266 lines
36 KiB
XML
266 lines
36 KiB
XML
<document id="13BAAA06459691CDB9EF7BEBF9C70471" ID-DOI="10.5281/zenodo.274528" ID-GBIF-Dataset="eeadc93a-b560-43f7-a6dd-40a2bee02685" ID-ISSN="1175-5326" ID-Zenodo-Dep="274528" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459982486851" checkinUser="plazi" docAuthor="Buckeridge, John S." docDate="2008" docId="10083F3FCA74FFB27CACB78514464BFC" docLanguage="en" docName="zt01897p052.pdf" docOrigin="Zootaxa 1897" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Tetraclitella judiciae Buckeridge, 2008, sp. nov." docType="treatment" docVersion="8" lastPageNumber="50" masterDocId="EC314747CA71FFB57C3BB460115E4A30" masterDocTitle="Two new species and a new subspecies of Tetraclitella (Cirripedia: Thoracica) from the Cainozoic of Australia and New Zealand and a consideration of the significance of tubiferous walls" masterLastPageNumber="52" masterPageNumber="43" pageNumber="48" updateTime="1698231993478" updateUser="plazi">
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<mods:titleInfo id="74932190670DA0D8AB3E41983A2B71D6">
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<mods:title id="85181E41C71195FE1CFB04916CB49587">Two new species and a new subspecies of Tetraclitella (Cirripedia: Thoracica) from the Cainozoic of Australia and New Zealand and a consideration of the significance of tubiferous walls</mods:title>
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<mods:name id="254ED5CECEFADD213BCEF89A36E3D584" type="personal">
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<mods:roleTerm id="3B1F68049BCD4BA8FD92E9A2C8492E80">Author</mods:roleTerm>
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<mods:namePart id="88A61ADB4172B6D99DE2ABA4D6CF935A">Buckeridge, John S.</mods:namePart>
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<mods:typeOfResource id="058FE5E9DD6C23777B57152B634F493D">text</mods:typeOfResource>
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<mods:title id="B71477969BA47F36A6933CFE736B8215">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="981B4EDAA31498428CA2384BBB253DB9">
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<mods:date id="5F4AF5F93D0B99B3F325DA0103ACDC35">2008</mods:date>
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<mods:detail id="B5F9C52833732AE1ABF69465B9090B75" type="volume">
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<mods:number id="9A391511F66FAEE4EDC95891227A838D">1897</mods:number>
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<mods:classification id="418A87D5FCD274D3B1C3185309B384CE">journal article</mods:classification>
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<mods:identifier id="E8A42F4128758FC9E2F22D5C28AE18CA" type="DOI">10.5281/zenodo.274528</mods:identifier>
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<mods:identifier id="1D478C298CB984620BD90F8CD6290BBC" type="GBIF-Dataset">eeadc93a-b560-43f7-a6dd-40a2bee02685</mods:identifier>
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<mods:identifier id="D1D4AC2FA3F97BACB2C1394917184E35" type="ISSN">1175-5326</mods:identifier>
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<treatment id="10083F3FCA74FFB27CACB78514464BFC" ID-DOI="http://doi.org/10.5281/zenodo.5675339" ID-GBIF-Taxon="119378818" ID-Zenodo-Dep="5675339" LSID="urn:lsid:zoobank.org:act:CB6E33FB-389A-4859-8E2D-960B24624435" httpUri="http://treatment.plazi.org/id/10083F3FCA74FFB27CACB78514464BFC" lastPageId="7" lastPageNumber="50" pageId="5" pageNumber="48">
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<subSubSection id="D0BBDDA2CA74FFB07CACB78510064E12" pageId="5" pageNumber="48" type="nomenclature">
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<paragraph id="981E8E29CA74FFB07CACB78510AD49CE" blockId="5.[151,499,996,1058]" box="[151,499,996,1023]" pageId="5" pageNumber="48">
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<heading id="C3563945CA74FFB07CACB78510AD49CE" bold="true" box="[151,499,996,1023]" fontSize="11" level="1" pageId="5" pageNumber="48" reason="1">
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||
<emphasis id="AAD5523BCA74FFB07CACB78510AD49CE" bold="true" box="[151,499,996,1023]" pageId="5" pageNumber="48">
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<taxonomicName id="5FA1F5AACA74FFB07CACB78510D049CF" box="[151,398,997,1023]" class="Maxillopoda" family="Tetraclitidae" genus="Tetraclitella" kingdom="Animalia" order="Sessilia" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="judiciae" status="sp. nov.">
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<emphasis id="AAD5523BCA74FFB07CACB78510D049CF" bold="true" box="[151,398,997,1023]" italics="true" pageId="5" pageNumber="48">Tetraclitella judiciae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="B1E6EF40CA74FFB07DAEB78410AD49CE" box="[405,499,996,1022]" pageId="5" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="981E8E29CA74FFB07CACB06810064E12" blockId="5.[151,499,996,1058]" box="[151,344,1032,1058]" pageId="5" pageNumber="48">
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<figureCitation id="009A92ACCA74FFB07CACB068105E4E12" box="[151,256,1032,1058]" captionStart="FIGURE 1" captionStartId="3.[151,255,1767,1791]" captionTargetBox="[238,1327,187,1725]" captionTargetId="figure@3.[238,1357,185,1736]" captionTargetPageId="3" captionText="FIGURE 1. Tetraclitella spp. A – E: Tetraclitella judiciae sp. nov. A: RM 318 – latus (exterior); B: RM 318 – latus (interior), the central, large, ellipsoidal structure is a grain of calcite, that if removed may have damaged the plate. It is not part of the barnacle; C: P 313914 – detail showing eroded radius of left latus (central cavity on image), and ellipsoidal pattern produced by eroded parietal pores intersecting the exterior surface; D: P 313914 – detail showing basal section through the shell wall, with inner row of large, elongate pores and outer multi-porous zone; E: P 313914 – detail showing parietal pores that have been apically infilled; F: Tetraclitella purpurascens purpurascens (Wood, 1815): Basal view of parietes exposing edge of sheath (upper, unpatterned zone) and multi-porous zone that lacks a distinct inner row or larger, elongate pores." httpUri="https://zenodo.org/record/274529/files/figure.png" pageId="5" pageNumber="48">Figures 1</figureCitation>
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A–E; 3
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</paragraph>
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</subSubSection>
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<subSubSection id="D0BBDDA2CA74FFB07CACB03713264E9E" pageId="5" pageNumber="48" type="description">
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<paragraph id="981E8E29CA74FFB07CACB03713264E9E" blockId="5.[151,1130,1111,1199]" pageId="5" pageNumber="48">
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<bibRef id="41011ADFCA74FFB07CACB03713264E9E" pageId="5" pageNumber="48" type="journal article">
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<author id="401C66CCCA74FFB07CACB03715394E5E" box="[151,1127,1111,1135]" pageId="5" pageNumber="48">
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1983:?
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<emphasis id="AAD5523BCA74FFB07CDDB03810394E5F" box="[230,359,1112,1135]" italics="true" pageId="5" pageNumber="48">Tetraclitella</emphasis>
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sp. cf.
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<emphasis id="AAD5523BCA74FFB07D8EB038133E4E5F" box="[437,608,1112,1135]" italics="true" pageId="5" pageNumber="48">T. purpurascens</emphasis>
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(Wood, 1815): Buckeridge, 77–78, (not pl.5c,d)
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</author>
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.
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<year id="EC09CF7ACA74FFB07CACB017118E4EBE" box="[151,208,1143,1166]" pageId="5" pageNumber="48">1985</year>
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:?
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<title id="EAA5651FCA74FFB07CDDB01813EF4EBE" box="[230,689,1143,1167]" jopScore="0.04015339" pageId="5" pageNumber="48" type="_" typeScore="47">
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<emphasis id="AAD5523BCA74FFB07CDDB01810394EBF" box="[230,359,1144,1167]" italics="true" pageId="5" pageNumber="48">Tetraclitella</emphasis>
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sp. cf.
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<emphasis id="AAD5523BCA74FFB07D8EB018133E4EBF" box="[437,608,1144,1167]" italics="true" pageId="5" pageNumber="48">T. purpurascens</emphasis>
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(Wood
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</title>
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,
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<part id="EC0524B4CA74FFB07E87B01713A84EBE" box="[700,758,1143,1166]" pageId="5" pageNumber="48" type="volume">1815</part>
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): Buckeridge,
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<pagination id="A16870DDCA74FFB07FAAB01712AE4EBE" box="[913,1008,1143,1166]" pageId="5" pageNumber="48" type="pagination">144–146</pagination>
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, fig.4a,b. 2008:
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<emphasis id="AAD5523BCA74FFB07CE1B0F810054E9F" box="[218,347,1176,1199]" italics="true" pageId="5" pageNumber="48">Tetraclitella</emphasis>
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sp.:
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<journalOrPublisher id="426084E5CA74FFB07DB4B0F7132A4E9E" box="[399,628,1175,1198]" jopScore="0.046330847" pageId="5" pageNumber="48">Buckeridge, 70, fig.4</journalOrPublisher>
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.
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</bibRef>
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</paragraph>
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</subSubSection>
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<subSubSection id="D0BBDDA2CA74FFB07CACB0BF10E94F12" pageId="5" pageNumber="48" type="diagnosis">
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<paragraph id="981E8E29CA74FFB07CACB0BF10E94F12" blockId="5.[151,1436,1247,2036]" pageId="5" pageNumber="48">
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<emphasis id="AAD5523BCA74FFB07CACB0BF10474EC9" bold="true" box="[151,281,1247,1273]" pageId="5" pageNumber="48">Diagnosis:</emphasis>
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Low conic
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<taxonomicName id="5FA1F5AACA74FFB07D9DB080136A4EC9" box="[422,564,1248,1273]" class="Maxillopoda" family="Tetraclitidae" genus="Tetraclitella" kingdom="Animalia" order="Sessilia" pageId="5" pageNumber="48" phylum="Arthropoda" rank="genus">
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<emphasis id="AAD5523BCA74FFB07D9DB080136A4EC9" box="[422,564,1248,1273]" italics="true" pageId="5" pageNumber="48">Tetraclitella</emphasis>
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</taxonomicName>
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with orifice large, radii thin-walled, paries with inner ring of larger, elongate, thick-walled pores.
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</paragraph>
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</subSubSection>
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<subSubSection id="D0BBDDA2CA74FFB07CFDB14F14D04FAA" pageId="5" pageNumber="48" type="distribution">
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<paragraph id="981E8E29CA74FFB07CFDB14F12AC4F42" blockId="5.[151,1436,1247,2036]" pageId="5" pageNumber="48">
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<emphasis id="AAD5523BCA74FFB07CFDB14F10874F79" bold="true" box="[198,473,1327,1353]" pageId="5" pageNumber="48">Distribution and age:</emphasis>
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Southern Victoria,
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<collectingCountry id="E0B6CEB9CA74FFB07EF2B15012674F7A" box="[713,825,1328,1354]" name="Australia" pageId="5" pageNumber="48">Australia</collectingCountry>
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. Late lower Miocene (Batesfordian Stage to Early Pliocene (Kalimnan Stage). For Victorian stages refer to Singleton, 1941.
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</paragraph>
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<paragraph id="981E8E29CA74FFB07CFDB11F14D04FAA" blockId="5.[151,1436,1247,2036]" box="[198,1422,1407,1434]" pageId="5" pageNumber="48">
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<emphasis id="AAD5523BCA74FFB07CFDB11F10754FA9" bold="true" box="[198,299,1407,1433]" pageId="5" pageNumber="48">Holotyp</emphasis>
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e: P313914, a single specimen (
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<figureCitation id="009A92ACCA74FFB07EA5B1E013A34FAA" box="[670,765,1408,1434]" captionStart="FIGURE 1" captionStartId="3.[151,255,1767,1791]" captionTargetBox="[238,1327,187,1725]" captionTargetId="figure@3.[238,1357,185,1736]" captionTargetPageId="3" captionText="FIGURE 1. Tetraclitella spp. A – E: Tetraclitella judiciae sp. nov. A: RM 318 – latus (exterior); B: RM 318 – latus (interior), the central, large, ellipsoidal structure is a grain of calcite, that if removed may have damaged the plate. It is not part of the barnacle; C: P 313914 – detail showing eroded radius of left latus (central cavity on image), and ellipsoidal pattern produced by eroded parietal pores intersecting the exterior surface; D: P 313914 – detail showing basal section through the shell wall, with inner row of large, elongate pores and outer multi-porous zone; E: P 313914 – detail showing parietal pores that have been apically infilled; F: Tetraclitella purpurascens purpurascens (Wood, 1815): Basal view of parietes exposing edge of sheath (upper, unpatterned zone) and multi-porous zone that lacks a distinct inner row or larger, elongate pores." httpUri="https://zenodo.org/record/274529/files/figure.png" pageId="5" pageNumber="48">Figure 1</figureCitation>
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C–E;
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<figureCitation id="009A92ACCA74FFB07F7DB1E012F64FAA" box="[838,936,1408,1434]" captionStart="FIGURE 3" captionStartId="6.[151,255,1868,1892]" captionTargetBox="[523,1066,1321,1835]" captionTargetId="figure@6.[517,1070,1317,1842]" captionTargetPageId="6" captionText="FIGURE 3. Tetraclitella judiciae sp. nov. P 313914 (holotype) – shell with opercular valves absent (the opercular opening is the large central quadrangular structure); carina at top. The dark, “ round ” cavity (c. 1.0 mm in diameter) to right of the opercular opening is an eroded radius, and it is here where the recrystallized shell fabric is suggestive of having had radial pores." httpUri="https://zenodo.org/record/274531/files/figure.png" pageId="5" pageNumber="48">Figure 3</figureCitation>
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) from the Batesford Limestone, Victoria.
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</paragraph>
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</subSubSection>
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<subSubSection id="D0BBDDA2CA74FFB37CFDB1C710F14A82" lastPageId="6" lastPageNumber="49" pageId="5" pageNumber="48" type="materials_examined">
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<paragraph id="981E8E29CA74FFB37CFDB1C710F14A82" blockId="5.[151,1436,1247,2036]" lastBlockId="6.[151,1437,152,1258]" lastPageId="6" lastPageNumber="49" pageId="5" pageNumber="48">
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<emphasis id="AAD5523BCA74FFB07CFDB1C713094FF1" bold="true" box="[198,599,1447,1473]" pageId="5" pageNumber="48">Material: Batesford Limestone:</emphasis>
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P313914 (
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<figureCitation id="009A92ACCA74FFB07EE7B1C8121B4FF2" box="[732,837,1448,1474]" captionStart="FIGURE 3" captionStartId="6.[151,255,1868,1892]" captionTargetBox="[523,1066,1321,1835]" captionTargetId="figure@6.[517,1070,1317,1842]" captionTargetPageId="6" captionText="FIGURE 3. Tetraclitella judiciae sp. nov. P 313914 (holotype) – shell with opercular valves absent (the opercular opening is the large central quadrangular structure); carina at top. The dark, “ round ” cavity (c. 1.0 mm in diameter) to right of the opercular opening is an eroded radius, and it is here where the recrystallized shell fabric is suggestive of having had radial pores." httpUri="https://zenodo.org/record/274531/files/figure.png" pageId="5" pageNumber="48">Figure 3</figureCitation>
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), a complete shell comprising four compartments (but lacking opercula) from the Batesford Limestone, Victoria, carino-rostral length:
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<quantity id="5F5923CCCA74FFB07843B1B015BA4FDA" box="[1144,1252,1488,1514]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" pageId="5" pageNumber="48" unit="mm" value="15.5">15.5 mm</quantity>
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. [The specimen was removed from the old Magistrates' Court building, situated at the corner of Russell and Latrobe Streets, Melbourne, on
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<date id="EC1FA8E9CA74FFB07D72B24210E54C0C" box="[329,443,1567,1596]" pageId="5" pageNumber="48">6th March</date>
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, 2006 by J. Buckeridge. The precise locality of the site from where the block of limestone was quarried is unknown, although the Limestone Quarry is close to the Moorabool River, near Geelong, Victoria. Date removed from the quarry, circa 1912 (see Buckeridge, 2008)]; RG77640091 (figure 4a,b in Buckeridge, 1985), an articulated rostrum and latus (rostrum incomplete – apical part and much of the right radius absent), length
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<quantity id="5F5923CCCA74FFB07DC2B2A213004CEC" box="[505,606,1730,1756]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="5" pageNumber="48" unit="mm" value="4.5">4.5 mm</quantity>
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, collected in 1977 from
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<quantity id="5F5923CCCA74FFB07F9FB2A212BE4CEC" box="[932,992,1730,1756]" metricMagnitude="0" metricUnit="m" metricValue="9.8" pageId="5" pageNumber="48" unit="m" value="9.8">9.8m</quantity>
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above the quarry floor, Batesford; R77640090, an incomplete latus, as previous but from
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<quantity id="5F5923CCCA74FFB07F18B28A12014D34" box="[803,863,1770,1796]" metricMagnitude="0" metricUnit="m" metricValue="8.2" pageId="5" pageNumber="48" unit="m" value="8.2">8.2m</quantity>
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above the quarry floor; R77640095, a?latus, as previous but from
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<quantity id="5F5923CCCA74FFB07D49B37210E24D1C" box="[370,444,1810,1836]" metricMagnitude="1" metricUnit="m" metricValue="1.59" pageId="5" pageNumber="48" unit="m" value="15.9">15.9m</quantity>
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above the quarry floor; R77640099, a?latus, as previous but from 21.0m above the quarry floor; RM318 (
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<figureCitation id="009A92ACCA74FFB07D9AB35A135A4D64" box="[417,516,1850,1876]" captionStart="FIGURE 1" captionStartId="3.[151,255,1767,1791]" captionTargetBox="[238,1327,187,1725]" captionTargetId="figure@3.[238,1357,185,1736]" captionTargetPageId="3" captionText="FIGURE 1. Tetraclitella spp. A – E: Tetraclitella judiciae sp. nov. A: RM 318 – latus (exterior); B: RM 318 – latus (interior), the central, large, ellipsoidal structure is a grain of calcite, that if removed may have damaged the plate. It is not part of the barnacle; C: P 313914 – detail showing eroded radius of left latus (central cavity on image), and ellipsoidal pattern produced by eroded parietal pores intersecting the exterior surface; D: P 313914 – detail showing basal section through the shell wall, with inner row of large, elongate pores and outer multi-porous zone; E: P 313914 – detail showing parietal pores that have been apically infilled; F: Tetraclitella purpurascens purpurascens (Wood, 1815): Basal view of parietes exposing edge of sheath (upper, unpatterned zone) and multi-porous zone that lacks a distinct inner row or larger, elongate pores." httpUri="https://zenodo.org/record/274529/files/figure.png" pageId="5" pageNumber="48">Figure 1</figureCitation>
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A,B), length
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<quantity id="5F5923CCCA74FFB07E9FB35A13A54D64" box="[676,763,1850,1876]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="5" pageNumber="48" unit="mm" value="2.4">2.4mm</quantity>
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: a latus collected from a loose block of moderately well cemented limestone in the Batesford Quarry,
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<geoCoordinate id="FD95E8EECA74FFB07E90B302127A4D4C" box="[683,804,1890,1916]" direction="south" orientation="latitude" pageId="5" pageNumber="48" precision="92" value="-38.108334">38°06.5’S</geoCoordinate>
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,
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<geoCoordinate id="FD95E8EECA74FFB07F0BB30212E74D4C" box="[816,953,1890,1916]" direction="east" orientation="longitude" pageId="5" pageNumber="48" precision="92" value="144.28833">144°17.3’E</geoCoordinate>
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, on
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<date id="EC1FA8E9CA74FFB07FD1B30215354D4C" box="[1002,1131,1890,1916]" pageId="5" pageNumber="48" value="1930-01">30 January</date>
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, 2007 by JSB. Age: Early Miocene. Specimens with the prefix “P” are held in the
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<typeStatus id="471A308BCA74FFB07F76B3EA12214D94" box="[845,895,1930,1956]" pageId="5" pageNumber="48">type</typeStatus>
|
||
collection of the Museum of Victoria, Melbourne; those with the prefix “R” are held in the Palaeontology Collection, Bureau of Mineral Resources, Canberra, ACT,
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<collectingCountry id="E0B6CEB9CA74FFB07D63B3BA10994DC4" box="[344,455,2010,2036]" name="Australia" pageId="5" pageNumber="48">Australia</collectingCountry>
|
||
; those with “RM” are held in the Palaeobiology Collection of Earth & Oceanic Systems,
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<collectionCode id="FEB016ECCA77FFB37CE6B4F810784A82" box="[221,294,152,178]" pageId="6" pageNumber="49">RMIT</collectionCode>
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University.
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</paragraph>
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</subSubSection>
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<subSubSection id="D0BBDDA2CA77FFB37CFDB4DF10B04902" pageId="6" pageNumber="49" type="description">
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<paragraph id="981E8E29CA77FFB37CFDB4DF127E4872" blockId="6.[151,1437,152,1258]" pageId="6" pageNumber="49">
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||
<emphasis id="AAD5523BCA77FFB37CFDB4DF103F4AE9" bold="true" box="[198,353,191,217]" pageId="6" pageNumber="49">Description:</emphasis>
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Shell of moderate size, possessing a medium-low profile; orifice large, rhomboidal (carinorostral length in
|
||
<typeStatus id="471A308BCA77FFB37D62B488109A4B32" box="[345,452,232,258]" pageId="6" pageNumber="49" type="holotype">holotype</typeStatus>
|
||
:
|
||
<quantity id="5F5923CCCA77FFB37DF4B488134E4B32" box="[463,528,232,258]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="6" pageNumber="49" unit="mm" value="7.0">7mm</quantity>
|
||
); exterior with up to six narrow, well-spaced longitudinal ridges separated by slightly more than their own widths, ridges crossed by fine, transverse growth lines;
|
||
<typeStatus id="471A308BCA77FFB378A8B57015A74B1A" box="[1171,1273,272,298]" pageId="6" pageNumber="49" type="holotype">holotype</typeStatus>
|
||
with exterior surface corroded and/or corraded in part, exposing very slightly raised parietal pores, demonstrating an appearance commensurate with that observed in the chitinous lined pores of living tetraclitids (
|
||
<figureCitation id="009A92ACCA77FFB3793AB500143D4B4A" box="[1281,1379,352,378]" captionStart="FIGURE 1" captionStartId="3.[151,255,1767,1791]" captionTargetBox="[238,1327,187,1725]" captionTargetId="figure@3.[238,1357,185,1736]" captionTargetPageId="3" captionText="FIGURE 1. Tetraclitella spp. A – E: Tetraclitella judiciae sp. nov. A: RM 318 – latus (exterior); B: RM 318 – latus (interior), the central, large, ellipsoidal structure is a grain of calcite, that if removed may have damaged the plate. It is not part of the barnacle; C: P 313914 – detail showing eroded radius of left latus (central cavity on image), and ellipsoidal pattern produced by eroded parietal pores intersecting the exterior surface; D: P 313914 – detail showing basal section through the shell wall, with inner row of large, elongate pores and outer multi-porous zone; E: P 313914 – detail showing parietal pores that have been apically infilled; F: Tetraclitella purpurascens purpurascens (Wood, 1815): Basal view of parietes exposing edge of sheath (upper, unpatterned zone) and multi-porous zone that lacks a distinct inner row or larger, elongate pores." httpUri="https://zenodo.org/record/274529/files/figure.png" pageId="6" pageNumber="49">Figure 1</figureCitation>
|
||
C,E;
|
||
<figureCitation id="009A92ACCA77FFB37CACB5E811A04B92" box="[151,254,392,418]" captionStart="FIGURE 3" captionStartId="6.[151,255,1868,1892]" captionTargetBox="[523,1066,1321,1835]" captionTargetId="figure@6.[517,1070,1317,1842]" captionTargetPageId="6" captionText="FIGURE 3. Tetraclitella judiciae sp. nov. P 313914 (holotype) – shell with opercular valves absent (the opercular opening is the large central quadrangular structure); carina at top. The dark, “ round ” cavity (c. 1.0 mm in diameter) to right of the opercular opening is an eroded radius, and it is here where the recrystallized shell fabric is suggestive of having had radial pores." httpUri="https://zenodo.org/record/274531/files/figure.png" pageId="6" pageNumber="49">Figure 3</figureCitation>
|
||
); radii narrow, slightly depressed with respect to paries, relatively flat, more susceptible to being eroded than with paries, such that sub-vertical cavities may form at the radii sites (see
|
||
<figureCitation id="009A92ACCA77FFB378BDB5D015B74BFA" box="[1158,1257,432,458]" captionStart="FIGURE 3" captionStartId="6.[151,255,1868,1892]" captionTargetBox="[523,1066,1321,1835]" captionTargetId="figure@6.[517,1070,1317,1842]" captionTargetPageId="6" captionText="FIGURE 3. Tetraclitella judiciae sp. nov. P 313914 (holotype) – shell with opercular valves absent (the opercular opening is the large central quadrangular structure); carina at top. The dark, “ round ” cavity (c. 1.0 mm in diameter) to right of the opercular opening is an eroded radius, and it is here where the recrystallized shell fabric is suggestive of having had radial pores." httpUri="https://zenodo.org/record/274531/files/figure.png" pageId="6" pageNumber="49">Figure 3</figureCitation>
|
||
). Sheath weak, extending for approximately half height of paries; pores rounded to oblate in section, becoming significantly larger, flattened, with thickened walls adjacent to the sheath (see
|
||
<typeStatus id="471A308BCA77FFB37FACB660155F482A" box="[919,1025,512,538]" pageId="6" pageNumber="49" type="holotype">holotype</typeStatus>
|
||
,
|
||
<figureCitation id="009A92ACCA77FFB37837B6601532482A" box="[1036,1132,512,538]" captionStart="FIGURE 1" captionStartId="3.[151,255,1767,1791]" captionTargetBox="[238,1327,187,1725]" captionTargetId="figure@3.[238,1357,185,1736]" captionTargetPageId="3" captionText="FIGURE 1. Tetraclitella spp. A – E: Tetraclitella judiciae sp. nov. A: RM 318 – latus (exterior); B: RM 318 – latus (interior), the central, large, ellipsoidal structure is a grain of calcite, that if removed may have damaged the plate. It is not part of the barnacle; C: P 313914 – detail showing eroded radius of left latus (central cavity on image), and ellipsoidal pattern produced by eroded parietal pores intersecting the exterior surface; D: P 313914 – detail showing basal section through the shell wall, with inner row of large, elongate pores and outer multi-porous zone; E: P 313914 – detail showing parietal pores that have been apically infilled; F: Tetraclitella purpurascens purpurascens (Wood, 1815): Basal view of parietes exposing edge of sheath (upper, unpatterned zone) and multi-porous zone that lacks a distinct inner row or larger, elongate pores." httpUri="https://zenodo.org/record/274529/files/figure.png" pageId="6" pageNumber="49">Figure 1</figureCitation>
|
||
D), less so in other specimens (
|
||
<figureCitation id="009A92ACCA77FFB37CDFB64810184872" box="[228,326,552,578]" captionStart="FIGURE 1" captionStartId="3.[151,255,1767,1791]" captionTargetBox="[238,1327,187,1725]" captionTargetId="figure@3.[238,1357,185,1736]" captionTargetPageId="3" captionText="FIGURE 1. Tetraclitella spp. A – E: Tetraclitella judiciae sp. nov. A: RM 318 – latus (exterior); B: RM 318 – latus (interior), the central, large, ellipsoidal structure is a grain of calcite, that if removed may have damaged the plate. It is not part of the barnacle; C: P 313914 – detail showing eroded radius of left latus (central cavity on image), and ellipsoidal pattern produced by eroded parietal pores intersecting the exterior surface; D: P 313914 – detail showing basal section through the shell wall, with inner row of large, elongate pores and outer multi-porous zone; E: P 313914 – detail showing parietal pores that have been apically infilled; F: Tetraclitella purpurascens purpurascens (Wood, 1815): Basal view of parietes exposing edge of sheath (upper, unpatterned zone) and multi-porous zone that lacks a distinct inner row or larger, elongate pores." httpUri="https://zenodo.org/record/274529/files/figure.png" pageId="6" pageNumber="49">Figure 1</figureCitation>
|
||
B); nature of basis, if present, unknown.
|
||
</paragraph>
|
||
<paragraph id="981E8E29CA77FFB37CFDB62F10B04902" blockId="6.[151,1437,152,1258]" pageId="6" pageNumber="49">
|
||
<emphasis id="AAD5523BCA77FFB37CFDB62F13084859" bold="true" box="[198,598,591,617]" pageId="6" pageNumber="49">Comparison with other species:</emphasis>
|
||
<taxonomicName id="5FA1F5AACA77FFB37E5BB630120B4859" box="[608,853,592,617]" class="Maxillopoda" family="Tetraclitidae" genus="Tetraclitella" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="judiciae" status="sp. nov.">
|
||
<emphasis id="AAD5523BCA77FFB37E5BB630120B4859" box="[608,853,592,617]" italics="true" pageId="6" pageNumber="49">Tetraclitella judiciae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="B1E6EF40CA77FFB37F64B63012E5485A" box="[863,955,592,618]" pageId="6" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is easily distinguished from
|
||
<taxonomicName id="5FA1F5AACA77FFB3792DB63014C24859" box="[1302,1436,592,617]" class="Maxillopoda" family="Tetraclitidae" genus="Tetraclitella" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="depressa">
|
||
<emphasis id="AAD5523BCA77FFB3792DB63014C24859" box="[1302,1436,592,617]" italics="true" pageId="6" pageNumber="49">T. depressa</emphasis>
|
||
</taxonomicName>
|
||
by the former’s narrower radii and higher conic shell; it can be distinguished from both subspecies of
|
||
<taxonomicName id="5FA1F5AACA77FFB37977B61810504889" class="Maxillopoda" family="Tetraclitidae" genus="Tetraclitella" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="purpurascens">
|
||
<emphasis id="AAD5523BCA77FFB37977B61810504889" italics="true" pageId="6" pageNumber="49">T. purpurascens</emphasis>
|
||
</taxonomicName>
|
||
by possessing much thinner radii (in no specimens of
|
||
<taxonomicName id="5FA1F5AACA77FFB37F90B6C015774889" box="[939,1065,672,697]" class="Maxillopoda" family="Tetraclitidae" genus="Tetraclitella" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="judiciae">
|
||
<emphasis id="AAD5523BCA77FFB37F90B6C015774889" box="[939,1065,672,697]" italics="true" pageId="6" pageNumber="49">T. judiciae</emphasis>
|
||
</taxonomicName>
|
||
have complete radii been preserved, and this is considered to reflect a more delicate, thinner structure). The inner row of larger parietal pores is reminiscent of both
|
||
<taxonomicName id="5FA1F5AACA77FFB37DD3B69013294939" box="[488,631,752,777]" class="Maxillopoda" family="Tetraclitidae" genus="Tesseropora" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="49" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AAD5523BCA77FFB37DD3B69013294939" box="[488,631,752,777]" italics="true" pageId="6" pageNumber="49">Tesseropora</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="5FA1F5AACA77FFB37E89B69012624939" box="[690,828,752,777]" class="Maxillopoda" family="Tetraclitidae" genus="Tesseroplax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="49" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AAD5523BCA77FFB37E89B69012624939" box="[690,828,752,777]" italics="true" pageId="6" pageNumber="49">Tesseroplax</emphasis>
|
||
</taxonomicName>
|
||
, but these genera are distinguished by possessing a single multitubiferous paries.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D0BBDDA2CA77FFB27CFDB75F13264B4C" lastPageId="7" lastPageNumber="50" pageId="6" pageNumber="49" type="discussion">
|
||
<paragraph id="981E8E29CA77FFB37CFDB75F15454E7A" blockId="6.[151,1437,152,1258]" pageId="6" pageNumber="49">
|
||
<emphasis id="AAD5523BCA77FFB37CFDB75F101C4969" bold="true" box="[198,322,831,857]" pageId="6" pageNumber="49">Remarks:</emphasis>
|
||
The presence of horizontal pores within the radii is an important characteristic of
|
||
<taxonomicName id="5FA1F5AACA77FFB3793DB72014CA4969" box="[1286,1428,832,857]" class="Maxillopoda" family="Tetraclitidae" genus="Tetraclitella" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="49" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AAD5523BCA77FFB3793DB72014CA4969" box="[1286,1428,832,857]" italics="true" pageId="6" pageNumber="49">Tetraclitella</emphasis>
|
||
</taxonomicName>
|
||
. Pores are not clearly distinguishable in any of the fossil species described in this paper, although the radii in all specimens are poorly preserved. [For example, in
|
||
<taxonomicName id="5FA1F5AACA77FFB37F30B7F012EF4999" box="[779,945,912,937]" class="Maxillopoda" family="Tetraclitidae" genus="Tetraclitella" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="nodicostata" status="sp. nov.">
|
||
<emphasis id="AAD5523BCA77FFB37F30B7F012EF4999" box="[779,945,912,937]" italics="true" pageId="6" pageNumber="49">T. nodicostata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="B1E6EF40CA77FFB37F82B7F0154A499A" box="[953,1044,912,938]" pageId="6" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(specimen A177), there are remnants of a radius on the carinal side of the right latus and although there are no definite pores, there is a subtle change in shell texture, both here and on other radii, which is suggestive of pores]. All specimens of
|
||
<taxonomicName id="5FA1F5AACA77FFB3797CB78011964E11" class="Maxillopoda" family="Tetraclitidae" genus="Tetraclitella" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="judiciae">
|
||
<emphasis id="AAD5523BCA77FFB3797CB78011964E11" italics="true" pageId="6" pageNumber="49">T. judiciae</emphasis>
|
||
</taxonomicName>
|
||
have undergone some diagenesis – in the
|
||
<typeStatus id="471A308BCA77FFB37E8EB06812414E12" box="[693,799,1032,1058]" pageId="6" pageNumber="49" type="holotype">holotype</typeStatus>
|
||
, weak dissolution and crystallisation has occurred and the fine, delicate nature of radial pores would not have survived this process.
|
||
</paragraph>
|
||
<paragraph id="981E8E29CA77FFB37CFDB038158F4EDA" blockId="6.[151,1437,152,1258]" pageId="6" pageNumber="49">
|
||
Well-developed radii are an important characteristic of
|
||
<taxonomicName id="5FA1F5AACA77FFB37F77B03812844E41" box="[844,986,1112,1137]" class="Maxillopoda" family="Tetraclitidae" genus="Tetraclitella" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="49" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AAD5523BCA77FFB37F77B03812844E41" box="[844,986,1112,1137]" italics="true" pageId="6" pageNumber="49">Tetraclitella</emphasis>
|
||
</taxonomicName>
|
||
, as is the low profile and multitubiferous paries.
|
||
<collectingCountry id="E0B6CEB9CA77FFB37D25B0E010614EAA" box="[286,319,1152,1178]" name="American Samoa" pageId="6" pageNumber="49">As</collectingCountry>
|
||
noted, the
|
||
<typeStatus id="471A308BCA77FFB37DFEB0E013724EAA" box="[453,556,1152,1178]" pageId="6" pageNumber="49" type="holotype">holotype</typeStatus>
|
||
differs from modern
|
||
<taxonomicName id="5FA1F5AACA77FFB37F16B0E012E54EA9" box="[813,955,1152,1177]" class="Maxillopoda" family="Tetraclitidae" genus="Tetraclitella" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="49" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AAD5523BCA77FFB37F16B0E012E54EA9" box="[813,955,1152,1177]" italics="true" pageId="6" pageNumber="49">Tetraclitella</emphasis>
|
||
</taxonomicName>
|
||
by possession of an inner row of larger, elongate pores; nonetheless, this may be an ontogenetic feature, as the specimen is large and eroded. The inner pore arrangement however, is similar to what could be expected in an ancestral
|
||
<taxonomicName id="5FA1F5AACA77FFB37807B0B015954ED9" box="[1084,1227,1232,1257]" class="Maxillopoda" family="Tetraclitidae" genus="Tesseropora" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="49" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AAD5523BCA77FFB37807B0B015954ED9" box="[1084,1227,1232,1257]" italics="true" pageId="6" pageNumber="49">Tesseropora</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<caption id="CCDEDEA1CA77FFB37CACB32C10454DFB" httpUri="https://zenodo.org/record/274531/files/figure.png" pageId="6" pageNumber="49" targetBox="[523,1066,1321,1835]" targetPageId="6">
|
||
<paragraph id="981E8E29CA77FFB37CACB32C10454DFB" blockId="6.[151,1436,1868,1995]" pageId="6" pageNumber="49">
|
||
<emphasis id="AAD5523BCA77FFB37CACB32C10464D54" bold="true" box="[151,280,1868,1892]" pageId="6" pageNumber="49">FIGURE 3.</emphasis>
|
||
<taxonomicName id="5FA1F5AACA77FFB37D24B32D10A24D54" box="[287,508,1869,1892]" class="Maxillopoda" family="Tetraclitidae" genus="Tetraclitella" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="judiciae" status="sp. nov.">
|
||
<emphasis id="AAD5523BCA77FFB37D24B32D10A24D54" box="[287,508,1869,1892]" italics="true" pageId="6" pageNumber="49">Tetraclitella judiciae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="AAD5523BCA77FFB37E39B32C13094D54" bold="true" box="[514,599,1868,1892]" pageId="6" pageNumber="49">
|
||
<taxonomicNameLabel id="B1E6EF40CA77FFB37E39B32C13094D54" box="[514,599,1868,1892]" pageId="6" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
P313914 (holotype)–shell with opercular valves absent (the opercular opening is the large central quadrangular structure); carina at top. The dark, “round” cavity (c. 1.0 mm in diameter) to right of the opercular opening is an eroded radius, and it is here where the recrystallized shell fabric is suggestive of having had radial pores.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="981E8E29CA76FFB27CFDB4F813264B4C" blockId="7.[151,1436,152,460]" pageId="7" pageNumber="50">
|
||
[The history of the
|
||
<typeStatus id="471A308BCA76FFB27D94B4F8134B4A82" box="[431,533,152,178]" pageId="7" pageNumber="50" type="holotype">holotype</typeStatus>
|
||
is worthy of note. It was found exposed in the limestone wall of “Building 20” fronting Russell Street, Central Melbourne, currently occupied by both the Chancellor and Vice Chancellor of RMIT University. The building, originally the Magistrates’ Law Courts, is one of Melbourne’s grandest; not surprisingly it is “heritage listed” meaning that it cannot be damaged. The saga that culminated in the extraction of this unique specimen was beamed around the world on Sky Television on
|
||
<date id="EC1FA8E9CA76FFB278ACB55A14154B64" box="[1175,1355,312,340]" pageId="7" pageNumber="50" value="2006-03-07">7th March 2006</date>
|
||
, and is fully documented in Buckeridge (2008)].
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D0BBDDA2CA76FFB27CFDB5EA14464BFC" pageId="7" pageNumber="50" type="etymology">
|
||
<paragraph id="981E8E29CA76FFB27CFDB5EA14464BFC" blockId="7.[151,1436,152,460]" pageId="7" pageNumber="50">
|
||
<emphasis id="AAD5523BCA76FFB27CFDB5EA10094B94" bold="true" box="[198,343,394,420]" pageId="7" pageNumber="50">Etymology:</emphasis>
|
||
Noun in apposition, [Latin
|
||
<emphasis id="AAD5523BCA76FFB27EA7B5EB138F4B94" box="[668,721,395,420]" italics="true" pageId="7" pageNumber="50">iuris</emphasis>
|
||
= law (court)] in reference to purpose for which the building in which it was found in was built, and the caution exercised during the extraction of the
|
||
<typeStatus id="471A308BCA76FFB27891B5D2144B4BFC" box="[1194,1301,434,460]" pageId="7" pageNumber="50" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |