134 lines
17 KiB
XML
134 lines
17 KiB
XML
<document id="B7675B725FD310D04F536A7F070225E8" ENCODING="UTF-8" ID-GBIF-Dataset="4cb112e9-a4d9-4bff-8a2b-3104bf2db199" ModsDocID="z01462p001" checkinTime="1247402249539" checkinUser="thomas" docAuthor="Raphael Covain & Sonia Fisch-Muller" docDate="2007" docId="205E9A70ED76D6F2499852B3C3F818A7" docLanguage="en" docName="2007_Covain_Fisch-Muller_gg1.xml" docOrigin="Zootaxa 1462" docSource="http://www.zoobank.org/urn:lsid:zoobank.org:pub:D1F13841-BD7B-4D00-B57D-9CBEC187B83C" docTitle="Harttia Steindachner 1877" docType="treatment" docVersion="13" lastPageNumber="29" masterDocId="FA6C6E047A967CDD773AC3443913C4A8" masterDocTitle="The genera of the Neotropical armored catfish subfamily Loricariinae (Siluriformes: Loricariidae): a practical key and synopsis." masterLastPageNumber="40" masterPageNumber="1" pageNumber="29" updateTime="1701314601160" updateUser="plazi">
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<mods:titleInfo id="021EB89292B7C4DAFB4F264F04E1052D">
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<mods:title id="237F3A2275E4BADE60E6A34A5EE72743">The genera of the Neotropical armored catfish subfamily Loricariinae (Siluriformes: Loricariidae): a practical key and synopsis.</mods:title>
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<mods:name id="C338D52851CAC255B43142CF1A8E9E04" type="personal">
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<mods:roleTerm id="E630A4657469C548372314420C7D8A9F">Author</mods:roleTerm>
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<mods:namePart id="E7591CF4E4A3323A12D31C1E7124B954">Raphael Covain</mods:namePart>
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<mods:name id="0A97A80916CA59752C75E951A41C5305" type="personal">
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<mods:namePart id="5988A769BA06D67CDCFD7C60DF3B11BD">Sonia Fisch-Muller</mods:namePart>
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<mods:typeOfResource id="740982D544ED3DEE2AE0451FB5FB4A0B">text</mods:typeOfResource>
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<mods:title id="221C5F8A79B2C1CA0EB9867C1BA9EE45">Zootaxa</mods:title>
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<mods:part id="E773F0F33B0A67EE43C0280841804BC0">
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<mods:date id="62943FE61BFD9C4245A39D7EAA45D94A">2007</mods:date>
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<mods:detail id="B2B653CFB1526CBBA5333608639EEDA6" type="volume">
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<mods:number id="0E215314329E6315E8DE04175A9ECECF">1462</mods:number>
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<mods:url id="71455B49F2BB5EDE5B526EDDAF2F60E2">http://www.zoobank.org/urn:lsid:zoobank.org:pub:D1F13841-BD7B-4D00-B57D-9CBEC187B83C</mods:url>
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<mods:classification id="27ACE3F38ABAE95C84C1BB1BA13DED99">journal article</mods:classification>
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<mods:identifier id="A5F8F99583AD941ABE53613C8EFBDEFB" type="Plazi-Custom">z01462p001</mods:identifier>
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<mods:identifier id="28C873ADB1C521554348AE07EB4421E4" type="ZooBank">D1F13841-BD7B-4D00-B57D-9CBEC187B83C</mods:identifier>
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<treatment id="205E9A70ED76D6F2499852B3C3F818A7" ID-DOI="http://doi.org/10.5281/zenodo.6236649" ID-GBIF-Taxon="100117576" ID-Zenodo-Dep="6236649" LSID="urn:lsid:plazi:treatment:205E9A70ED76D6F2499852B3C3F818A7" httpUri="http://treatment.plazi.org/id/205E9A70ED76D6F2499852B3C3F818A7" lastPageNumber="29" pageNumber="29">
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<subSubSection id="957ACC724C7D2EC26F99F6708C2C8FE6" type="nomenclature">
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<paragraph id="BC2747ED9B2A89490C7EBEF975050CB5" pageNumber="29">
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<taxonomicName id="F57FE5814537EA638309D28855992791" ID-CoL="4TXW" ID-ENA="503137" LSID-ZBK="urn:lsid:zoobank.org:act:BFDF34D7-CAA6-4D83-B3D7-672132E03918" class="Actinopterygii" family="Loricariidae" genus="Harttia" kingdom="Animalia" lsidName-ZBK="Harttia Steindachner 1877:668 [110]" order="Siluriformes" pageNumber="29" phylum="Chordata" rank="genus">Harttia Steindachner, 1877</taxonomicName>
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.
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</paragraph>
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<paragraph id="2317AE13D7FC88186A4EB7A4FF4A2E7C" pageNumber="29">
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Type species:
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<taxonomicName id="C2FD306F3FE16B306EE9D3881A0B7100" LSID-ZBK="urn:lsid:zoobank.org:act:BD840882-3D9C-424F-BA34-108D33732D35" class="Actinopterygii" family="Loricariidae" genus="Harttia" kingdom="Animalia" lsidName-ZBK="Harttia loricariformis Steindachner 1877:669 [111]" order="Siluriformes" pageNumber="29" phylum="Chordata" rank="species" species="loricariformis">Harttia loricariformis Steindachner, 1877</taxonomicName>
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.
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</paragraph>
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<paragraph id="4361B7BB84C1A9E92E678FDEEA23D4C9" pageNumber="29">
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<materialsCitation id="61A394FC237B7D6E1AEFC4A52D8B5E9D" ID-GBIF-Occurrence="923893101" collectionCode="NMW" country="Brazil" location="Rio Paraiba do Sul." specimenCode="NMW 46346" typeStatus="Lectotype">
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<typeStatus id="117363679C19FB2E768FD75CA774E105">Lectotype</typeStatus>
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:
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<specimenCode id="93011A03AEF395329CA4F6BB75823819">
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<collectionCode id="5B28509BBAED81B43E5DDDACCEF671DF">NMW</collectionCode>
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46346
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</specimenCode>
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,
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<collectingCountry id="4771D06E5E47C12E7EE7054C86892667">Brazil</collectingCountry>
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,
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<location id="774533DA407873A0102518BAA0804E95" LSID="urn:lsid:plazi:treatment:205E9A70ED76D6F2499852B3C3F818A7:774533DA407873A0102518BAA0804E95" country="Brazil" name="Rio Paraiba do Sul.">
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Rio
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<normalizedToken id="E3013FB5AA49E9B301E9A4CB4462FFCE" originalValue="Paraìba">Paraiba</normalizedToken>
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do Sul.
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</location>
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</materialsCitation>
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</paragraph>
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<paragraph id="B33F43D099401509ECEF445835C2A513" pageNumber="29">Gender: feminine.</paragraph>
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</subSubSection>
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<subSubSection id="90EBC62136C77258256724232389C0FA" type="discussion">
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<paragraph id="28C62B1F0E9788FFE9DEA26D68D3D230" lastPageNumber="30" pageNumber="29">
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Distribution primarily includes rivers draining the Guyana Shield, coastal rivers in northeastern Brazil, and the Amazon basin. These rheophilic fishes are found in the upper courses of rivers over rocky and sandy bottoms. Casatti & Castro (2006) characterized ecomorphological trends in fishes living in riffles of the Rio
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<normalizedToken id="5CB6C738351F81619AF78376B69B5C23" originalValue="São">Sao</normalizedToken>
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Francisco. They split the fish communities into three groups comprising nektonic, nektobenthic, and benthic species. Among the latter,
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<taxonomicName id="9EAC3B3946ACED0A4D070433BB2C4015" LSID-ZBK="urn:lsid:zoobank.org:act:BFDF34D7-CAA6-4D83-B3D7-672132E03918" class="Actinopterygii" family="Loricariidae" genus="Harttia" kingdom="Animalia" lsidName-ZBK="Harttia Steindachner 1877:668 [110]" order="Siluriformes" pageNumber="29" phylum="Chordata" rank="genus">Harttia</taxonomicName>
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sp. is supposed to be able to exploit areas with the strongest current, because of its extremely depressed body and long caudal peduncle, comparing to other species. This fact was also empirically noted by Le Bail et al. (2000). Sexual dimorphism includes hypertrophied odontodes on the pectoral spines and along the margins of the snout in mature males. Representatives of this genus seem to be open brooders (Dotzer & Weidner 2003). Recent evidence has suggested that
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<taxonomicName id="1645FC8CA87023E646EF3F54D610B9C0" pageNumber="29" rank="tribe" tribe="Harttiini">Harttiini</taxonomicName>
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could represent a paraphyletic assemblage. Using molecular data, MontoyaBurgos et al. (1998) demonstrated
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<taxonomicName id="A613A6477092A3952325219CDD710C8D" LSID-ZBK="urn:lsid:zoobank.org:act:BFDF34D7-CAA6-4D83-B3D7-672132E03918" class="Actinopterygii" family="Loricariidae" genus="Harttia" kingdom="Animalia" lsidName-ZBK="Harttia Steindachner 1877:668 [110]" order="Siluriformes" pageNumber="29" phylum="Chordata" rank="genus">Harttia</taxonomicName>
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to be sister to two sister clades, one consisting of
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<taxonomicName id="25AAF0159A1E1DC987F06A9539E72E5E" LSID-ZBK="urn:lsid:zoobank.org:act:F67EBCDF-C072-49E3-99DF-AE3E2436805F" class="Actinopterygii" family="Loricariidae" genus="Farlowella" kingdom="Animalia" lsidName-ZBK="Farlowella Eigenmann & Eigenmann 1889:32" order="Siluriformes" pageNumber="29" phylum="Chordata" rank="genus">Farlowella</taxonomicName>
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and
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<taxonomicName id="655167E869F622BAABDF768CDDC4FCF2" class="Actinopterygii" family="Loricariidae" genus="Sturisoma" kingdom="Animalia" order="Siluriformes" pageNumber="29" phylum="Chordata" rank="genus">Sturisoma</taxonomicName>
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, two representatives of the
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<taxonomicName id="D0734B8E29B794B41B6ECEEFDCCECAD5" pageNumber="29" rank="tribe" tribe="Harttiini">Harttiini</taxonomicName>
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, and the second including the representatives of the
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<taxonomicName id="C3BC1F0A9D8851B274DCBDF2A96C50E9" pageNumber="29" rank="tribe" tribe="Loricariini">Loricariini</taxonomicName>
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. Moreover,
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<taxonomicName id="EBF1DCCED9E5C0120F3E62933C4D5184" LSID-ZBK="urn:lsid:zoobank.org:act:BFDF34D7-CAA6-4D83-B3D7-672132E03918" class="Actinopterygii" family="Loricariidae" genus="Harttia" kingdom="Animalia" lsidName-ZBK="Harttia Steindachner 1877:668 [110]" order="Siluriformes" pageNumber="29" phylum="Chordata" rank="genus">Harttia</taxonomicName>
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is in need of revision. For example, the synonymy of
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<taxonomicName id="4274DCC25E612B57C1C001E8AB7E7A1C" LSID-ZBK="urn:lsid:zoobank.org:act:E76DC3E5-AF20-4B2C-9727-40587836C25D" class="Actinopterygii" family="Loricariidae" genus="Cteniloricaria" kingdom="Animalia" lsidName-ZBK="Cteniloricaria Isbrücker & Nijssen 1979:88, 91" order="Siluriformes" pageNumber="29" phylum="Chordata" rank="genus">Cteniloricaria</taxonomicName>
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with
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<taxonomicName id="7C85CF209A521E09401D92C9F5F2508E" LSID-ZBK="urn:lsid:zoobank.org:act:BFDF34D7-CAA6-4D83-B3D7-672132E03918" class="Actinopterygii" family="Loricariidae" genus="Harttia" kingdom="Animalia" lsidName-ZBK="Harttia Steindachner 1877:668 [110]" order="Siluriformes" pageNumber="29" phylum="Chordata" rank="genus">Harttia</taxonomicName>
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(Rapp Py-Daniel & Oliveira 2001) is questionable (Covain et al. 2006) because it rests solely on the characteristics of
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<taxonomicName id="022C68284076259AC144C0C7088838F4" class="Actinopterygii" family="Loricariidae" genus="Harttia" kingdom="Animalia" order="Siluriformes" pageNumber="29" phylum="Chordata" rank="species" species="fowleri">Harttia fowleri (Pellegrin, 1908)</taxonomicName>
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without considering the type species of
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<taxonomicName id="C1E699A00E92AD73E53BE3814178E110" LSID-ZBK="urn:lsid:zoobank.org:act:E76DC3E5-AF20-4B2C-9727-40587836C25D" class="Actinopterygii" family="Loricariidae" genus="Cteniloricaria" kingdom="Animalia" lsidName-ZBK="Cteniloricaria Isbrücker & Nijssen 1979:88, 91" order="Siluriformes" pageNumber="29" phylum="Chordata" rank="genus">Cteniloricaria</taxonomicName>
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. Likewise, the genus
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<taxonomicName id="75F57732EDFADDFE0F32A742F73D9E4F" LSID-ZBK="urn:lsid:zoobank.org:act:72000A6F-6D08-46EB-B661-D34C7C55A75B" authority="Isbrücker, 2001" authorityName="Isbrücker" authorityYear="2001" class="Actinopterygii" family="Loricariidae" genus="Quiritixys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName-ZBK="Quiritixys Isbrücker 2001:21" order="Siluriformes" pageNumber="29" phylum="Chordata" rank="genus">
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Quiritixys
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<normalizedToken id="307022939212ACD9A8C663F9C7F18B5A" originalValue="Isbrücker">Isbruecker</normalizedToken>
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, 2001
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</taxonomicName>
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, also placed in synonymy by Ferraris (2003) is possibly valid. The description of
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<taxonomicName id="43055C8BF9C98B10037C5F83B61B7240" LSID-ZBK="urn:lsid:zoobank.org:act:72000A6F-6D08-46EB-B661-D34C7C55A75B" authority="Isbrücker, 2001" authorityName="Isbrücker" authorityYear="2001" class="Actinopterygii" family="Loricariidae" genus="Quiritixys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName-ZBK="Quiritixys Isbrücker 2001:21" order="Siluriformes" pageNumber="29" phylum="Chordata" rank="genus">Quiritixys</taxonomicName>
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is based on the unusual sexual dimorphism of
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<taxonomicName id="D76B97C278FC5E6ECD72784D57F3D02F" LSID-ZBK="urn:lsid:zoobank.org:act:86D4654E-1E33-440A-9956-EE9CB967527B" class="Actinopterygii" family="Loricariidae" genus="Harttia" kingdom="Animalia" lsidName-ZBK="Harttia leiopleura Oyakawa 1993:5" order="Siluriformes" pageNumber="29" phylum="Chordata" rank="species" species="leiopleura">Harttia leiopleura Oyakawa, 1993</taxonomicName>
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. This feature alone is insufficient to define a genus because it only concerns mature males and is most of the time seasonal in
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<taxonomicName id="CD88AA6F2ED7B2E54E3DF5429E197FC7" pageNumber="29" rank="subFamily" subFamily="Loricariinae">Loricariinae</taxonomicName>
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. This means that the majority of individuals of the species (juveniles, females, and non breeding males) cannot be diagnosed by this single criterion because they do not possess this feature. Nevertheless, the addition of some other features by Oyakawa (1993), such as the absence of the subpreopercle, supports the validity of
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<taxonomicName id="B4537BD20E255106E1037817F6C0D6FA" LSID-ZBK="urn:lsid:zoobank.org:act:72000A6F-6D08-46EB-B661-D34C7C55A75B" authority="Isbrücker, 2001" authorityName="Isbrücker" authorityYear="2001" class="Actinopterygii" family="Loricariidae" genus="Quiritixys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName-ZBK="Quiritixys Isbrücker 2001:21" order="Siluriformes" pageNumber="29" phylum="Chordata" rank="genus">Quiritixys</taxonomicName>
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. Other species such as
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<taxonomicName id="F2614A85DC95A69160C54EE1D24E4588" LSID-ZBK="urn:lsid:zoobank.org:act:7E43BBB0-96F2-41FC-9187-CC38E5129762" class="Actinopterygii" family="Loricariidae" genus="Harttia" kingdom="Animalia" lsidName-ZBK="Harttia novalimensis Oyakawa 1993:8" order="Siluriformes" pageNumber="29" phylum="Chordata" rank="species" species="novalimensis">H. novalimensis</taxonomicName>
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could also belong to
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<taxonomicName id="732968A1CDF5AE147BDAA208CE70C108" LSID-ZBK="urn:lsid:zoobank.org:act:72000A6F-6D08-46EB-B661-D34C7C55A75B" authority="Isbrücker, 2001" authorityName="Isbrücker" authorityYear="2001" class="Actinopterygii" family="Loricariidae" genus="Quiritixys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName-ZBK="Quiritixys Isbrücker 2001:21" order="Siluriformes" pageNumber="29" phylum="Chordata" rank="genus">Quiritixys</taxonomicName>
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because this species also lacks the subpreopercle, but its sexual dimorphism is undescribed. Nevertheless, Langeani et al. (2001) noted well developed odontodes on the posterior body of this species.
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<taxonomicName id="6DFC8AB8A05D42A223A63B840C9E8B89" LSID-ZBK="urn:lsid:zoobank.org:act:BFDF34D7-CAA6-4D83-B3D7-672132E03918" class="Actinopterygii" family="Loricariidae" genus="Harttia" kingdom="Animalia" lsidName-ZBK="Harttia Steindachner 1877:668 [110]" order="Siluriformes" pageNumber="29" phylum="Chordata" rank="genus">Harttia</taxonomicName>
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also exhibits considerable karyotypic diversity with chromosome numbers between 2n = 52 and 2n = 58 in the four species characterized (Kavalco et al. 2005, Centofante et al. 2006). Kavalco et al. (2005) also reported differences in karyotypic formula, symmetry, nucleolar organizing regions (NOR), and diploid number (2n = 52 versus 2n = 56) between two different populations (respectively Grande Stream and Paraitinga River) of
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<taxonomicName id="1BB3069F5E520C9F8D7CE47D791A21C4" LSID-ZBK="urn:lsid:zoobank.org:act:BD840882-3D9C-424F-BA34-108D33732D35" class="Actinopterygii" family="Loricariidae" genus="Harttia" kingdom="Animalia" lsidName-ZBK="Harttia loricariformis Steindachner 1877:669 [111]" order="Siluriformes" pageNumber="29" phylum="Chordata" rank="species" species="loricariformis">H. loricariformis</taxonomicName>
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from the
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<normalizedToken id="70119F122BDB749B79BB962D075F5DE9" originalValue="Paraíba">Paraiba</normalizedToken>
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do Sul basin. These authors hypothesized the sedentary habits of some species to explain these differences. Centofante et al. (2006) characterized a heteromorphic XX/XY1Y2 sex chromosome
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<pageStartToken id="2BD1B927C38A4942B61DFB2329011BF2" pageNumber="30">system</pageStartToken>
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in
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<taxonomicName id="DC741896887905691034FDD40409EC9D" LSID-ZBK="urn:lsid:zoobank.org:act:230783DC-FCEB-4D45-9339-DF4E7AEEB71F" class="Actinopterygii" family="Loricariidae" genus="Harttia" kingdom="Animalia" lsidName-ZBK="Harttia carvalhoi Miranda-Ribeiro 1939:11" order="Siluriformes" pageNumber="29" phylum="Chordata" rank="species" species="carvalhoi">H. carvalhoi</taxonomicName>
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. Currently,
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<taxonomicName id="9EBC7FD8523DE3212D2961862DBCA3E6" LSID-ZBK="urn:lsid:zoobank.org:act:BFDF34D7-CAA6-4D83-B3D7-672132E03918" class="Actinopterygii" family="Loricariidae" genus="Harttia" kingdom="Animalia" lsidName-ZBK="Harttia Steindachner 1877:668 [110]" order="Siluriformes" pageNumber="29" phylum="Chordata" rank="genus">Harttia</taxonomicName>
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comprises 22 species (Provenzano et al. 2005). Partial keys are available for species occurring in Atlantic coastal drainages (Oyakawa 1993, Langeani et al. 2001), Amazon and Guianas drainages (Rapp Py-Daniel & Oliveira 2001), and Guianas drainages (Covain et al. 2006).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |