271 lines
23 KiB
XML
271 lines
23 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1145.96724" ID-Pensoft-Pub="1313-2970-1145-1" ID-Pensoft-UUID="ACBBD138B7375B0D9F63CE792A82F653" ID-ZooBank="830628C276CD4641BFC6144CD775ED6B" ModsDocID="1313-2970-1145-1" checkinTime="1675506069289" checkinUser="pensoft" docAuthor="Hedin, Marshal & Milne, Marc A." docDate="2023" docId="209BAACAF37451E4871F0AC8359AAFFA" docLanguage="en" docName="ZooKeys 1145: 1-130" docOrigin="ZooKeys 1145" docPubDate="2023-02-03" docSource="http://dx.doi.org/10.3897/zookeys.1145.96724" docTitle="Nesticus jemisinae Hedin & Milne 2023, sp. nov." docType="treatment" docUuid="EC278B87-636E-41F5-AA3E-59599C83AB18" docUuidSource="ZooBank" docVersion="1" id="ACBBD138B7375B0D9F63CE792A82F653" lastPageNumber="1" masterDocId="ACBBD138B7375B0D9F63CE792A82F653" masterDocTitle="New species in old mountains: integrative taxonomy reveals ten new species and extensive short-range endemism in Nesticus spiders (Araneae, Nesticidae) from the southern Appalachian Mountains" masterLastPageNumber="130" masterPageNumber="1" pageNumber="1" updateTime="1675506069289" updateUser="pensoft">
|
||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo>
|
||
<mods:title>New species in old mountains: integrative taxonomy reveals ten new species and extensive short-range endemism in Nesticus spiders (Araneae, Nesticidae) from the southern Appalachian Mountains</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Hedin, Marshal</mods:namePart>
|
||
<mods:affiliation>Department of Biology, San Diego State University, San Diego, California 92182 - 4614, USA</mods:affiliation>
|
||
<mods:nameIdentifier type="email">mhedin@sdsu.edu</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Milne, Marc A.</mods:namePart>
|
||
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-1943-0161</mods:nameIdentifier>
|
||
<mods:affiliation>Department of Biology, University of Indianapolis, Indianapolis, Indiana 46227, USA</mods:affiliation>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:relatedItem type="host">
|
||
<mods:titleInfo>
|
||
<mods:title>ZooKeys</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2023</mods:date>
|
||
<mods:detail type="pubDate">
|
||
<mods:number>2023-02-03</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="volume">
|
||
<mods:number>1145</mods:number>
|
||
</mods:detail>
|
||
<mods:extent unit="page">
|
||
<mods:start>1</mods:start>
|
||
<mods:end>130</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location>
|
||
<mods:url>http://dx.doi.org/10.3897/zookeys.1145.96724</mods:url>
|
||
</mods:location>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1145.96724</mods:identifier>
|
||
<mods:identifier type="Pensoft-Pub">1313-2970-1145-1</mods:identifier>
|
||
<mods:identifier type="ZooBank">830628C276CD4641BFC6144CD775ED6B</mods:identifier>
|
||
<mods:identifier type="Pensoft-UUID">ACBBD138B7375B0D9F63CE792A82F653</mods:identifier>
|
||
</mods:mods>
|
||
<treatment LSID="urn:lsid:zoobank.org:act:EC278B87-636E-41F5-AA3E-59599C83AB18" httpUri="http://treatment.plazi.org/id/209BAACAF37451E4871F0AC8359AAFFA" lastPageNumber="1" pageId="0" pageNumber="1">
|
||
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<taxonomicName LSID="https://zoobank.org/EC278B87-636E-41F5-AA3E-59599C83AB18" authority="Hedin & Milne, 2023" authorityName="Hedin & Milne" authorityYear="2023" class="Arachnida" family="Nesticidae" genus="Nesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nesticus jemisinae" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="jemisinae" status="sp. nov.">Nesticus jemisinae</taxonomicName>
|
||
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="description">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Nesticus jemisinae sp. nov. Tennessee, Marion Co., Rainbow Cave, ♂ holotype, SDSU _ TAC 000662, ventral (A), dorsal (B). Rainbow Cave, ♀ paratype, SDSU _ TAC 000663, ventral (C), dorsal (D). Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1145.96724.figure9" httpUri="https://binary.pensoft.net/fig/804460" pageId="0" pageNumber="1">Fig. 9A-D</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
|
||
<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
Type material:
|
||
<materialsCitation collectingDate="2021-10-20" collectorName="K. S. Zigler" country="USA" county="Marion Co." location="Rainbow Cave" specimenCount="♂" stateProvince="Tennessee" typeStatus="Holotype">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">
|
||
<typeStatus>Holotype</typeStatus>
|
||
</emphasis>
|
||
:
|
||
<collectingCountry name="United States of America">USA</collectingCountry>
|
||
-
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<collectingRegion country="United States of America" name="Tennessee">Tennessee</collectingRegion>
|
||
,
|
||
<collectingCounty>Marion Co.</collectingCounty>
|
||
</emphasis>
|
||
•
|
||
<specimenCount>♂</specimenCount>
|
||
<typeStatus>holotype</typeStatus>
|
||
;
|
||
<location LSID="urn:lsid:plazi:treatment:209BAACAF37451E4871F0AC8359AAFFA:A92EED2387B29BE5A44B0E7FF9F3B34F" country="USA" county="Marion Co." name="Rainbow Cave" stateProvince="Tennessee">Rainbow Cave</location>
|
||
(TMN20);
|
||
<collectingDate value="2021-10-20">20 Oct. 2021</collectingDate>
|
||
;
|
||
<collectorName>K.S. Zigler</collectorName>
|
||
leg.; SDSU_TAC000662
|
||
</materialsCitation>
|
||
</emphasis>
|
||
;
|
||
<materialsCitation collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" collectorName="K. S. Zigler" country="United States of America" county="Marion Co." location="Rainbow Cave" specimenCount="♀" stateProvince="Tennessee" typeStatus="Paratypes">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">
|
||
<typeStatus>Paratypes</typeStatus>
|
||
</emphasis>
|
||
: •
|
||
<specimenCount>♀</specimenCount>
|
||
<typeStatus>paratype</typeStatus>
|
||
; data as for holotype; SDSU_TAC000663
|
||
</materialsCitation>
|
||
; •
|
||
<materialsCitation collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" collectorName="Non, Marion Co" country="United States of America" county="Marion Co." location="Rainbow Cave" specimenCount="2" specimenCount-female="2" stateProvince="Tennessee" typeStatus="paratypes">
|
||
<specimenCount>♂</specimenCount>
|
||
,
|
||
<specimenCount type="female">2♀</specimenCount>
|
||
<typeStatus>paratypes</typeStatus>
|
||
; data as for holotype; SDSU_TAC000664;
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<collectorName>Non</collectorName>
|
||
type material
|
||
</emphasis>
|
||
: -
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<collectorName>Marion Co</collectorName>
|
||
.
|
||
</emphasis>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation collectingDate="2013-11-10" collectorName="K. S. Zigler" country="United States of America" location="Rainbow Cave" specimenCount="♀" stateProvince="Tennessee" typeStatus="paratype">
|
||
<specimenCount>♀</specimenCount>
|
||
;
|
||
<location LSID="urn:lsid:plazi:treatment:209BAACAF37451E4871F0AC8359AAFFA:644C878FD14B4AB9EDDD6F4F5BB0DE07" country="United States of America" name="Rainbow Cave" stateProvince="Tennessee">Rainbow Cave</location>
|
||
(TMN20);
|
||
<collectingDate value="2013-11-10">10 Nov. 2013</collectingDate>
|
||
;
|
||
<collectorName>K.S. Zigler</collectorName>
|
||
leg.
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation collectingDate="2021-10-20" collectorName="K. S. Zigler" country="United States of America" location="Rainbow Cave" specimenCount="1" stateProvince="Tennessee" typeStatus="paratype">
|
||
KSZ 14-248. • 5 imm;
|
||
<location LSID="urn:lsid:plazi:treatment:209BAACAF37451E4871F0AC8359AAFFA:ED10D84E4733279555484D6DC72AEEAD" country="United States of America" name="Rainbow Cave" stateProvince="Tennessee">Rainbow Cave</location>
|
||
(TMN20);
|
||
<collectingDate value="2021-10-20">20 Oct. 2021</collectingDate>
|
||
;
|
||
<collectorName>K.S. Zigler</collectorName>
|
||
leg.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
|
||
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Easily distinguished from other members of the
|
||
<taxonomicName authorityName="Gertsch" authorityYear="1984" class="Arachnida" family="Nesticidae" genus="Nesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nesticus archeri" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="archeri">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Nesticus archeri</emphasis>
|
||
</taxonomicName>
|
||
group. Distinctly small-bodied
|
||
<taxonomicName authorityName="Hedin & Milne" authorityYear="2023" class="Arachnida" family="Nesticidae" genus="Nesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nesticus jemisinae" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="jemisinae">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Nesticus jemisinae</emphasis>
|
||
</taxonomicName>
|
||
possesses well-developed eyes, different from the eyeless
|
||
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="N. cressleri" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="cressleri">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">N. cressleri</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="N. stygius" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="stygius">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">N. stygius</emphasis>
|
||
</taxonomicName>
|
||
. Males possess a relatively simple paracymbium, contrasting with the multiple apophyses of the complex paracymbium of
|
||
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="N. archeri" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="archeri">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">N. archeri</emphasis>
|
||
</taxonomicName>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Nesticus archeri A Alabama, Clay Co., vicinity Mill Shoal Creek, ♂ specimen MCH # 2132, ventral palp B Alabama, Clay Co., vicinity Mill Shoal Creek, ♀ specimen MCH # 2129, epigynum, ventral view C dorsal view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1145.96724.figure10" httpUri="https://binary.pensoft.net/fig/804461" pageId="0" pageNumber="1">10A</figureCitation>
|
||
) and
|
||
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="N. pecki" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="pecki">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">N. pecki</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.904.1.1" author="Hedin, M" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="1 - 19" refId="B17" refString="Hedin, M, Dellinger, B, 2005. Descriptions of a new species and previously unknown males of Nesticus (Araneae: Nesticidae) from caves in eastern North America, with comments on species rarity. Zootaxa 904: 1 - 19, DOI: https://doi.org/10.11646/zootaxa.904.1.1" title="Descriptions of a new species and previously unknown males of Nesticus (Araneae: Nesticidae) from caves in eastern North America, with comments on species rarity." url="https://doi.org/10.11646/zootaxa.904.1.1" volume="904" year="2005">Hedin and Dellinger 2005</bibRefCitation>
|
||
: figs 17, 18). The distinctive tegular apophysis is dark and sinuous, extending under the median apophysis. Female
|
||
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="N. jemisinae" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="jemisinae">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">N. jemisinae</emphasis>
|
||
</taxonomicName>
|
||
may also be distinguished from the latter species by the epigynum. The epigynum of
|
||
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="N. archeri" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="archeri">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">N. archeri</emphasis>
|
||
</taxonomicName>
|
||
is subtriangular with large anterior fovea and a narrow median septum (Fig.
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Nesticus archeri A Alabama, Clay Co., vicinity Mill Shoal Creek, ♂ specimen MCH # 2132, ventral palp B Alabama, Clay Co., vicinity Mill Shoal Creek, ♀ specimen MCH # 2129, epigynum, ventral view C dorsal view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1145.96724.figure10" httpUri="https://binary.pensoft.net/fig/804461" pageId="0" pageNumber="1">10B, C</figureCitation>
|
||
),
|
||
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="N. pecki" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="pecki">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">N. pecki</emphasis>
|
||
</taxonomicName>
|
||
possesses a broad posteriorly-broadening median septum (Fig.
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Nesticus pecki epigynum variation, ventral views A Tennessee, Marion Co., Monteagle Saltpeter Cave, specimen # 1625 B Tennessee, Fentress Co., Hurricane Maze Cave, MLN 13 - 063 (SDSU _ G 2100) C Tennessee, White Co., Haskell Sims Cave, MLN 14 - 004 (SDSU _ G 3697). Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1145.96724.figure11" httpUri="https://binary.pensoft.net/fig/804462" pageId="0" pageNumber="1">11A-C</figureCitation>
|
||
), and
|
||
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="N. jemisinae" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="jemisinae">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">N. jemisinae</emphasis>
|
||
</taxonomicName>
|
||
possesses a posteriorly-pointed median septum with the spermathecae expanded into small bulbs distally (Fig.
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Nesticus jemisinae sp. nov. Tennessee, Marion Co., Rainbow Cave, ♂ holotype, SDSU _ TAC 000662, ventral (A), dorsal (B). Rainbow Cave, ♀ paratype, SDSU _ TAC 000663, ventral (C), dorsal (D). Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1145.96724.figure9" httpUri="https://binary.pensoft.net/fig/804460" pageId="0" pageNumber="1">9C, D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="description">
|
||
<paragraph pageId="0" pageNumber="1">Description of ♂ holotype</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
(SDSU_TAC000662; Fig.
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Nesticus jemisinae sp. nov. Tennessee, Marion Co., Rainbow Cave, ♂ holotype, SDSU _ TAC 000662, ventral (A), dorsal (B). Rainbow Cave, ♀ paratype, SDSU _ TAC 000663, ventral (C), dorsal (D). Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1145.96724.figure9" httpUri="https://binary.pensoft.net/fig/804460" pageId="0" pageNumber="1">9A, B</figureCitation>
|
||
).
|
||
</emphasis>
|
||
Carapace and appendages are a dusky yellow. Abdomen mottled gray with darker patches between lighter parts. Eyes ringed with black and equally well-developed except for AME, which are significantly reduced. Carapace 1.18 long, 1.09 wide. Total body length 2.43. Leg I total length 9.50 (2.74, 0.46, 2.73, 2.56, 1.01), leg I / CW ratio 8.72, leg formula 1423. Paracymbium of palp relatively simple with a large proximally directed ventral process and a dark sinuous distal process. Dorsal process of paracymbium largely reduced to a shallow pocket on the distal edge. Palp tegular apophysis dark, long, narrow, pointed, and extends under median apophysis. Median apophysis elongated towards base of palp and angled proximally at tip.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="♂ variation">
|
||
<paragraph pageId="0" pageNumber="1">♂ Variation.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">No significant genitalic variation was noted in the material examined.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="description">
|
||
<paragraph pageId="0" pageNumber="1">Description of ♀ paratype</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
(SDSU_TAC000663; Fig.
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Nesticus jemisinae sp. nov. Tennessee, Marion Co., Rainbow Cave, ♂ holotype, SDSU _ TAC 000662, ventral (A), dorsal (B). Rainbow Cave, ♀ paratype, SDSU _ TAC 000663, ventral (C), dorsal (D). Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1145.96724.figure9" httpUri="https://binary.pensoft.net/fig/804460" pageId="0" pageNumber="1">9C, D</figureCitation>
|
||
).
|
||
</emphasis>
|
||
Color of carapace, appendages, and abdomen as in male. Eyes as in male. CL 1.20, CW 0.97. Total body length 2.61. Leg I total length 7.96 (2.30, 0.44, 2.29, 1.98, 0.95), Ieg I / CW ratio 8.21, leg formula 1423. Epigynum width approximately half the width of the abdomen. Median septum pointed posteriorly and flanked by fovea along posterior margin. Internal foveal pockets visible from ventral inspection without dissection extending anteriorly angled outwards. Thin, elongate spermathecae curve slightly on outside margins of epigynum, extending anteriorly beyond foveal pockets, expanded into small bulbs distally.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="♀ variation">
|
||
<paragraph pageId="0" pageNumber="1">♀ Variation.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">No significant genitalic variation was noted in the material examined.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="distribution">
|
||
<paragraph pageId="0" pageNumber="1">Distribution and natural history.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Known only from Rainbow Cave, located near Pocket Creek, a tributary to the Little Sequatchie River (Fig.
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Distribution of the Nesticus archeri group, including Nesticus archeri, N. pecki, N. jemisinae, N. cressleri, and N. stygius. State boundaries and major cities shown for geographic context. Dashed lines circumscribe the distributions of N. pecki and N. stygius." figureDoi="10.3897/zookeys.1145.96724.figure8" httpUri="https://binary.pensoft.net/fig/804459" pageId="0" pageNumber="1">8</figureCitation>
|
||
). This cave is approximately 200 meters in length; spiders were collected ~ 50 meters from the cave entrance on the cave ceiling and walls, in total darkness.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="etymology">
|
||
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
The specific name is a matronym in honor of N. K. Jemisin whose 'Broken
|
||
<normalizedToken originalValue="Earth’">Earth'</normalizedToken>
|
||
book series features a subterranean colony, including scientists who study caves.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="remarks">
|
||
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<taxonomicName authorityName="Hedin & Milne" authorityYear="2023" class="Arachnida" family="Nesticidae" genus="Nesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nesticus jemisinae" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="jemisinae">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Nesticus jemisinae</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. is a relictual, single-site endemic whose morphology is quite distinct from that of other members of the species group. This species is nested within a diverged nuclear and mitochondrial subclade of the
|
||
<taxonomicName authorityName="Gertsch" authorityYear="1984" class="Arachnida" family="Nesticidae" genus="Nesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nesticus archeri" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="archeri">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Nesticus archeri</emphasis>
|
||
</taxonomicName>
|
||
group, sister to
|
||
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="N. pecki" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="pecki">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">N. pecki</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="N. archeri" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="archeri">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">N. archeri</emphasis>
|
||
</taxonomicName>
|
||
(Figs
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. UCE concatenated maximum likelihood tree. Distant outgroups removed (for graphical purposes), specimen numbers correspond to those in Suppl. material 3 (with detailed location provided in Suppl. material 1). Node numbers correspond to bootstrap (bold text) / gCF / sCF. Only bootstrap values below 100 shown, all others 100. gCF and sCF values rounded to nearest integer; sCF values below 38 highlighted with red text." figureDoi="10.3897/zookeys.1145.96724.figure3" httpUri="https://binary.pensoft.net/fig/804454" pageId="0" pageNumber="1">3</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. UCE ASTRAL species tree. Distant outgroups removed (for graphical purposes), specimen numbers correspond to those in Suppl. material 3 (with detailed location provided in Suppl. material 1). Included are quartet support values (rounded to nearest integer), and local posterior probability values (if less than 1.0, red text). Branch lengths in coalescent units for internal branches only, terminal branch lengths arbitrary. Internal branch to Nesticus cressleri truncated for graphical purposes." figureDoi="10.3897/zookeys.1145.96724.figure4" httpUri="https://binary.pensoft.net/fig/804455" pageId="0" pageNumber="1">4</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. COI IQT gene tree. Distant outgroups trimmed. Specimen numbers correspond to those in Suppl. material 3 (with detailed location provided in Suppl. material 1). Sequences from UCE capture denoted with " NODE ". Populations with a " Nesticus cooperi - like " male paracymbium denoted by small circles. Not all tip bootstrap values shown (see Suppl. material 4 for. tre file with all bootstrap values)." figureDoi="10.3897/zookeys.1145.96724.figure6" httpUri="https://binary.pensoft.net/fig/804457" pageId="0" pageNumber="1">6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |