256 lines
31 KiB
XML
256 lines
31 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.210.3042" ID-PMC="PMC3406452" ID-Pensoft-Pub="1313-2970-210-19" ID-Pensoft-UUID="BD1FF0348E7DFFF87D580B20FFF06438" ID-PubMed="22859895" ID-Zenodo-Dep="577401" ID-ZooBank="DF32BC38E8394DC48EA0FB6508EC7B2B" ModsDocID="1313-2970-210-19" checkinTime="1451248849322" checkinUser="pensoft" docAuthor="Valdes, Pavel" docDate="2012" docId="EC13C1FE2B893787FFC4F0A331A0B47A" docLanguage="en" docName="ZooKeys 210: 19-67" docOrigin="ZooKeys 210" docPubDate="2012-07-24" docSource="http://dx.doi.org/10.3897/zookeys.210.3042" docTitle="Semiardistomis labialis" docType="treatment" docVersion="5" id="BD1FF0348E7DFFF87D580B20FFF06438" lastPageNumber="46" masterDocId="BD1FF0348E7DFFF87D580B20FFF06438" masterDocTitle="Notes about morphological features of the Western Hemisphere subtribe Ardistomina, and revision of genus Semiardistomis Kult (Coleoptera, Carabidae, Scaritinae, Clivinini)" masterLastPageNumber="67" masterPageNumber="19" pageNumber="45" updateTime="1668154243518" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Notes about morphological features of the Western Hemisphere subtribe Ardistomina, and revision of genus Semiardistomis Kult (Coleoptera, Carabidae, Scaritinae, Clivinini)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Valdes, Pavel</mods:namePart>
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<mods:affiliation>Gertrudis 365 apto 5 e / D'Strampes y Goicuria, Cp 10500, Habana, Cuba</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2012</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2012-07-24</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>210</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>19</mods:start>
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<mods:end>67</mods:end>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.210.3042</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.210.3042</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-210-19</mods:identifier>
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<mods:identifier type="ZooBank">DF32BC38E8394DC48EA0FB6508EC7B2B</mods:identifier>
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<mods:identifier type="Pensoft-UUID">BD1FF0348E7DFFF87D580B20FFF06438</mods:identifier>
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<mods:identifier type="Zenodo-Dep">577401</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152036407" LSID="urn:lsid:plazi:treatment:EC13C1FE2B893787FFC4F0A331A0B47A" httpUri="http://treatment.plazi.org/id/EC13C1FE2B893787FFC4F0A331A0B47A" lastPageId="28" lastPageNumber="46" pageId="26" pageNumber="45">
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<subSubSection pageId="26" pageNumber="45" type="nomenclature">
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<paragraph pageId="26" pageNumber="45">
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<taxonomicName LSID="EC13C1FE-2B89-3787-FFC4-F0A331A0B47A" authority="(Chaudoir, 1837)" baseAuthorityName="Chaudoir" baseAuthorityYear="1837" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Semiardistomis labialis" order="Coleoptera" pageId="26" pageNumber="45" phylum="Arthropoda" rank="species" species="labialis">Semiardistomis labialis (Chaudoir, 1837)</taxonomicName>
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<figureCitation captionStart="Figures 1–2" captionStartId="F1" captionText="Figures 1 - 2. Semiardistomis labialis (Chaudoir). 1 Dorsal aspect 2 Ventral aspect. Legend: a 1 a 2 a 3 antennomeres 1 - 3, respectively. Scale bar 1 mm." httpUri="https://binary.pensoft.net/fig/19659" pageId="26" pageNumber="45">Figs 1, 2</figureCitation>
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<figureCitation captionStart="Figures 3–5" captionStartId="F2" captionText="Figures 3 - 5. 3 - 4 Labrum (dorsal aspect) 3 Semiardistomis glabratus (Putzeys) 4 Semiardistomis labialis (Chaudoir) 5 RightMaxilla (dorsal aspect) of Semiardistomis labialis (Chaudoir). Legend: Labrum: ds dorsal seta es epipharyngeal seta lp labral dentiform projection. Maxilla: c cardo g 1 galeomere 1 g 2 galeomere 2 lac lacinia mp 2 mp 3 mp 4 maxillary palpomeres 2 - 4 respectively pf palpifer s stipes. Scale bar 0.1 mm." httpUri="https://binary.pensoft.net/fig/19660" pageId="26" pageNumber="45">4</figureCitation>
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<figureCitation captionStart="Figures 6–12" captionStartId="F3" captionText="Figures 6 - 12. Mandibles ofArdistomina, Clivinina and Dyschiriina with length / width ratios: A left, dorsal aspect B right, dorsal aspect C left, ventral aspect and D right, ventral aspect) 6 Semiardistomis labialis (Chaudoir) 7 Ardistomis fasciolatus Putzeys 8 Aspidoglossa mexicana (Chaudoir) 9 Oxydrepanus rufus (Putzeys) 10 Clivina dentipes Dejean 11 Schizogenius arimao Darlington 12 Dyschiriodes larochellei Bousquet. Legend: Occlusal teeth: m molar pm premolar rt retinacular tt terebral. Ventral surface: vg ventral groove mt microtrichia. Scale bar 0.1 mm." httpUri="https://binary.pensoft.net/fig/19661" pageId="26" pageNumber="45">6</figureCitation>
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<figureCitation captionStart="Figures 13–21" captionStartId="F4" captionText="Figures 13 - 21. Labium ofClivinini (ventral aspects): A mentum and palpi B ligula C mental carina, lateral aspect. 13 - 14 Genus Semiardistomis: 13 S. glabratus (Putzeys) 14 Semiardistomis labialis (Chaudoir) 15 Ardistomis fasciolatus Putzeys 16 Aspidoglossa mexicana (Chaudoir) 17 Clivina dentipes Dejean 18 Oxydrepanus rufus (Putzeys) 19 Schizogenius arimao Darlington 20 Ancus excavaticeps (Putzeys) 21 Nyctosyles planicollis (Reiche). Legend: as apical setae of glossal sclerite gs glossal sclerite lp 2 labial palpomere 2 lp 3 labial palpomere 3 ll lateral lobes mc median carina m mentum mt mental tooth pas preapical seta of glossal sclerite pg paraglossa. Scale bar 0.1 mm." httpUri="https://binary.pensoft.net/fig/19662" pageId="26" pageNumber="45">14</figureCitation>
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<figureCitation captionStart="Figures 22–24" captionStartId="F5" captionText="Figures 22 - 24. 22 Body extremity (lateral aspect) of Semiardistomis labialis (Chaudoir) 23 - 24 Protarsus (ventral aspect) of Semiardistomis 23 Semiardistomis labialis (Chaudoir) 24 Semiardistomis glabratus (Putzeys). Legend: app abdominal epipleural projectionof abdominal sternum VII ep elytral plica. Scale bar 1 mm." httpUri="https://binary.pensoft.net/fig/19663" pageId="26" pageNumber="45">22, 23</figureCitation>
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<figureCitation captionStart="Figures 25–26" captionStartId="F6" captionText="Figures 25 - 26. Male genitalia of genus Semiardistomis: A phallus and parameres in dorsal aspect B phallus and endophallus everted in lateral aspect 25 Semiardistomis labialis (Chaudoir) 26 Semiardistomis glabratus (Putzeys). Legend: ba basal apophysis bo basal opening bp basal projection bs basal sclerite ll left lobe rl right lobe. Scale bar 0.1 mm." httpUri="https://binary.pensoft.net/fig/19664" pageId="26" pageNumber="45">25</figureCitation>
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<figureCitation captionStart="Figures 27–30" captionStartId="F7" captionText="Figures 27 - 30. Female genitalia (ventral aspect). of Ardistomina 27 Semiardistomis glabratus (Putzeys) 28 Semiardistomis labialis (Chaudoir) 29 Ardistomis fasciolatus Putzeys 30 Aspidoglossa mexicana (Chaudoir). Legend: bc bursa copulatrix gc gonocoxa (gonocoxite 1 and 2, fused) lt laterotergite mt mediotergite sp spermatheca spd spermathecal duct spg spermathecal gland spgd spermathecal gland duct. Scale bar 0.1 mm." httpUri="https://binary.pensoft.net/fig/19665" pageId="26" pageNumber="45">28</figureCitation>
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<figureCitation captionStart="Figure 59" captionStartId="F21" captionText="Figure 59. Distribution map of members of the labialis species-group in Central America." httpUri="https://binary.pensoft.net/fig/19679" pageId="26" pageNumber="45">59</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="26" pageNumber="45" type="reference_group">
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<paragraph pageId="26" pageNumber="45">
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<taxonomicName authorityName="Chaudoir" authorityYear="1837" class="Hexapoda" family="Carabidae" genus="Clivina" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="26" pageNumber="45" phylum="Arthropoda" rank="species" species="labialis">Clivina labialis</taxonomicName>
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Chaudoir, 1837: 18.
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</paragraph>
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<paragraph pageId="26" pageNumber="45">
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<taxonomicName baseAuthorityName="Chaudoir" baseAuthorityYear="1837" class="Hexapoda" family="Carabidae" genus="Ardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="26" pageNumber="45" phylum="Arthropoda" rank="species" species="labialis">Ardistomis labialis</taxonomicName>
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(Chaudoir):
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<bibRefCitation author="Putzeys, JAAH" journalOrPublisher="Memoires de la Societe Royale des Sciences de Liege" pageId="34" pageNumber="53" pagination="521 - 663" refId="B31" refString="Putzeys, JAAH, 1846. Monographie des Clivina et genres voisins, precedee d'un tableau synoptique des genres de la tribu des scaritides. Memoires de la Societe Royale des Sciences de Liege 2: 521 - 663" title="Monographie des Clivina et genres voisins, precedee d'un tableau synoptique des genres de la tribu des scaritides." volume="2" year="1846">Putzeys 1846</bibRefCitation>
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: 648;
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<bibRefCitation author="Bates, HW" editor="Godman, FD" journalOrPublisher="Coleoptera. Volume 1, Part 1. Taylor and Francis, London" pageId="34" pageNumber="53" pagination="1 - 40" refId="B4" refString="Bates, HW, 1881. Insecta, Coleoptera, Carabidae, Cicindelidae. In: Godman, FD, Salvin, O, Eds., Biologia Centrali- Americana. Coleoptera. Volume 1, Part 1. Taylor and Francis, London: 1 - 40" title="Insecta, Coleoptera, Carabidae, Cicindelidae." volumeTitle="Biologia Centrali- Americana." year="1881">Bates 1881</bibRefCitation>
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: 35;
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<bibRefCitation author="Lorenz, W" journalOrPublisher="Paussinae, Cicindelinae, Rhysodinae). W. Lorenz, Tutzing" pageId="34" pageNumber="53" refId="B28" refString="Lorenz, W, 2005. Systematic list of extant ground beetles of the world (Insecta, Coleoptera "Geadephaga": Trachypachidae and Carabidae, incl. Paussinae, Cicindelinae, Rhysodinae). W. Lorenz, Tutzing, 530 pp." title="Systematic list of extant ground beetles of the world (Insecta, Coleoptera " Geadephaga ": Trachypachidae and Carabidae, incl." year="2005">Lorenz 2005</bibRefCitation>
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: 146.
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</paragraph>
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<paragraph pageId="26" pageNumber="45">
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<taxonomicName class="Insecta" family="Carabidae" genus="Ardistomus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="26" pageNumber="45" phylum="Arthropoda" rank="species" species="labialis">Ardistomus labialis</taxonomicName>
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(Chaudoir):
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<bibRefCitation author="Csiki, E" editor="Junk, W" journalOrPublisher="Volume II. Carabidae II, W. Junk, Berlin" pageId="34" pageNumber="53" pagination="739 - 1022" refId="B10" refString="Csiki, E, 1931. Carabidae: Harpalinae V (Pars 115). In: Junk, W, Shenkling, S, Eds., Coleopterorum catalogus. Volume II. Carabidae II, W. Junk, Berlin: 739 - 1022" title="Carabidae: Harpalinae V (Pars 115)." volumeTitle="Coleopterorum catalogus." year="1931">Csiki 1927</bibRefCitation>
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: 548;
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<bibRefCitation author="Blackwelder, RE" journalOrPublisher="Smithsonian Institution, United States National Museum Bulletin" pageId="34" pageNumber="53" pagination="1 - 188" refId="B5" refString="Blackwelder, RE, 1944. Checklist of the coleopterous insects of Mexico, Central America, The West Indies, and South America. Part 1. Smithsonian Institution, United States National Museum Bulletin 185: 1 - 188" title="Checklist of the coleopterous insects of Mexico, Central America, The West Indies, and South America. Part 1." volume="185" year="1944">Blackwelder 1944</bibRefCitation>
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: 27.
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</paragraph>
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<paragraph pageId="26" pageNumber="45">
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<taxonomicName baseAuthorityName="Chaudoir" baseAuthorityYear="1837" class="Hexapoda" family="Carabidae" genus="Ardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="26" pageNumber="45" phylum="Arthropoda" rank="species" species="labialis" subGenus="Semiardistomis">Ardistomis (Semiardistomis) labialis</taxonomicName>
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(Chaudoir):
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<bibRefCitation author="Kult, K" journalOrPublisher="Arthropoda" pageId="34" pageNumber="53" pagination="299 - 325" refId="B20" refString="Kult, K, 1950. New Neotropical species of the group Ardistominina (Carabidae, Col.). Arthropoda 1: 299 - 325" title="New Neotropical species of the group Ardistominina (Carabidae, Col.)." volume="1" year="1950">Kult 1950</bibRefCitation>
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: 301.
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</paragraph>
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<paragraph pageId="26" pageNumber="45">
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<taxonomicName class="Hexapoda" family="Carabidae" genus="Ardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="26" pageNumber="45" phylum="Arthropoda" rank="variety" species="labialis" subGenus="Semiardistomis" variety="picipes">Ardistomis (Semiardistomis) labialis var. picipes</taxonomicName>
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Bates, 1881: 35.
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</paragraph>
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<paragraph pageId="26" pageNumber="45">
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<taxonomicName class="Hexapoda" family="Carabidae" genus="Ardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="26" pageNumber="45" phylum="Arthropoda" rank="variety" species="labialis" subGenus="Semiardistomis" variety="nanus">Ardistomis (Semiardistomis) labialis var. nanus</taxonomicName>
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Bates, 1881: 35.
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</paragraph>
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<paragraph pageId="26" pageNumber="45">
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<taxonomicName class="Hexapoda" family="Carabidae" genus="Ardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="26" pageNumber="45" phylum="Arthropoda" rank="variety" species="labialis" subGenus="Semiardistomis" variety="dilatatus">Ardistomis (Semiardistomis) labialis var. dilatatus</taxonomicName>
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Bates, 1881: 35.
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</paragraph>
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<paragraph pageId="26" pageNumber="45">
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<taxonomicName authorityName="Putzeys" authorityYear="1846" class="Hexapoda" family="Carabidae" genus="Ardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="26" pageNumber="45" phylum="Arthropoda" rank="species" species="tuspanensis">Ardistomis tuspanensis</taxonomicName>
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<bibRefCitation author="Putzeys, JAAH" journalOrPublisher="Memoires de la Societe Royale des Sciences de Liege" pageId="34" pageNumber="53" pagination="521 - 663" refId="B31" refString="Putzeys, JAAH, 1846. Monographie des Clivina et genres voisins, precedee d'un tableau synoptique des genres de la tribu des scaritides. Memoires de la Societe Royale des Sciences de Liege 2: 521 - 663" title="Monographie des Clivina et genres voisins, precedee d'un tableau synoptique des genres de la tribu des scaritides." volume="2" year="1846">Putzeys 1846</bibRefCitation>
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: 649 syn. n.
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</paragraph>
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<paragraph pageId="26" pageNumber="45">
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<taxonomicName baseAuthorityName="Chaudoir" baseAuthorityYear="1837" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="26" pageNumber="45" phylum="Arthropoda" rank="species" species="labialis">Semiardistomis labialis</taxonomicName>
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(Chaudoir);
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<bibRefCitation author="Erwin, TL" journalOrPublisher="Pensoft, Sofia-Moscow" pageId="34" pageNumber="53" refId="B15" refString="Erwin, TL, 2011. A Treatise on the Western Hemisphere Caraboidea (Coleoptera): Their classification, distributions, and ways of life Volume III (Carabidae - loxomeriformes, melaeniformes). Pensoft, Sofia-Moscow, 412 pp." title="A Treatise on the Western Hemisphere Caraboidea (Coleoptera): Their classification, distributions, and ways of life Volume III (Carabidae - loxomeriformes, melaeniformes)." year="2011">Erwin 2011</bibRefCitation>
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: 235.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="26" pageNumber="45" type="type material">
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<paragraph pageId="26" pageNumber="45">Type material.</paragraph>
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<paragraph pageId="26" pageNumber="45">
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Lectotype, here designated, at MHNP, glued on point, labeled: green paper handwritten "Labialis Chaud Mexico Dej."/ handwritten in box "labialis Chaud Mexique Chevrol"/ handwritten
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<normalizedToken originalValue="“38”">"38"</normalizedToken>
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/ "Lectotype Ardistomis labialis Chd. Det George E. Ball 72"/ "Lectotype Clivina labialis Chd. Des. S.W. Nichols 1984"/ printed in red paper
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<normalizedToken originalValue="“lectotype”">"lectotype"</normalizedToken>
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/ printed in circle "Lectotype
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</paragraph>
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<paragraph pageId="26" pageNumber="45">Paralectotypes, here designated: two exemplars at MHNP labeled: handwritten in box "labialis Chaud Mexique Chevrol"; handwritten "Paralectotype Clivina labialis Chd. Des. S.W. Nichols 1984".</paragraph>
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<paragraph pageId="26" pageNumber="45">
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Lectotype of
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<taxonomicName authorityName="Putzeys" authorityYear="1846" class="Hexapoda" family="Carabidae" genus="Ardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="26" pageNumber="45" phylum="Arthropoda" rank="species" species="tuspanensis">
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<emphasis italics="true" pageId="26" pageNumber="45">Ardistomis tuspanensis</emphasis>
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</taxonomicName>
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, here designated, at IRSBN labeled: green paper handwritten " Tuspanensis Tuspan (Chev.)"/ printed "Soc. Ent. Belg. Coll. Putzeys"/ handwritten "P. Basilewsky Ardistomus tuspanensis Putz".
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</paragraph>
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</subSubSection>
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<subSubSection pageId="27" pageNumber="46" type="type area">
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<paragraph pageId="27" pageNumber="46">
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<pageBreakToken pageId="27" pageNumber="46" start="start">Type</pageBreakToken>
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area.
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</paragraph>
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<paragraph pageId="27" pageNumber="46">
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Given by
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<bibRefCitation author="Chaudoir, M de" journalOrPublisher="Bulletin de la Societe Imperiale des Naturalistes de Moscou" pageId="34" pageNumber="53" pagination="3 - 48" refId="B8" refString="Chaudoir, M de, 1837. Genres nouveaux et especes nouvelles de Coleopteres de la famille des Carabiques. Bulletin de la Societe Imperiale des Naturalistes de Moscou 10: 3 - 48" title="Genres nouveaux et especes nouvelles de Coleopteres de la famille des Carabiques." volume="10" year="1837">Chaudoir (1837</bibRefCitation>
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: 19) as
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<normalizedToken originalValue="“Mexique”">"Mexique"</normalizedToken>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="27" pageNumber="46" type="diagnosis">
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<paragraph pageId="27" pageNumber="46">Diagnosis.</paragraph>
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<paragraph pageId="27" pageNumber="46">Body piceous with green reflections. Elytral surface completely smooth, striae continuously impressed in basal 1/8, punctures deeply impressed in basal 2/3, 3 setiferous pores on interval 3. Abdominal sternum VII with 4+4 setiferous punctures.</paragraph>
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</subSubSection>
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<subSubSection pageId="27" pageNumber="46" type="habitus">
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<paragraph pageId="27" pageNumber="46">Habitus.</paragraph>
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<paragraph pageId="27" pageNumber="46">
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dorsal and ventral aspects, respectively, as in
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<figureCitation captionStart="Figures 1–2" captionStartId="F1" captionText="Figures 1 - 2. Semiardistomis labialis (Chaudoir). 1 Dorsal aspect 2 Ventral aspect. Legend: a 1 a 2 a 3 antennomeres 1 - 3, respectively. Scale bar 1 mm." httpUri="https://binary.pensoft.net/fig/19659" pageId="27" pageNumber="46">Figs 1, 2</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="27" pageNumber="46" type="mouthparts">
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<paragraph pageId="27" pageNumber="46">Mouthparts.</paragraph>
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<paragraph pageId="27" pageNumber="46">
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<emphasis bold="true" pageId="27" pageNumber="46">Labrum</emphasis>
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, dorsal aspect, as in
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<figureCitation captionStart="Figures 3–5" captionStartId="F2" captionText="Figures 3 - 5. 3 - 4 Labrum (dorsal aspect) 3 Semiardistomis glabratus (Putzeys) 4 Semiardistomis labialis (Chaudoir) 5 RightMaxilla (dorsal aspect) of Semiardistomis labialis (Chaudoir). Legend: Labrum: ds dorsal seta es epipharyngeal seta lp labral dentiform projection. Maxilla: c cardo g 1 galeomere 1 g 2 galeomere 2 lac lacinia mp 2 mp 3 mp 4 maxillary palpomeres 2 - 4 respectively pf palpifer s stipes. Scale bar 0.1 mm." httpUri="https://binary.pensoft.net/fig/19660" pageId="27" pageNumber="46">Fig. 4</figureCitation>
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.
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<emphasis bold="true" pageId="27" pageNumber="46">Labium</emphasis>
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, as in
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<figureCitation captionStart="Figures 13–21" captionStartId="F4" captionText="Figures 13 - 21. Labium ofClivinini (ventral aspects): A mentum and palpi B ligula C mental carina, lateral aspect. 13 - 14 Genus Semiardistomis: 13 S. glabratus (Putzeys) 14 Semiardistomis labialis (Chaudoir) 15 Ardistomis fasciolatus Putzeys 16 Aspidoglossa mexicana (Chaudoir) 17 Clivina dentipes Dejean 18 Oxydrepanus rufus (Putzeys) 19 Schizogenius arimao Darlington 20 Ancus excavaticeps (Putzeys) 21 Nyctosyles planicollis (Reiche). Legend: as apical setae of glossal sclerite gs glossal sclerite lp 2 labial palpomere 2 lp 3 labial palpomere 3 ll lateral lobes mc median carina m mentum mt mental tooth pas preapical seta of glossal sclerite pg paraglossa. Scale bar 0.1 mm." httpUri="https://binary.pensoft.net/fig/19662" pageId="27" pageNumber="46">Fig. 14A</figureCitation>
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; ligula, enlarged, as in
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<figureCitation captionStart="Figures 13–21" captionStartId="F4" captionText="Figures 13 - 21. Labium ofClivinini (ventral aspects): A mentum and palpi B ligula C mental carina, lateral aspect. 13 - 14 Genus Semiardistomis: 13 S. glabratus (Putzeys) 14 Semiardistomis labialis (Chaudoir) 15 Ardistomis fasciolatus Putzeys 16 Aspidoglossa mexicana (Chaudoir) 17 Clivina dentipes Dejean 18 Oxydrepanus rufus (Putzeys) 19 Schizogenius arimao Darlington 20 Ancus excavaticeps (Putzeys) 21 Nyctosyles planicollis (Reiche). Legend: as apical setae of glossal sclerite gs glossal sclerite lp 2 labial palpomere 2 lp 3 labial palpomere 3 ll lateral lobes mc median carina m mentum mt mental tooth pas preapical seta of glossal sclerite pg paraglossa. Scale bar 0.1 mm." httpUri="https://binary.pensoft.net/fig/19662" pageId="27" pageNumber="46">Fig. 14B</figureCitation>
|
||
..
|
||
</paragraph>
|
||
<paragraph pageId="27" pageNumber="46">
|
||
<emphasis bold="true" pageId="27" pageNumber="46">Legs:</emphasis>
|
||
male protarsus as in
|
||
<figureCitation captionStart="Figures 22–24" captionStartId="F5" captionText="Figures 22 - 24. 22 Body extremity (lateral aspect) of Semiardistomis labialis (Chaudoir) 23 - 24 Protarsus (ventral aspect) of Semiardistomis 23 Semiardistomis labialis (Chaudoir) 24 Semiardistomis glabratus (Putzeys). Legend: app abdominal epipleural projectionof abdominal sternum VII ep elytral plica. Scale bar 1 mm." httpUri="https://binary.pensoft.net/fig/19663" pageId="27" pageNumber="46">Fig. 23</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph pageId="27" pageNumber="46">
|
||
<emphasis bold="true" pageId="27" pageNumber="46">Male genitalia:</emphasis>
|
||
as in
|
||
<figureCitation captionStart="Figures 25–26" captionStartId="F6" captionText="Figures 25 - 26. Male genitalia of genus Semiardistomis: A phallus and parameres in dorsal aspect B phallus and endophallus everted in lateral aspect 25 Semiardistomis labialis (Chaudoir) 26 Semiardistomis glabratus (Putzeys). Legend: ba basal apophysis bo basal opening bp basal projection bs basal sclerite ll left lobe rl right lobe. Scale bar 0.1 mm." httpUri="https://binary.pensoft.net/fig/19664" pageId="27" pageNumber="46">Fig. 25</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph pageId="27" pageNumber="46">
|
||
<emphasis bold="true" pageId="27" pageNumber="46">Female genitalia:</emphasis>
|
||
as in
|
||
<figureCitation captionStart="Figures 27–30" captionStartId="F7" captionText="Figures 27 - 30. Female genitalia (ventral aspect). of Ardistomina 27 Semiardistomis glabratus (Putzeys) 28 Semiardistomis labialis (Chaudoir) 29 Ardistomis fasciolatus Putzeys 30 Aspidoglossa mexicana (Chaudoir). Legend: bc bursa copulatrix gc gonocoxa (gonocoxite 1 and 2, fused) lt laterotergite mt mediotergite sp spermatheca spd spermathecal duct spg spermathecal gland spgd spermathecal gland duct. Scale bar 0.1 mm." httpUri="https://binary.pensoft.net/fig/19665" pageId="27" pageNumber="46">Fig. 28</figureCitation>
|
||
<emphasis bold="true" pageId="27" pageNumber="46">.</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="27" pageNumber="46" type="measurements and variation">
|
||
<paragraph pageId="27" pageNumber="46">Measurements and variation.</paragraph>
|
||
<paragraph pageId="27" pageNumber="46">
|
||
Variation of measurements (mm) and ratios for
|
||
<taxonomicName baseAuthorityName="Chaudoir" baseAuthorityYear="1837" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="27" pageNumber="46" phylum="Arthropoda" rank="species" species="labialis">
|
||
<emphasis italics="true" pageId="27" pageNumber="46">Semiardistomis labialis</emphasis>
|
||
</taxonomicName>
|
||
from Mexico (n=11) are: HL = 0,53-0,58-0,68; PL = 0,90-1,11-1,25; PW = 1,00-1,23-1,40; EL = 2,65-3,10-3,40; EW = 1,70-1,94-2,20;
|
||
<emphasis bold="true" pageId="27" pageNumber="46">SBL =</emphasis>
|
||
4,10-4,80-5,23; PW/EW = 0,63; PW/PL = 1,10; PL/EL = 0,36; EW/EL = 0,63.
|
||
</paragraph>
|
||
<paragraph pageId="27" pageNumber="46">
|
||
Variation of measurements (mm) and ratios for
|
||
<taxonomicName baseAuthorityName="Chaudoir" baseAuthorityYear="1837" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="27" pageNumber="46" phylum="Arthropoda" rank="species" species="labialis">
|
||
<emphasis italics="true" pageId="27" pageNumber="46">Semiardistomis labialis</emphasis>
|
||
</taxonomicName>
|
||
from Costa Rica (n=11) are: HL = 0,54-0,60-0,68; PL 1,00-1,07-1,20; PW 1,10-1,21-1,36; EL 2,60-2,86-3,20; EW 1,68-1,86-2,16;
|
||
<emphasis bold="true" pageId="27" pageNumber="46">SBL</emphasis>
|
||
4,14-4,54-5,08; PW/EW 0,65; PW/PL 1,13; PL/EL 0,38; EW/EL 0,65.
|
||
</paragraph>
|
||
<paragraph pageId="27" pageNumber="46">
|
||
Variation of measurements (mm) and ratios for
|
||
<taxonomicName baseAuthorityName="Chaudoir" baseAuthorityYear="1837" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="27" pageNumber="46" phylum="Arthropoda" rank="species" species="labialis">
|
||
<emphasis italics="true" pageId="27" pageNumber="46">Semiardistomis labialis</emphasis>
|
||
</taxonomicName>
|
||
identified in collections as
|
||
<taxonomicName baseAuthorityName="Putzeys" baseAuthorityYear="1846" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="27" pageNumber="46" phylum="Arthropoda" rank="species" species="tuspanensis">
|
||
<emphasis italics="true" pageId="27" pageNumber="46">Semiardistomis tuspanensis</emphasis>
|
||
</taxonomicName>
|
||
(included syntypes) (n= 5) are: HL = 0,55-0,60-0,68; PL 1,03-1,08-1,23; PW 1,08-1,18-1,30; EL 2,65-2,80-3,00; EW 1,70-1,75-1,85;
|
||
<emphasis bold="true" pageId="27" pageNumber="46">SBL</emphasis>
|
||
4,33-4,48-4,63; PW/EW 0,67; PW/PL 1,09; PL/EL 0,39; EW/EL 0,63.
|
||
</paragraph>
|
||
<paragraph pageId="27" pageNumber="46">
|
||
<bibRefCitation author="Bates, HW" editor="Godman, FD" journalOrPublisher="Coleoptera. Volume 1, Part 1. Taylor and Francis, London" pageId="34" pageNumber="53" pagination="1 - 40" refId="B4" refString="Bates, HW, 1881. Insecta, Coleoptera, Carabidae, Cicindelidae. In: Godman, FD, Salvin, O, Eds., Biologia Centrali- Americana. Coleoptera. Volume 1, Part 1. Taylor and Francis, London: 1 - 40" title="Insecta, Coleoptera, Carabidae, Cicindelidae." volumeTitle="Biologia Centrali- Americana." year="1881">Bates (1881)</bibRefCitation>
|
||
pointed the high variability of
|
||
<emphasis italics="true" pageId="27" pageNumber="46">labialis</emphasis>
|
||
populations in Central America. The polymorphic condition of characters, such as body measurements and punctuation of elytra striae, have arose different designations that includes
|
||
<normalizedToken originalValue="Bates’">Bates'</normalizedToken>
|
||
varieties and
|
||
<taxonomicName baseAuthorityName="Putzeys" baseAuthorityYear="1846" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="27" pageNumber="46" phylum="Arthropoda" rank="species" species="tuspanensis">
|
||
<emphasis italics="true" pageId="27" pageNumber="46">Semiardistomis tuspanensis</emphasis>
|
||
</taxonomicName>
|
||
(Putzeys). Body sizes show little variation between geographic regions and since for each measured sample group, distance between extremes measurements is high, this character is useless to identify forms proposed. About elytra disc punctured striae, forms varies in low grades without following any clear geographic pattern having striae punctures weekly to strongly produced and striae almost disappeared (except stria number one) to moderately impressed.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="27" pageNumber="46" type="geographical distribution">
|
||
<paragraph pageId="27" pageNumber="46">Geographical distribution</paragraph>
|
||
<paragraph pageId="27" pageNumber="46">
|
||
(
|
||
<figureCitation captionStart="Figure 59" captionStartId="F21" captionText="Figure 59. Distribution map of members of the labialis species-group in Central America." httpUri="https://binary.pensoft.net/fig/19679" pageId="27" pageNumber="46">Fig. 59</figureCitation>
|
||
)
|
||
<emphasis bold="true" pageId="27" pageNumber="46">.</emphasis>
|
||
The known range of this species extends in Middle America from central Costa Rica northward to northwestern Mexico (slightly north of the Tropic of Cancer).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="27" pageNumber="46" type="activity">
|
||
<paragraph pageId="27" pageNumber="46">Activity.</paragraph>
|
||
<paragraph pageId="27" pageNumber="46">Most label data indicate "attracted to lights". Tenerals were collected in November.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="28" lastPageNumber="47" pageId="27" pageNumber="46" type="material examined">
|
||
<paragraph pageId="27" pageNumber="46">Material examined.</paragraph>
|
||
<paragraph lastPageId="28" lastPageNumber="47" pageId="27" pageNumber="46">
|
||
MEXICO. Sinaloa,
|
||
<normalizedToken originalValue="Culiacán">Culiacan</normalizedToken>
|
||
Apr 1969 (USNM, 9) Veracruz, Cordoba Nov. 1966 (USNM, 7) Oaxaca Tehuantepec Dec. 1964 (USNM, 1) Chiapas Tapilulas May 1974 (USNM, 9) Colima Volcano (USNM, 5) Jalapa (IRSNB, 2) Sa. De Durango (IRSNB, 4) Guerrero Chiapas Jul 2005 (PBPC, 2). BELIZE. Sibun River at Gracy Rock Jun 1974 (USNM, 7). GUATEMALA. Alta Verapaz May 1973 (USNM, 40) Sacapulas 4500ft Dec. 1947 (USNM, 3). HONDURAS. Dept. Con
|
||
<pageBreakToken pageId="28" pageNumber="47" start="start">ayagua</pageBreakToken>
|
||
Rancho Chiquito May 1964 (UASM, 8), Belize district Jun 1968 (USNM, 18), 5 mi E. Choluteca Jul 1965 (USNM, 22), 16 mi W. Sabana Grande Jul 1965 (USNM, 7), Pespire Jul 1965 (USNM, 7), San Marcus Colon Jul. 1965 (USNM, 10). EL SALVADOR. Ch. Del Guayabo May 1975 (USNM, 1). NICARAGUA. Rivas Rio Canas Gordas Jun 1964 (UASM, 18). COSTA RICA. Guanacaste Santa Rosa NP Jun 2004 (ADVA, 34), Guanacaste NP Jun 1990 (USNM, 9), Guanacaste Santa Cruz Est. Bosque Diria 150- 250 m Nov 1998 (USNM, 18), Guanacaste Santa Rosa 300 m Mar 1990 (USNM, 5).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |