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<document id="48EBDC14D67B18EB398C04A95F5A09A4" ID-CLB-Dataset="128835" ID-DOI="10.5962/bhl.part.117806" ID-GBIF-Dataset="3ddb85a2-5038-40de-b8a0-75cfb6bbef9f" ID-ISSN="0035-418X" ID-Zenodo-Dep="5828501" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1641578156393" checkinUser="carolina" docAuthor="Hong, Yong" docDate="2011" docId="BF3E87AFFF84361640DBF985FD20573E" docLanguage="en" docName="RevSuisseZool.118.2.19-26.pdf.imf" docOrigin="Revue suisse de Zoologie 118 (2)" docSource="http://www.biodiversitylibrary.org/part/117806" docStyle="DocumentStyle:0846B485C0C87F6EB7648DEEBF34B77F.2:RevSuisseZool.2011-2014.journal_article" docStyleId="0846B485C0C87F6EB7648DEEBF34B77F" docStyleName="RevSuisseZool.2011-2014.journal_article" docStyleVersion="2" docTitle="Amynthas samgaki Hong 2011, sp. n." docType="treatment" docVersion="8" lastPageNumber="229" masterDocId="4307FFD7FF873610404CFFADFFE45424" masterDocTitle="Two new species of Amynthas (Clitellata: Megascolecidae) from lettuce fields of Mt. Taebaek, Korea" masterLastPageNumber="230" masterPageNumber="223" pageNumber="226" updateTime="1699296750336" updateUser="ExternalLinkService">
<mods:mods id="05761DCDFAACB3F0FCC3B790299C511F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C644CC1ECF91A1551595CE4377B69A3B">
<mods:title id="F4C17A7EC0AC4542F089663CCF90B4B4">Two new species of Amynthas (Clitellata: Megascolecidae) from lettuce fields of Mt. Taebaek, Korea</mods:title>
</mods:titleInfo>
<mods:name id="C2EC54C013C74D493D5E927335635511" type="personal">
<mods:role id="DA10B0F16B3381F9E09B725BC6CD6025">
<mods:roleTerm id="BCB35203DC052DFFF79288FD629B806D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0CC567AB6D06206B0ADE6F3DB1C30B8C">Hong, Yong</mods:namePart>
</mods:name>
<mods:typeOfResource id="F7366089566FF5CE42D0EDE318D8B424">text</mods:typeOfResource>
<mods:relatedItem id="2C7E6CFB8AC35F19BB6DA9770D5303A5" type="host">
<mods:titleInfo id="7BDF422F095B506567CC0F23866BBEC9">
<mods:title id="5D5846513B6A5D0D4F121F16C57926D7">Revue suisse de Zoologie</mods:title>
</mods:titleInfo>
<mods:part id="7496CFEE24A109C94F55061B7E0180F0">
<mods:date id="5D598BE980AB88AFFD1B356F79F16BED">2011</mods:date>
<mods:detail id="E01E6E96FF292643A845CCFB32AB54A0" type="pubDate">
<mods:number id="03FCCDD8AFF6AA0A099AB97D7BE3C77E">2011-06-30</mods:number>
</mods:detail>
<mods:detail id="5110635D59C1103D9E49DD52497075F6" type="volume">
<mods:number id="FCDEFF57E5584D8B4105F7D8CD92850F">118</mods:number>
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<mods:number id="CE98A578190E942BDE3BF937AD1168EA">2</mods:number>
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<mods:start id="7EAF461C10EAAE173BEC309072F82FA2">223</mods:start>
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<mods:url id="72F471C39FEFDBB45467E71B507400CC">http://www.biodiversitylibrary.org/part/117806</mods:url>
</mods:location>
<mods:classification id="F69716E21AC7598142D3F7F7A56AA695">journal article</mods:classification>
<mods:identifier id="F4B6E3B1A0D030011B592B8E96F5C471" type="CLB-Dataset">128835</mods:identifier>
<mods:identifier id="B3D8467871505D6AF0AE9A392874A912" type="DOI">10.5962/bhl.part.117806</mods:identifier>
<mods:identifier id="CC7F684B2858F49B8CA2F7884092F25D" type="GBIF-Dataset">3ddb85a2-5038-40de-b8a0-75cfb6bbef9f</mods:identifier>
<mods:identifier id="5E485FC20A2987E7EAE05E25D86E91F0" type="ISSN">0035-418X</mods:identifier>
<mods:identifier id="AB6BE278E82C71999949FBCD2CC14696" type="Zenodo-Dep">5828501</mods:identifier>
</mods:mods>
<treatment id="BF3E87AFFF84361640DBF985FD20573E" ID-DOI="http://doi.org/10.5281/zenodo.7023165" ID-GBIF-Taxon="199832990" ID-Zenodo-Dep="7023165" LSID="urn:lsid:plazi:treatment:BF3E87AFFF84361640DBF985FD20573E" httpUri="http://treatment.plazi.org/id/BF3E87AFFF84361640DBF985FD20573E" lastPageId="6" lastPageNumber="229" pageId="3" pageNumber="226">
<subSubSection id="7F8D6532FF84361340DBF985FE4F5264" box="[151,427,1576,1600]" pageId="3" pageNumber="226" type="nomenclature">
<paragraph id="372836B9FF84361340DBF985FE4F5264" blockId="3.[151,1073,1576,1600]" box="[151,427,1576,1600]" pageId="3" pageNumber="226">
<heading id="6C6081D5FF84361340DBF985FE4F5264" box="[151,427,1576,1600]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="226" reason="2">
<taxonomicName id="F0974D3AFF84361340DBF985FE8E5264" authority="Hong, 2011" authorityName="Hong" authorityYear="2011" box="[151,362,1576,1600]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="3" pageNumber="226" phylum="Annelida" rank="species" species="samgaki" status="sp. nov.">
<emphasis id="05E3EAABFF84361340DBF985FE8E5264" bold="true" box="[151,362,1576,1600]" italics="true" pageId="3" pageNumber="226">Amynthas samgaki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1ED057D0FF843613413DF984FE4F5264" box="[369,427,1577,1600]" pageId="3" pageNumber="226" rank="species">sp. n.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="7F8D6532FF843613438DF985FBD55264" box="[961,1073,1576,1600]" pageId="3" pageNumber="226" type="description">
<paragraph id="372836B9FF843613438DF985FBD55264" blockId="3.[151,1073,1576,1600]" box="[961,1073,1576,1600]" pageId="3" pageNumber="226">
<figureCitation id="AFAC2A3CFF843613438DF985FBE75264" box="[961,1027,1576,1600]" captionStart="FIG" captionStartId="4.[569,601,1582,1603]" captionTargetBox="[315,875,242,1497]" captionTargetId="figure-27@4.[287,904,229,1514]" captionTargetPageId="4" captionText="FIG. 2 Amynthas samgaki sp. n. (A) Ventral view. (B) Spermathecae and diverticulum. Scale bars = 4 mm (A), 2 mm (B)." figureDoi="http://doi.org/10.5281/zenodo.5828505" httpUri="https://zenodo.org/record/5828505/files/figure.png" pageId="3" pageNumber="226">Figs 2</figureCitation>
A-B
</paragraph>
</subSubSection>
<subSubSection id="7F8D6532FF84361540AFF9E1FBF55560" lastPageId="5" lastPageNumber="228" pageId="3" pageNumber="226" type="materials_examined">
<paragraph id="372836B9FF84361540AFF9E1FBF55560" blockId="3.[151,1073,1611,1659]" lastBlockId="5.[151,1074,227,325]" lastPageId="5" lastPageNumber="228" pageId="3" pageNumber="226">
MATERIAL:
<materialsCitation id="87FF3CE4FF8436154115F9E6FCED54DC" ID-GBIF-Occurrence="3893495303" collectingDate="2006-06-29" collectionCode="NIBRIV" collectorName="Y. Hong" country="South Korea" elevation="700" lastPageId="5" lastPageNumber="228" latitude="37.234566" location="Samsu-dong" longLatPrecision="1" longitude="128.98586" municipality="Taebaek" pageId="3" pageNumber="226" specimenCode="NIBRIV0000224642" specimenCount="1" stateProvince="Gangwon province" typeStatus="holotype">
<typeStatus id="E82C881BFF8436134115F9E6FE575245" box="[345,435,1611,1633]" pageId="3" pageNumber="226" type="holotype">Holotype</typeStatus>
;
<specimenCount id="2191FD30FF8436134184F9E6FDFF5245" box="[456,539,1611,1633]" pageId="3" pageNumber="226">clitellate</specimenCount>
(
<specimenCode id="67319EC2FF843613427EF9E1FD195245" box="[562,765,1611,1633]" collectionCode="NIBRIV" pageId="3" pageNumber="226">NIBRIV0000224642</specimenCode>
);
<collectingCountry id="4F807629FF8436134357F9E1FCB25245" box="[795,854,1612,1633]" name="South Korea" pageId="3" pageNumber="226">Korea</collectingCountry>
,
<collectingRegion id="F553F85BFF8436134327F9E6FBCF5245" box="[875,1067,1611,1633]" country="South Korea" name="Gangwon" pageId="3" pageNumber="226">Gangwon province</collectingRegion>
,
<collectingMunicipality id="D74CACC3FF84361340DBF9C8FF03525F" box="[151,231,1637,1659]" pageId="3" pageNumber="226">Taebaek</collectingMunicipality>
municipality,
<location id="32486062FF8436134139F9C8FE09525F" LSID="urn:lsid:plazi:treatment:BF3E87AFFF84361640DBF985FD20573E:32486062FF8436134139F9C8FE09525F" box="[373,493,1637,1659]" country="South Korea" latitude="37.234566" longLatPrecision="1" longitude="128.98586" municipality="Taebaek" name="Samsu-dong" pageId="3" pageNumber="226" stateProvince="Gangwon province">Samsu-dong</location>
,
<location id="32486062FF84361341B7F9C8FD8D525F" LSID="urn:lsid:plazi:treatment:BF3E87AFFF84361640DBF985FD20573E:32486062FF84361341B7F9C8FD8D525F" box="[507,617,1637,1659]" country="South Korea" latitude="37.234566" longLatPrecision="1" longitude="128.98586" municipality="Taebaek" name="Hasam 2 - ri" pageId="3" pageNumber="226" stateProvince="Gangwon province">Hasam 2-ri</location>
,
<location id="32486062FF843613423BF9C8FD15525F" LSID="urn:lsid:plazi:treatment:BF3E87AFFF84361640DBF985FD20573E:32486062FF843613423BF9C8FD15525F" box="[631,753,1637,1659]" country="South Korea" latitude="37.234566" longLatPrecision="1" longitude="128.98586" municipality="Taebaek" name="Mt. Taebaek" pageId="3" pageNumber="226" stateProvince="Gangwon province">Mt. Taebaek</location>
(
<geoCoordinate id="52A3507EFF843613434DF9C8FC62525E" box="[769,902,1637,1658]" degrees="37" direction="north" minutes="14. 074" orientation="latitude" pageId="3" pageNumber="226" precision="1" value="37.234566">37° 14.074’N</geoCoordinate>
,
<geoCoordinate id="52A3507EFF84361343D9F9C8FBC0525E" box="[917,1060,1637,1659]" degrees="128" direction="east" minutes="59. 151" orientation="longitude" pageId="3" pageNumber="226" precision="1" value="128.98586">128° 59.151’E</geoCoordinate>
),
<quantity id="F06F9B5CFF82361540DBFF4EFEE054DC" box="[151,260,227,249]" metricMagnitude="2" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="5" pageNumber="228" unit="m" value="700.0" valueMax="800.0" valueMin="600.0">
<elevation id="BCBAD18AFF82361540DBFF4EFEE054DC" box="[151,260,227,249]" metricMagnitude="2" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="5" pageNumber="228" unit="m" value="700.0" valueMax="800.0" valueMin="600.0">600-800 m</elevation>
</quantity>
, soil and litter layers,
<date id="43291079FF82361541A0FF4EFD9754DD" box="[492,627,227,249]" pageId="5" pageNumber="228" value="2006-06-29">
<collectingDate id="536DE991FF82361541A0FF4EFD9754DD" box="[492,627,227,249]" pageId="5" pageNumber="228" value="2006-06-29">29 June 2006</collectingDate>
</date>
,
<collectorName id="9A62536FFF82361542CEFF4EFD3154DC" box="[642,725,227,248]" pageId="5" pageNumber="228">Y. Hong</collectorName>
coll.
</materialsCitation>
–
<materialsCitation id="87FF3CE4FF8236154364FF4EFC265536" ID-GBIF-Occurrence="3893495301" collectingDate="2006-06-29" collectionCode="NI, MHNG" collectorName="Y. Hong" country="South Korea" elevation="700" lastPageId="5" lastPageNumber="228" latitude="37.234566" location="Samsu-dong" longLatPrecision="1" longitude="128.98586" municipality="Taebaek" pageId="5" pageNumber="228" specimenCode="NI- BRIV0000224643, MHNG INVE 75338" specimenCount="2" stateProvince="Gangwon province" typeStatus="paratype">
<typeStatus id="E82C881BFF8236154364FF4EFC6154DC" box="[808,901,227,248]" pageId="5" pageNumber="228" type="paratype">Paratypes</typeStatus>
;
<specimenCount id="2191FD30FF82361543DAFF4EFC4654DC" box="[918,930,227,248]" pageId="5" pageNumber="228">1</specimenCount>
clitellate (
<specimenCode id="67319EC2FF823615445CFF4EFEAD5536" pageId="5" pageNumber="228">NIBRIV0000224643</specimenCode>
),
<specimenCount id="2191FD30FF8236154113FF51FE8F5535" box="[351,363,252,273]" pageId="5" pageNumber="228">1</specimenCount>
clitellate (
<specimenCode id="67319EC2FF823615419AFF51FD425536" box="[470,678,252,274]" pageId="5" pageNumber="228">MHNG INVE 75338</specimenCode>
); same data as for holotype.
</materialsCitation>
–
<location id="32486062FF8236154393FF50FBD65536" LSID="urn:lsid:plazi:treatment:BF3E87AFFF84361640DBF985FD20573E:32486062FF8236154393FF50FBD65536" box="[991,1074,253,274]" county="Mt. Taebaek" latitude="37.234566" longLatPrecision="1" longitude="128.98586" municipality="Hasam" name="Nontype" pageId="5" pageNumber="228">Nontype</location>
material;
<materialsCitation id="87FF3CE4FF82361540BAFEBBFBF55560" ID-GBIF-Occurrence="3893495305" collectingDate="2007-07-04" collectorName="Y. Hong" elevation="700" latitude="37.234566" location="Samsu-dong" longLatPrecision="1" longitude="128.98586" municipality="Taebaek" pageId="5" pageNumber="228" specimenCount="1">
<specimenCount id="2191FD30FF82361540BAFEBBFEE6550F" box="[246,258,278,299]" pageId="5" pageNumber="228">1</specimenCount>
clitellate specimen,
<collectingMunicipality id="D74CACC3FF823615419DFEB8FDC5550F" box="[465,545,277,299]" pageId="5" pageNumber="228">Taebaek</collectingMunicipality>
municipality,
<location id="32486062FF82361542FDFEB8FCCD550F" LSID="urn:lsid:plazi:treatment:BF3E87AFFF84361640DBF985FD20573E:32486062FF82361542FDFEB8FCCD550F" box="[689,809,277,299]" county="Mt. Taebaek" latitude="37.234566" longLatPrecision="1" longitude="128.98586" municipality="Taebaek" name="Samsu-dong" pageId="5" pageNumber="228">Samsu-dong</location>
,
<location id="32486062FF8236154375FEBBFC4C550F" LSID="urn:lsid:plazi:treatment:BF3E87AFFF84361640DBF985FD20573E:32486062FF8236154375FEBBFC4C550F" box="[825,936,277,299]" latitude="37.234566" longLatPrecision="1" longitude="128.98586" municipality="Taebaek" name="Hasam 2 - ri" pageId="5" pageNumber="228">Hasam 2-ri</location>
,
<location id="32486062FF82361543FBFEBBFBD6550F" LSID="urn:lsid:plazi:treatment:BF3E87AFFF84361640DBF985FD20573E:32486062FF82361543FBFEBBFBD6550F" box="[951,1074,277,299]" latitude="37.234566" longLatPrecision="1" longitude="128.98586" municipality="Taebaek" name="Mt. Taebaek" pageId="5" pageNumber="228">Mt. Taebaek</location>
(
<geoCoordinate id="52A3507EFF82361540D3FE82FEC65560" box="[159,290,303,324]" degrees="37" direction="north" minutes="14. 074" orientation="latitude" pageId="5" pageNumber="228" precision="1" value="37.234566">37° 14.074’N</geoCoordinate>
,
<geoCoordinate id="52A3507EFF8236154163FE82FE585560" box="[303,444,303,325]" degrees="128" direction="east" minutes="59. 151" orientation="longitude" pageId="5" pageNumber="228" precision="1" value="128.98586">128° 59.151’E</geoCoordinate>
),
<quantity id="F06F9B5CFF823615419DFE82FDDE5560" box="[465,570,303,325]" metricMagnitude="2" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="5" pageNumber="228" unit="m" value="700.0" valueMax="800.0" valueMin="600.0">
<elevation id="BCBAD18AFF823615419DFE82FDDE5560" box="[465,570,303,325]" metricMagnitude="2" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="5" pageNumber="228" unit="m" value="700.0" valueMax="800.0" valueMin="600.0">600-800 m</elevation>
</quantity>
, soil and litter layers,
<date id="43291079FF823615435FFE82FC605560" box="[787,900,303,325]" pageId="5" pageNumber="228" value="2007-07-04">
<collectingDate id="536DE991FF823615435FFE82FC605560" box="[787,900,303,325]" pageId="5" pageNumber="228" value="2007-07-04">4 July 2007</collectingDate>
</date>
,
<collectorName id="9A62536FFF82361543DCFE82FC045560" box="[912,992,303,324]" pageId="5" pageNumber="228">Y. Hong</collectorName>
coll.
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="63E86631FF8336144275F983FEBD525F" ID-DOI="http://doi.org/10.5281/zenodo.5828505" ID-Zenodo-Dep="5828505" httpUri="https://zenodo.org/record/5828505/files/figure.png" pageId="4" pageNumber="227" startId="4.[569,601,1582,1603]" targetBox="[315,875,242,1497]" targetPageId="4">
<paragraph id="372836B9FF8336144275F983FEBD525F" blockId="4.[136,1059,1582,1660]" pageId="4" pageNumber="227">
FIG. 2
<taxonomicName id="F0974D3AFF83361440C4F9E1FEDB5245" authority="Hong, 2011" authorityName="Hong" authorityYear="2011" box="[136,319,1612,1633]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="227" phylum="Annelida" rank="species" species="samgaki" status="sp. nov.">
<emphasis id="05E3EAABFF83361440C4F9E1FEDB5245" box="[136,319,1612,1633]" italics="true" pageId="4" pageNumber="227">Amynthas samgaki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1ED057D0FF8336144104F9E0FE9A5246" box="[328,382,1613,1634]" pageId="4" pageNumber="227" rank="species">sp. n.</taxonomicNameLabel>
(A) Ventral view. (B) Spermathecae and diverticulum. Scale bars =
<quantity id="F06F9B5CFF83361440C4F9CBFF24525F" box="[136,192,1638,1659]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="227" unit="mm" value="4.0">4 mm</quantity>
(A),
<quantity id="F06F9B5CFF83361440B8F9CBFEC8525F" box="[244,300,1638,1659]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="227" unit="mm" value="2.0">2 mm</quantity>
(B).
</paragraph>
</caption>
<subSubSection id="7F8D6532FF82361540AFFEF5FF0855AB" pageId="5" pageNumber="228" type="etymology">
<paragraph id="372836B9FF82361540AFFEF5FF0855AB" blockId="5.[151,1073,344,400]" pageId="5" pageNumber="228">
ETYMOLOGY: The species is named after Sim Samgak, whose farm is its
<typeStatus id="E82C881BFF823615444FFEF5FBD55554" box="[1027,1073,344,368]" pageId="5" pageNumber="228">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="7F8D6532FF82361540AFFE0EFD1555DE" pageId="5" pageNumber="228" type="diagnosis">
<paragraph id="372836B9FF82361540AFFE0EFD1555DE" blockId="5.[151,1073,418,506]" pageId="5" pageNumber="228">DIAGNOSIS: Paired spermathecal pores in 5/6, 6/7; male field with large eggshaped raised pads with transverse seminal groove across pad center, 2.5-3.0 mm distance between male pores, testes sacs ventral, paired.</paragraph>
</subSubSection>
<subSubSection id="7F8D6532FF82361540AFFDBDFDAB5123" pageId="5" pageNumber="228" type="description">
<paragraph id="372836B9FF82361540AFFDBDFE0156C3" blockId="5.[151,1074,528,1288]" pageId="5" pageNumber="228">
DESCRIPTION: Dimensions
<quantity id="F06F9B5CFF823615424EFDBDFD765603" box="[514,658,528,552]" metricMagnitude="-2" metricUnit="m" metricValue="9.375" metricValueMax="11.05" metricValueMin="7.7" pageId="5" pageNumber="228" unit="mm" value="93.75" valueMax="110.5" valueMin="77.0">77-110.5 mm</quantity>
by
<quantity id="F06F9B5CFF82361542F3FDBDFCD85603" box="[703,828,528,552]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" metricValueMax="4.7" metricValueMin="4.5" pageId="5" pageNumber="228" unit="mm" value="4.6" valueMax="4.7" valueMin="4.5">4.5-4.7 mm</quantity>
at segment X, 6.
<quantity id="F06F9B5CFF82361543B6FDBDFF245663" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="6.5" metricValueMin="0.0" pageId="5" pageNumber="228" unit="mm" value="3.25" valueMax="6.5" valueMin="0.0">0-6.5 mm</quantity>
at segment XXX,
<quantity id="F06F9B5CFF82361541C5FD9DFDE15663" box="[393,517,560,584]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="6.5" metricValueMin="5.5" pageId="5" pageNumber="228" unit="mm" value="6.0" valueMax="6.5" valueMin="5.5">5.5-6.5 mm</quantity>
at clitellum; segments 64-79. Setae regularly distributed around segmental equators, numbering 24-32 at VII, 52-63 at XX, 7 between male pores, setal formula AA:AB:ZZ:YZ= 2.5:2:2:2 at XIII. Female pore single in XIV, on
<quantity id="F06F9B5CFF82361540BAFD3DFE975683" box="[246,371,656,680]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="9.0" metricValueMin="6.0" pageId="5" pageNumber="228" unit="mm" value="0.75" valueMax="0.9" valueMin="0.6">0.6-0.9 mm</quantity>
circle or oval. Prostomium epilobic with tongue open, clitellum coffee color, formalin preservation. First dorsal pore at 12/13. Clitellum annular XIV-XVI; setae invisible externally.
</paragraph>
<paragraph id="372836B9FF82361540AFFD5DFE4957A3" blockId="5.[151,1074,528,1288]" pageId="5" pageNumber="228">Male field with large egg-shaped raised pads, narrow end lateral, with transverse seminal groove from center of pad to just inside lateral edge, male pore at medial end of groove. Three to four concentric arcs around lateral end of pad, 2.5-3.0 mm distance between male pores. Paired spermathecal pores in 5/6 and 6/7, lateral, minute. Genital markings lacking.</paragraph>
<paragraph id="372836B9FF82361540AFFC3DFEFC50A3" blockId="5.[151,1074,528,1288]" pageId="5" pageNumber="228">Septa 5/6-7/8 thick, 8/9, 9/10 absent, 10/11-12/13 thickened. Gizzard large in VIII-X. Intestine begins in XV, lymph glands not found. Typhlosole medium sized from XXVI. Intestinal cecum originating in XXVII and extending anteriorly about to XXIV, bent down to ventral body wall, simple finger-shaped. Oesophageal hearts four pairs in X-XIII; X, XII and XIII large. Male sexual system holandric, testes and funnels in ventrally paired sacs in X-XI. Seminal vesicles two pairs in XI-XII medium sized. Prostates in XVIII, extending through XVII-XIX, long stout duct, glands divided into small lobes.</paragraph>
<paragraph id="372836B9FF82361540AFFB3DFDAB5123" blockId="5.[151,1074,528,1288]" pageId="5" pageNumber="228">Ovaries in XIII. Paired spermathecae in VI, VII; each ampulla roughly ovate, flattened, wrinkled on surfaces, slender duct as long as ampulla; diverticulum chamber club-shaped, white; slender stalk as long as ampulla, no nephridia on spermathecae ducts. Genital marking glands not found.</paragraph>
</subSubSection>
<subSubSection id="7F8D6532FF82361640AFFAB3FD20573E" lastPageId="6" lastPageNumber="229" pageId="5" pageNumber="228" type="discussion">
<paragraph id="372836B9FF82361540AFFAB3FD525251" blockId="5.[151,1074,1309,1653]" pageId="5" pageNumber="228">
REMARKS:
<taxonomicName id="F0974D3AFF823615411AFAB0FDFA5111" authority="Hong, 2011" authorityName="Hong" authorityYear="2011" box="[342,542,1309,1333]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="228" phylum="Annelida" rank="species" species="samgaki" status="sp. nov.">
<emphasis id="05E3EAABFF823615411AFAB0FDFA5111" box="[342,542,1309,1333]" italics="true" pageId="5" pageNumber="228">Amynthas samgaki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1ED057D0FF8236154269FAB3FDBA5111" box="[549,606,1310,1333]" pageId="5" pageNumber="228" rank="species">sp. n.</taxonomicNameLabel>
keys to the
<emphasis id="05E3EAABFF8236154291FAB3FCC85111" box="[733,812,1310,1333]" italics="true" pageId="5" pageNumber="228">morrisi</emphasis>
group in
<bibRefCitation id="53064B48FF82361543DEFAB0FF3A5171" author="SIMS, R. W. &amp; EASTON, E. G." pageId="5" pageNumber="228" pagination="169 - 268" refId="ref3685" refString="SIMS, R. W. &amp; EASTON, E. G. 1972. A numerical revision of the earthworm genus Pheretima auct. (Megascolecidae: Oligochaeta) with the recognition of new genera and an appendix on the earthworms collected by the Royal Society North Borneo Expedition. The Biological Journal of the Linnean Society, London 4: 169 - 268." type="journal article" year="1972">Sims &amp; Easton (1972)</bibRefCitation>
, which is composed of 30 species.
<bibRefCitation id="53064B48FF8236154216FA90FD335171" author="CHEN, Y." box="[602,727,1341,1365]" pageId="5" pageNumber="228" pagination="178 - 296" refId="ref3176" refString="CHEN, Y. 1933. A preliminary survey of the earthworms of the Lower Yangtze valley. Contributions of the Biology Laboratory Sciences Society of China (Zoology), Nanking 9: 178 - 296." type="journal article" year="1933">Chen (1933</bibRefCitation>
;
<bibRefCitation id="53064B48FF82361542AAFA90FCFF5171" author="CHEN, Y." box="[742,795,1341,1365]" pageId="5" pageNumber="228" pagination="269 - 306" refId="ref3214" refString="CHEN, Y. 1936. On the terrestrial Oligochaeta from Szechwan II with the notes on Gates' types. Contributions of the Biology Laboratory Sciences Society of China (Zoology), Nanking 11: 269 - 306." type="journal article" year="1936">1936</bibRefCitation>
;
<bibRefCitation id="53064B48FF8236154365FA90FCBA5171" author="CHEN, Y." box="[809,862,1341,1365]" pageId="5" pageNumber="228" pagination="375 - 427" refId="ref3254" refString="CHEN, Y. 1938. Oligochaeta from Hainan, Kwangtung. Contributions from Biological Laboratory of Science Society of China (Zoological series) 12: 375 - 427." type="journal article" year="1938">1938</bibRefCitation>
;
<bibRefCitation id="53064B48FF8236154321FA90FC465171" author="CHEN, Y." box="[877,930,1341,1365]" pageId="5" pageNumber="228" pagination="83 - 141" refId="ref3285" refString="CHEN, Y. 1946. On the terrestrial Oligochaeta from Szechwan III. Journal of the West China Border Research Society 16: 83 - 141." type="journal article" year="1946">1946</bibRefCitation>
) recorded 12 species of the
<emphasis id="05E3EAABFF823615417FFAF3FE665151" box="[307,386,1374,1397]" italics="true" pageId="5" pageNumber="228">morrisi</emphasis>
group from
<collectingCountry id="4F807629FF8236154240FAF0FDA85151" box="[524,588,1373,1397]" name="China" pageId="5" pageNumber="228">China</collectingCountry>
. Among them, none has male pores on discs or similar pads.
<emphasis id="05E3EAABFF8236154105FAD0FDF051B1" box="[329,532,1405,1429]" italics="true" pageId="5" pageNumber="228">
<taxonomicName id="F0974D3AFF8236154105FAD0FE5651B1" authorityName="Kinberg" authorityYear="1867" box="[329,434,1405,1429]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="228" phylum="Annelida" rank="genus">Amynthas</taxonomicName>
oculatus
</emphasis>
(
<bibRefCitation id="53064B48FF823615426AFAD0FD4751B1" author="CHEN, Y." box="[550,675,1405,1429]" pageId="5" pageNumber="228" pagination="375 - 427" refId="ref3254" refString="CHEN, Y. 1938. Oligochaeta from Hainan, Kwangtung. Contributions from Biological Laboratory of Science Society of China (Zoological series) 12: 375 - 427." type="journal article" year="1938">Chen, 1938</bibRefCitation>
) has a large pair of postsetal genital papillae but the male pores lie outside these. Two species of the
<emphasis id="05E3EAABFF823615431EFA30FC065191" box="[850,994,1437,1461]" italics="true" pageId="5" pageNumber="228">canaliculatus</emphasis>
-group,
<emphasis id="05E3EAABFF82361540DBFA10FEC051F1" box="[151,292,1469,1493]" italics="true" pageId="5" pageNumber="228">A. jangbogoi</emphasis>
and
<emphasis id="05E3EAABFF8236154111FA10FE0E51F1" box="[349,490,1469,1493]" italics="true" pageId="5" pageNumber="228">A. jindoensis</emphasis>
, also have male discs with seminal grooves, and the pairs of spermathecal pores are intrasegmental on VI and VII, rather than in 5/6/7. In
<emphasis id="05E3EAABFF82361540DBFA50FEC75231" box="[151,291,1533,1557]" italics="true" pageId="5" pageNumber="228">A. jangbogoi</emphasis>
the disc is bowling-pin shaped. The spermathecal diverticulum stalk of the latter has numerous tight kinks, as opposed to straight. I include this comparison because the
<emphasis id="05E3EAABFF823615416EF990FE565271" box="[290,434,1597,1621]" italics="true" pageId="5" pageNumber="228">canaliculatus</emphasis>
-group could include morrisi-group species with independently evolved intrasegmental spermathecal pores.
</paragraph>
<paragraph id="372836B9FF8136164098FF4EFDE056BE" blockId="6.[136,1059,227,795]" pageId="6" pageNumber="229">
The following Korean
<emphasis id="05E3EAABFF8136164196FF49FDCD54DF" box="[474,553,228,251]" italics="true" pageId="6" pageNumber="229">morrisi</emphasis>
-group species are known:
<emphasis id="05E3EAABFF813616432CFF4EFBC754DF" box="[864,1059,227,251]" italics="true" pageId="6" pageNumber="229">A. fibulus fibulus</emphasis>
(Kobayashi, 1936),
<emphasis id="05E3EAABFF8136164136FEAEFD9A553F" box="[378,638,259,283]" italics="true" pageId="6" pageNumber="229">A. fibulus ranunculus</emphasis>
(Kobayashi, 1936),
<emphasis id="05E3EAABFF81361643CDFEAEFBC6553F" box="[897,1058,259,283]" italics="true" pageId="6" pageNumber="229">A. kobayashii</emphasis>
(Kobayashi, 1938),
<emphasis id="05E3EAABFF8136164128FE8EFE03551F" box="[356,487,291,315]" italics="true" pageId="6" pageNumber="229">A. koreanus</emphasis>
(Kobayashi, 1938),
<emphasis id="05E3EAABFF8136164282FE8EFC93551F" box="[718,887,291,315]" italics="true" pageId="6" pageNumber="229">A. geojeinsulae</emphasis>
(Song &amp; Paik, 1970),
<emphasis id="05E3EAABFF813616409BFEEEFEB1557F" box="[215,341,323,347]" italics="true" pageId="6" pageNumber="229">A. draconis</emphasis>
Hong &amp; James, 2001,
<emphasis id="05E3EAABFF8136164216FEEEFCE2557F" box="[602,774,323,347]" italics="true" pageId="6" pageNumber="229">A. naejangensis</emphasis>
Hong &amp; James, 2001,
<emphasis id="05E3EAABFF8136164447FEEEFF01555F" italics="true" pageId="6" pageNumber="229">A. assimilis</emphasis>
Hong &amp; Kim 2002, and
<emphasis id="05E3EAABFF813616424BFECEFD26555F" box="[519,706,355,379]" italics="true" pageId="6" pageNumber="229">A. gyeryongensis</emphasis>
Hong &amp; Kim, 2002.
<emphasis id="05E3EAABFF81361643F5FECEFF0155BF" italics="true" pageId="6" pageNumber="229">
<taxonomicName id="F0974D3AFF81361643F5FECEFBC6555F" authorityName="Kinberg" authorityYear="1867" box="[953,1058,355,379]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="6" pageNumber="229" phylum="Annelida" rank="genus">Amynthas</taxonomicName>
assimilis
</emphasis>
and
<emphasis id="05E3EAABFF8136164154FE2EFE2455BF" box="[280,448,387,411]" italics="true" pageId="6" pageNumber="229">A. naejangensis</emphasis>
have male discs with seminal grooves. In
<emphasis id="05E3EAABFF81361643CDFE2EFC1F55BF" box="[897,1019,387,411]" italics="true" pageId="6" pageNumber="229">A. assimilis</emphasis>
the disc and groove are oriented diagonally.
<emphasis id="05E3EAABFF813616427BFE0EFCD6559F" box="[567,818,419,443]" italics="true" pageId="6" pageNumber="229">
<taxonomicName id="F0974D3AFF813616427BFE0EFD44559F" authorityName="Kinberg" authorityYear="1867" box="[567,672,419,443]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="6" pageNumber="229" phylum="Annelida" rank="genus">Amynthas</taxonomicName>
naejangensis
</emphasis>
is larger and has more segments (
<quantity id="F06F9B5CFF813616414CFE6EFE6955FE" box="[256,397,451,475]" metricMagnitude="-1" metricUnit="m" metricValue="1.345" metricValueMax="1.53" metricValueMin="1.16" pageId="6" pageNumber="229" unit="mm" value="134.5" valueMax="153.0" valueMin="116.0">116-153 mm</quantity>
vs
<quantity id="F06F9B5CFF81361641F0FE6EFDD855FE" box="[444,572,451,475]" metricMagnitude="-2" metricUnit="m" metricValue="9.35" metricValueMax="11.0" metricValueMin="7.7" pageId="6" pageNumber="229" unit="mm" value="93.5" valueMax="110.0" valueMin="77.0">77-110 mm</quantity>
, 96-117 vs 64-79 respectively) but is very similar in most respects. The following details of
<emphasis id="05E3EAABFF81361642D3FE4EFCAC55DF" box="[671,840,483,507]" italics="true" pageId="6" pageNumber="229">A. naejangensis</emphasis>
are to be contrasted to the description of
<taxonomicName id="F0974D3AFF813616412BFDAEFDD4563F" authority="Hong, 2011" authorityName="Hong" authorityYear="2011" box="[359,560,515,539]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="6" pageNumber="229" phylum="Annelida" rank="species" species="samgaki" status="sp. nov.">
<emphasis id="05E3EAABFF813616412BFDAEFDD4563F" box="[359,560,515,539]" italics="true" pageId="6" pageNumber="229">Amynthas samgaki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1ED057D0FF813616427BFDAEFD95563E" box="[567,625,515,538]" pageId="6" pageNumber="229" rank="species">sp. n.</taxonomicNameLabel>
: testes sacs dorsally united in X and XI, and enclosing seminal vesicles of XI, lymph glands present, male discs encroaching on adjacent segments, dorsal setal gap present, as indicated in the setal formula YZ:ZZ = 2.5:4. Clearly the two species are quite similar and could be regionally-differentiated descendants of a common ancestor.
</paragraph>
<paragraph id="372836B9FF8136164098FD0EFD20573E" blockId="6.[136,1059,227,795]" pageId="6" pageNumber="229">
<taxonomicName id="F0974D3AFF8136164098FD0EFE46569F" authority="Hong, 2011" authorityName="Hong" authorityYear="2011" box="[212,418,675,699]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="6" pageNumber="229" phylum="Annelida" rank="species" species="samgaki" status="sp. nov.">
<emphasis id="05E3EAABFF8136164098FD0EFE46569F" box="[212,418,675,699]" italics="true" pageId="6" pageNumber="229">Amynthas samgaki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1ED057D0FF81361641E3FD0EFE0A569E" box="[431,494,675,698]" pageId="6" pageNumber="229" rank="species">sp. n.</taxonomicNameLabel>
is probably an endemic species, but more distributional and ecological data are needed to establish its relationships to soils and land use patterns. Further investigations are needed on its phylogenetic relationships with other
<taxonomicName id="F0974D3AFF813616408AFCAEFE60573F" box="[198,388,771,795]" pageId="6" pageNumber="229">
<emphasis id="05E3EAABFF813616408AFCAEFECB573F" box="[198,303,771,795]" italics="true" pageId="6" pageNumber="229">Amynthas</emphasis>
species
</taxonomicName>
, particularly
<emphasis id="05E3EAABFF8136164258FCAEFD59573F" box="[532,701,771,795]" italics="true" pageId="6" pageNumber="229">A. naejangensis</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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