1FF68FAA9D462A007" box="[151,275,1364,1389]" direction="west" orientation="longitude" pageId="19" pageNumber="526" precision="92" value="-14.258333">14°15.5'W</geoCoordinate>
): 17 specs (
<collectionCode id="4BDFAE44FFF5FFC1FE5FFAA9D491A006" box="[416,480,1364,1388]" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="19" pageNumber="526">ZMH</collectionCode>
<collectionCode id="4BDFAE44FFF5FFC1FE14FAA9D774A006" box="[491,517,1364,1388]" pageId="19" pageNumber="526">P-</collectionCode>
17193), coll. 'Meteor 9c', station 120, 0 
<date id="59701041FFF5FFC1FC29FAA9D123A007" box="[982,1106,1364,1389]" pageId="19" pageNumber="526" value="1967-07-01">1 Jul 1967</date>
, det. G. Hartmann-Schröder, 1979. 
<taxonomicName id="EACE4D02FFF5FFC1FF21FA8AD675A0FA" authority="Kudenov &amp; Harris, 1995" authorityName="Kudenov &amp; Harris" authorityYear="1995" box="[222,772,1399,1424]" class="Polychaeta" family="Syllidae" genus="Opisthodonta" kingdom="Animalia" order="Phyllodocida" pageId="19" pageNumber="526" phylum="Annelida" rank="species" species="mitchelli">
<emphasis id="1FBAEA93FFF5FFC1FF21FA8AD493A0FA" box="[222,482,1399,1424]" italics="true" pageId="19" pageNumber="526">Opisthodonta mitchelli</emphasis>
<bibRefCitation id="495F4B70FFF5FFC1FE15FA8AD675A0FA" author="Kudenov" box="[490,772,1399,1424]" pageId="19" pageNumber="526" refString="Kudenov, J. D. &amp; Harris, L. H. (1995) Family Syllidae Grube, 1850. In: Blake, J. A., Hilbig, B. &amp; Scott, P. H. (Eds.), Taxonomic Atlas of the Benthic Fauna of the Santa Maria Basin and Western Santa Barbara Channel. Vol. 5. The Annelida, part 2 - Polychaeta: Phyllodocida (Syllidae and Scale-Bearing Families), Amphinomida and Eunicida. Santa Barbara Museum of Natural History, Santa Barbara, pp. 1 - 97." type="book chapter" year="1995">Kudenov &amp; Harris, 1995</bibRefCitation>
</taxonomicName>
— 
<collectingCountry id="55D97611FFF5FFC1FCDDFA8AD62DA0FA" box="[802,860,1399,1424]" name="United States of America" pageId="19" pageNumber="526">USA</collectingCountry>
, California, Santa Maria Basin, off Point Arguello (
<geoCoordinate id="48FA5046FFF5FFC1FF60FA61D455A0DF" box="[159,292,1436,1461]" direction="north" orientation="latitude" pageId="19" pageNumber="526" precision="15" value="34.504166">34°30'15&quot;N</geoCoordinate>
<geoCoordinate id="48FA5046FFF5FFC1FED2FA61D4B8A0DF" box="[301,457,1436,1461]" direction="west" orientation="longitude" pageId="19" pageNumber="526" precision="15" value="-120.59278">120°35'34&quot;W</geoCoordinate>
), 
<quantity id="EA369B64FFF5FFC1FE20FA61D739A0DE" box="[479,584,1436,1461]" metricMagnitude="1" metricUnit="m" metricValue="5.85" metricValueMax="6.3" metricValueMin="5.4" pageId="19" pageNumber="526" unit="m" value="58.5" valueMax="63.0" valueMin="54.0">54–63 m</quantity>
: 1 spec. (
<typeStatus id="F2758823FFF5FFC1FD48FA61D66CA0DF" box="[695,797,1436,1461]" pageId="19" pageNumber="526" type="holotype">holotype</typeStatus>
, 
<collectionCode id="4BDFAE44FFF5FFC1FCD4FA61D60DA0DE" LSID="urn:lsid:biocol.org:col:34871" box="[811,892,1436,1460]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="19" pageNumber="526">USNM</collectionCode>
<date id="59701041FFF5FFC1FC7AFA61D6ACA0DE" box="[901,989,1436,1460]" pageId="19" pageNumber="526" value="19-09-17">170919</date>
), coll. 
<collectionCode id="4BDFAE44FFF5FFC1FBD2FA61D11CA0DE" box="[1069,1133,1436,1460]" pageId="19" pageNumber="526">SAIC</collectionCode>
/ Mbc BLM / MMS, R/V Diaphus DS, 
<date id="59701041FFF5FFC1FED1FA3DD4C5A0B2" box="[302,436,1471,1496]" pageId="19" pageNumber="526" value="1984-07-11">11 Jul 1984</date>
, det. 
<bibRefCitation id="495F4B70FFF5FFC1FE0EFA42D67DA0B2" author="Kudenov" box="[497,780,1471,1496]" pageId="19" pageNumber="526" refString="Kudenov, J. D. &amp; Harris, L. H. (1995) Family Syllidae Grube, 1850. In: Blake, J. A., Hilbig, B. &amp; Scott, P. H. (Eds.), Taxonomic Atlas of the Benthic Fauna of the Santa Maria Basin and Western Santa Barbara Channel. Vol. 5. The Annelida, part 2 - Polychaeta: Phyllodocida (Syllidae and Scale-Bearing Families), Amphinomida and Eunicida. Santa Barbara Museum of Natural History, Santa Barbara, pp. 1 - 97." type="book chapter" year="1995">Kudenov &amp; Harris, 1995</bibRefCitation>
; off Purisima Point (
<geoCoordinate id="48FA5046FFF5FFC1FC08FA42D10AA0B2" box="[1015,1147,1471,1496]" direction="north" orientation="latitude" pageId="19" pageNumber="526" precision="15" value="34.770554">34°46'14&quot;N</geoCoordinate>
<geoCoordinate id="48FA5046FFF5FFC1FB7CFA42D06EA0B2" box="[1155,1311,1471,1496]" direction="west" orientation="longitude" pageId="19" pageNumber="526" precision="15" value="-120.83278">120°49'58&quot;W</geoCoordinate>
), 
<quantity id="EA369B64FFF5FFC1FACCFA42D5C0A096" metricMagnitude="2" metricUnit="m" metricValue="1.0725" metricValueMax="1.23" metricValueMin="0.915" pageId="19" pageNumber="526" unit="m" value="107.25" valueMax="123.0" valueMin="91.5">91.5–123 m</quantity>
: 1 spec. (
<typeStatus id="F2758823FFF5FFC1FEE9FA19D40AA096" box="[278,379,1508,1532]" pageId="19" pageNumber="526" type="paratype">paratype</typeStatus>
, 
<collectionCode id="4BDFAE44FFF5FFC1FE79FA19D4A6A096" LSID="urn:lsid:biocol.org:col:34871" box="[390,471,1508,1532]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="19" pageNumber="526">USNM</collectionCode>
<date id="59701041FFF5FFC1FE22FA19D744A096" box="[477,565,1508,1532]" pageId="19" pageNumber="526" value="20-09-17">170920</date>
), coll. 
<collectionCode id="4BDFAE44FFF5FFC1FD80FA19D7CEA096" box="[639,703,1508,1532]" pageId="19" pageNumber="526">SAIC</collectionCode>
/ Mbc BLM / MMS, R/V Diaphus DS, 
<date id="59701041FFF5FFC1FB88FA19D18DA096" box="[1143,1276,1508,1533]" pageId="19" pageNumber="526" value="1984-07-23">23 Jul 1984</date>
, det. 
<bibRefCitation id="495F4B70FFF5FFC1FACAFA19D432A34A" author="Kudenov" pageId="19" pageNumber="526" refString="Kudenov, J. D. &amp; Harris, L. H. (1995) Family Syllidae Grube, 1850. In: Blake, J. A., Hilbig, B. &amp; Scott, P. H. (Eds.), Taxonomic Atlas of the Benthic Fauna of the Santa Maria Basin and Western Santa Barbara Channel. Vol. 5. The Annelida, part 2 - Polychaeta: Phyllodocida (Syllidae and Scale-Bearing Families), Amphinomida and Eunicida. Santa Barbara Museum of Natural History, Santa Barbara, pp. 1 - 97." type="book chapter" year="1995">Kudenov &amp; Harris, 1995</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="65D4650AFFF5FFC4FF38F9D1D7C9A4AC" lastPageId="22" lastPageNumber="529" pageId="19" pageNumber="526" type="description">
<paragraph id="2D713681FFF5FFC4FF38F9D1D7C9A4AC" blockId="19.[151,1437,356,2037]" lastBlockId="22.[151,1437,151,627]" lastPageId="22" lastPageNumber="529" pageId="19" pageNumber="526">
<emphasis id="1FBAEA93FFF5FFC1FF38F9D1D428A32F" bold="true" box="[199,345,1580,1605]" pageId="19" pageNumber="526">Description.</emphasis>
Medium-sized, fragile body, specimens examined mostly anterior fragments, with few chaetigers after proventricle segments; largest fragment 
<quantity id="EA369B64FFF5FFC1FD52F9B2D679A302" box="[685,776,1615,1640]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="19" pageNumber="526" unit="mm" value="2.6">2.6 mm</quantity>
long, 
<quantity id="EA369B64FFF5FFC1FCA6F9B2D6B3A302" box="[857,962,1615,1640]" metricMagnitude="-4" metricUnit="m" metricValue="5.2" pageId="19" pageNumber="526" unit="mm" value="0.52">0.52 mm</quantity>
wide, with 26 chaetigers; body without pigmentation patterns, dorsally with transverse rows of tufts of cilia, at least on midbody (
<figureCitation id="B5F52A04FFF5FFC1FB47F989D07CA3E6" box="[1208,1293,1652,1677]" captionStart="FIGURE 13" captionStartId="22.[151,250,1934,1956]" captionTargetBox="[156,1429,686,1906]" captionTargetId="figure@22.[152,1435,681,1912]" captionTargetPageId="22" captionText="FIGURE 13. Opisthodonta morena, SEM. (A) Anterior body, fronto-ventral view; (B) midbody, dorsal view; (C) anterior body, ventro-lateral view; (D) tufts of cilia on dorsum, midboy. Scale bars: A – B = 100 µm; C = 50 µm; D = 20 µm." httpUri="https://zenodo.org/record/233505/files/figure.png" pageId="19" pageNumber="526">Fig. 13</figureCitation>
B, D). Palps foliaceous, broader than prostomium, frequently ventrally folded (
<figureCitation id="B5F52A04FFF5FFC1FC6FF96AD696A3DA" box="[912,999,1687,1712]" captionStart="FIGURE 11" captionStartId="20.[151,250,1951,1973]" captionTargetBox="[184,1403,205,1926]" captionTargetId="figure@20.[165,1421,193,1930]" captionTargetPageId="20" captionText="FIGURE 11. Opisthodonta morena. (A) anterior body, dorsal view; (B) falcigers, anterior body; (C) falcigers, mid- and posterior body; (D) dorsal simple chaeta; (E) ventral simple chaeta; (F) aciculae, anterior body; (G) acicula, posterior body. Scale bars: A = 200 µm; B – G = 10 µm." httpUri="https://zenodo.org/record/233503/files/figure.png" pageId="19" pageNumber="526">Figs 11</figureCitation>
A; 12A–B; 13A). Prostomium oval to pentagonal, with 2 pairs of eyes in trapezoidal arrangement and 1 pair of anterior eyespots (
<figureCitation id="B5F52A04FFF5FFC1FB2CF941D057A3BE" box="[1235,1318,1724,1749]" captionStart="FIGURE 11" captionStartId="20.[151,250,1951,1973]" captionTargetBox="[184,1403,205,1926]" captionTargetId="figure@20.[165,1421,193,1930]" captionTargetPageId="20" captionText="FIGURE 11. Opisthodonta morena. (A) anterior body, dorsal view; (B) falcigers, anterior body; (C) falcigers, mid- and posterior body; (D) dorsal simple chaeta; (E) ventral simple chaeta; (F) aciculae, anterior body; (G) acicula, posterior body. Scale bars: A = 200 µm; B – G = 10 µm." httpUri="https://zenodo.org/record/233503/files/figure.png" pageId="19" pageNumber="526">Fig. 11</figureCitation>
A); lateral antennae inserted in front of anterior pair of eyes, near anterior margin of prostomium, up to twice as long as combined length of prostomium and palps; median antenna inserted centrally, almost twice as long as lateral antennae (
<figureCitation id="B5F52A04FFF5FFC1FEF6F8DAD42CA22A" box="[265,349,1831,1856]" captionStart="FIGURE 11" captionStartId="20.[151,250,1951,1973]" captionTargetBox="[184,1403,205,1926]" captionTargetId="figure@20.[165,1421,193,1930]" captionTargetPageId="20" captionText="FIGURE 11. Opisthodonta morena. (A) anterior body, dorsal view; (B) falcigers, anterior body; (C) falcigers, mid- and posterior body; (D) dorsal simple chaeta; (E) ventral simple chaeta; (F) aciculae, anterior body; (G) acicula, posterior body. Scale bars: A = 200 µm; B – G = 10 µm." httpUri="https://zenodo.org/record/233503/files/figure.png" pageId="19" pageNumber="526">Figs 11</figureCitation>
A; 12A–B); nuchal organs as paired, transverse, dorso-lateral rows of cilia at posterior margin of prostomium (
<figureCitation id="B5F52A04FFF5FFC1FED1F8B1D4F1A20E" box="[302,384,1868,1893]" captionStart="FIGURE 12" captionStartId="21.[151,250,1594,1616]" captionTargetBox="[156,1428,198,1565]" captionTargetId="figure@21.[152,1435,193,1572]" captionTargetPageId="21" captionText="FIGURE 12. Opisthodonta morena, SEM. (A) Anterior body, dorsal view; (B) prostomium, dorsal view; (C) posterior body, ventral view; (D) anterior body parapodia, dorsal view. Scale bars: A, C = 100 µm; B = 50 µm; D = 20 µm." httpUri="https://zenodo.org/record/233504/files/figure.png" pageId="19" pageNumber="526">Fig. 12</figureCitation>
A–B). Peristomium dorsally short, frequently covered by prostomium and chaetiger 1; dorsal peristomial cirri of approximately same size as median antenna, ventral peristomial cirri with approximately same size as lateral antennae or slightly longer (
<figureCitation id="B5F52A04FFF5FFC1FD70F869D796A2C6" box="[655,743,1940,1965]" captionStart="FIGURE 11" captionStartId="20.[151,250,1951,1973]" captionTargetBox="[184,1403,205,1926]" captionTargetId="figure@20.[165,1421,193,1930]" captionTargetPageId="20" captionText="FIGURE 11. Opisthodonta morena. (A) anterior body, dorsal view; (B) falcigers, anterior body; (C) falcigers, mid- and posterior body; (D) dorsal simple chaeta; (E) ventral simple chaeta; (F) aciculae, anterior body; (G) acicula, posterior body. Scale bars: A = 200 µm; B – G = 10 µm." httpUri="https://zenodo.org/record/233503/files/figure.png" pageId="19" pageNumber="526">Figs 11</figureCitation>
A; 12A–B). Dorsal cirri of anterior chaetigers, specially of chaetiger 1, long and slender, sometimes conspicuously longer than median antenna (
<figureCitation id="B5F52A04FFF5FFC1FB6BF845D19BA2BA" box="[1172,1258,1976,2001]" captionStart="FIGURE 11" captionStartId="20.[151,250,1951,1973]" captionTargetBox="[184,1403,205,1926]" captionTargetId="figure@20.[165,1421,193,1930]" captionTargetPageId="20" captionText="FIGURE 11. Opisthodonta morena. (A) anterior body, dorsal view; (B) falcigers, anterior body; (C) falcigers, mid- and posterior body; (D) dorsal simple chaeta; (E) ventral simple chaeta; (F) aciculae, anterior body; (G) acicula, posterior body. Scale bars: A = 200 µm; B – G = 10 µm." httpUri="https://zenodo.org/record/233503/files/figure.png" pageId="19" pageNumber="526">Fig. 11</figureCitation>
A); dorsal cirri alternating long, almost twice as long as body width, and short, slightly longer than half body width at corresponding chaetiger (
<figureCitation id="B5F52A04FFF3FFC7FE45F968D760A3C4" box="[442,529,1685,1710]" captionStart="FIGURE 11" captionStartId="20.[151,250,1951,1973]" captionTargetBox="[184,1403,205,1926]" captionTargetId="figure@20.[165,1421,193,1930]" captionTargetPageId="20" captionText="FIGURE 11. Opisthodonta morena. (A) anterior body, dorsal view; (B) falcigers, anterior body; (C) falcigers, mid- and posterior body; (D) dorsal simple chaeta; (E) ventral simple chaeta; (F) aciculae, anterior body; (G) acicula, posterior body. Scale bars: A = 200 µm; B – G = 10 µm." httpUri="https://zenodo.org/record/233503/files/figure.png" pageId="21" pageNumber="528">Figs 11</figureCitation>
A; 12A; 13B); ventral cirri large, ovate, partially fused to parapodial lobes on anterior segments (
<figureCitation id="B5F52A04FFF3FFC7FE8BF945D4B9A3BB" box="[372,456,1720,1745]" captionStart="FIGURE 13" captionStartId="22.[151,250,1934,1956]" captionTargetBox="[156,1429,686,1906]" captionTargetId="figure@22.[152,1435,681,1912]" captionTargetPageId="22" captionText="FIGURE 13. Opisthodonta morena, SEM. (A) Anterior body, fronto-ventral view; (B) midbody, dorsal view; (C) anterior body, ventro-lateral view; (D) tufts of cilia on dorsum, midboy. Scale bars: A – B = 100 µm; C = 50 µm; D = 20 µm." httpUri="https://zenodo.org/record/233505/files/figure.png" pageId="21" pageNumber="528">Fig. 13</figureCitation>
A, C); from proventricle segments onwards, ventral cirri short, conical, shorter than parapodial lobes and free from them (
<figureCitation id="B5F52A04FFF3FFC7FDC5F926D7FDA399" box="[570,652,1755,1780]" captionStart="FIGURE 12" captionStartId="21.[151,250,1594,1616]" captionTargetBox="[156,1428,198,1565]" captionTargetId="figure@21.[152,1435,193,1572]" captionTargetPageId="21" captionText="FIGURE 12. Opisthodonta morena, SEM. (A) Anterior body, dorsal view; (B) prostomium, dorsal view; (C) posterior body, ventral view; (D) anterior body parapodia, dorsal view. Scale bars: A, C = 100 µm; B = 50 µm; D = 20 µm." httpUri="https://zenodo.org/record/233504/files/figure.png" pageId="21" pageNumber="528">Fig. 12</figureCitation>
C). Antennae, peristomial and dorsal cirri throughout similar, long and slender, with short cirrophores (
<figureCitation id="B5F52A04FFF3FFC7FDECF900D71CA27C" box="[531,621,1789,1814]" captionStart="FIGURE 11" captionStartId="20.[151,250,1951,1973]" captionTargetBox="[184,1403,205,1926]" captionTargetId="figure@20.[165,1421,193,1930]" captionTargetPageId="20" captionText="FIGURE 11. Opisthodonta morena. (A) anterior body, dorsal view; (B) falcigers, anterior body; (C) falcigers, mid- and posterior body; (D) dorsal simple chaeta; (E) ventral simple chaeta; (F) aciculae, anterior body; (G) acicula, posterior body. Scale bars: A = 200 µm; B – G = 10 µm." httpUri="https://zenodo.org/record/233503/files/figure.png" pageId="21" pageNumber="528">Figs 11</figureCitation>
A; 12C; 13B). Anterior body parapodia with ~20–25 falcigers each, decreasing to ~5–10 falcigers per parapodium from midbody onwards; falcigers with shafts and blades spinulated, blades with spines slightly distally directed, bidentate; on anterior body, 2–4 dorsalmost falcigers with longer blades, ~30–22 µm long, teeth of even size or distal tooth slightly larger, rounded space between teeth; blades of remaining falcigers of anterior parapodia ~17–13 µm long, subdistal tooth of same size or slightly larger than distal tooth (
<figureCitation id="B5F52A04FFF3FFC7FF1DF856D44BA2A9" box="[226,314,1963,1988]" captionStart="FIGURE 11" captionStartId="20.[151,250,1951,1973]" captionTargetBox="[184,1403,205,1926]" captionTargetId="figure@20.[165,1421,193,1930]" captionTargetPageId="20" captionText="FIGURE 11. Opisthodonta morena. (A) anterior body, dorsal view; (B) falcigers, anterior body; (C) falcigers, mid- and posterior body; (D) dorsal simple chaeta; (E) ventral simple chaeta; (F) aciculae, anterior body; (G) acicula, posterior body. Scale bars: A = 200 µm; B – G = 10 µm." httpUri="https://zenodo.org/record/233503/files/figure.png" pageId="21" pageNumber="528">Figs 11</figureCitation>
B; 12D); on midbody parapodia, 1–2 dorsalmost falcigers with blades 22–20 µm long, blades of remaining falcigers ~15 µm long, subdistal tooth progressively larger and more hooked towards posterior body (
<figureCitation id="B5F52A04FFF0FFC4FF60FF6AD583A5DA" box="[159,242,151,176]" captionStart="FIGURE 11" captionStartId="20.[151,250,1951,1973]" captionTargetBox="[184,1403,205,1926]" captionTargetId="figure@20.[165,1421,193,1930]" captionTargetPageId="20" captionText="FIGURE 11. Opisthodonta morena. (A) anterior body, dorsal view; (B) falcigers, anterior body; (C) falcigers, mid- and posterior body; (D) dorsal simple chaeta; (E) ventral simple chaeta; (F) aciculae, anterior body; (G) acicula, posterior body. Scale bars: A = 200 µm; B – G = 10 µm." httpUri="https://zenodo.org/record/233503/files/figure.png" pageId="22" pageNumber="529">Fig. 11</figureCitation>
C). Dorsal simple chaetae only present on posteriormost parapodia, thin, slightly spinulated subdistally, bidentate, teeth minute, of even size, with rounded space between teeth (
<figureCitation id="B5F52A04FFF0FFC4FC1FFF47D143A5B9" box="[992,1074,186,211]" captionStart="FIGURE 11" captionStartId="20.[151,250,1951,1973]" captionTargetBox="[184,1403,205,1926]" captionTargetId="figure@20.[165,1421,193,1930]" captionTargetPageId="20" captionText="FIGURE 11. Opisthodonta morena. (A) anterior body, dorsal view; (B) falcigers, anterior body; (C) falcigers, mid- and posterior body; (D) dorsal simple chaeta; (E) ventral simple chaeta; (F) aciculae, anterior body; (G) acicula, posterior body. Scale bars: A = 200 µm; B – G = 10 µm." httpUri="https://zenodo.org/record/233503/files/figure.png" pageId="22" pageNumber="529">Fig. 11</figureCitation>
D); ventral simple chaetae only present on posteriormost parapodia, sigmoid, bidentate, subdistal tooth larger (
<figureCitation id="B5F52A04FFF0FFC4FBF7FF20D129A59C" box="[1032,1112,221,246]" captionStart="FIGURE 11" captionStartId="20.[151,250,1951,1973]" captionTargetBox="[184,1403,205,1926]" captionTargetId="figure@20.[165,1421,193,1930]" captionTargetPageId="20" captionText="FIGURE 11. Opisthodonta morena. (A) anterior body, dorsal view; (B) falcigers, anterior body; (C) falcigers, mid- and posterior body; (D) dorsal simple chaeta; (E) ventral simple chaeta; (F) aciculae, anterior body; (G) acicula, posterior body. Scale bars: A = 200 µm; B – G = 10 µm." httpUri="https://zenodo.org/record/233503/files/figure.png" pageId="22" pageNumber="529">Fig. 11</figureCitation>
E); dorsal and ventral simple chaetae sometimes present from anterior body, in juveniles. Anterior body parapodia with up to 4 aciculae each (
<figureCitation id="B5F52A04FFF0FFC4FF60FEDFD59FA451" box="[159,238,290,315]" captionStart="FIGURE 11" captionStartId="20.[151,250,1951,1973]" captionTargetBox="[184,1403,205,1926]" captionTargetId="figure@20.[165,1421,193,1930]" captionTargetPageId="20" captionText="FIGURE 11. Opisthodonta morena. (A) anterior body, dorsal view; (B) falcigers, anterior body; (C) falcigers, mid- and posterior body; (D) dorsal simple chaeta; (E) ventral simple chaeta; (F) aciculae, anterior body; (G) acicula, posterior body. Scale bars: A = 200 µm; B – G = 10 µm." httpUri="https://zenodo.org/record/233503/files/figure.png" pageId="22" pageNumber="529">Fig. 11</figureCitation>
F), aciculae per parapodium diminishing towards posterior body, from midbody onwards only 1 acicula per parapodium (
<figureCitation id="B5F52A04FFF0FFC4FED2FEB8D40DA434" box="[301,380,325,350]" captionStart="FIGURE 11" captionStartId="20.[151,250,1951,1973]" captionTargetBox="[184,1403,205,1926]" captionTargetId="figure@20.[165,1421,193,1930]" captionTargetPageId="20" captionText="FIGURE 11. Opisthodonta morena. (A) anterior body, dorsal view; (B) falcigers, anterior body; (C) falcigers, mid- and posterior body; (D) dorsal simple chaeta; (E) ventral simple chaeta; (F) aciculae, anterior body; (G) acicula, posterior body. Scale bars: A = 200 µm; B – G = 10 µm." httpUri="https://zenodo.org/record/233503/files/figure.png" pageId="22" pageNumber="529">Fig. 11</figureCitation>
G); aciculae progressively thinner towards posterior body, distally tapering, with rounded tips protruding from parapodial lobes (
<figureCitation id="B5F52A04FFF0FFC4FDDEFE9AD705A4EA" box="[545,628,359,384]" captionStart="FIGURE 12" captionStartId="21.[151,250,1594,1616]" captionTargetBox="[156,1428,198,1565]" captionTargetId="figure@21.[152,1435,193,1572]" captionTargetPageId="21" captionText="FIGURE 12. Opisthodonta morena, SEM. (A) Anterior body, dorsal view; (B) prostomium, dorsal view; (C) posterior body, ventral view; (D) anterior body parapodia, dorsal view. Scale bars: A, C = 100 µm; B = 50 µm; D = 20 µm." httpUri="https://zenodo.org/record/233504/files/figure.png" pageId="22" pageNumber="529">Fig. 12</figureCitation>
D), more evidently on posterior body. Pharynx long and wide, through 8–12 segments, with large, conical to pyriform tooth near midlength or slightly anteriorly; proventricle through 7–8 segments, with ~23 muscle cell rows (
<figureCitation id="B5F52A04FFF0FFC4FDBBFE50D7E2A4AC" box="[580,659,429,454]" captionStart="FIGURE 11" captionStartId="20.[151,250,1951,1973]" captionTargetBox="[184,1403,205,1926]" captionTargetId="figure@20.[165,1421,193,1930]" captionTargetPageId="20" captionText="FIGURE 11. Opisthodonta morena. (A) anterior body, dorsal view; (B) falcigers, anterior body; (C) falcigers, mid- and posterior body; (D) dorsal simple chaeta; (E) ventral simple chaeta; (F) aciculae, anterior body; (G) acicula, posterior body. Scale bars: A = 200 µm; B – G = 10 µm." httpUri="https://zenodo.org/record/233503/files/figure.png" pageId="22" pageNumber="529">Fig. 11</figureCitation>
A).
</paragraph>
</subSubSection>
<caption id="79B16609FFF2FFC6FF68F862D759A298" httpUri="https://zenodo.org/record/233503/files/figure.png" pageId="20" pageNumber="527" targetBox="[184,1403,205,1926]" targetPageId="20">
<paragraph id="2D713681FFF2FFC6FF68F862D759A298" blockId="20.[151,1437,1951,2034]" pageId="20" pageNumber="527">
<emphasis id="1FBAEA93FFF2FFC6FF68F862D453A2DE" bold="true" box="[151,290,1951,1973]" pageId="20" pageNumber="527">FIGURE 11.</emphasis>
<taxonomicName id="EACE4D02FFF2FFC6FED2F862D77FA2DF" box="[301,526,1951,1973]" class="Polychaeta" family="Syllidae" genus="Opisthodonta" kingdom="Animalia" order="Phyllodocida" pageId="20" pageNumber="527" phylum="Annelida" rank="species" species="morena">
<emphasis id="1FBAEA93FFF2FFC6FED2F862D77FA2DF" box="[301,526,1951,1973]" italics="true" pageId="20" pageNumber="527">Opisthodonta morena</emphasis>
</taxonomicName>
. (A) anterior body, dorsal view; (B) falcigers, anterior body; (C) falcigers, mid- and posterior body; (D) dorsal simple chaeta; (E) ventral simple chaeta; (F) aciculae, anterior body; (G) acicula, posterior body. Scale bars: A = 200 µm; B–G = 10 µm.
</paragraph>
</caption>
<caption id="79B16609FFF3FFC7FF68F9C7D1A7A304" httpUri="https://zenodo.org/record/233504/files/figure.png" pageId="21" pageNumber="528" targetBox="[156,1428,198,1565]" targetPageId="21">
<paragraph id="2D713681FFF3FFC7FF68F9C7D1A7A304" blockId="21.[151,1436,1594,1646]" pageId="21" pageNumber="528">
<emphasis id="1FBAEA93FFF3FFC7FF68F9C7D450A325" bold="true" box="[151,289,1594,1616]" pageId="21" pageNumber="528">FIGURE 12.</emphasis>
<emphasis id="1FBAEA93FFF3FFC7FED7F9C7D77DA33A" box="[296,524,1594,1616]" italics="true" pageId="21" pageNumber="528">
<taxonomicName id="EACE4D02FFF3FFC7FED7F9C7D778A33A" box="[296,521,1594,1616]" class="Polychaeta" family="Syllidae" genus="Opisthodonta" kingdom="Animalia" order="Phyllodocida" pageId="21" pageNumber="528" phylum="Annelida" rank="species" species="morena">Opisthodonta morena</taxonomicName>
,
</emphasis>
SEM. (A) Anterior body, dorsal view; (B) prostomium, dorsal view; (C) posterior body, ventral view; (D) anterior body parapodia, dorsal view. Scale bars: A, C = 100 µm; B = 50 µm; D = 20 µm.
</paragraph>
</caption>
<subSubSection id="65D4650AFFF0FFC5FF38FE32D7E6A592" lastPageId="23" lastPageNumber="530" pageId="22" pageNumber="529" type="discussion">
<paragraph id="2D713681FFF0FFC4FF38FE32D193A719" blockId="22.[151,1437,151,627]" pageId="22" pageNumber="529">
<emphasis id="1FBAEA93FFF0FFC4FF38FE32D44AA482" bold="true" box="[199,315,463,488]" pageId="22" pageNumber="529">Remarks.</emphasis>
<taxonomicName id="EACE4D02FFF0FFC4FEB8FE32D730A482" box="[327,577,463,488]" class="Polychaeta" family="Syllidae" genus="Opisthodonta" kingdom="Animalia" order="Phyllodocida" pageId="22" pageNumber="529" phylum="Annelida" rank="species" species="morena">
<emphasis id="1FBAEA93FFF0FFC4FEB8FE32D730A482" box="[327,577,463,488]" italics="true" pageId="22" pageNumber="529">Opisthodonta morena</emphasis>
</taxonomicName>
closely resembles 
<taxonomicName id="EACE4D02FFF0FFC4FCDBFE2DD6DFA482" box="[804,942,463,488]" class="Polychaeta" family="Syllidae" genus="Opisthodonta" kingdom="Animalia" order="Phyllodocida" pageId="22" pageNumber="529" phylum="Annelida" rank="species" species="mitchelli">
<emphasis id="1FBAEA93FFF0FFC4FCDBFE2DD6DFA482" box="[804,942,463,488]" italics="true" pageId="22" pageNumber="529">O. mitchelli</emphasis>
</taxonomicName>
, from California (
<collectingCountry id="55D97611FFF0FFC4FB75FE2DD1B6A482" box="[1162,1223,464,488]" name="United States of America" pageId="22" pageNumber="529">USA</collectingCountry>
), but both can be differentiated mostly by posterior body features: in 
<taxonomicName id="EACE4D02FFF0FFC4FD28FE0ED62AA761" box="[727,859,498,523]" class="Polychaeta" family="Syllidae" genus="Opisthodonta" kingdom="Animalia" order="Phyllodocida" pageId="22" pageNumber="529" phylum="Annelida" rank="species" species="mitchelli">
<emphasis id="1FBAEA93FFF0FFC4FD28FE0ED62AA761" box="[727,859,498,523]" italics="true" pageId="22" pageNumber="529">O. mitchelli</emphasis>
</taxonomicName>
aciculae have tips slightly bent, and falciger blades have a finer spinulation and the subdistal tooth conspicuously larger than the distal tooth, which resembles an enlarged spine in some chaetae; on the other hand, in 
<taxonomicName id="EACE4D02FFF0FFC4FCF2FDC5D6F9A73A" box="[781,904,568,592]" class="Polychaeta" family="Syllidae" genus="Opisthodonta" kingdom="Animalia" order="Phyllodocida" pageId="22" pageNumber="529" phylum="Annelida" rank="species" species="morena">
<emphasis id="1FBAEA93FFF0FFC4FCF2FDC5D6F9A73A" box="[781,904,568,592]" italics="true" pageId="22" pageNumber="529">O. morena</emphasis>
</taxonomicName>
all aciculae are straight (
<figureCitation id="B5F52A04FFF0FFC4FB50FDCAD188A73A" box="[1199,1273,567,592]" captionStart="FIGURE 11" captionStartId="20.[151,250,1951,1973]" captionTargetBox="[184,1403,205,1926]" captionTargetId="figure@20.[165,1421,193,1930]" captionTargetPageId="20" captionText="FIGURE 11. Opisthodonta morena. (A) anterior body, dorsal view; (B) falcigers, anterior body; (C) falcigers, mid- and posterior body; (D) dorsal simple chaeta; (E) ventral simple chaeta; (F) aciculae, anterior body; (G) acicula, posterior body. Scale bars: A = 200 µm; B – G = 10 µm." httpUri="https://zenodo.org/record/233503/files/figure.png" pageId="22" pageNumber="529">Fig 11</figureCitation>
F–G), and the falciger blades have coarser spinulation and less pronounced difference in sizes between the teeth.
</paragraph>
<caption id="79B16609FFF0FFC4FF68F873D058A2A9" httpUri="https://zenodo.org/record/233505/files/figure.png" pageId="22" pageNumber="529" targetBox="[156,1429,686,1906]" targetPageId="22">
<paragraph id="2D713681FFF0FFC4FF68F873D058A2A9" blockId="22.[151,1436,1934,1987]" pageId="22" pageNumber="529">
<emphasis id="1FBAEA93FFF0FFC4FF68F873D452A2CE" bold="true" box="[151,291,1934,1956]" pageId="22" pageNumber="529">FIGURE 13.</emphasis>
<emphasis id="1FBAEA93FFF0FFC4FED3F873D762A2CF" box="[300,531,1934,1957]" italics="true" pageId="22" pageNumber="529">
<taxonomicName id="EACE4D02FFF0FFC4FED3F873D77FA2CF" box="[300,526,1934,1957]" class="Polychaeta" family="Syllidae" genus="Opisthodonta" kingdom="Animalia" order="Phyllodocida" pageId="22" pageNumber="529" phylum="Annelida" rank="species" species="morena">Opisthodonta morena</taxonomicName>
,
</emphasis>
SEM. (A) Anterior body, fronto-ventral view; (B) midbody, dorsal view; (C) anterior body, ventro-lateral view; (D) tufts of cilia on dorsum, midboy. Scale bars: A–B = 100 µm; C = 50 µm; D = 20 µm.
</paragraph>
</caption>
<paragraph id="2D713681FFF1FFC5FF38FF6AD7E6A592" blockId="23.[151,1436,151,357]" pageId="23" pageNumber="530">
Brazilian specimens of 
<taxonomicName id="EACE4D02FFF1FFC5FE27FF65D723A5DA" box="[472,594,152,176]" class="Polychaeta" family="Syllidae" genus="Opisthodonta" kingdom="Animalia" order="Phyllodocida" pageId="23" pageNumber="530" phylum="Annelida" rank="species" species="morena">
<emphasis id="1FBAEA93FFF1FFC5FE27FF65D723A5DA" box="[472,594,152,176]" italics="true" pageId="23" pageNumber="530">O. morena</emphasis>
</taxonomicName>
are virtually indistinguishible from those from close to the 
<typeStatus id="F2758823FFF1FFC5FAF2FF65D04CA5DA" box="[1293,1341,152,176]" pageId="23" pageNumber="530">type</typeStatus>
locality (Josephine Banks, Atlantic Ocean). As the species is reported from nearly all over the world (see below), it is a possible case of a complex of sibling-species.
</paragraph>
</subSubSection>
<subSubSection id="65D4650AFFF1FFC5FF38FEF9D636A477" box="[199,839,260,285]" pageId="23" pageNumber="530" type="materials_examined">
<paragraph id="2D713681FFF1FFC5FF38FEF9D636A477" blockId="23.[151,1436,151,357]" box="[199,839,260,285]" pageId="23" pageNumber="530">
<emphasis id="1FBAEA93FFF1FFC5FF38FEF9D414A477" bold="true" box="[199,357,260,285]" pageId="23" pageNumber="530">
<typeStatus id="F2758823FFF1FFC5FF38FEF9D473A477" box="[199,258,260,285]" pageId="23" pageNumber="530">Type</typeStatus>
locality.
</emphasis>
Atlantic Ocean: Portugal—Madeira Island
</paragraph>
</subSubSection>
<subSubSection id="65D4650AFFF1FFC5FF38FEDAD1A9A40E" pageId="23" pageNumber="530" type="distribution">
<paragraph id="2D713681FFF1FFC5FF38FEDAD1A9A40E" blockId="23.[151,1436,151,357]" pageId="23" pageNumber="530">
<emphasis id="1FBAEA93FFF1FFC5FF38FEDAD411A42A" bold="true" box="[199,352,295,320]" pageId="23" pageNumber="530">Distribution.</emphasis>
Pacific Ocean: 
<collectingCountry id="55D97611FFF1FFC5FDE7FEDAD70FA42A" box="[536,638,295,320]" name="Australia" pageId="23" pageNumber="530">Australia</collectingCountry>
(New South 
<collectingCountry id="55D97611FFF1FFC5FCE5FEDAD612A42A" box="[794,867,295,320]" name="United Kingdom" pageId="23" pageNumber="530">Wales</collectingCountry>
). Indic Ocean: 
<collectingCountry id="55D97611FFF1FFC5FBEDFEDAD108A42A" box="[1042,1145,295,320]" name="Australia" pageId="23" pageNumber="530">Australia</collectingCountry>
(Western 
<collectingCountry id="55D97611FFF1FFC5FB10FEDAD02BA42A" box="[1263,1370,295,320]" name="Australia" pageId="23" pageNumber="530">Australia</collectingCountry>
), Red Sea. Mediterranean Sea. Atlantic Ocean: Madeira Island. First occurrence for the Brazilian coast.
</paragraph>
</subSubSection>
</treatment>
</document>