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<document id="CDE2554C6D0C630F455C859B6CDD69D4" ID-ISSN="2201-4349" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629578942507" checkinUser="felipe" docAuthor="Munari, L." docDate="2004" docId="03FDBD16FFDE6C5BFC66B9B4FD270D38" docLanguage="en" docName="RecAustMus.56.1.29-56.pdf" docOrigin="Records of the Australian Museum 56 (1)" docStyle="DocumentStyle:7863F7301B7BEE922F541F6F9B5AEEC6.2:RecAustMus.2008-2018.journal_article.0cover" docStyleId="7863F7301B7BEE922F541F6F9B5AEEC6" docStyleName="RecAustMus.2008-2018.journal_article.0cover" docStyleVersion="2" docTitle="Thitena Munari 2004, n.gen." docType="treatment" docVersion="2" lastPageNumber="51" masterDocId="FFC4C56EFFCB6C4DFF99B92CFFB80E10" masterDocTitle="Beach Flies (Diptera: Tethinidae: Tethininae) From Australia and Papua New Guinea, with Descriptions of Two New Genera and Ten New Species" masterLastPageNumber="56" masterPageNumber="29" pageNumber="50" updateTime="1699079709040" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="87E9D196120386CFEF0CC124B2044310">Beach Flies (Diptera: Tethinidae: Tethininae) From Australia and Papua New Guinea, with Descriptions of Two New Genera and Ten New Species</mods:title>
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<mods:namePart id="C93E6A83BC7833C14F43AF4F18851732">Munari, L.</mods:namePart>
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<mods:title id="65274A83620FE608CD3307E36C85F451">Records of the Australian Museum</mods:title>
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<mods:date id="EF54BCBDC9926B21B829C0FBDD58DB06">2004</mods:date>
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<treatment id="03FDBD16FFDE6C5BFC66B9B4FD270D38" LSID="urn:lsid:plazi:treatment:03FDBD16FFDE6C5BFC66B9B4FD270D38" httpUri="http://treatment.plazi.org/id/03FDBD16FFDE6C5BFC66B9B4FD270D38" lastPageId="22" lastPageNumber="51" pageId="21" pageNumber="50">
<subSubSection id="C34E5F8BFFDE6C58FC66B9B4FB100EA0" box="[1023,1192,152,176]" pageId="21" pageNumber="50" type="nomenclature">
<paragraph id="8BEB0C00FFDE6C58FC66B9B4FB100EA0" blockId="21.[1023,1192,152,176]" box="[1023,1192,152,176]" pageId="21" pageNumber="50">
<heading id="D0A3BB6CFFDE6C58FC66B9B4FB100EA0" box="[1023,1192,152,176]" centered="true" fontSize="11" level="2" pageId="21" pageNumber="50" reason="2">
<emphasis id="B920D012FFDE6C58FC66B9B4FB100EA0" bold="true" box="[1023,1192,152,176]" pageId="21" pageNumber="50">
<taxonomicName id="4C547783FFDE6C58FC66B9B4FBE10EA0" ID-CoL="7W3G" authority="Munari, 2004" authorityName="Munari" authorityYear="2004" box="[1023,1113,152,176]" class="Insecta" family="Canacidae" genus="Thitena" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="50" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B920D012FFDE6C58FC66B9B4FBE10EA0" bold="true" box="[1023,1113,152,176]" italics="true" pageId="21" pageNumber="50">Thitena</emphasis>
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<taxonomicNameLabel id="A2136D69FFDE6C58FBF8B9B4FB100EA0" box="[1121,1192,152,176]" pageId="21" pageNumber="50" rank="genus">n.gen.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C34E5F8BFFDE6C58FC83B9E0FA880EF4" box="[794,1328,204,228]" pageId="21" pageNumber="50" type="type_taxon">
<paragraph id="8BEB0C00FFDE6C58FC83B9E0FA880EF4" blockId="21.[794,1328,204,228]" box="[794,1328,204,228]" pageId="21" pageNumber="50">
<emphasis id="B920D012FFDE6C58FC83B9E0FC110EF4" bold="true" box="[794,937,204,228]" pageId="21" pageNumber="50">
<typeStatus id="54EFB2A2FFDE6C58FC83B9E0FCEB0EF4" box="[794,851,204,228]" pageId="21" pageNumber="50">Type</typeStatus>
species
</emphasis>
.
<taxonomicName id="4C547783FFDE6C58FC2EB9E1FB310EF4" ID-CoL="56HWJ" authority="Munari, 2004" authorityName="Munari" authorityYear="2004" box="[951,1161,205,228]" class="Insecta" family="Canacidae" genus="Thitena" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="50" phylum="Arthropoda" rank="species" species="cadaverina" status="sp. nov.">
<emphasis id="B920D012FFDE6C58FC2EB9E1FB310EF4" box="[951,1161,205,228]" italics="true" pageId="21" pageNumber="50">Thitena cadaverina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2136D69FFDE6C58FB09B9E1FB7B0EF4" box="[1168,1219,205,228]" pageId="21" pageNumber="50" rank="species">n.sp.</taxonomicNameLabel>
(
<figureCitation id="136F1085FFDE6C58FB4BB9E1FA9C0EF4" box="[1234,1316,205,228]" captionStart="Fig" captionStartId="21.[121,156,1975,1996]" captionTargetBox="[121,1417,873,2001]" captionTargetId="figure-528@21.[454,1445,861,2002]" captionTargetPageId="21" captionText="Fig. 29. Thitena cadaverina n.gen. and n.sp., habitus. (Drawing by G. DEste)." pageId="21" pageNumber="50">Fig. 29</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34E5F8BFFDE6C5BFC83B82CFDC10F18" lastPageId="22" lastPageNumber="51" pageId="21" pageNumber="50" type="diagnosis">
<paragraph id="8BEB0C00FFDE6C5BFC83B82CFDC10F18" blockId="21.[794,1422,256,837]" lastBlockId="22.[166,794,153,264]" lastPageId="22" lastPageNumber="51" pageId="21" pageNumber="50">
<emphasis id="B920D012FFDE6C58FC83B82CFC310F08" bold="true" box="[794,905,256,280]" pageId="21" pageNumber="50">Diagnosis</emphasis>
. Robust, yellowish brown flies with legs bearing strong setae and long and thick villosity which is particularly developed in male; head roundish; face depressed, bearing shining, foldlike tubercle above vibrissal angle; cephalic chaetotaxy typical of subfamily; frons without discernible golden patch on both sides of ocellar triangle, only with slight infuscation; arista microscopically pubescent; eye roundish, with few, sparse interfacetal setulae; longest diameter of eye over 3.5× as long as genal height. Thorax very slightly flattened dorsally, with chaetotaxy typical of subfamily; disc of scutellum pubescent (
<figureCitation id="136F1085FFDE6C58FB23BB0AFAB00C2D" box="[1210,1288,550,573]" captionStart="Fig" captionStartId="22.[839,874,492,513]" captionTargetBox="[1013,1289,165,460]" captionTargetId="figure-11@22.[1013,1289,141,461]" captionTargetPageId="22" captionText="Fig. 30. Thitena cadaverina n.gen. and n.sp., scutellum dorsal view. (Drawing by G. DEste)." pageId="21" pageNumber="50">Fig. 30</figureCitation>
); all thoracic setae distinctly thinner than those in all other
<taxonomicName id="4C547783FFDE6C58FA8CBB6FFA320C4A" authorityName="Hendel" authorityYear="1916" box="[1301,1418,579,602]" class="Insecta" family="Canacidae" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="50" phylum="Arthropoda" rank="subFamily" subFamily="Tethininae">Tethininae</taxonomicName>
. Legs very robust, strongly villose, particularly forelegs in male (an apomorphic character state with respect to the ground plan of the
<taxonomicName id="4C547783FFDE6C58FC70BBB7FBD80CA2" authorityName="Hendel" authorityYear="1916" box="[1001,1120,667,690]" class="Insecta" family="Tethinidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="50" phylum="Arthropoda" rank="family">Tethinidae</taxonomicName>
); tarsi of male stumpy, with tarsomeres noticeably short (in particular basitarsomeres), setulose, and flattened; true ctenidium absent; tibiae with rows of strong setae. Wing typical of subfamily; halter pale yellow. Abdomen rather robust, with tergites and sternites well sclerotized; male terminalia (
<figureCitation id="136F1085FFDE6C58FB08BA02FAB10D55" box="[1169,1289,814,837]" captionStart="Figs" captionStartId="23.[147,192,863,884]" captionTargetBox="[191,1354,120,831]" captionTargetId="figure-488@23.[191,1354,120,831]" captionTargetPageId="23" captionText="Figs. 3132. Terminalia of male. Thitena cadaverina n.gen. and n.sp.: (31) lateral view, (32) posterior view. Scale bar: 0.1 mm." pageId="21" pageNumber="50">Figs. 3132</figureCitation>
) with very long and slender anterior surstylus, and large posterior surstylus externally covered with dense microtrichia; both surstyli subequal in length and reciprocally parallel. Spermathecae of female spherical, smooth.
</paragraph>
</subSubSection>
<caption id="DF2B5C88FFDE6C58FFE0BE9BFCC009DC" box="[121,888,1975,1996]" pageId="21" pageNumber="50" startId="21.[121,156,1975,1996]" targetBox="[121,1417,873,2001]" targetPageId="21">
<paragraph id="8BEB0C00FFDE6C58FFE0BE9BFCC009DC" blockId="21.[121,888,1975,1996]" box="[121,888,1975,1996]" pageId="21" pageNumber="50">
Fig. 29.
<taxonomicName id="4C547783FFDE6C58FF52BE9BFE3309DC" authority="Munari, 2004" authorityName="Munari" authorityYear="2004" box="[203,395,1975,1996]" class="Insecta" family="Canacidae" genus="Thitena" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="50" phylum="Arthropoda" rank="species" species="cadaverina" status="n.gen. and n.sp.">
<emphasis id="B920D012FFDE6C58FF52BE9BFE3309DC" box="[203,395,1975,1996]" italics="true" pageId="21" pageNumber="50">Thitena cadaverina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2136D69FFDE6C58FE08BE9BFD9609DC" box="[401,558,1975,1996]" pageId="21" pageNumber="50" rank="species">n.gen. and n.sp.</taxonomicNameLabel>
, habitus. (Drawing by G. DEste).
</paragraph>
</caption>
<subSubSection id="C34E5F8BFFDD6C5BFF3FB808FE3F0CC6" pageId="22" pageNumber="51" type="discussion">
<paragraph id="8BEB0C00FFDD6C5BFF3FB808FE3F0CC6" blockId="22.[166,794,292,726]" pageId="22" pageNumber="51">
<emphasis id="B920D012FFDD6C5BFF3FB808FEB70F2C" bold="true" box="[166,271,292,316]" pageId="22" pageNumber="51">Remarks</emphasis>
. For some morphological and chaetotactic traits as well as for the general habitus, the new genus would seem to be related to
<taxonomicName id="4C547783FFDD6C5BFE0DB873FDE00F66" authorityName="Malloch" authorityYear="1914" box="[404,600,351,374]" class="Insecta" family="Canacidae" genus="Pseudorhicnoessa" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B920D012FFDD6C5BFE0DB873FDE00F66" box="[404,600,351,374]" italics="true" pageId="22" pageNumber="51">Pseudorhicnoessa</emphasis>
</taxonomicName>
, differing from it particularly for the following characters besides those given in the key: larger size; body and legs distinctly more robust and setulose; frons with numerous additional setulae; thoracic setae distinctly thinner; true ctenidium of fore femur absent; fore tibia with row of strong setae; mid and hind tibiae with different chaetotactic pattern; tarsi of male stumpy, with tarsomeres noticeably short (in particular basitarsomeres), setulose, and flattened; crossvein
<emphasis id="B920D012FFDD6C5BFD4FBB66FCA10C71" box="[726,793,586,609]" italics="true" pageId="22" pageNumber="51">dm-cu</emphasis>
longer, about as long as half of last section of CuA
<subScript id="17D00E45FFDD6C5BFC93BB5FFCAA0C92" attach="right" box="[778,786,627,642]" fontSize="6" pageId="22" pageNumber="51">1</subScript>
; abdomen with sternites well sclerotized; male terminalia with anterior and posterior surstyli subequal in length and reciprocally parallel.
</paragraph>
</subSubSection>
<caption id="DF2B5C88FFDD6C5BFCDEB8C0FBD50C0C" pageId="22" pageNumber="51" startId="22.[839,874,492,513]" targetBox="[1013,1289,165,460]" targetPageId="22">
<paragraph id="8BEB0C00FFDD6C5BFCDEB8C0FBD50C0C" blockId="22.[839,1467,492,540]" pageId="22" pageNumber="51">
Fig. 30.
<taxonomicName id="4C547783FFDD6C5BFC07B8C0FBD80C11" authority="Munari, 2004" authorityName="Munari" authorityYear="2004" box="[926,1120,492,513]" class="Insecta" family="Canacidae" genus="Thitena" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="51" phylum="Arthropoda" rank="species" species="cadaverina" status="n.gen. and n.sp.">
<emphasis id="B920D012FFDD6C5BFC07B8C0FBD80C11" box="[926,1120,492,513]" italics="true" pageId="22" pageNumber="51">Thitena cadaverina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2136D69FFDD6C5BFBF0B8C0FAB20C11" box="[1129,1290,492,513]" pageId="22" pageNumber="51" rank="species">n.gen. and n.sp.</taxonomicNameLabel>
, scutellum dorsal view. (Drawing by G. DEste).
</paragraph>
</caption>
<subSubSection id="C34E5F8BFFDD6C5BFF3FBBDFFD270D38" pageId="22" pageNumber="51" type="etymology">
<paragraph id="8BEB0C00FFDD6C5BFF3FBBDFFD270D38" blockId="22.[166,794,755,808]" pageId="22" pageNumber="51">
<emphasis id="B920D012FFDD6C5BFF3FBBDFFE9A0D1B" bold="true" box="[166,290,755,779]" pageId="22" pageNumber="51">Etymology</emphasis>
. The generic epithet derives from an anagram of the generic name
<taxonomicName id="4C547783FFDD6C5BFE1ABA3DFE6B0D38" box="[387,467,785,808]" class="Insecta" family="Canacidae" genus="Tethina" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B920D012FFDD6C5BFE1ABA3DFE6B0D38" box="[387,467,785,808]" italics="true" pageId="22" pageNumber="51">Tethina</emphasis>
</taxonomicName>
. Gender feminine.
</paragraph>
</subSubSection>
</treatment>
</document>