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<document id="F8A02F4CB1334A4369A392877D658000" ID-DOI="10.11646/zootaxa.3713.1.1" ID-GBIF-Dataset="935605ac-a0e6-4963-8264-b4b0ccb8568b" ID-ISSN="1175-5326" ID-Zenodo-Dep="284803" ID-ZooBank="C5F0BC11-92C0-4B30-9DB3-200882AC8950" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460679115380" checkinUser="plazi" docAuthor="Huber, Bernhard A." docDate="2013" docId="B20287EDFFF2FF8CB990C6FAFAEE39C5" docLanguage="en" docName="zt03713p160.pdf" docOrigin="Zootaxa 3713 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Smeringopina attuleh Huber, 2013, new species" docType="treatment" docVersion="8" lastPageNumber="23" masterDocId="4E3BFF95FFE4FF9AB907C624FFF93F1D" masterDocTitle="Revision and cladistic analysis of the Guineo-Congolian spider genus Smeringopina Kraus (Araneae, Pholcidae" masterLastPageNumber="160" masterPageNumber="1" pageNumber="23" updateTime="1698350282054" updateUser="plazi">
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<mods:title id="EEE6A004F01FF6A3EB47F298D8ABC282">Revision and cladistic analysis of the Guineo-Congolian spider genus Smeringopina Kraus (Araneae, Pholcidae</mods:title>
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<mods:namePart id="738FEF8F5ED0CCA31202FEF07EEC8AA0">Huber, Bernhard A.</mods:namePart>
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<mods:date id="EFB262AE9EA838FDE7BFB84ED03E9C8C">2013</mods:date>
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<treatment id="B20287EDFFF2FF8CB990C6FAFAEE39C5" ID-DOI="http://doi.org/10.5281/zenodo.6161999" ID-GBIF-Taxon="119578765" ID-Zenodo-Dep="6161999" LSID="urn:lsid:plazi:treatment:B20287EDFFF2FF8CB990C6FAFAEE39C5" httpUri="http://treatment.plazi.org/id/B20287EDFFF2FF8CB990C6FAFAEE39C5" lastPageNumber="23" pageId="22" pageNumber="23">
<subSubSection id="72B16570FFF2FF8CB990C6FAFAEE39C5" pageId="22" pageNumber="23" type="nomenclature">
<paragraph id="3A1436FBFFF2FF8CB990C6FAFDCC3FE5" blockId="22.[151,577,222,282]" box="[151,565,222,248]" pageId="22" pageNumber="23">
<heading id="615C8197FFF2FF8CB990C6FAFDCC3FE5" bold="true" box="[151,565,222,248]" fontSize="11" level="1" pageId="22" pageNumber="23" reason="1">
<emphasis id="08DFEAE9FFF2FF8CB990C6FAFDCC3FE5" bold="true" box="[151,565,222,248]" pageId="22" pageNumber="23">
<taxonomicName id="FDAB4D78FFF2FF8CB990C6FAFE673FE5" ID-CoL="4XVFP" box="[151,414,222,248]" class="Arachnida" family="Pholcidae" genus="Smeringopina" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="attuleh" status="sp. nov.">
<emphasis id="08DFEAE9FFF2FF8CB990C6FAFE673FE5" bold="true" box="[151,414,222,248]" italics="true" pageId="22" pageNumber="23">Smeringopina attuleh</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13EC5792FFF2FF8CB8A2C6FAFDCC3FE5" box="[421,565,222,248]" pageId="22" pageNumber="23" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="3A1436FBFFF2FF8CB990C725FDB83E04" blockId="22.[151,577,222,282]" box="[151,577,257,282]" pageId="22" pageNumber="23">
<figureCitation id="A2902A7EFFF2FF8CB990C725FF0A3E04" box="[151,243,257,282]" captionStart="FIGURES 17 31" captionStartId="60.[151,264,1832,1854]" captionTargetBox="[185,1409,179,1826]" captionTargetId="figure@60.[185,1409,179,1826]" captionTargetPageId="60" captionText="FIGURES 17 31. Smeringopina, webs and alive specimens in their natural habitats. 17. S. guineensis, web (Diéké, Guinea). 18. S. kribi, web with nematocerous flies (S Kribi, Cameroon). 19. S. kala, male (Mt. Kala, Cameroon). 20. S. bamenda, pair (Bamenda, Cameroon). 21 22. S. nyasoso, male and female with eggsac (Mt. Koupé, Cameroon). 23. S. mbouda, male (near Lake Oku, Cameroon). 24. S. attuleh, female with eggsac (Essotah, Cameroon). 25. S. kinguele, female with juveniles (Monts de Cristal, Gabon). 26. S. tebe, male (N Tébé, Gabon). 27. S. lekoni, female with eggsac (at Lekoni River, Gabon). 28. S. moudouma, mating pair (Moudouma, Gabon). 29 30. S. fang, male (Djidji, Gabon) and female with eggsac (Mayebout, Gabon). 31. S. ankasa, female with eggsac (Ankasa N. P., Ghana)." httpUri="https://zenodo.org/record/284806/files/figure.png" pageId="22" pageNumber="23">Figs. 24</figureCitation>
,
<figureCitation id="A2902A7EFFF2FF8CB9F8C725FE9A3E04" box="[255,355,257,281]" captionStart="FIGURES 294 303" captionStartId="92.[151,264,1931,1953]" captionTargetBox="[200,1402,193,1896]" captionTargetId="figure@92.[172,1416,193,1909]" captionTargetPageId="92" captionText="FIGURES 294 303. Smeringopina attuleh n. sp. (294 298) and S. mbouda n. sp. (299 303). 294 295, 299 300. Males, dorsal, lateral, and ventral views. 296 298, 301 303. Left male palps, prolateral, dorsal, and retrolateral views. Arrow points at retrolateral ridge bordering whitish ventral area." httpUri="https://zenodo.org/record/284839/files/figure.png" pageId="22" pageNumber="23">294298</figureCitation>
,
<figureCitation id="A2902A7EFFF2FF8CB868C725FE2D3E07" box="[367,468,257,282]" captionStart="FIGURES 304 317" captionStartId="93.[151,264,1901,1923]" captionTargetBox="[168,1380,204,1880]" captionTargetId="figure@93.[151,1395,193,1883]" captionTargetPageId="93" captionText="FIGURES 304 317. Smeringopina attuleh n. sp. (304 306), S. mbouda n. sp. (307 309), and S. fon (310 317). 304 305, 307 308, 310 311. Female abdomens, ventral and lateral views. 306, 309, 312. Cleared female genitalia, dorsal views. 313 314. Male, dorsal and ventral views. 315 317. Left male palp, prolateral, dorsal, and retrolateral views. Arrow points at retrolateral ridge bordering whitish ventral area." httpUri="https://zenodo.org/record/284840/files/figure.png" pageId="22" pageNumber="23">304306</figureCitation>
,
<figureCitation id="A2902A7EFFF2FF8CB8D8C725FDB83E04" box="[479,577,257,281]" captionStart="FIGURES 318 323" captionStartId="94.[151,264,1493,1515]" captionTargetBox="[151,1427,198,1460]" captionTargetId="figure@94.[151,1436,193,1472]" captionTargetPageId="94" captionText="FIGURES 318 323. Smeringopina attuleh n. sp. 318 319. Left procursus, prolateral and retrolateral views. 320. Male chelicerae, frontal view. 321. Left embolus, prolateral view. 322. Epigynum, ventral view. 323. Cleared female genitalia, dorsal view. Scale lines: 0.2 (321), 0.5 (318 320, 322 323)." httpUri="https://zenodo.org/record/284841/files/figure.png" pageId="22" pageNumber="23">318323</figureCitation>
</paragraph>
<paragraph id="3A1436FBFFF2FF8CB990C76EFCBE3E42" blockId="22.[151,839,329,351]" box="[151,839,329,351]" pageId="22" pageNumber="23">
<emphasis id="08DFEAE9FFF2FF8CB990C76EFE8D3E42" box="[151,372,329,351]" italics="true" pageId="22" pageNumber="23">
<taxonomicName id="FDAB4D78FFF2FF8CB990C76EFEA33E42" authority="Cam" authorityName="Cam" box="[151,346,329,351]" class="Arachnida" family="Pholcidae" genus="Smeringopina" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">Smeringopina Cam</taxonomicName>
40
</emphasis>
: Dimitrov, Astrin &amp; Huber 2013 (DNA data).
</paragraph>
<paragraph id="3A1436FBFFF2FF8CB990C7A9FC593ED5" blockId="22.[151,1436,396,1752]" pageId="22" pageNumber="23">
<typeStatus id="E5108859FFF2FF8CB990C7A9FF2B3EB8" box="[151,210,397,421]" pageId="22" pageNumber="23">Type</typeStatus>
. ♂
<typeStatus id="E5108859FFF2FF8CB9FDC7A8FEA53EB8" box="[250,348,396,421]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
from
<collectingCountry id="42BC766BFFF2FF8CB8A6C7A8FDE03EB9" box="[417,537,396,420]" name="Cameroon" pageId="22" pageNumber="23">Cameroon</collectingCountry>
, Southwest Region, near Dschang,
<taxonomicName id="FDAB4D78FFF2FF8CBAAFC7A8FC063EB8" box="[936,1023,396,421]" class="Arachnida" family="Pholcidae" genus="Smeringopina" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="95" phylum="Arthropoda" rank="species" species="attuleh" status="sp. nov.">Attuleh</taxonomicName>
, “site 1” (
<geoCoordinate id="5F9F503CFFF2FF8CBD77C7A8FB223EB8" box="[1136,1243,396,421]" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="92" value="5.4616666">5°27.7N</geoCoordinate>
,
<geoCoordinate id="5F9F503CFFF2FF8CBDE0C7A8FAB63EB8" box="[1255,1359,396,421]" direction="east" orientation="longitude" pageId="22" pageNumber="23" precision="92" value="9.941667">9°56.5E</geoCoordinate>
),
<quantity id="FD539B1EFFF2FF8CBC63C7A8FF543EDA" metricMagnitude="3" metricUnit="m" metricValue="1.88" pageId="22" pageNumber="23" unit="m" value="1880.0">1880 m</quantity>
a.s.l., near ground,
<date id="4E15103BFFF2FF8CB88DC78BFDF93ED5" box="[394,512,431,456]" pageId="22" pageNumber="23" value="2009-04-20">20.iv.2009</date>
(B.A. Huber), in ZFMK (Ar 10216).
</paragraph>
<paragraph id="3A1436FBFFF2FF8CB9C0C7F0FE233DBD" blockId="22.[151,1436,396,1752]" pageId="22" pageNumber="23">
Other material examined.
<collectingCountry id="42BC766BFFF2FF8CB8F1C7F0FD623EF1" box="[502,667,468,492]" name="Cameroon" pageId="22" pageNumber="23">CAMEROON</collectingCountry>
:
<emphasis id="08DFEAE9FFF2FF8CBBAEC7F0FC8C3EF1" box="[681,885,468,493]" italics="true" pageId="22" pageNumber="23">Southwest Region</emphasis>
: near Dschang,
<taxonomicName id="FDAB4D78FFF2FF8CBD31C7F0FB743EF0" box="[1078,1165,468,493]" class="Arachnida" family="Pholcidae" genus="Smeringopina" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="95" phylum="Arthropoda" rank="species" species="attuleh" status="sp. nov.">Attuleh</taxonomicName>
, “site 1”, same data as
<typeStatus id="E5108859FFF2FF8CB990C7D3FF053D0D" box="[151,252,503,528]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
,
<specimenCount id="2CADFD72FFF2FF8CB800C7DCFED43D0D" box="[263,301,504,528]" pageId="22" pageNumber="23" type="generic">3♂</specimenCount>
2♀ in ZFMK (Ar 10217); same data, 1♀ 4 juvs. in pure ethanol, in ZFMK (Cam 126);
<taxonomicName id="FDAB4D78FFF2FF8CBC01C7D3FAA43D0D" box="[1286,1373,503,528]" class="Arachnida" family="Pholcidae" genus="Smeringopina" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="95" phylum="Arthropoda" rank="species" species="attuleh" status="sp. nov.">Attuleh</taxonomicName>
, “site 2” (
<geoCoordinate id="5F9F503CFFF2FF8CB9C4C438FED73D28" box="[195,302,540,565]" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="92" value="5.465">5°27.9N</geoCoordinate>
,
<geoCoordinate id="5F9F503CFFF2FF8CB83BC438FE5D3D28" box="[316,420,540,565]" direction="east" orientation="longitude" pageId="22" pageNumber="23" precision="92" value="9.941667">9°56.5E</geoCoordinate>
),
<quantity id="FD539B1EFFF2FF8CB8BDC438FDEB3D29" box="[442,530,540,564]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="22" pageNumber="23" unit="m" value="1800.0">1800 m</quantity>
a.s.l., near ground,
<date id="4E15103BFFF2FF8CBBFFC438FC943D28" box="[760,877,540,565]" pageId="22" pageNumber="23" value="2009-04-20">20.iv.2009</date>
(B.A. &amp; J.C. Huber), 4♀ in ZFMK (Ar 10218); same data, 3♀ in pure ethanol, in ZFMK (Cam 122). Near Dschang,
<taxonomicName id="FDAB4D78FFF2FF8CBAA1C41BFC043D45" box="[934,1021,575,600]" class="Arachnida" family="Pholcidae" genus="Smeringopina" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="146" phylum="Arthropoda" rank="species" species="essotah" status="sp. nov.">Essotah</taxonomicName>
(
<geoCoordinate id="5F9F503CFFF2FF8CBD09C41BFB803D45" box="[1038,1145,575,600]" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="92" value="5.425">5°25.5N</geoCoordinate>
,
<geoCoordinate id="5F9F503CFFF2FF8CBD81C41BFB173D45" box="[1158,1262,575,600]" direction="east" orientation="longitude" pageId="22" pageNumber="23" precision="92" value="9.911667">9°54.7E</geoCoordinate>
),
<quantity id="FD539B1EFFF2FF8CBC04C41BFAA03D4A" box="[1283,1369,575,600]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="22" pageNumber="23" unit="m" value="1600.0">1600 m</quantity>
a.s.l., near ground,
<date id="4E15103BFFF2FF8CB836C440FE5E3D60" box="[305,423,612,637]" pageId="22" pageNumber="23" value="2009-04-21">21.iv.2009</date>
(B.A. Huber),
<specimenCount id="2CADFD72FFF2FF8CBB5DC440FD7A3D61" box="[602,643,612,636]" pageId="22" pageNumber="23" type="generic">2♂</specimenCount>
1♀ (2 vials) in ZFMK (Ar 10219-20); same data, 2♀ 3 juvs. in pure ethanol, in ZFMK (Cam 96).
</paragraph>
<paragraph id="3A1436FBFFF2FF8CB9C0C488FBDD3DD8" blockId="22.[151,1436,396,1752]" box="[199,1060,684,709]" pageId="22" pageNumber="23">
Etymology. The name is a noun in apposition, derived from the
<typeStatus id="E5108859FFF2FF8CBA94C489FC3A3DD8" box="[915,963,685,709]" pageId="22" pageNumber="23">type</typeStatus>
locality.
</paragraph>
<paragraph id="3A1436FBFFF2FF8CB9C0C4EBFA613C65" blockId="22.[151,1436,396,1752]" pageId="22" pageNumber="23">
Diagnosis. Distinguished from most congeners by distinctive male cheliceral armature (very densely packed modified hairs on distinctively shaped apophyses;
<figureCitation id="A2902A7EFFF2FF8CBBDBC4D0FCC73C11" box="[732,830,756,781]" captionStart="FIGURES 318 323" captionStartId="94.[151,264,1493,1515]" captionTargetBox="[151,1427,198,1460]" captionTargetId="figure@94.[151,1436,193,1472]" captionTargetPageId="94" captionText="FIGURES 318 323. Smeringopina attuleh n. sp. 318 319. Left procursus, prolateral and retrolateral views. 320. Male chelicerae, frontal view. 321. Left embolus, prolateral view. 322. Epigynum, ventral view. 323. Cleared female genitalia, dorsal view. Scale lines: 0.2 (321), 0.5 (318 320, 322 323)." httpUri="https://zenodo.org/record/284841/files/figure.png" pageId="22" pageNumber="23">Fig. 320</figureCitation>
) and procursus with ventral indentation occupied by complex membranous structures (
<figureCitation id="A2902A7EFFF2FF8CBB14C533FD8F3C2D" box="[531,630,791,816]" captionStart="FIGURES 318 323" captionStartId="94.[151,264,1493,1515]" captionTargetBox="[151,1427,198,1460]" captionTargetId="figure@94.[151,1436,193,1472]" captionTargetPageId="94" captionText="FIGURES 318 323. Smeringopina attuleh n. sp. 318 319. Left procursus, prolateral and retrolateral views. 320. Male chelicerae, frontal view. 321. Left embolus, prolateral view. 322. Epigynum, ventral view. 323. Cleared female genitalia, dorsal view. Scale lines: 0.2 (321), 0.5 (318 320, 322 323)." httpUri="https://zenodo.org/record/284841/files/figure.png" pageId="22" pageNumber="23">Fig. 319</figureCitation>
); from the very similar
<taxonomicName id="FDAB4D78FFF2FF8CBA78C53CFC0F3C2D" box="[895,1014,791,816]" class="Arachnida" family="Pholcidae" genus="Smeringopina" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="mbouda">
<emphasis id="08DFEAE9FFF2FF8CBA78C53CFC0F3C2D" box="[895,1014,791,816]" italics="true" pageId="22" pageNumber="23">S. mbouda</emphasis>
</taxonomicName>
by shorter male cheliceral apophyses (
<figureCitation id="A2902A7EFFF2FF8CB998C518FEFD3C49" box="[159,260,828,853]" captionStart="FIGURES 318 323" captionStartId="94.[151,264,1493,1515]" captionTargetBox="[151,1427,198,1460]" captionTargetId="figure@94.[151,1436,193,1472]" captionTargetPageId="94" captionText="FIGURES 318 323. Smeringopina attuleh n. sp. 318 319. Left procursus, prolateral and retrolateral views. 320. Male chelicerae, frontal view. 321. Left embolus, prolateral view. 322. Epigynum, ventral view. 323. Cleared female genitalia, dorsal view. Scale lines: 0.2 (321), 0.5 (318 320, 322 323)." httpUri="https://zenodo.org/record/284841/files/figure.png" pageId="22" pageNumber="23">Fig. 320</figureCitation>
), simple embolus (
<figureCitation id="A2902A7EFFF2FF8CB8E7C518FDBD3C49" box="[480,580,828,853]" captionStart="FIGURES 318 323" captionStartId="94.[151,264,1493,1515]" captionTargetBox="[151,1427,198,1460]" captionTargetId="figure@94.[151,1436,193,1472]" captionTargetPageId="94" captionText="FIGURES 318 323. Smeringopina attuleh n. sp. 318 319. Left procursus, prolateral and retrolateral views. 320. Male chelicerae, frontal view. 321. Left embolus, prolateral view. 322. Epigynum, ventral view. 323. Cleared female genitalia, dorsal view. Scale lines: 0.2 (321), 0.5 (318 320, 322 323)." httpUri="https://zenodo.org/record/284841/files/figure.png" pageId="22" pageNumber="23">Fig. 321</figureCitation>
), dorso-distal process on procursus (
<figureCitation id="A2902A7EFFF2FF8CBAEAC518FBAB3C49" box="[1005,1106,828,853]" captionStart="FIGURES 318 323" captionStartId="94.[151,264,1493,1515]" captionTargetBox="[151,1427,198,1460]" captionTargetId="figure@94.[151,1436,193,1472]" captionTargetPageId="94" captionText="FIGURES 318 323. Smeringopina attuleh n. sp. 318 319. Left procursus, prolateral and retrolateral views. 320. Male chelicerae, frontal view. 321. Left embolus, prolateral view. 322. Epigynum, ventral view. 323. Cleared female genitalia, dorsal view. Scale lines: 0.2 (321), 0.5 (318 320, 322 323)." httpUri="https://zenodo.org/record/284841/files/figure.png" pageId="22" pageNumber="23">Fig. 319</figureCitation>
), and anterior epigynal plate with distinct median process (in lateral view;
<figureCitation id="A2902A7EFFF2FF8CBB91C57BFD0D3C65" box="[662,756,863,888]" captionStart="FIGURES 304 317" captionStartId="93.[151,264,1901,1923]" captionTargetBox="[168,1380,204,1880]" captionTargetId="figure@93.[151,1395,193,1883]" captionTargetPageId="93" captionText="FIGURES 304 317. Smeringopina attuleh n. sp. (304 306), S. mbouda n. sp. (307 309), and S. fon (310 317). 304 305, 307 308, 310 311. Female abdomens, ventral and lateral views. 306, 309, 312. Cleared female genitalia, dorsal views. 313 314. Male, dorsal and ventral views. 315 317. Left male palp, prolateral, dorsal, and retrolateral views. Arrow points at retrolateral ridge bordering whitish ventral area." httpUri="https://zenodo.org/record/284840/files/figure.png" pageId="22" pageNumber="23">Fig. 305</figureCitation>
) and more angular ventral edges (in ventral view;
<figureCitation id="A2902A7EFFF2FF8CBC2DC57BFA723C65" box="[1322,1419,863,888]" captionStart="FIGURES 318 323" captionStartId="94.[151,264,1493,1515]" captionTargetBox="[151,1427,198,1460]" captionTargetId="figure@94.[151,1436,193,1472]" captionTargetPageId="94" captionText="FIGURES 318 323. Smeringopina attuleh n. sp. 318 319. Left procursus, prolateral and retrolateral views. 320. Male chelicerae, frontal view. 321. Left embolus, prolateral view. 322. Epigynum, ventral view. 323. Cleared female genitalia, dorsal view. Scale lines: 0.2 (321), 0.5 (318 320, 322 323)." httpUri="https://zenodo.org/record/284841/files/figure.png" pageId="22" pageNumber="23">Fig. 322</figureCitation>
).
</paragraph>
<paragraph id="3A1436FBFFF2FF8CB9C0C5A0FA7C3AA5" blockId="22.[151,1436,396,1752]" pageId="22" pageNumber="23">
Male (
<typeStatus id="E5108859FFF2FF8CB817C5A0FE803C80" box="[272,377,900,925]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
). Total body length 4.6, carapace width 1.5. Leg 1: 33.7 (8.1 + 0.5 + 8.1 + 14.9 + 2.1), tibia 2: 5.2, tibia 3: 3.8, tibia 4: 6.0; tibia 1 L/d: 55. Distance PME-PME 135 µm, diameter PME 140 µm, distance PME- ALE 70 µm, distance AME-AME 55 µm, diameter AME 125 µm. Carapace ochre with dark brown triangular mark posteriorly, brown mark behind ocular area, brown lateral margins; ocular area not darkened, clypeus with pair of indistinct bands below eyes, sternum brown; legs ochre-brown, darker rings subdistally on femora and tibiae and in patella area, tips of femora and tibiae lighter; abdomen ochre-gray with dark pattern dorsally, laterally, and ventrally. Habitus as in
<figureCitation id="A2902A7EFFF2FF8CB8B2C278FDA43B68" box="[437,605,1116,1141]" captionStart="FIGURES 294 303" captionStartId="92.[151,264,1931,1953]" captionTargetBox="[200,1402,193,1896]" captionTargetId="figure@92.[172,1416,193,1909]" captionTargetPageId="92" captionText="FIGURES 294 303. Smeringopina attuleh n. sp. (294 298) and S. mbouda n. sp. (299 303). 294 295, 299 300. Males, dorsal, lateral, and ventral views. 296 298, 301 303. Left male palps, prolateral, dorsal, and retrolateral views. Arrow points at retrolateral ridge bordering whitish ventral area." httpUri="https://zenodo.org/record/284839/files/figure.png" pageId="22" pageNumber="23">Figs. 294295</figureCitation>
, ocular area slightly elevated, secondary eyes with indistinct pseudolenses; clypeus unmodified; deep thoracic pit and pair of shallow furrows diverging behind pit. Chelicerae as in
<figureCitation id="A2902A7EFFF2FF8CB990C280FF003BA1" box="[151,249,1188,1213]" captionStart="FIGURES 318 323" captionStartId="94.[151,264,1493,1515]" captionTargetBox="[151,1427,198,1460]" captionTargetId="figure@94.[151,1436,193,1472]" captionTargetPageId="94" captionText="FIGURES 318 323. Smeringopina attuleh n. sp. 318 319. Left procursus, prolateral and retrolateral views. 320. Male chelicerae, frontal view. 321. Left embolus, prolateral view. 322. Epigynum, ventral view. 323. Cleared female genitalia, dorsal view. Scale lines: 0.2 (321), 0.5 (318 320, 322 323)." httpUri="https://zenodo.org/record/284841/files/figure.png" pageId="22" pageNumber="23">Fig. 320</figureCitation>
, with lateral proximal apophyses connected to short rounded distal apophyses, the latter with ~25 large modified (cone-shaped) hairs on each side. Palps as in
<figureCitation id="A2902A7EFFF2FF8CBA27C2E3FC313BFD" box="[800,968,1223,1248]" captionStart="FIGURES 294 303" captionStartId="92.[151,264,1931,1953]" captionTargetBox="[200,1402,193,1896]" captionTargetId="figure@92.[172,1416,193,1909]" captionTargetPageId="92" captionText="FIGURES 294 303. Smeringopina attuleh n. sp. (294 298) and S. mbouda n. sp. (299 303). 294 295, 299 300. Males, dorsal, lateral, and ventral views. 296 298, 301 303. Left male palps, prolateral, dorsal, and retrolateral views. Arrow points at retrolateral ridge bordering whitish ventral area." httpUri="https://zenodo.org/record/284839/files/figure.png" pageId="22" pageNumber="23">Figs. 296298</figureCitation>
; coxa unmodified; trochanter with short conical retrolatero-ventral apophysis; femur with retrolateral flap and large whitish area ventrally, without prolateral modification; prolateral femur-patella joint strongly shifted toward ventrally (hidden by bulb in
<figureCitation id="A2902A7EFFF2FF8CBC77C32BFF3C3A50" captionStart="FIGURES 294 303" captionStartId="92.[151,264,1931,1953]" captionTargetBox="[200,1402,193,1896]" captionTargetId="figure@92.[172,1416,193,1909]" captionTargetPageId="92" captionText="FIGURES 294 303. Smeringopina attuleh n. sp. (294 298) and S. mbouda n. sp. (299 303). 294 295, 299 300. Males, dorsal, lateral, and ventral views. 296 298, 301 303. Left male palps, prolateral, dorsal, and retrolateral views. Arrow points at retrolateral ridge bordering whitish ventral area." httpUri="https://zenodo.org/record/284839/files/figure.png" pageId="22" pageNumber="23">Fig. 296</figureCitation>
); tarsus with some longer but barely stronger hairs dorsally; procursus with ventral indentation occupied by complex membranous structures, without hinge, with dorso-distal process; bulb with simple small embolus with small sclerotized branch (
<figureCitation id="A2902A7EFFF2FF8CB8C4C358FDD03A89" box="[451,553,1404,1429]" captionStart="FIGURES 318 323" captionStartId="94.[151,264,1493,1515]" captionTargetBox="[151,1427,198,1460]" captionTargetId="figure@94.[151,1436,193,1472]" captionTargetPageId="94" captionText="FIGURES 318 323. Smeringopina attuleh n. sp. 318 319. Left procursus, prolateral and retrolateral views. 320. Male chelicerae, frontal view. 321. Left embolus, prolateral view. 322. Epigynum, ventral view. 323. Cleared female genitalia, dorsal view. Scale lines: 0.2 (321), 0.5 (318 320, 322 323)." httpUri="https://zenodo.org/record/284841/files/figure.png" pageId="22" pageNumber="23">Fig. 321</figureCitation>
). Legs without spines and curved hairs, with few vertical hairs; retrolateral trichobothrium on tibia 1 at 1.5%; prolateral trichobothrium present on all tibiae; pseudosegments barely visible.
</paragraph>
<paragraph id="3A1436FBFFF2FF8CB9C0C3E0FCE83AC0" blockId="22.[151,1436,396,1752]" box="[199,785,1476,1501]" pageId="22" pageNumber="23">
Variation. Tibia
<quantity id="FD539B1EFFF2FF8CB87BC3E0FE513AC0" box="[380,424,1476,1501]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="22" pageNumber="23" unit="in" value="1.0">1 in</quantity>
4 other males: 8.6, 8.6, 8.7, 9.5.
</paragraph>
<paragraph id="3A1436FBFFF2FF8CB9C0C3C3FE9E3971" blockId="22.[151,1436,396,1752]" pageId="22" pageNumber="23">
Female. In general similar to male. Tibia
<quantity id="FD539B1EFFF2FF8CBBC4C3CCFD0D391D" box="[707,756,1511,1536]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="22" pageNumber="23" unit="in" value="1.0">1 in</quantity>
<specimenCount id="2CADFD72FFF2FF8CBA05C3CCFC81391D" box="[770,888,1511,1536]" pageId="22" pageNumber="23" type="female">7 females</specimenCount>
: 7.07.8 (mean 7.4). Epigynum anterior plate trapezoidal (
<figureCitation id="A2902A7EFFF2FF8CB824C028FE763939" box="[291,399,1548,1573]" captionStart="FIGURES 304 317" captionStartId="93.[151,264,1901,1923]" captionTargetBox="[168,1380,204,1880]" captionTargetId="figure@93.[151,1395,193,1883]" captionTargetPageId="93" captionText="FIGURES 304 317. Smeringopina attuleh n. sp. (304 306), S. mbouda n. sp. (307 309), and S. fon (310 317). 304 305, 307 308, 310 311. Female abdomens, ventral and lateral views. 306, 309, 312. Cleared female genitalia, dorsal views. 313 314. Male, dorsal and ventral views. 315 317. Left male palp, prolateral, dorsal, and retrolateral views. Arrow points at retrolateral ridge bordering whitish ventral area." httpUri="https://zenodo.org/record/284840/files/figure.png" pageId="22" pageNumber="23">Figs. 304</figureCitation>
,
<figureCitation id="A2902A7EFFF2FF8CB89DC028FE313939" box="[410,456,1548,1572]" captionStart="FIGURES 318 323" captionStartId="94.[151,264,1493,1515]" captionTargetBox="[151,1427,198,1460]" captionTargetId="figure@94.[151,1436,193,1472]" captionTargetPageId="94" captionText="FIGURES 318 323. Smeringopina attuleh n. sp. 318 319. Left procursus, prolateral and retrolateral views. 320. Male chelicerae, frontal view. 321. Left embolus, prolateral view. 322. Epigynum, ventral view. 323. Cleared female genitalia, dorsal view. Scale lines: 0.2 (321), 0.5 (318 320, 322 323)." httpUri="https://zenodo.org/record/284841/files/figure.png" pageId="22" pageNumber="23">322</figureCitation>
), with distinct median process (
<figureCitation id="A2902A7EFFF2FF8CBA2DC028FC703938" box="[810,905,1548,1573]" captionStart="FIGURES 304 317" captionStartId="93.[151,264,1901,1923]" captionTargetBox="[168,1380,204,1880]" captionTargetId="figure@93.[151,1395,193,1883]" captionTargetPageId="93" captionText="FIGURES 304 317. Smeringopina attuleh n. sp. (304 306), S. mbouda n. sp. (307 309), and S. fon (310 317). 304 305, 307 308, 310 311. Female abdomens, ventral and lateral views. 306, 309, 312. Cleared female genitalia, dorsal views. 313 314. Male, dorsal and ventral views. 315 317. Left male palp, prolateral, dorsal, and retrolateral views. Arrow points at retrolateral ridge bordering whitish ventral area." httpUri="https://zenodo.org/record/284840/files/figure.png" pageId="22" pageNumber="23">Fig. 305</figureCitation>
) variable in shape even within localities (either rounded or with pair of small processes), posterior edges angular; large flat posterior plate; internal genitalia as in
<figureCitation id="A2902A7EFFF2FF8CB990C070FF063970" box="[151,255,1620,1645]" captionStart="FIGURES 304 317" captionStartId="93.[151,264,1901,1923]" captionTargetBox="[168,1380,204,1880]" captionTargetId="figure@93.[151,1395,193,1883]" captionTargetPageId="93" captionText="FIGURES 304 317. Smeringopina attuleh n. sp. (304 306), S. mbouda n. sp. (307 309), and S. fon (310 317). 304 305, 307 308, 310 311. Female abdomens, ventral and lateral views. 306, 309, 312. Cleared female genitalia, dorsal views. 313 314. Male, dorsal and ventral views. 315 317. Left male palp, prolateral, dorsal, and retrolateral views. Arrow points at retrolateral ridge bordering whitish ventral area." httpUri="https://zenodo.org/record/284840/files/figure.png" pageId="22" pageNumber="23">Figs. 306</figureCitation>
and
<figureCitation id="A2902A7EFFF2FF8CB832C070FE9B3971" box="[309,354,1620,1644]" captionStart="FIGURES 318 323" captionStartId="94.[151,264,1493,1515]" captionTargetBox="[151,1427,198,1460]" captionTargetId="figure@94.[151,1436,193,1472]" captionTargetPageId="94" captionText="FIGURES 318 323. Smeringopina attuleh n. sp. 318 319. Left procursus, prolateral and retrolateral views. 320. Male chelicerae, frontal view. 321. Left embolus, prolateral view. 322. Epigynum, ventral view. 323. Cleared female genitalia, dorsal view. Scale lines: 0.2 (321), 0.5 (318 320, 322 323)." httpUri="https://zenodo.org/record/284841/files/figure.png" pageId="22" pageNumber="23">323</figureCitation>
.
</paragraph>
<paragraph id="3A1436FBFFF2FF8CB9C0C053FE3539A9" blockId="22.[151,1436,396,1752]" pageId="22" pageNumber="23">
Natural history.
<taxonomicName id="FDAB4D78FFF2FF8CB87CC05CFE1D398D" box="[379,484,1655,1680]" class="Arachnida" family="Pholcidae" genus="Smeringopina" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="attuleh">
<emphasis id="08DFEAE9FFF2FF8CB87CC05CFE1D398D" box="[379,484,1655,1680]" italics="true" pageId="22" pageNumber="23">S. attuleh</emphasis>
</taxonomicName>
was found mainly under large dead leaves on the ground, but also in small cavities in the ground and under roots.
</paragraph>
<paragraph id="3A1436FBFFF2FF8CB9C0C09BFAEE39C5" blockId="22.[151,1436,396,1752]" box="[199,1303,1727,1752]" pageId="22" pageNumber="23">
Distribution. Known from three neighboring localities in
<collectingCountry id="42BC766BFFF2FF8CBA4EC0E4FC3839C5" box="[841,961,1728,1752]" name="Cameroon" pageId="22" pageNumber="23">Cameroon</collectingCountry>
, Southwest Region (
<figureCitation id="A2902A7EFFF2FF8CBDAFC09BFAF339C5" box="[1192,1290,1727,1752]" captionStart="FIGURE 293" captionStartId="91.[151,250,1057,1079]" captionTargetBox="[151,1436,193,1035]" captionTargetId="figure@91.[151,1436,193,1036]" captionTargetPageId="91" captionText="FIGURE 293. Known distributions of the attuleh and ankasa species groups, and of S. fon and S. ngungu." httpUri="https://zenodo.org/record/284838/files/figure.png" pageId="22" pageNumber="23">Fig. 293</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>