221 lines
24 KiB
XML
221 lines
24 KiB
XML
<document id="231AED411AD31D7497A340E342DAFB01" ID-DOI="10.1080/00222930050020122" ID-ISSN="1464-5262" ID-Zenodo-Dep="4747979" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1620672552581" checkinUser="carolina" docAuthor="Gussmann S. M. & Scholtz, C. H." docDate="2000" docId="7B60F030FF90FFC001D1FCA5FC66DAA7" docLanguage="en" docName="JNATHIST.34.1045-1123.pdf.imf" docOrigin="Journal of Natural History 34" docStyle="DocumentStyle:B3570C21952C5CF44265488F0CE432D6.2:JNatHist.2003-2004.journal_article.2cover" docStyleId="B3570C21952C5CF44265488F0CE432D6" docStyleName="JNatHist.2003-2004.journal_article.2cover" docStyleVersion="2" docTitle="Namibiotrupes Krikken 1977" docType="treatment" docVersion="4" lastPageNumber="1061" masterDocId="87598848FF81FFD20079FFE2FFB0DF5E" masterDocTitle="Systematic revision of endemic southern African genera of Bolboceratinae (Coleoptera: Scarabaeoidea: Bolboceratidae)" masterLastPageNumber="1123" masterPageNumber="1045" pageNumber="1060" updateTime="1698914725550" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:mods id="1DF34127CE5A1CC4E8B0BC378AEA0B21" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="9F19A0F91DBAB6D34173444D1D3E74DD">
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<mods:title id="6BFBC395C1A349A8A1863308C92CFF7C">Systematic revision of endemic southern African genera of Bolboceratinae (Coleoptera: Scarabaeoidea: Bolboceratidae)</mods:title>
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<mods:namePart id="3465F17B8239D05B69165A18FF1733A1">Gussmann S. M.</mods:namePart>
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<mods:roleTerm id="4C6F42720F8B1E3E5453BCE08D078723">Author</mods:roleTerm>
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<mods:namePart id="CFB6E66EFC5EDD6C93BAD6F936339EE2">Scholtz, C. H.</mods:namePart>
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<mods:typeOfResource id="1A7D9E06B99F59F8BC166D7488D04541">text</mods:typeOfResource>
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<mods:titleInfo id="D597BF86AE707B5FE60A846E5E989CFD">
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<mods:title id="E33323C2A7C271C4A7016531C64B3B94">Journal of Natural History</mods:title>
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</mods:titleInfo>
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<mods:part id="789A367FDB19B85CBEFCD066BEC3D43E">
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<mods:date id="74B154B43C160356B629072D2C358C8F">2000</mods:date>
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<mods:detail id="629D77204F33DEA1C25654B22A1C3343" type="pubDate">
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<mods:number id="140228CF38E03B2890F8C3E23C2703AA">2000-12-31</mods:number>
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</mods:detail>
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<mods:detail id="24C263D4CA667531B2A2600ED59FAC1B" type="volume">
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<mods:number id="3AE2903AE55984C79D47BA9903D94370">34</mods:number>
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</mods:detail>
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<mods:extent id="A4BC9F700FCE912FBE06D21F5D5DA5B1" unit="page">
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<mods:start id="6EB384AD7C9B70CBBE51E1AC14C0C46B">1045</mods:start>
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<mods:end id="290519121C36B6EBF3B010E23371D355">1123</mods:end>
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<mods:classification id="5D6FD088842CD16EEC7A78478DEBDE4D">journal article</mods:classification>
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<mods:identifier id="23CA82A09C2160392F376EF7F80A41C4" type="DOI">10.1080/00222930050020122</mods:identifier>
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<mods:identifier id="5BCDF0F808D260E08D8EA1889D7DB322" type="ISSN">1464-5262</mods:identifier>
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<mods:identifier id="D190A2E43EAB0109DEF103D8278A02C2" type="Zenodo-Dep">4747979</mods:identifier>
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<treatment id="7B60F030FF90FFC001D1FCA5FC66DAA7" ID-DOI="http://doi.org/10.5281/zenodo.4747917" ID-Zenodo-Dep="4747917" LSID="urn:lsid:plazi:treatment:7B60F030FF90FFC001D1FCA5FC66DAA7" httpUri="http://treatment.plazi.org/id/7B60F030FF90FFC001D1FCA5FC66DAA7" lastPageId="18" lastPageNumber="1061" pageId="17" pageNumber="1060">
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<subSubSection id="BBD312ADFF90FFC301D1FCA5FD6CDC21" pageId="17" pageNumber="1060" type="nomenclature">
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<paragraph id="F3764126FF90FFC301D1FCA5FD01DC01" blockId="17.[79,893,839,1007]" box="[424,689,839,863]" pageId="17" pageNumber="1060">
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<heading id="A83EF64AFF90FFC301D1FCA5FD01DC01" box="[424,689,839,863]" centered="true" fontSize="10" level="2" pageId="17" pageNumber="1060" reason="3">
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<taxonomicName id="34C93AA5FF90FFC301D1FCA5FD01DC01" authority="Krikken" authorityName="Krikken" authorityYear="1977" box="[424,689,839,863]" class="Insecta" family="Bolboceratidae" genus="Namibiotrupes" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="1060" phylum="Arthropoda" rank="genus">
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<emphasis id="C1BD9D34FF90FFC301D1FCA5FDF9DC01" bold="true" box="[424,585,839,863]" italics="true" pageId="17" pageNumber="1060">Namibiotrupes</emphasis>
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Krikken
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="F3764126FF90FFC30107FC85FD6CDC21" blockId="17.[79,893,839,1007]" box="[382,732,871,895]" pageId="17" pageNumber="1060">(®gures 4a±g, 5a±h, 6a±f, 7, 8)</paragraph>
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</subSubSection>
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<subSubSection id="BBD312ADFF90FFC30036FC6BFD97DCC1" box="[79,551,905,927]" pageId="17" pageNumber="1060" type="reference_group">
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<paragraph id="F3764126FF90FFC30036FC6BFD97DCC1" blockId="17.[79,893,839,1007]" box="[79,551,905,927]" pageId="17" pageNumber="1060">
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<treatmentCitationGroup id="D3D96608FF90FFC30036FC6BFD97DCC1" box="[79,551,905,927]" pageId="17" pageNumber="1060">
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<treatmentCitation id="72686737FF90FFC30036FC6BFE0CDCC1" author="KRIKKEN, J." box="[79,444,905,927]" page="165" pageId="17" pageNumber="1060" year="1977">
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<taxonomicName id="34C93AA5FF90FFC30036FC6BFE0CDCC1" authority="Krikken, 1977 a: 165" authorityName="Krikken" authorityPageNumber="165" authorityYear="1977" box="[79,444,905,927]" class="Insecta" family="Bolboceratidae" genus="Namibiotrupes" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="1060" phylum="Arthropoda" rank="genus">
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<emphasis id="C1BD9D34FF90FFC30036FC6BFF50DCC1" box="[79,224,905,927]" italics="true" pageId="17" pageNumber="1060">Namibiotrupes</emphasis>
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<bibRefCitation id="97583CD7FF90FFC30093FC6BFE0CDCC1" author="KRIKKEN, J." box="[234,444,905,927]" pageId="17" pageNumber="1060" pagination="161" refId="ref44555" refString="KRIKKEN, J., 1977 a, New bolboceratine genera from southwestern Africa (Coleoptera: Geotrupidae), Zoologische Mededelingen, 52 (12), 161 ± 168." type="journal article" year="1977">Krikken, 1977a: 165</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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;
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<treatmentCitation id="72686737FF90FFC301B3FC6BFD90DCC1" author="KRIKKEN, J." box="[458,544,905,927]" page="31" pageId="17" pageNumber="1060" year="1984">
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<bibRefCitation id="97583CD7FF90FFC301B3FC6BFD90DCC1" author="KRIKKEN, J." box="[458,544,905,927]" pageId="17" pageNumber="1060" pagination="23" refId="ref44655" refString="KRIKKEN, J., 1984, A generic reclassi ® cation of the Afrotropical Bolboceratini (Coleoptera: Geotrupidae), Zoologische Mededelingen, 58 (3), 23 ± 45." type="journal article" year="1984">1984: 31</bibRefCitation>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="BBD312ADFF90FFC00036FC47FC66DAA7" lastPageId="18" lastPageNumber="1061" pageId="17" pageNumber="1060" type="materials_examined">
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<paragraph id="F3764126FF90FFC30036FC47FDA5DC8A" blockId="17.[79,893,839,1007]" pageId="17" pageNumber="1060">
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<typeStatus id="2C72FF84FF90FFC30036FC47FF32DCE4" box="[79,130,933,954]" pageId="17" pageNumber="1060">Type</typeStatus>
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species:
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<taxonomicName id="34C93AA5FF90FFC300A7FC46FDCBDCE4" authority="Krikken, 1977 a" authorityName="Krikken" authorityYear="1977" box="[222,635,932,954]" class="Insecta" family="Bolboceratidae" genus="Namibiotrupes" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="1060" phylum="Arthropoda" rank="species" species="penrithae">
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<emphasis id="C1BD9D34FF90FFC300A7FC46FE64DCE4" box="[222,468,932,954]" italics="true" pageId="17" pageNumber="1060">Namibiotrupes penrithae</emphasis>
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<bibRefCitation id="97583CD7FF90FFC301A5FC46FDCBDCE4" author="KRIKKEN, J." box="[476,635,932,954]" pageId="17" pageNumber="1060" pagination="161" refId="ref44555" refString="KRIKKEN, J., 1977 a, New bolboceratine genera from southwestern Africa (Coleoptera: Geotrupidae), Zoologische Mededelingen, 52 (12), 161 ± 168." type="journal article" year="1977">Krikken, 1977a</bibRefCitation>
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</taxonomicName>
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, by original designation.
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<taxonomicName id="34C93AA5FF90FFC30036FC5CFE1ADC8A" authority="Krikken, 1984: 32" authorityName="Krikken" authorityPageNumber="32" authorityYear="1984" box="[79,426,958,980]" class="Insecta" family="Bolboceratidae" genus="Bolboceratellus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="1060" phylum="Arthropoda" rank="genus">
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<emphasis id="C1BD9D34FF90FFC30036FC5CFF56DC8A" box="[79,230,958,980]" italics="true" pageId="17" pageNumber="1060">Bolboceratellus</emphasis>
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<bibRefCitation id="97583CD7FF90FFC30089FC5CFE1ADC8A" author="KRIKKEN, J." box="[240,426,958,980]" pageId="17" pageNumber="1060" pagination="23" refId="ref44655" refString="KRIKKEN, J., 1984, A generic reclassi ® cation of the Afrotropical Bolboceratini (Coleoptera: Geotrupidae), Zoologische Mededelingen, 58 (3), 23 ± 45." type="journal article" year="1984">Krikken, 1984: 32</bibRefCitation>
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</taxonomicName>
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.
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<emphasis id="C1BD9D34FF90FFC301C1FC5CFDA5DC8A" bold="true" box="[440,533,958,980]" pageId="17" pageNumber="1060">Syn. nov.</emphasis>
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</paragraph>
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<paragraph id="F3764126FF90FFC30036FC38FCC0DCB1" blockId="17.[79,893,839,1007]" box="[79,880,985,1007]" pageId="17" pageNumber="1060">
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<typeStatus id="2C72FF84FF90FFC30036FC38FF32DCB1" box="[79,130,986,1007]" pageId="17" pageNumber="1060">Type</typeStatus>
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species:
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<taxonomicName id="34C93AA5FF90FFC300A7FC3BFDDEDCB1" authority="PeAringuey, 1901" authorityName="PeAringuey" authorityYear="1901" box="[222,622,985,1007]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="1060" phylum="Arthropoda" rank="species" species="auspicatum">
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<emphasis id="C1BD9D34FF90FFC300A7FC3BFE73DCB1" box="[222,451,985,1007]" italics="true" pageId="17" pageNumber="1060">Bolboceras auspicatum</emphasis>
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<bibRefCitation id="97583CD7FF90FFC301B2FC3BFDDEDCB1" author="PEARINGUEY, L." box="[459,622,985,1007]" pageId="17" pageNumber="1060" pagination="1" refId="ref44791" refString="PEARINGUEY, L., 1901, Descriptive catalogue of the Coleoptera of South Africa (Lucanidae and Scarabaeidae), Transactions of the South African Philosophical Society, 12, 1 ± 920." type="journal article" year="1901">PeÂringuey, 1901</bibRefCitation>
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</taxonomicName>
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, by original designation.
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</paragraph>
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<paragraph id="F3764126FF90FFC30036FBCDFF7DDB18" blockId="17.[79,1035,1071,1701]" box="[79,205,1071,1094]" pageId="17" pageNumber="1060">
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<emphasis id="C1BD9D34FF90FFC30036FBCDFF7DDB18" box="[79,205,1071,1094]" italics="true" pageId="17" pageNumber="1060">Description</emphasis>
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</paragraph>
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<paragraph id="F3764126FF90FFC30001FBACFE06DB38" blockId="17.[79,1035,1071,1701]" box="[120,438,1102,1126]" pageId="17" pageNumber="1060">
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<emphasis id="C1BD9D34FF90FFC30001FBACFF4DDB38" box="[120,253,1102,1126]" italics="true" pageId="17" pageNumber="1060">Body length</emphasis>
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.
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<quantity id="3431ECC3FF90FFC30166FBACFE1EDB38" box="[287,430,1102,1126]" metricMagnitude="-3" metricUnit="m" metricValue="9.55" metricValueMax="13.5" metricValueMin="5.6" pageId="17" pageNumber="1060" unit="mm" value="9.55" valueMax="13.5" valueMin="5.6">5.6±13.5 mm</quantity>
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.
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</paragraph>
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<paragraph id="F3764126FF90FFC30001FB8CFD7CDBD8" blockId="17.[79,1035,1071,1701]" box="[120,716,1134,1158]" pageId="17" pageNumber="1060">
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<emphasis id="C1BD9D34FF90FFC30001FB8CFF73DBD8" box="[120,195,1134,1158]" italics="true" pageId="17" pageNumber="1060">Colour</emphasis>
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. Light to dark brown but forebody darker.
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</paragraph>
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<paragraph id="F3764126FF90FFC30001FB6CFD5DDA78" blockId="17.[79,1035,1071,1701]" pageId="17" pageNumber="1060">
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<emphasis id="C1BD9D34FF90FFC30001FB6CFF03DBF8" box="[120,179,1166,1190]" italics="true" pageId="17" pageNumber="1060">Head</emphasis>
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.
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<emphasis id="C1BD9D34FF90FFC300ACFB6CFEC5DBF8" box="[213,373,1166,1190]" italics="true" pageId="17" pageNumber="1060">Dorsal surface</emphasis>
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asetose.
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<emphasis id="C1BD9D34FF90FFC301A0FB6CFD43DBF8" box="[473,755,1166,1190]" italics="true" pageId="17" pageNumber="1060">Outer margin of mandible</emphasis>
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rounded (e.g. ®gure 5b).
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<emphasis id="C1BD9D34FF90FFC30036FB4CFF16DB98" box="[79,166,1198,1222]" italics="true" pageId="17" pageNumber="1060">Labrum</emphasis>
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declivous in anterior third, with or without feebly elevated transverse carina in anterior third; asetose; anterior margin straight or feebly sinuate.
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<emphasis id="C1BD9D34FF90FFC30313FB2CFC72DBB8" box="[874,962,1230,1254]" italics="true" pageId="17" pageNumber="1060">Clypeus</emphasis>
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,
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<emphasis id="C1BD9D34FF90FFC303AAFB2CFBB9DBB8" box="[979,1033,1230,1254]" italics="true" pageId="17" pageNumber="1060">frons</emphasis>
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with variable ornamentation (®gure 5b, d, f).
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<emphasis id="C1BD9D34FF90FFC30201FB0CFCA5DA58" box="[632,789,1262,1286]" italics="true" pageId="17" pageNumber="1060">Antennal club</emphasis>
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variable in size and shape but basal segment always setose over entire surface.
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</paragraph>
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<paragraph id="F3764126FF90FFC30001FACCFC04DA3B" blockId="17.[79,1035,1071,1701]" pageId="17" pageNumber="1060">
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<emphasis id="C1BD9D34FF90FFC30001FACCFF55DA1B" box="[120,229,1326,1349]" italics="true" pageId="17" pageNumber="1060">Pronotum</emphasis>
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. With steep anterior declivity, topped by straight or bi-angulate transverse ridge adjacent to antero-lateral concavities; lateral margin entire.
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</paragraph>
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<paragraph id="F3764126FF90FFC30001FA8FFBBBDADB" blockId="17.[79,1035,1071,1701]" box="[120,1035,1389,1414]" pageId="17" pageNumber="1060">
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<emphasis id="C1BD9D34FF90FFC30001FA8FFF54DADB" box="[120,228,1389,1413]" italics="true" pageId="17" pageNumber="1060">Scutellum</emphasis>
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. Elongate, basal width 1.13±1.27Ölength; deltoid, variabl y punctate.
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</paragraph>
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<paragraph id="F3764126FF90FFC30001FA6FFC44D95A" blockId="17.[79,1035,1071,1701]" pageId="17" pageNumber="1060">
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<emphasis id="C1BD9D34FF90FFC30001FA6FFF7BDAFB" box="[120,203,1421,1445]" italics="true" pageId="17" pageNumber="1060">Elytron</emphasis>
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. Anterior margin not upturned; striae moderately to strongly impressed, their punctures large and separated by two to less than one puncture diameters; intervals more or less ¯at, with variable punctation; sutural stria terminating at scutellum in a straight line, second stria reaching or almost reaching elytral base.
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</paragraph>
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<paragraph id="F3764126FF90FFC30001F9ECFC67D97A" blockId="17.[79,1035,1071,1701]" box="[120,983,1549,1573]" pageId="17" pageNumber="1060">
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<emphasis id="C1BD9D34FF90FFC30001F9ECFE07D97B" box="[120,439,1549,1573]" italics="true" pageId="17" pageNumber="1060">Prosternum behind procoxae</emphasis>
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(®gure 4b). With median longitudinal carina.
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</paragraph>
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<paragraph id="F3764126FF90FFC30001F9CCFE73D9FA" blockId="17.[79,1035,1071,1701]" pageId="17" pageNumber="1060">
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<emphasis id="C1BD9D34FF90FFC30001F9CCFEBAD91B" box="[120,266,1582,1605]" italics="true" pageId="17" pageNumber="1060">Metasternum</emphasis>
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(®gure 4a). Pyriform, bulbous; process with longitudinal, median suture terminating well before anterior declivity, widely separating mesocoxae; process roundedly declivous anteriad; meso-metasternal suture present, positioned halfway down the anterior declivity.
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</paragraph>
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<caption id="A7B611AEFF93FFC00036FCE5FCF9DC0F" ID-DOI="http://doi.org/10.5281/zenodo.4747989" ID-Zenodo-Dep="4747989" httpUri="https://zenodo.org/record/4747989/files/figure.png" pageId="18" pageNumber="1061" startId="18.[79,95,775,796]" targetBox="[177,937,145,747]" targetPageId="18">
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<paragraph id="F3764126FF93FFC00036FCE5FCF9DC0F" blockId="18.[79,1034,775,850]" pageId="18" pageNumber="1061">
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FIG. 4.
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<taxonomicName id="34C93AA5FF93FFC000D2FCE5FEDEDC42" box="[171,366,775,797]" class="Insecta" family="Bolboceratidae" genus="Namibiotrupes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="1061" phylum="Arthropoda" rank="species" species="undetermined">
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<emphasis id="C1BD9D34FF93FFC000D2FCE5FE8CDC43" box="[171,316,775,797]" italics="true" pageId="18" pageNumber="1061">Namibiotrupes</emphasis>
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spp.
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</taxonomicName>
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: metasternum (a), posterior end of prosternum frontal view (b), metafemur ventral view (c).
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<taxonomicName id="34C93AA5FF93FFC001B7FCC0FDE1DC66" authorityName="Krikken" authorityYear="1977" box="[462,593,802,824]" class="Insecta" family="Bolboceratidae" genus="Namibiotrupes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="1061" phylum="Arthropoda" rank="species" species="penrithae">
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<emphasis id="C1BD9D34FF93FFC001B7FCC0FDE1DC66" box="[462,593,802,824]" italics="true" pageId="18" pageNumber="1061">N. penrithae</emphasis>
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</taxonomicName>
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: aedeagus, ventral (d), dorsal view (e).
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<taxonomicName id="34C93AA5FF93FFC000F6FCDFFEA7DC0C" box="[143,279,829,850]" class="Insecta" family="Bolboceratidae" genus="Namibiotrupes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="1061" phylum="Arthropoda" rank="species" species="auspicatus">
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<emphasis id="C1BD9D34FF93FFC000F6FCDFFEA7DC0C" box="[143,279,829,850]" italics="true" pageId="18" pageNumber="1061">N. auspicatus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="34C93AA5FF93FFC00134FCDFFE01DC0C" authorityName="Gussmann S. M. & Scholtz" authorityYear="2000" baseAuthorityName="Gussmann S. M. & Scholtz" baseAuthorityYear="2000" box="[333,433,829,850]" class="Insecta" family="Bolboceratidae" genus="Namibiotrupes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="1061" phylum="Arthropoda" rank="species" species="zumpti">
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|
<emphasis id="C1BD9D34FF93FFC00134FCDFFE01DC0C" box="[333,433,829,850]" italics="true" pageId="18" pageNumber="1061">N. zumpti</emphasis>
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</taxonomicName>
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|
: aedeagus, ventral (f), dorsal view (g).
|
|
</paragraph>
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|
</caption>
|
|
<paragraph id="F3764126FF93FFC00001FC9AFC2ADB16" blockId="18.[79,1035,887,1530]" pageId="18" pageNumber="1061">
|
|
<emphasis id="C1BD9D34FF93FFC00001FC9AFF1DDCD1" box="[120,173,888,911]" italics="true" pageId="18" pageNumber="1061">Legs</emphasis>
|
|
.
|
|
<emphasis id="C1BD9D34FF93FFC000B7FC95FE99DCD1" box="[206,297,887,911]" italics="true" pageId="18" pageNumber="1061">Protibia</emphasis>
|
|
of normal shape, non-dilated, ®ve- to eight-dentate.
|
|
<emphasis id="C1BD9D34FF93FFC003D1FC95FFCDDCF3" italics="true" pageId="18" pageNumber="1061">Protibial spur</emphasis>
|
|
subparallel, with rounded tip, shorter than ®fth tarsomere.
|
|
<emphasis id="C1BD9D34FF93FFC00348FC74FC2BDCF3" box="[817,923,918,941]" italics="true" pageId="18" pageNumber="1061">Protarsus</emphasis>
|
|
with ®rst tarsomere shorter than ®fth tarsomere, shorter than tarsomeres 2, 3 and 4 together.
|
|
<emphasis id="C1BD9D34FF93FFC00036FC36FF7ADCB2" box="[79,202,980,1004]" italics="true" pageId="18" pageNumber="1061">Metafemur</emphasis>
|
|
with punctation and setation of ventral side as in ®gure 4c.
|
|
<emphasis id="C1BD9D34FF93FFC00306FC36FC5CDCB2" box="[895,1004,980,1004]" italics="true" pageId="18" pageNumber="1061">Metatibia</emphasis>
|
|
in lateral view variably shaped (see species descriptions) but always with subapical carina unlobed and with knoblike projections randomly spread over surface above subapical carina.
|
|
<emphasis id="C1BD9D34FF93FFC00163FBD3FE26DB16" box="[282,406,1073,1096]" italics="true" pageId="18" pageNumber="1061">Metatarsus</emphasis>
|
|
with few subapical setae on tarsomeres 2±4.
|
|
</paragraph>
|
|
<paragraph id="F3764126FF93FFC00001FBADFD58DB39" blockId="18.[79,1035,887,1530]" box="[120,744,1103,1127]" pageId="18" pageNumber="1061">
|
|
<emphasis id="C1BD9D34FF93FFC00001FBADFF56DB39" box="[120,230,1103,1127]" italics="true" pageId="18" pageNumber="1061">Underside</emphasis>
|
|
. With moderately dense and long setation.
|
|
</paragraph>
|
|
<paragraph id="F3764126FF93FFC00001FB8CFEFEDBFB" blockId="18.[79,1035,887,1530]" pageId="18" pageNumber="1061">
|
|
<emphasis id="C1BD9D34FF93FFC00001FB8CFEA7DBD8" box="[120,279,1134,1158]" italics="true" pageId="18" pageNumber="1061">Male genitalia</emphasis>
|
|
(®gure 4d±g). Aedeagus small, simple in structure, parallel-sided, with lobed parameres.
|
|
</paragraph>
|
|
<paragraph id="F3764126FF93FFC00001FB4EFDE0DB9A" blockId="18.[79,1035,887,1530]" box="[120,592,1196,1221]" pageId="18" pageNumber="1061">
|
|
<emphasis id="C1BD9D34FF93FFC00001FB4EFF4CDB9A" box="[120,252,1196,1220]" italics="true" pageId="18" pageNumber="1061">Distribution</emphasis>
|
|
(®gure 7). Southern Africa.
|
|
</paragraph>
|
|
<paragraph id="F3764126FF93FFC00001FB29FC66DAA7" blockId="18.[79,1035,887,1530]" pageId="18" pageNumber="1061">
|
|
<emphasis id="C1BD9D34FF93FFC00001FB29FF5CDBBD" box="[120,236,1227,1251]" italics="true" pageId="18" pageNumber="1061">Comments</emphasis>
|
|
.
|
|
<bibRefCitation id="97583CD7FF93FFC00177FB29FE77DBBD" author="KRIKKEN, J." box="[270,455,1227,1251]" pageId="18" pageNumber="1061" pagination="161" refId="ref44555" refString="KRIKKEN, J., 1977 a, New bolboceratine genera from southwestern Africa (Coleoptera: Geotrupidae), Zoologische Mededelingen, 52 (12), 161 ± 168." type="journal article" year="1977">Krikken (1977a</bibRefCitation>
|
|
,
|
|
<bibRefCitation id="97583CD7FF93FFC001A6FB29FDA4DBBD" author="KRIKKEN, J." box="[479,532,1227,1251]" pageId="18" pageNumber="1061" pagination="23" refId="ref44655" refString="KRIKKEN, J., 1984, A generic reclassi ® cation of the Afrotropical Bolboceratini (Coleoptera: Geotrupidae), Zoologische Mededelingen, 58 (3), 23 ± 45." type="journal article" year="1984">1984</bibRefCitation>
|
|
) erected both genera,
|
|
<taxonomicName id="34C93AA5FF93FFC00356FB29FC60DBBD" authorityName="Krikken" authorityYear="1977" box="[815,976,1227,1251]" class="Insecta" family="Bolboceratidae" genus="Namibiotrupes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="1061" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="C1BD9D34FF93FFC00356FB29FC60DBBD" box="[815,976,1227,1251]" italics="true" pageId="18" pageNumber="1061">Namibiotrupes</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="34C93AA5FF93FFC00036FB08FF47DA5C" authorityName="Krikken" authorityYear="1984" box="[79,247,1258,1282]" class="Insecta" family="Bolboceratidae" genus="Bolboceratellus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="1061" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="C1BD9D34FF93FFC00036FB08FF47DA5C" box="[79,247,1258,1282]" italics="true" pageId="18" pageNumber="1061">Bolboceratellus</emphasis>
|
|
</taxonomicName>
|
|
, and maintained that obvious similarities between the two genera might be due to parallelism and plesiomorphism. We argue, however, that the identical and unique shape of the metasternum, the identical shape of the posterior end of the prosternum, the entirely setose surface of the basal segment of the antennal club, the strong similarities in pronotal ornamentation, the large and densely spaced strial punctures of the elytron, the subparallel and short protibial spur, and lastly and importantly the identical shape and structure of the aedeagi of all three species in both genera indicate true relationship and not convergent similarity. We therefore have synonymized
|
|
<taxonomicName id="34C93AA5FF93FFC00238FA00FD59DAA4" authorityName="Krikken" authorityYear="1984" box="[577,745,1506,1530]" class="Insecta" family="Bolboceratidae" genus="Bolboceratellus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="1061" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="C1BD9D34FF93FFC00238FA00FD59DAA4" box="[577,745,1506,1530]" italics="true" pageId="18" pageNumber="1061">Bolboceratellus</emphasis>
|
|
</taxonomicName>
|
|
with
|
|
<taxonomicName id="34C93AA5FF93FFC00354FA00FC7EDAA4" authorityName="Krikken" authorityYear="1977" box="[813,974,1506,1530]" class="Insecta" family="Bolboceratidae" genus="Namibiotrupes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="1061" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="C1BD9D34FF93FFC00354FA00FC7EDAA4" box="[813,974,1506,1530]" italics="true" pageId="18" pageNumber="1061">Namibiotrupes</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |