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<document id="B889D06EC18A6C4D08335058CB339B6F" ID-DOI="10.1080/00222930802610378" ID-ISSN="1464-5262" ID-Zenodo-Dep="4590612" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1615293246825" checkinUser="carolina" docAuthor="Liu, Xingyue, Hayashi, Fumio &amp; Yang, Ding" docDate="2009" docId="CA6F87DD4616FFADFE5B3EF11D23FD80" docLanguage="en" docName="JNATHIST.43.5-6.355-372.pdf" docOrigin="Journal of Natural History 43 (5 - 6)" docSource="http://dx.doi.org/10.1080/00222930802610378" docStyle="DocumentStyle:DE8A86AD4006EFA56F95B2AAEFBF2813.5:JNatHist.2008-2012.journal_article.2cover" docStyleId="DE8A86AD4006EFA56F95B2AAEFBF2813" docStyleName="JNatHist.2008-2012.journal_article.2cover" docStyleVersion="5" docTitle="Protohermes congruens Liu &amp; Hayashi &amp; Yang 2009, sp. nov." docType="treatment" docVersion="3" lastPageNumber="361" masterDocId="3656FFA54613FFA5FFE73D581F0DFF85" masterDocTitle="Systematics of the Protohermes parcus species group (Megaloptera: Corydalidae), with notes on its phylogeny and biogeography" masterLastPageNumber="372" masterPageNumber="355" pageNumber="358" updateTime="1698881889615" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="F40F8E29BA710539FA6F5C529A7A8F24" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0CE09244586D2E1AB88BFC97A3057DDB">
<mods:title id="DA4EE4323D3AD0BE695ED264E7F83B6C">Systematics of the Protohermes parcus species group (Megaloptera: Corydalidae), with notes on its phylogeny and biogeography</mods:title>
</mods:titleInfo>
<mods:name id="DD54FDFC299B8AECE1F718DF64DA81E3" type="personal">
<mods:role id="EAD7B457954D245AFE4C769D2AFB7C5A">
<mods:roleTerm id="E1FC1C9079E9DCBA0D8F18D2F336BFDD">Author</mods:roleTerm>
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<mods:namePart id="3A844C6C64CD7CE694C836751FB4FC93">Liu, Xingyue</mods:namePart>
</mods:name>
<mods:name id="EE8856A48D5AA68495B5C41926B5059F" type="personal">
<mods:role id="A217F4A53E3A35AD9AF9D339C49F6A67">
<mods:roleTerm id="F283E4A0596DA875062CD23820B8C039">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="866D212A204A68537F4EA468455EE772">Hayashi, Fumio</mods:namePart>
</mods:name>
<mods:name id="CEC9BDF1B1C681D0F0A653B8E9A80BB4" type="personal">
<mods:role id="68776343E29BCEFC2D25EFF88FA7DAB9">
<mods:roleTerm id="B531186C2C2AD9903ABF09C167D171D7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="91D74B2F7036C1B73D61197C3F42CE51">Yang, Ding</mods:namePart>
</mods:name>
<mods:typeOfResource id="076BC8D638CB46E22A720D11FBCD20EE">text</mods:typeOfResource>
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<mods:titleInfo id="EA1515932BED7250B604655A4F83A394">
<mods:title id="8970EFDD227F78AFC38A6BEB3599989D">Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part id="00FFCC7135DC28B7DC5A720124BA6284">
<mods:date id="DE137577D2A21D35FFA02CE66D7A92D9">2009</mods:date>
<mods:detail id="09A6ACA3863759090AC945463041796F" type="pubDate">
<mods:number id="79151ABECAD4D816474FCA827686F05E">2009-02-28</mods:number>
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<mods:detail id="48DF75D1179035C3D55E52CB4EAEA7CE" type="volume">
<mods:number id="57DA72C1D70200AAA879A6DCDCC93054">43</mods:number>
</mods:detail>
<mods:detail id="00C12D59E05BF336869012829FAA66C3" type="issue">
<mods:number id="BF195002F006B1575D23B54BDF0F90A9">5 - 6</mods:number>
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<mods:start id="FC6A96C442FA3FD5F3B8A5C5A878B853">355</mods:start>
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<mods:url id="AD02733FD77E9B29E98C6B537680FE9E">http://dx.doi.org/10.1080/00222930802610378</mods:url>
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<mods:classification id="DB61F61091FED76280B239615AA435C1">journal article</mods:classification>
<mods:identifier id="F2726F20AC150CF0C9BE89C87C344619" type="DOI">10.1080/00222930802610378</mods:identifier>
<mods:identifier id="929013C90EE8B2D049C4E87D7E532A1B" type="ISSN">1464-5262</mods:identifier>
<mods:identifier id="C3B3ACDF36F86EF43AFFCA94EF5866AD" type="Zenodo-Dep">4590612</mods:identifier>
</mods:mods>
<treatment id="CA6F87DD4616FFADFE5B3EF11D23FD80" LSID="urn:lsid:plazi:treatment:CA6F87DD4616FFADFE5B3EF11D23FD80" httpUri="http://treatment.plazi.org/id/CA6F87DD4616FFADFE5B3EF11D23FD80" lastPageId="8" lastPageNumber="361" pageId="5" pageNumber="358">
<subSubSection id="0ADC65404616FFA0FE5B3EF11C28FC44" box="[444,805,937,961]" pageId="5" pageNumber="358" type="nomenclature">
<paragraph id="427936CB4616FFA0FE5B3EF11C28FC44" blockId="5.[444,805,937,993]" box="[444,805,937,961]" pageId="5" pageNumber="358">
<heading id="193181A74616FFA0FE5B3EF11C28FC44" box="[444,805,937,961]" centered="true" fontSize="36" level="2" pageId="5" pageNumber="358" reason="7">
<taxonomicName id="85C64D484616FFA0FE5B3EF11DCDFC44" ID-CoL="4N5H4" authority="Liu &amp; Hayashi &amp; Yang, 2009" authorityName="Liu &amp; Hayashi &amp; Yang" authorityYear="2009" box="[444,704,937,961]" class="Insecta" family="Corydalidae" genus="Protohermes" kingdom="Animalia" order="Megaloptera" pageId="5" pageNumber="358" phylum="Arthropoda" rank="species" species="congruens" status="sp. nov.">
<emphasis id="70B2EAD94616FFA0FE5B3EF11DCDFC44" bold="true" box="[444,704,937,961]" italics="true" pageId="5" pageNumber="358">Protohermes congruens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6B8157A24616FFA0FD2E3EF11C28FC44" box="[713,805,937,961]" pageId="5" pageNumber="358" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0ADC65404616FFADFDF63E911D23FD80" lastPageId="8" lastPageNumber="361" pageId="5" pageNumber="358" type="description">
<paragraph id="427936CB4616FFA0FDF63E911DC2FC64" blockId="5.[444,805,937,993]" box="[529,719,969,993]" pageId="5" pageNumber="358">
(
<figureCitation id="DAFD2A4E4616FFA0FDFE3E911D8FFC64" box="[537,642,969,993]" captionStart="Figures 1–5" captionStartId="6.[213,289,947,969]" captionTargetBox="[213,1170,165,910]" captionTargetId="figure-164@6.[213,1170,165,910]" captionTargetPageId="6" captionText="Figures 1–5. Habitus photographs of the species in P. parcus group. (1) Protohermes congruens sp. nov., male holotype; (2) Protohermes flavinervus sp. nov., male holotype; (3) Protohermes parcus Yang and Yang, male holotype; (4) Protohermes pennyi sp. nov., male holotype; (5) Protohermes subparcus Liu and Yang, male holotype." figureDoi="http://doi.org/10.5281/zenodo.4590614" httpUri="https://zenodo.org/record/4590614/files/figure.png" pageId="5" pageNumber="358">Figures 1</figureCitation>
,
<figureCitation id="DAFD2A4E4616FFA0FD763E911DC5FC64" box="[657,712,969,993]" captionStart="Figures 6–11" captionStartId="7.[146,224,835,857]" captionTargetBox="[151,1103,171,793]" captionTargetId="figure-137@7.[146,1104,165,798]" captionTargetPageId="7" captionText="Figures 6–11. Protohermes congruens sp. nov. (6) Male head and prothorax, dorsal; (7) male genitalia, dorsal view; (8) male genitalia, ventral view; (9) male tenth sternum, ventral view; (10) female abdominal apex, lateral view; (11) female eighth sternum, ventral view. Notes: L, lateral lobe; Gs9, ninth gonostylus; Gx9, ninth gonocoxite; Pdm, dorsomedial process; S8, eighth sternum; S9, ninth sternum; T9, ninth tergum; T10, tenth tergum. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4590616" httpUri="https://zenodo.org/record/4590616/files/figure.png" pageId="5" pageNumber="358">6–11</figureCitation>
)
</paragraph>
<paragraph id="427936CB4616FFA0FF7539421E06FBB7" blockId="5.[146,267,1050,1074]" box="[146,267,1050,1074]" pageId="5" pageNumber="358">
<emphasis id="70B2EAD94616FFA0FF7539421E06FBB7" box="[146,267,1050,1074]" italics="true" pageId="5" pageNumber="358">Etymology</emphasis>
</paragraph>
<paragraph id="427936CB4616FFA0FF75391B1EF7FBFE" blockId="5.[146,1103,1091,1147]" pageId="5" pageNumber="358">
The specific epithet ‘‘
<taxonomicName id="85C64D484616FFA0FE66391B1EE2FBDE" authorityName="Liu &amp; Hayashi &amp; Yang" authorityYear="2009" box="[385,495,1091,1115]" class="Insecta" family="Corydalidae" genus="Protohermes" kingdom="Animalia" order="Megaloptera" pageId="5" pageNumber="358" phylum="Arthropoda" rank="species" species="congruens">
<emphasis id="70B2EAD94616FFA0FE66391B1EE2FBDE" box="[385,495,1091,1115]" italics="true" pageId="5" pageNumber="358">congruens</emphasis>
</taxonomicName>
’’ refers to the similar appearance among
<taxonomicName id="85C64D484616FFA0FC27391B1B25FBDE" box="[960,1064,1091,1115]" class="Insecta" family="Corydalidae" genus="Protohermes" kingdom="Animalia" order="Megaloptera" pageId="5" pageNumber="358" phylum="Arthropoda" rank="species" species="parcus">
<emphasis id="70B2EAD94616FFA0FC27391B1B25FBDE" box="[960,1064,1091,1115]" italics="true" pageId="5" pageNumber="358">P. parcus</emphasis>
</taxonomicName>
,
<taxonomicName id="85C64D484616FFA0FBD1391B1FF2FBFE" class="Insecta" family="Corydalidae" genus="Protohermes" kingdom="Animalia" order="Megaloptera" pageId="5" pageNumber="358" phylum="Arthropoda" rank="species" species="subparcus">
<emphasis id="70B2EAD94616FFA0FBD1391B1FF2FBFE" italics="true" pageId="5" pageNumber="358">P. subparcus</emphasis>
</taxonomicName>
, and the new species.
</paragraph>
<paragraph id="427936CB4616FFA0FF7539E71FF2FB52" blockId="5.[146,255,1215,1239]" box="[146,255,1215,1239]" pageId="5" pageNumber="358">
<emphasis id="70B2EAD94616FFA0FF7539E71FF2FB52" box="[146,255,1215,1239]" italics="true" pageId="5" pageNumber="358">Diagnosis</emphasis>
</paragraph>
<paragraph id="427936CB4616FFA0FF7539BF1D5FFAE4" blockId="5.[146,1103,1255,1377]" pageId="5" pageNumber="358">Head without any markings; pronotum with three pairs of slender vittae and a pair of small subtriangular markings; male ninth sternum subtrapezoidal, shallowly incised medially, with posterolateral corners acutely pointed; male tenth sternum with short and digitiform lateral lobes.</paragraph>
<paragraph id="427936CB4616FFA0FF7538FC1E37FA39" blockId="5.[146,314,1444,1468]" box="[146,314,1444,1468]" pageId="5" pageNumber="358">
<emphasis id="70B2EAD94616FFA0FF7538FC1E37FA39" box="[146,314,1444,1468]" italics="true" pageId="5" pageNumber="358">Type materials</emphasis>
</paragraph>
<paragraph id="427936CB4616FFA0FF7538951D78F9A3" blockId="5.[146,1102,1485,1574]" pageId="5" pageNumber="358">
<materialsCitation id="F2AE3C964616FFA0FF7538951DFDFA60" box="[146,752,1485,1509]" collectingDate="1972-05-30" collectionCode="NSM" country="Nepal" location="NEPAL" pageId="5" pageNumber="358" specimenCount="1" typeStatus="holotype">
<typeStatus id="9D7D88694616FFA0FF7538951E0CFA60" box="[146,257,1485,1509]" pageId="5" pageNumber="358" type="holotype">Holotype</typeStatus>
: male,
<collectingCountry id="3AD1765B4616FFA0FEB638951EBFFA60" box="[337,434,1485,1509]" name="Nepal" pageId="5" pageNumber="358">NEPAL</collectingCountry>
, L.T.,
<date id="3678100B4616FFA0FE1838951D99FA60" box="[511,660,1485,1509]" pageId="5" pageNumber="358" value="1972-05-30">
<collectingDate id="263CE9E34616FFA0FE1838951D99FA60" box="[511,660,1485,1509]" pageId="5" pageNumber="358" value="1972-05-30">30 May 1972</collectingDate>
</date>
(
<collectionCode id="24D7AE0E4616FFA0FD4338951DE5FA60" box="[676,744,1485,1509]" country="Canada" httpUri="http://grbio.org/cool/zeys-ppzt" name="Nova Scotia Museum of Natural History" pageId="5" pageNumber="358">NSM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="F2AE3C964616FFA0FD1B38951EB1F980" collectingDate="1972-06-01" collectionCode="NSM" country="Nepal" location="NEPAL" pageId="5" pageNumber="358" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<typeStatus id="9D7D88694616FFA0FD1B38951C62FA60" box="[764,879,1485,1509]" pageId="5" pageNumber="358" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="3AD1765B4616FFA0FC9D38951CD1FA60" box="[890,988,1485,1509]" name="Nepal" pageId="5" pageNumber="358">NEPAL</collectingCountry>
:
<specimenCount id="54C0FD424616FFA0FC0F38951B46FA60" box="[1000,1099,1485,1509]" pageId="5" pageNumber="358" type="female">1 female</specimenCount>
, L.T.,
<date id="3678100B4616FFA0FF3238B51E52F980" box="[213,351,1517,1541]" pageId="5" pageNumber="358" value="1972-06-01">
<collectingDate id="263CE9E34616FFA0FF3238B51E52F980" box="[213,351,1517,1541]" pageId="5" pageNumber="358" value="1972-06-01">1 June 1972</collectingDate>
</date>
(
<collectionCode id="24D7AE0E4616FFA0FE9638B51EB9F980" box="[369,436,1517,1541]" country="Canada" httpUri="http://grbio.org/cool/zeys-ppzt" name="Nova Scotia Museum of Natural History" pageId="5" pageNumber="358">NSM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="F2AE3C964616FFA0FE2D38B51C11F980" box="[458,796,1517,1541]" collectingDate="1972-05-30" collectionCode="NSM" country="Nepal" pageId="5" pageNumber="358" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="54C0FD424616FFA0FE2D38B51D17F980" box="[458,538,1517,1541]" pageId="5" pageNumber="358" type="male">1 male</specimenCount>
,
<date id="3678100B4616FFA0FDCF38B51DB2F980" box="[552,703,1517,1541]" pageId="5" pageNumber="358" value="1972-05-30">
<collectingDate id="263CE9E34616FFA0FDCF38B51DB2F980" box="[552,703,1517,1541]" pageId="5" pageNumber="358" value="1972-05-30">30 May 1972</collectingDate>
</date>
(
<collectionCode id="24D7AE0E4616FFA0FD3638B51C19F980" box="[721,788,1517,1541]" country="Canada" httpUri="http://grbio.org/cool/zeys-ppzt" name="Nova Scotia Museum of Natural History" pageId="5" pageNumber="358">NSM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="F2AE3C964616FFA0FCCD38B51FE8F9A3" collectingDate="1972-06-01" collectionCode="NSM" country="Nepal" pageId="5" pageNumber="358" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="54C0FD424616FFA0FCCD38B51C77F980" box="[810,890,1517,1541]" pageId="5" pageNumber="358" type="male">1 male</specimenCount>
, J.E.,
<date id="3678100B4616FFA0FC2338B51B43F980" box="[964,1102,1517,1541]" pageId="5" pageNumber="358" value="1972-06-01">
<collectingDate id="263CE9E34616FFA0FC2338B51B43F980" box="[964,1102,1517,1541]" pageId="5" pageNumber="358" value="1972-06-01">1 June 1972</collectingDate>
</date>
(
<collectionCode id="24D7AE0E4616FFA0FF7D3B561FD0F9A3" box="[154,221,1550,1574]" country="Canada" httpUri="http://grbio.org/cool/zeys-ppzt" name="Nova Scotia Museum of Natural History" pageId="5" pageNumber="358">NSM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="F2AE3C964616FFA0FF143B561D7CF9A3" box="[243,625,1550,1574]" collectingDate="1972-06-02" collectionCode="NSM" country="Nepal" pageId="5" pageNumber="358" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="54C0FD424616FFA0FF143B561E4FF9A3" box="[243,322,1550,1574]" pageId="5" pageNumber="358" type="male">1 male</specimenCount>
, J.E.,
<date id="3678100B4616FFA0FE6C3B561D19F9A3" box="[395,532,1550,1574]" pageId="5" pageNumber="358" value="1972-06-02">
<collectingDate id="263CE9E34616FFA0FE6C3B561D19F9A3" box="[395,532,1550,1574]" pageId="5" pageNumber="358" value="1972-06-02">2 June 1972</collectingDate>
</date>
(
<collectionCode id="24D7AE0E4616FFA0FDC23B561D64F9A3" box="[549,617,1550,1574]" country="Canada" httpUri="http://grbio.org/cool/zeys-ppzt" name="Nova Scotia Museum of Natural History" pageId="5" pageNumber="358">NSM</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="427936CB4616FFA0FF753B311FC1F904" blockId="5.[146,204,1641,1665]" box="[146,204,1641,1665]" pageId="5" pageNumber="358">
<emphasis id="70B2EAD94616FFA0FF753B311FC1F904" box="[146,204,1641,1665]" italics="true" pageId="5" pageNumber="358">Male</emphasis>
</paragraph>
<paragraph id="427936CB4616FFA3FF753BCA1E8FFB5B" blockId="5.[146,1061,1682,1706]" lastBlockId="6.[213,1169,1125,1246]" lastPageId="6" lastPageNumber="359" pageId="5" pageNumber="358">
Body length
<quantity id="853E9B2E4616FFA0FEC03BCA1E90F92F" box="[295,413,1682,1706]" metricMagnitude="-2" metricUnit="m" metricValue="2.9" metricValueMax="3.0" metricValueMin="2.8" pageId="5" pageNumber="358" unit="mm" value="29.0" valueMax="30.0" valueMin="28.0">28–30 mm</quantity>
; forewing length
<quantity id="853E9B2E4616FFA0FD823BCA1DD1F92F" box="[613,732,1682,1706]" metricMagnitude="-2" metricUnit="m" metricValue="4.05" metricValueMax="4.2" metricValueMin="3.9" pageId="5" pageNumber="358" unit="mm" value="40.5" valueMax="42.0" valueMin="39.0">39–42 mm</quantity>
, hindwing length
<quantity id="853E9B2E4616FFA0FC4D3BCA1B2DF92F" box="[938,1056,1682,1706]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" metricValueMax="3.7" metricValueMin="3.5" pageId="5" pageNumber="358" unit="mm" value="36.0" valueMax="37.0" valueMin="35.0">35–37 mm</quantity>
.
<emphasis id="70B2EAD94615FFA3FF32393D1E1CFBF8" box="[213,273,1125,1149]" italics="true" pageId="6" pageNumber="359">Head</emphasis>
. Yellow without any markings; post-ocular spine short and blunt. Compound eyes grayish brown; ocelli pale yellow, medially margined black. Antennae blackish brown, with scape and pedicel yellow. Mouthparts yellow; mandibles with apices reddish brown.
</paragraph>
<caption id="16B966434615FFA3FF323EEB1C87FBA5" ID-DOI="http://doi.org/10.5281/zenodo.4590614" ID-Zenodo-Dep="4590614" httpUri="https://zenodo.org/record/4590614/files/figure.png" pageId="6" pageNumber="359" startId="6.[213,289,947,969]" targetBox="[213,1170,165,910]" targetPageId="6">
<paragraph id="427936CB4615FFA3FF323EEB1C87FBA5" blockId="6.[213,1169,947,1056]" pageId="6" pageNumber="359">
Figures 1–5. Habitus photographs of the species in
<taxonomicName id="85C64D484615FFA3FCC63EEB1C8AFC4D" box="[801,903,947,968]" class="Insecta" family="Corydalidae" genus="Protohermes" kingdom="Animalia" order="Megaloptera" pageId="6" pageNumber="359" phylum="Arthropoda" rank="species" species="parcus">
<emphasis id="70B2EAD94615FFA3FCC63EEB1C8AFC4D" box="[801,903,947,968]" italics="true" pageId="6" pageNumber="359">P. parcus</emphasis>
</taxonomicName>
group. (1)
<taxonomicName id="85C64D484615FFA3FBF53EEB1E35FC60" authority="Liu &amp; Hayashi &amp; Yang, 2009" authorityName="Liu &amp; Hayashi &amp; Yang" authorityYear="2009" class="Insecta" family="Corydalidae" genus="Protohermes" kingdom="Animalia" order="Megaloptera" pageId="6" pageNumber="359" phylum="Arthropoda" rank="species" species="congruens" status="sp. nov.">
<emphasis id="70B2EAD94615FFA3FBF53EEB1E35FC60" italics="true" pageId="6" pageNumber="359">Protohermes congruens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6B8157A24615FFA3FEA53E881E95FC63" box="[322,408,976,998]" pageId="6" pageNumber="359" rank="species">sp. nov.</taxonomicNameLabel>
, male holotype; (2)
<taxonomicName id="85C64D484615FFA3FD973E881C6FFC60" authority="Liu &amp; Hayashi &amp; Yang, 2009" authorityName="Liu &amp; Hayashi &amp; Yang" authorityYear="2009" box="[624,866,976,997]" class="Insecta" family="Corydalidae" genus="Protohermes" kingdom="Animalia" order="Megaloptera" pageId="6" pageNumber="359" phylum="Arthropoda" rank="species" species="flavinervus" status="sp. nov.">
<emphasis id="70B2EAD94615FFA3FD973E881C6FFC60" box="[624,866,976,997]" italics="true" pageId="6" pageNumber="359">Protohermes flavinervus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6B8157A24615FFA3FC8A3E881CCEFC63" box="[877,963,976,998]" pageId="6" pageNumber="359" rank="species">sp. nov.</taxonomicNameLabel>
, male holotype; (3)
<taxonomicName id="85C64D484615FFA3FF323EB51D5EFB86" authority="Yang and Yang" authorityName="Yang and Yang" box="[213,595,1005,1027]" class="Insecta" family="Corydalidae" genus="Protohermes" kingdom="Animalia" order="Megaloptera" pageId="6" pageNumber="359" phylum="Arthropoda" rank="species" species="parcus">
<emphasis id="70B2EAD94615FFA3FF323EB51E93FB87" box="[213,414,1005,1026]" italics="true" pageId="6" pageNumber="359">Protohermes parcus</emphasis>
Yang and Yang
</taxonomicName>
, male holotype; (4)
<taxonomicName id="85C64D484615FFA3FCCD3EB51CFEFB87" authority="Liu &amp; Hayashi &amp; Yang, 2009" authorityName="Liu &amp; Hayashi &amp; Yang" authorityYear="2009" box="[810,1011,1005,1026]" class="Insecta" family="Corydalidae" genus="Protohermes" kingdom="Animalia" order="Megaloptera" pageId="6" pageNumber="359" phylum="Arthropoda" rank="species" species="pennyi" status="sp. nov.">
<emphasis id="70B2EAD94615FFA3FCCD3EB51CFEFB87" box="[810,1011,1005,1026]" italics="true" pageId="6" pageNumber="359">Protohermes pennyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6B8157A24615FFA3FC1A3EB51B5EFB86" box="[1021,1107,1005,1027]" pageId="6" pageNumber="359" rank="species">sp. nov.</taxonomicNameLabel>
, male holotype; (5)
<taxonomicName id="85C64D484615FFA3FE8639531DE8FBA5" authority="Liu and Yang" authorityName="Liu and Yang" box="[353,741,1034,1056]" class="Insecta" family="Corydalidae" genus="Protohermes" kingdom="Animalia" order="Megaloptera" pageId="6" pageNumber="359" phylum="Arthropoda" rank="species" species="subparcus">
<emphasis id="70B2EAD94615FFA3FE8639531D44FBA5" box="[353,585,1035,1056]" italics="true" pageId="6" pageNumber="359">Protohermes subparcus</emphasis>
Liu and Yang
</taxonomicName>
, male holotype.
</paragraph>
</caption>
<paragraph id="427936CB4615FFA3FF32384F1D21F997" blockId="6.[213,1169,1303,1554]" pageId="6" pageNumber="359">
<emphasis id="70B2EAD94615FFA3FF32384F1E2AFAAA" box="[213,295,1303,1327]" italics="true" pageId="6" pageNumber="359">Thorax</emphasis>
. Yellow; pronotum with two pairs of narrow blackish vittae on lateral margins, anterior pair with apices curved laterally, posterior pair separated into two pairs of slender vittae and a pair of small subtriangular markings. Thoracic pilosity pale yellow, much longer on meso- and metathorax. Legs yellow with yellowish short dense setae; tarsal claws reddish brown. Wings hyaline, immaculate; veins yellow to yellowish brown, with crossveins much darker. Rs nine-branched, last branch bifurcate; 9–15 crossveins between R
<subScript id="DE42348E4615FFA3FD7638BC1D96FA71" attach="left" box="[657,667,1508,1524]" fontSize="7" pageId="6" pageNumber="359">1</subScript>
and Rs; M
<subScript id="DE42348E4615FFA3FCD638BC1C5DFA71" attach="left" box="[817,848,1506,1524]" fontSize="7" pageId="6" pageNumber="359">1+2</subScript>
four-branched, M
<subScript id="DE42348E4615FFA3FBD738BC1B42FA71" attach="left" box="[1072,1103,1506,1524]" fontSize="7" pageId="6" pageNumber="359">3+4</subScript>
twobranched; 1A three-branched.
</paragraph>
<paragraph id="427936CB4615FFA2FF323B131E28FA9C" blockId="6.[213,1169,1610,1699]" lastBlockId="7.[146,1102,1086,1305]" lastPageId="7" lastPageNumber="360" pageId="6" pageNumber="359">
<emphasis id="70B2EAD94615FFA3FF323B131E4DF9E6" box="[213,320,1611,1635]" italics="true" pageId="6" pageNumber="359">Abdomen</emphasis>
. Brown with venter yellow. Ninth tergum (
<figureCitation id="DAFD2A4E4615FFA3FC953B121CDEF9E7" box="[882,979,1610,1634]" captionStart="Figures 6–11" captionStartId="7.[146,224,835,857]" captionTargetBox="[151,1103,171,793]" captionTargetId="figure-137@7.[146,1104,165,798]" captionTargetPageId="7" captionText="Figures 6–11. Protohermes congruens sp. nov. (6) Male head and prothorax, dorsal; (7) male genitalia, dorsal view; (8) male genitalia, ventral view; (9) male tenth sternum, ventral view; (10) female abdominal apex, lateral view; (11) female eighth sternum, ventral view. Notes: L, lateral lobe; Gs9, ninth gonostylus; Gx9, ninth gonocoxite; Pdm, dorsomedial process; S8, eighth sternum; S9, ninth sternum; T9, ninth tergum; T10, tenth tergum. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4590616" httpUri="https://zenodo.org/record/4590616/files/figure.png" pageId="6" pageNumber="359">Figure 7</figureCitation>
) broad, nearly hexagonal, with an arcuate anterior incision. Ninth sternum (
<figureCitation id="DAFD2A4E4615FFA3FC2A3B331B23F906" box="[973,1070,1643,1667]" captionStart="Figures 6–11" captionStartId="7.[146,224,835,857]" captionTargetBox="[151,1103,171,793]" captionTargetId="figure-137@7.[146,1104,165,798]" captionTargetPageId="7" captionText="Figures 6–11. Protohermes congruens sp. nov. (6) Male head and prothorax, dorsal; (7) male genitalia, dorsal view; (8) male genitalia, ventral view; (9) male tenth sternum, ventral view; (10) female abdominal apex, lateral view; (11) female eighth sternum, ventral view. Notes: L, lateral lobe; Gs9, ninth gonostylus; Gx9, ninth gonocoxite; Pdm, dorsomedial process; S8, eighth sternum; S9, ninth sternum; T9, ninth tergum; T10, tenth tergum. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4590616" httpUri="https://zenodo.org/record/4590616/files/figure.png" pageId="6" pageNumber="359">Figure 8</figureCitation>
) broad, subtrapezoidal, medially with feebly developed longitudinal incision; posterior margin nearly truncate, posterolateral corner acutely pointed. Ninth gonostylus unguiform, much shorter than ninth tergum, slightly curved inward. Tenth tergum (
<figureCitation id="DAFD2A4E4614FFA2FF1F39271E72FB12" box="[248,383,1151,1175]" captionStart="Figures 6–11" captionStartId="7.[146,224,835,857]" captionTargetBox="[151,1103,171,793]" captionTargetId="figure-137@7.[146,1104,165,798]" captionTargetPageId="7" captionText="Figures 6–11. Protohermes congruens sp. nov. (6) Male head and prothorax, dorsal; (7) male genitalia, dorsal view; (8) male genitalia, ventral view; (9) male tenth sternum, ventral view; (10) female abdominal apex, lateral view; (11) female eighth sternum, ventral view. Notes: L, lateral lobe; Gs9, ninth gonostylus; Gx9, ninth gonocoxite; Pdm, dorsomedial process; S8, eighth sternum; S9, ninth sternum; T9, ninth tergum; T10, tenth tergum. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4590616" httpUri="https://zenodo.org/record/4590616/files/figure.png" pageId="7" pageNumber="360">Figures 7–8</figureCitation>
) flattened, subquadrate, slightly shorter than ninth tergum, straightly directed posteriorly, with distal margin oblique and rounded. Tenth sternum (
<figureCitation id="DAFD2A4E4614FFA2FEE339981E68FB5D" box="[260,357,1216,1240]" captionStart="Figures 6–11" captionStartId="7.[146,224,835,857]" captionTargetBox="[151,1103,171,793]" captionTargetId="figure-137@7.[146,1104,165,798]" captionTargetPageId="7" captionText="Figures 6–11. Protohermes congruens sp. nov. (6) Male head and prothorax, dorsal; (7) male genitalia, dorsal view; (8) male genitalia, ventral view; (9) male tenth sternum, ventral view; (10) female abdominal apex, lateral view; (11) female eighth sternum, ventral view. Notes: L, lateral lobe; Gs9, ninth gonostylus; Gx9, ninth gonocoxite; Pdm, dorsomedial process; S8, eighth sternum; S9, ninth sternum; T9, ninth tergum; T10, tenth tergum. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4590616" httpUri="https://zenodo.org/record/4590616/files/figure.png" pageId="7" pageNumber="360">Figure 9</figureCitation>
) extremely small; dorsomedial process moderately developed; posterior margin medially with a small incision; lateral lobes short digitiform with rounded tip.
</paragraph>
<caption id="16B966434614FFA2FF753E1B1E00FC6E" ID-DOI="http://doi.org/10.5281/zenodo.4590616" ID-Zenodo-Dep="4590616" httpUri="https://zenodo.org/record/4590616/files/figure.png" pageId="7" pageNumber="360" startId="7.[146,224,835,857]" targetBox="[151,1103,171,793]" targetPageId="7">
<paragraph id="427936CB4614FFA2FF753E1B1E00FC6E" blockId="7.[146,1104,835,1003]" pageId="7" pageNumber="360">
Figures 6–11.
<taxonomicName id="85C64D484614FFA2FEC83E1B1D2DFCDD" authority="Liu &amp; Hayashi &amp; Yang, 2009" authorityName="Liu &amp; Hayashi &amp; Yang" authorityYear="2009" box="[303,544,835,856]" class="Insecta" family="Corydalidae" genus="Protohermes" kingdom="Animalia" order="Megaloptera" pageId="7" pageNumber="360" phylum="Arthropoda" rank="species" species="congruens" status="sp. nov.">
<emphasis id="70B2EAD94614FFA2FEC83E1B1D2DFCDD" box="[303,544,835,856]" italics="true" pageId="7" pageNumber="360">Protohermes congruens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6B8157A24614FFA2FDCB3E1B1D8FFCDC" box="[556,642,835,857]" pageId="7" pageNumber="360" rank="species">sp. nov.</taxonomicNameLabel>
(6) Male head and prothorax, dorsal; (7) male genitalia, dorsal view; (8) male genitalia, ventral view; (9) male tenth sternum, ventral view; (10) female abdominal apex, lateral view; (11) female eighth sternum, ventral view. Notes: L, lateral lobe; Gs9, ninth gonostylus; Gx9, ninth gonocoxite; Pdm, dorsomedial process; S8, eighth sternum; S9, ninth sternum; T9, ninth tergum; T10, tenth tergum. Scale bars: 1 mm.
</paragraph>
</caption>
<paragraph id="427936CB4614FFA2FF7538071FEFFAF2" blockId="7.[146,226,1375,1399]" box="[146,226,1375,1399]" pageId="7" pageNumber="360">
<emphasis id="70B2EAD94614FFA2FF7538071FEFFAF2" box="[146,226,1375,1399]" italics="true" pageId="7" pageNumber="360">Female</emphasis>
</paragraph>
<paragraph id="427936CB4614FFA2FF7538D01CA0FA25" blockId="7.[146,941,1416,1440]" box="[146,941,1416,1440]" pageId="7" pageNumber="360">
Body length
<quantity id="853E9B2E4614FFA2FEC038D01E78FA25" box="[295,373,1416,1440]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="7" pageNumber="360" unit="mm" value="32.0">32 mm</quantity>
; forewing length
<quantity id="853E9B2E4614FFA2FDD938D01D81FA25" box="[574,652,1416,1440]" metricMagnitude="-2" metricUnit="m" metricValue="5.6" pageId="7" pageNumber="360" unit="mm" value="56.0">56 mm</quantity>
, hindwing length
<quantity id="853E9B2E4614FFA2FCBD38D01CA5FA25" box="[858,936,1416,1440]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="360" unit="mm" value="50.0">50 mm</quantity>
.
</paragraph>
<paragraph id="427936CB4614FFA2FF7538BE1FDBF925" blockId="7.[146,1103,1510,1696]" pageId="7" pageNumber="360">
<emphasis id="70B2EAD94614FFA2FF7538BE1FF0FA7B" box="[146,253,1510,1534]" italics="true" pageId="7" pageNumber="360">Abdomen</emphasis>
. Eighth sternum (
<figureCitation id="DAFD2A4E4614FFA2FE0938BE1D9DFA7B" box="[494,656,1510,1534]" captionStart="Figures 6–11" captionStartId="7.[146,224,835,857]" captionTargetBox="[151,1103,171,793]" captionTargetId="figure-137@7.[146,1104,165,798]" captionTargetPageId="7" captionText="Figures 6–11. Protohermes congruens sp. nov. (6) Male head and prothorax, dorsal; (7) male genitalia, dorsal view; (8) male genitalia, ventral view; (9) male tenth sternum, ventral view; (10) female abdominal apex, lateral view; (11) female eighth sternum, ventral view. Notes: L, lateral lobe; Gs9, ninth gonostylus; Gx9, ninth gonocoxite; Pdm, dorsomedial process; S8, eighth sternum; S9, ninth sternum; T9, ninth tergum; T10, tenth tergum. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4590616" httpUri="https://zenodo.org/record/4590616/files/figure.png" pageId="7" pageNumber="360">Figures 10–11</figureCitation>
) subtrapezoidal in lateral view; in ventral view anterior margin moderately incised, posterior margin truncate. Ninth gonocoxite broad, posterior portion subquadrate, ventrally slightly incised, with a small digitiform gonostylus at tip. Tenth tergum short, with posterior margin medially incised, leaving thick, digitiform dorsal, and semicircular ventral lobes.
</paragraph>
<paragraph id="427936CB461BFFADFF323DF11E56FF44" blockId="8.[213,347,169,193]" box="[213,347,169,193]" pageId="8" pageNumber="361">
<emphasis id="70B2EAD9461BFFADFF323DF11E56FF44" box="[213,347,169,193]" italics="true" pageId="8" pageNumber="361">Distribution</emphasis>
</paragraph>
<paragraph id="427936CB461BFFADFF323D891CB7FF6C" blockId="8.[213,954,209,233]" box="[213,954,209,233]" pageId="8" pageNumber="361">
<collectingCountry id="3AD1765B461BFFADFF323D891E10FF6C" box="[213,285,209,233]" name="Nepal" pageId="8" pageNumber="361">Nepal</collectingCountry>
, without exact geographical data (longitude and latitude).
</paragraph>
<paragraph id="427936CB461BFFADFF323C7A1E35FEBF" blockId="8.[213,312,290,314]" box="[213,312,290,314]" pageId="8" pageNumber="361">
<emphasis id="70B2EAD9461BFFADFF323C7A1E35FEBF" box="[213,312,290,314]" italics="true" pageId="8" pageNumber="361">Remarks</emphasis>
</paragraph>
<paragraph id="427936CB461BFFADFF323C131D23FD80" blockId="8.[213,1169,331,517]" pageId="8" pageNumber="361">
The new species somewhat resembles
<taxonomicName id="85C64D48461BFFADFD9C3C131DEFFEE6" box="[635,738,331,355]" class="Insecta" family="Corydalidae" genus="Protohermes" kingdom="Animalia" order="Megaloptera" pageId="8" pageNumber="361" phylum="Arthropoda" rank="species" species="parcus">
<emphasis id="70B2EAD9461BFFADFD9C3C131DEFFEE6" box="[635,738,331,355]" italics="true" pageId="8" pageNumber="361">P. parcus</emphasis>
</taxonomicName>
and
<taxonomicName id="85C64D48461BFFADFCFE3C131CA8FEE6" box="[793,933,331,355]" class="Insecta" family="Corydalidae" genus="Protohermes" kingdom="Animalia" order="Megaloptera" pageId="8" pageNumber="361" phylum="Arthropoda" rank="species" species="subparcus">
<emphasis id="70B2EAD9461BFFADFCFE3C131CA8FEE6" box="[793,933,331,355]" italics="true" pageId="8" pageNumber="361">P. subparcus</emphasis>
</taxonomicName>
in having the similar appearance, but it can be easily separated from the latter two species by the male ninth sternum with a shallow longitudinal incision and the male tenth sternum with the digitiform lateral lobes. In
<taxonomicName id="85C64D48461BFFADFDDC3CF41DABFE41" box="[571,678,428,452]" class="Insecta" family="Corydalidae" genus="Protohermes" kingdom="Animalia" order="Megaloptera" pageId="8" pageNumber="361" phylum="Arthropoda" rank="species" species="parcus">
<emphasis id="70B2EAD9461BFFADFDDC3CF41DABFE41" box="[571,678,428,452]" italics="true" pageId="8" pageNumber="361">P. parcus</emphasis>
</taxonomicName>
and
<taxonomicName id="85C64D48461BFFADFD043CF41C7FFE41" box="[739,882,428,452]" class="Insecta" family="Corydalidae" genus="Protohermes" kingdom="Animalia" order="Megaloptera" pageId="8" pageNumber="361" phylum="Arthropoda" rank="species" species="subparcus">
<emphasis id="70B2EAD9461BFFADFD043CF41C7FFE41" box="[739,882,428,452]" italics="true" pageId="8" pageNumber="361">P. subparcus</emphasis>
</taxonomicName>
, the male ninth sternum possesses a deep longitudinal incision at middle and the lateral lobes of the male tenth sternum are subuliform.
</paragraph>
</subSubSection>
</treatment>
</document>
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