248 lines
58 KiB
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248 lines
58 KiB
XML
<document id="41D87DCB099F5431BB0D1DB4685CF35F" ID-DOI="10.11646/zootaxa.3577.1.2" ID-GBIF-Dataset="4e469dc8-3ed0-4b28-8879-85e3cca36766" ID-ISSN="1175-5326" ID-Zenodo-Dep="213482" ID-ZooBank="4BFAE75F-7BC0-4D5D-B1D1-DA2CC01F40C9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460559501141" checkinUser="plazi" docAuthor="Fuhrmann, Juares" docDate="2012" docId="096F054CFF98547EFF25FE63FAB0EA50" docLanguage="en" docName="zt03577p057.pdf" docOrigin="Zootaxa 3577 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.3577.1.2" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Compsodactylus, new genus" docType="treatment" docVersion="12" lastPageNumber="45" masterDocId="F5567D34FF99547CFFB2FFB9FFE5E90C" masterDocTitle="Compsodactylus, a new South American genus with one new species and two new combinations (Coleoptera: Scarabaeidae: Melolonthinae)" masterLastPageNumber="57" masterPageNumber="43" pageNumber="44" updateTime="1698320037866" updateUser="plazi">
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<mods:title id="D06F70EB6CF6420F6E73BAB525E2A8E7">Compsodactylus, a new South American genus with one new species and two new combinations (Coleoptera: Scarabaeidae: Melolonthinae)</mods:title>
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<mods:name id="63670194128A8D1C29556E32B7B394D0" type="personal">
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<mods:namePart id="CDDB0C6E71C393867C10470DE1CE93E5">Fuhrmann, Juares</mods:namePart>
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<mods:title id="44A18CE522111C51E2D0009E6C8B90DB">Zootaxa</mods:title>
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<mods:part id="917F601CE1A74A3382CB9F47DD0BB554">
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<mods:date id="306FC13ED189B22C410F7B9C356C420F">2012</mods:date>
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<mods:number id="9A84DF24B525DA311DC383122E286BDA">2012-12-10</mods:number>
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<mods:number id="B3F5D345DCF339A945FEB5C0EB4FDFC4">3577</mods:number>
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<mods:classification id="752E0090CF2C9DB699DAA649A546C6C9">journal article</mods:classification>
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<mods:identifier id="7091150FC4A86EAD1BB196961B920A5E" type="DOI">10.11646/zootaxa.3577.1.2</mods:identifier>
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<treatment id="096F054CFF98547EFF25FE63FAB0EA50" ID-DOI="http://doi.org/10.5281/zenodo.5874045" ID-GBIF-Taxon="119556605" ID-Zenodo-Dep="5874045" LSID="urn:lsid:plazi:treatment:096F054CFF98547EFF25FE63FAB0EA50" httpUri="http://treatment.plazi.org/id/096F054CFF98547EFF25FE63FAB0EA50" lastPageId="2" lastPageNumber="45" pageId="1" pageNumber="44">
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<subSubSection id="C9DCE7D1FF98547DFF25FE63FE00E8F8" box="[151,485,474,500]" pageId="1" pageNumber="44" type="nomenclature">
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<paragraph id="8179B45AFF98547DFF25FE63FE00E8F8" blockId="1.[151,485,474,533]" box="[151,485,474,500]" pageId="1" pageNumber="44">
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<heading id="DA310336FF98547DFF25FE63FE00E8F8" bold="true" box="[151,485,474,500]" fontSize="11" level="1" pageId="1" pageNumber="44" reason="1">
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<emphasis id="B3B26848FF98547DFF25FE63FE00E8F8" bold="true" box="[151,485,474,500]" pageId="1" pageNumber="44">
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<taxonomicName id="46C6CFD9FF98547DFF25FE63FEB9E8F8" authority="Fuhrmann 2012" box="[151,348,474,500]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="44" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B3B26848FF98547DFF25FE63FEB9E8F8" bold="true" box="[151,348,474,500]" italics="true" pageId="1" pageNumber="44">Compsodactylus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A881D533FF98547DFED1FE63FE00E8F8" box="[355,485,474,500]" pageId="1" pageNumber="44" rank="genus">new genus</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C9DCE7D1FF98547DFF25FE44FEFAEB19" box="[151,287,509,533]" pageId="1" pageNumber="44" type="description">
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<paragraph id="8179B45AFF98547DFF25FE44FEFAEB19" blockId="1.[151,485,474,533]" box="[151,287,509,533]" pageId="1" pageNumber="44">
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(
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<figureCitation id="19FDA8DFFF98547DFF2DFE44FEF2EB19" box="[159,279,509,533]" captionStart-0="FIGURES 1 – 9. 1, 2, 4" captionStart-1="FIGURES 10 – 16. 10" captionStart-2="FIGURES 17 – 27. 17 – 26" captionStart-3="FIGURES 28 – 39. 28 – 35, 37" captionStart-4="FIGURES 40 – 45" captionStart-5="FIGURES 46 – 51" captionStart-6="FIGURES 52 – 62. 52 – 60" captionStart-7="FIGURES 63 – 75. 63 – 69" captionStart-8="FIGURE 76" captionStartId-0="5.[151,264,1908,1931]" captionStartId-1="6.[151,264,1904,1927]" captionStartId-2="7.[151,264,1904,1927]" captionStartId-3="8.[151,264,1904,1927]" captionStartId-4="9.[151,264,1707,1730]" captionStartId-5="10.[151,264,1865,1888]" captionStartId-6="11.[151,264,1811,1834]" captionStartId-7="12.[151,264,1813,1836]" captionStartId-8="13.[151,250,1462,1485]" captionTargetBox-0="[157,1432,204,1879]" captionTargetBox-1="[153,1434,203,1882]" captionTargetBox-2="[154,1429,202,1882]" captionTargetBox-3="[157,1420,196,1882]" captionTargetBox-4="[222,1342,193,1665]" captionTargetBox-5="[202,1405,201,1824]" captionTargetBox-6="[191,1382,200,1761]" captionTargetBox-7="[188,1396,202,1785]" captionTargetBox-8="[153,1435,193,1440]" captionTargetId-0="figure@5.[151,1436,195,1884]" captionTargetId-1="figure@6.[151,1436,193,1883]" captionTargetId-2="figure@7.[151,1436,193,1883]" captionTargetId-3="figure@8.[151,1436,193,1883]" captionTargetId-4="figure@9.[222,1343,193,1667]" captionTargetId-5="figure@10.[166,1410,193,1828]" captionTargetId-6="figure@11.[188,1382,193,1762]" captionTargetId-7="figure@12.[186,1399,193,1787]" captionTargetId-8="figure@13.[151,1436,193,1441]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionTargetPageId-4="9" captionTargetPageId-5="10" captionTargetPageId-6="11" captionTargetPageId-7="12" captionTargetPageId-8="13" captionText-0="FIGURES 1 – 9. 1, 2, 4, Compsodactylus martinezi male habitus (dorsal, lateral, head dorsolateral); 3, C. martinezi female habitus (lateral; arrow shows the posterior margin bordered); 5, 6, C. parvulus holotype female habitus (dorsal, lateral); 7, 8, C. scabrosus holotype male habitus (dorsal, lateral); 9, C. scabrosus female habitus (lateral)." captionText-1="FIGURES 10 – 16. 10, Compsodactylus scabrosus head (dorsal); 11, C. parvulus head (dorsal); 12 – 16, C. martinezi; 12, head (dorsal); 13, epipharynx with details; 14, antenna; 15, hypopharynx; 16, labium." captionText-2="FIGURES 17 – 27. 17 – 26, Compsodactylus martinezi; 17 – 19, left mandible (dorsal, ventral, molar); 20 – 22, right mandible (dorsal, ventral, molar); 23 – 24 maxilla (dorsal, ventral); 25 – 26 galea (ventral, internal); 27, C. scabrosus galea (internal)." captionText-3="FIGURES 28 – 39. 28 – 35, 37, Compsodactylus martinezi; 28 – 30, prothorax (frontal, posterior, ventral); 31, detail of hypomeron posterior arm; 32, male protarsomere I (internal); 33, male protibia and protarsus; 34, male mesotibia and mesotarsus; 35, male metafemur, metatibia, and metatarsus; 36, C. parvulus female metaleg; 37, C. martinezi male tibial apex; 38, C. parvulus female tibial apex; 39, C. scabrosus male tibial apex." captionText-4="FIGURES 40 – 45. Compsodactylus martinezi pterothorax; 40, anterior, right without mesoalinotum, limit of meso-metathorax in blue, limit of mesoalinotum-postnotum in green; 41, posterior, right without mesoscutelum; 42, metanotum internal, left with mesopostnotum complex; 43, internodorsal, left without metepisternum-epimeron and associated region; 44, lateral; 45, internolateral. afm, anterior flange of metendosternite; alc, alacrista; alm, anterior lobe of metanotum; apn, postnotal apodeme; awp, anterior notal wing process; ba 2 – 3, meso-metabasalare; dm 2 – 3, dorsal mese-metepimeral arm; em 2 – 3, mesemetepimeron; es 2 – 3, mese-metepisternum; fu 2 – 3, meso-metafurca; its, intersegmentar sclerite; klm, metakatepisternal loop; ls 2 – 3, lateral scutellar arm; mfm, medial flange of metendosternite; msb, metendosternite bridge; mwp, medial notal wing process; pe 2 – 3, meso-metaprealare; ph 1 – 3, first, second, and third phragma; pl 2 – 3, lateral of meso-metapostnotum; pla, pleural arm; pn 2 – 3, meso-metapostnotum; po 2 – 3, meso-metapostalare; pwp, posterior notal wing process; sc 2 – 3, mesometascutum; su 3, metasubalare; st 2 – 3, meso-metascutellum; vt 2 – 3, meso-metaventrite; wp 2 – 3, pleural wing process of mesothorax and metathorax." captionText-5="FIGURES 46 – 51. Compsodactylus martinezi; 46, pterothorax dorsal, right without mesoalinotum, limit of meso-metathorax in blue; 47, mesepisternum-epimeron and associated (internal); 48, mesoalinotum internal, internoexternal limit in red; 49 – 51, abdomen male (lateral, ventral), female (ventral). aap, anterior abdominal process; im 7 – 8, intersegmentar membrane VII / VIII; lpf, lateral phragma flange; pcp, prealare contact point; ppg, propygidium; pyg, pygidium; smk, scutellar medial keel; us 2 – 8, urosternite II – VIII; ut 1 – 8, urotergite I – VIII; v 1 – 6, ventrite I – VI. Other abbreviations on the last plate." captionText-6="FIGURES 52 – 62. 52 – 60, Compsodactylus martinezi; 52 – 53, wing (distended, folded); 54, wing articulation and base; 55 – 56, first axillary (dorsal, ventral); 57, head of first axillary (anterior); 58 – 59, second axillary (dorsal, ventral); 60, third axillary (dorsal); 61, C. scabrosus wing distended; 62, C. martinezi posterior view of elytral medial contact section. Ax 1, 2, 3, first, second, third axillary; HP, humeral plate; med, medial plate (with two pieces). Wing base and venal: prefix B, basivenalia; suffix A, anterior; suffix P, posterior; b, BMP-BCu brace; PC, precosta; C, costa; R, radius; M, media; Cu, cubitus; A anal; J, jugal. Wing fold (red): af, anal fold; jf, jugal fold; lf, longitudinal fold; mf, medial fold; tf, transversal fold. Other abbreviations on the last plate." captionText-7="FIGURES 63 – 75. 63 – 69, Compsodactylus martinezi; 63, male genital ring; 64, aedeagus (lateral); 65, parameres (dorsal); 66, parameres (ventral); 67 – 69, endophallus (lateral), ventral basis detail, ornamentation detail; 70 – 71, C. scabrosus; 70, aedeagus (lateral); 71, parameres (dorsal); 72 – 73, C. martinezi; 72, female terminalia and internal associated region (ventral); 73, female external genitalia; 74, C. parvulus female external genitalia; 75, C. scabrosus female external genitalia. acg, accessory gland; bcx, bursa copulatrix; dgx, gonocoxite distal piece; gst, gonostylus; ovd, oviduct; pgx, gonocoxite proximal piece; ppt, paraproct; sga, spiculum gastrale caudal piece; sgc, spiculum gastrale caudal piece; spt, spermatecae; stg, spermatecae gland; ut 9, urosternite; tem, temones." captionText-8="FIGURE 76. Compsodactylus species geographic distribution." httpUri-0="https://zenodo.org/record/213483/files/figure.png" httpUri-1="https://zenodo.org/record/213484/files/figure.png" httpUri-2="https://zenodo.org/record/213485/files/figure.png" httpUri-3="https://zenodo.org/record/213486/files/figure.png" httpUri-4="https://zenodo.org/record/213487/files/figure.png" httpUri-5="https://zenodo.org/record/213488/files/figure.png" httpUri-6="https://zenodo.org/record/213489/files/figure.png" httpUri-7="https://zenodo.org/record/213490/files/figure.png" httpUri-8="https://zenodo.org/record/213492/files/figure.png" pageId="1" pageNumber="44">Figs. 1–76</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C9DCE7D1FF98547DFF25FDFCFCB7EB51" box="[151,850,581,606]" pageId="1" pageNumber="44" type="type_taxon">
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<paragraph id="8179B45AFF98547DFF25FDFCFCB7EB51" blockId="1.[151,1437,581,2013]" box="[151,850,581,606]" pageId="1" pageNumber="44">
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<emphasis id="B3B26848FF98547DFF25FDFCFECEEB52" bold="true" box="[151,299,581,606]" pageId="1" pageNumber="44">
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<typeStatus id="5E7D0AF8FF98547DFF25FDFCFF37EB51" box="[151,210,581,605]" pageId="1" pageNumber="44">Type</typeStatus>
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species
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</emphasis>
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.
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<taxonomicName id="46C6CFD9FF98547DFE8BFDFFFD75EB51" authority="Frey, 1972" authorityName="Frey" authorityYear="1972" box="[313,656,581,606]" class="Insecta" family="Melolonthidae" genus="Dicrania" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="44" phylum="Arthropoda" rank="species" species="martinezi">
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<emphasis id="B3B26848FF98547DFE8BFDFFFDEAEB51" box="[313,527,582,605]" italics="true" pageId="1" pageNumber="44">Dicrania martinezi</emphasis>
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<bibRefCitation id="E557C9ABFF98547DFDA4FDFFFD75EB51" author="Frey" box="[534,656,581,606]" firstAuthor="Frey" pageId="1" pageNumber="44" pagination="67 - 85" refId="ref6164" refString="Frey, G. (1972) Bestimmungstabelle der Gattung Dicrania Serv. (Coleoptera - Melolonthinae - Macrodactylini). Entomologische Arbeiten aus dem Museum G. Frey, 23, 67 - 85." type="journal article" year="1972">Frey, 1972</bibRefCitation>
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</taxonomicName>
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, here designated.
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</paragraph>
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</subSubSection>
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<subSubSection id="C9DCE7D1FF98547DFF75FDD3FD99EBA9" pageId="1" pageNumber="44" type="etymology">
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<paragraph id="8179B45AFF98547DFF75FDD3FD99EBA9" blockId="1.[151,1437,581,2013]" pageId="1" pageNumber="44">
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<emphasis id="B3B26848FF98547DFF75FDD3FEA3EB8E" bold="true" box="[199,326,618,642]" pageId="1" pageNumber="44">Etymology</emphasis>
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.
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<emphasis id="B3B26848FF98547DFEEEFDD3FE27EB8D" box="[348,450,618,641]" italics="true" pageId="1" pageNumber="44">Kompsos</emphasis>
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(Greek), elegant;
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<emphasis id="B3B26848FF98547DFD14FDD0FCE6EB8D" box="[678,771,617,641]" italics="true" pageId="1" pageNumber="44">daktylos</emphasis>
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(Greek), finger; in reference to distinct form of male metatibia; the name is masculine in gender.
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</paragraph>
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</subSubSection>
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<subSubSection id="C9DCE7D1FF98547DFF75FD08FED3EA39" pageId="1" pageNumber="44" type="diagnosis">
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<paragraph id="8179B45AFF98547DFF75FD08FED3EA39" blockId="1.[151,1437,581,2013]" pageId="1" pageNumber="44">
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<emphasis id="B3B26848FF98547DFF75FD08FEDFEBC6" bold="true" box="[199,314,689,714]" pageId="1" pageNumber="44">Diagnosis</emphasis>
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. Mentum longer than wide; posterior area of frons and anterior area of pronotum with denticle-like setae; protibiae with 2 teeth, spur developed; external and internal margin of elytra bordered; matecoxae with short projection over trochanter basis; male metatibiae with internoapical spine, lacking spurs; abdominal spiracular area VII narrowed.
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</paragraph>
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</subSubSection>
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<subSubSection id="C9DCE7D1FF98547DFF75FCF8FAD5EE98" pageId="1" pageNumber="44" type="description">
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<paragraph id="8179B45AFF98547DFF75FCF8FCC8ED75" blockId="1.[151,1437,581,2013]" pageId="1" pageNumber="44">
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<emphasis id="B3B26848FF98547DFF75FCF8FEB4EA56" bold="true" box="[199,337,833,858]" pageId="1" pageNumber="44">Description</emphasis>
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. Head (
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<figureCitation id="19FDA8DFFF98547DFE1FFCF8FE18EA55" box="[429,509,833,857]" captionStart="FIGURES 1 – 9. 1, 2, 4" captionStartId="5.[151,264,1908,1931]" captionTargetBox="[157,1432,204,1879]" captionTargetId="figure@5.[151,1436,195,1884]" captionTargetPageId="5" captionText="FIGURES 1 – 9. 1, 2, 4, Compsodactylus martinezi male habitus (dorsal, lateral, head dorsolateral); 3, C. martinezi female habitus (lateral; arrow shows the posterior margin bordered); 5, 6, C. parvulus holotype female habitus (dorsal, lateral); 7, 8, C. scabrosus holotype male habitus (dorsal, lateral); 9, C. scabrosus female habitus (lateral)." httpUri="https://zenodo.org/record/213483/files/figure.png" pageId="1" pageNumber="44">Figs. 4</figureCitation>
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,
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<figureCitation id="19FDA8DFFF98547DFDB9FCF8FDB0EA55" box="[523,597,833,857]" captionStart="FIGURES 10 – 16. 10" captionStartId="6.[151,264,1904,1927]" captionTargetBox="[153,1434,203,1882]" captionTargetId="figure@6.[151,1436,193,1883]" captionTargetPageId="6" captionText="FIGURES 10 – 16. 10, Compsodactylus scabrosus head (dorsal); 11, C. parvulus head (dorsal); 12 – 16, C. martinezi; 12, head (dorsal); 13, epipharynx with details; 14, antenna; 15, hypopharynx; 16, labium." httpUri="https://zenodo.org/record/213484/files/figure.png" pageId="1" pageNumber="44">10–12</figureCitation>
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). Epistomal suture poorly defined. Front with denticle-like setae. Clypeus trapezoid; anterior angles acute, upturned. Labrum thin, slightly emarginated. Epipharynx (
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<figureCitation id="19FDA8DFFF98547DFB52FCDCFADDEA71" box="[1248,1336,869,893]" captionStart="FIGURES 10 – 16. 10" captionStartId="6.[151,264,1904,1927]" captionTargetBox="[153,1434,203,1882]" captionTargetId="figure@6.[151,1436,193,1883]" captionTargetPageId="6" captionText="FIGURES 10 – 16. 10, Compsodactylus scabrosus head (dorsal); 11, C. parvulus head (dorsal); 12 – 16, C. martinezi; 12, head (dorsal); 13, epipharynx with details; 14, antenna; 15, hypopharynx; 16, labium." httpUri="https://zenodo.org/record/213484/files/figure.png" pageId="1" pageNumber="44">Fig. 13</figureCitation>
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) setose; posterior area covered by seta-like processes (
|
||
<figureCitation id="19FDA8DFFF98547DFD72FC30FCF3EAAD" box="[704,790,905,929]" captionStart="FIGURES 10 – 16. 10" captionStartId="6.[151,264,1904,1927]" captionTargetBox="[153,1434,203,1882]" captionTargetId="figure@6.[151,1436,193,1883]" captionTargetPageId="6" captionText="FIGURES 10 – 16. 10, Compsodactylus scabrosus head (dorsal); 11, C. parvulus head (dorsal); 12 – 16, C. martinezi; 12, head (dorsal); 13, epipharynx with details; 14, antenna; 15, hypopharynx; 16, labium." httpUri="https://zenodo.org/record/213484/files/figure.png" pageId="1" pageNumber="44">Fig. 13</figureCitation>
|
||
details); medial and posteromedial area with sensilla. Mandible (
|
||
<figureCitation id="19FDA8DFFF98547DFEA6FC14FE7AEAC9" box="[276,415,941,965]" captionStart="FIGURES 17 – 27. 17 – 26" captionStartId="7.[151,264,1904,1927]" captionTargetBox="[154,1429,202,1882]" captionTargetId="figure@7.[151,1436,193,1883]" captionTargetPageId="7" captionText="FIGURES 17 – 27. 17 – 26, Compsodactylus martinezi; 17 – 19, left mandible (dorsal, ventral, molar); 20 – 22, right mandible (dorsal, ventral, molar); 23 – 24 maxilla (dorsal, ventral); 25 – 26 galea (ventral, internal); 27, C. scabrosus galea (internal)." httpUri="https://zenodo.org/record/213485/files/figure.png" pageId="1" pageNumber="44">Figs. 17–22</figureCitation>
|
||
) with prosthecae greatly developed; incisor rounded; molar (
|
||
<figureCitation id="19FDA8DFFF98547DFBD7FC14FB14EAC9" box="[1125,1265,941,965]" captionStart="FIGURES 17 – 27. 17 – 26" captionStartId="7.[151,264,1904,1927]" captionTargetBox="[154,1429,202,1882]" captionTargetId="figure@7.[151,1436,193,1883]" captionTargetPageId="7" captionText="FIGURES 17 – 27. 17 – 26, Compsodactylus martinezi; 17 – 19, left mandible (dorsal, ventral, molar); 20 – 22, right mandible (dorsal, ventral, molar); 23 – 24 maxilla (dorsal, ventral); 25 – 26 galea (ventral, internal); 27, C. scabrosus galea (internal)." httpUri="https://zenodo.org/record/213485/files/figure.png" pageId="1" pageNumber="44">Figs. 19, 22</figureCitation>
|
||
) striatoserrate, with posterior depression. Maxilla (
|
||
<figureCitation id="19FDA8DFFF98547DFD96FC68FD4FEAE5" box="[548,682,977,1001]" captionStart="FIGURES 17 – 27. 17 – 26" captionStartId="7.[151,264,1904,1927]" captionTargetBox="[154,1429,202,1882]" captionTargetId="figure@7.[151,1436,193,1883]" captionTargetPageId="7" captionText="FIGURES 17 – 27. 17 – 26, Compsodactylus martinezi; 17 – 19, left mandible (dorsal, ventral, molar); 20 – 22, right mandible (dorsal, ventral, molar); 23 – 24 maxilla (dorsal, ventral); 25 – 26 galea (ventral, internal); 27, C. scabrosus galea (internal)." httpUri="https://zenodo.org/record/213485/files/figure.png" pageId="1" pageNumber="44">Figs. 23, 24</figureCitation>
|
||
) with lacinia toothed, galea (
|
||
<figureCitation id="19FDA8DFFF98547DFC5FFC68FB96EAE5" box="[1005,1139,977,1001]" captionStart="FIGURES 17 – 27. 17 – 26" captionStartId="7.[151,264,1904,1927]" captionTargetBox="[154,1429,202,1882]" captionTargetId="figure@7.[151,1436,193,1883]" captionTargetPageId="7" captionText="FIGURES 17 – 27. 17 – 26, Compsodactylus martinezi; 17 – 19, left mandible (dorsal, ventral, molar); 20 – 22, right mandible (dorsal, ventral, molar); 23 – 24 maxilla (dorsal, ventral); 25 – 26 galea (ventral, internal); 27, C. scabrosus galea (internal)." httpUri="https://zenodo.org/record/213485/files/figure.png" pageId="1" pageNumber="44">Figs. 25–27</figureCitation>
|
||
) 7–8-toothed; palpi with 4 palpomeres, IV dorsally striate. Hypopharynx (
|
||
<figureCitation id="19FDA8DFFF98547DFD67FC4CFCCAED01" box="[725,815,1013,1037]" captionStart="FIGURES 10 – 16. 10" captionStartId="6.[151,264,1904,1927]" captionTargetBox="[153,1434,203,1882]" captionTargetId="figure@6.[151,1436,193,1883]" captionTargetPageId="6" captionText="FIGURES 10 – 16. 10, Compsodactylus scabrosus head (dorsal); 11, C. parvulus head (dorsal); 12 – 16, C. martinezi; 12, head (dorsal); 13, epipharynx with details; 14, antenna; 15, hypopharynx; 16, labium." httpUri="https://zenodo.org/record/213484/files/figure.png" pageId="1" pageNumber="44">Fig. 15</figureCitation>
|
||
) laterally setose posteriorly covered with seta-like processes, medial area with sensilla. Labium (
|
||
<figureCitation id="19FDA8DFFF98547DFD1BFBA0FCD6ED3D" box="[681,819,1049,1073]" captionStart="FIGURES 10 – 16. 10" captionStartId="6.[151,264,1904,1927]" captionTargetBox="[153,1434,203,1882]" captionTargetId="figure@6.[151,1436,193,1883]" captionTargetPageId="6" captionText="FIGURES 10 – 16. 10, Compsodactylus scabrosus head (dorsal); 11, C. parvulus head (dorsal); 12 – 16, C. martinezi; 12, head (dorsal); 13, epipharynx with details; 14, antenna; 15, hypopharynx; 16, labium." httpUri="https://zenodo.org/record/213484/files/figure.png" pageId="1" pageNumber="44">Figs. 15, 16</figureCitation>
|
||
) with prementum prominent and fused with mentum; mentum longer than wide, with longitudinal groove, palpi with 3 palpomeres. Antenna with 9 antennomeres (
|
||
<figureCitation id="19FDA8DFFF98547DFADDFB84FF53ED75" captionStart="FIGURES 10 – 16. 10" captionStartId="6.[151,264,1904,1927]" captionTargetBox="[153,1434,203,1882]" captionTargetId="figure@6.[151,1436,193,1883]" captionTargetPageId="6" captionText="FIGURES 10 – 16. 10, Compsodactylus scabrosus head (dorsal); 11, C. parvulus head (dorsal); 12 – 16, C. martinezi; 12, head (dorsal); 13, epipharynx with details; 14, antenna; 15, hypopharynx; 16, labium." httpUri="https://zenodo.org/record/213484/files/figure.png" pageId="1" pageNumber="44">Fig. 14</figureCitation>
|
||
); club as long as or slightly shorter than pedicel-funicle.
|
||
</paragraph>
|
||
<paragraph id="8179B45AFF98547DFF75FB3FFEC1EC21" blockId="1.[151,1437,581,2013]" pageId="1" pageNumber="44">
|
||
Pronotum (
|
||
<figureCitation id="19FDA8DFFF98547DFEF5FB3CFE34ED91" box="[327,465,1157,1181]" captionStart="FIGURES 28 – 39. 28 – 35, 37" captionStartId="8.[151,264,1904,1927]" captionTargetBox="[157,1420,196,1882]" captionTargetId="figure@8.[151,1436,193,1883]" captionTargetPageId="8" captionText="FIGURES 28 – 39. 28 – 35, 37, Compsodactylus martinezi; 28 – 30, prothorax (frontal, posterior, ventral); 31, detail of hypomeron posterior arm; 32, male protarsomere I (internal); 33, male protibia and protarsus; 34, male mesotibia and mesotarsus; 35, male metafemur, metatibia, and metatarsus; 36, C. parvulus female metaleg; 37, C. martinezi male tibial apex; 38, C. parvulus female tibial apex; 39, C. scabrosus male tibial apex." httpUri="https://zenodo.org/record/213486/files/figure.png" pageId="1" pageNumber="44">Figs. 28–30</figureCitation>
|
||
) wider than long, maximum width at middle; all margins bordered; anterior margin with external membrane. Prosternal posterior process small, tubercle-like. Furca broad, truncate. Proleg (
|
||
<figureCitation id="19FDA8DFFF98547DFA88FB10FA6BEDCD" box="[1338,1422,1193,1217]" captionStart="FIGURES 28 – 39. 28 – 35, 37" captionStartId="8.[151,264,1904,1927]" captionTargetBox="[157,1420,196,1882]" captionTargetId="figure@8.[151,1436,193,1883]" captionTargetPageId="8" captionText="FIGURES 28 – 39. 28 – 35, 37, Compsodactylus martinezi; 28 – 30, prothorax (frontal, posterior, ventral); 31, detail of hypomeron posterior arm; 32, male protarsomere I (internal); 33, male protibia and protarsus; 34, male mesotibia and mesotarsus; 35, male metafemur, metatibia, and metatarsus; 36, C. parvulus female metaleg; 37, C. martinezi male tibial apex; 38, C. parvulus female tibial apex; 39, C. scabrosus male tibial apex." httpUri="https://zenodo.org/record/213486/files/figure.png" pageId="1" pageNumber="44">Fig. 33</figureCitation>
|
||
). Tibia with 2 teeth; spur present at middle of tibial length. Tarsus as long as or slightly shorter than tibia; male tarsomere I (
|
||
<figureCitation id="19FDA8DFFF98547DFE95FB48FE9CEC05" box="[295,377,1265,1289]" captionStart="FIGURES 28 – 39. 28 – 35, 37" captionStartId="8.[151,264,1904,1927]" captionTargetBox="[157,1420,196,1882]" captionTargetId="figure@8.[151,1436,193,1883]" captionTargetPageId="8" captionText="FIGURES 28 – 39. 28 – 35, 37, Compsodactylus martinezi; 28 – 30, prothorax (frontal, posterior, ventral); 31, detail of hypomeron posterior arm; 32, male protarsomere I (internal); 33, male protibia and protarsus; 34, male mesotibia and mesotarsus; 35, male metafemur, metatibia, and metatarsus; 36, C. parvulus female metaleg; 37, C. martinezi male tibial apex; 38, C. parvulus female tibial apex; 39, C. scabrosus male tibial apex." httpUri="https://zenodo.org/record/213486/files/figure.png" pageId="1" pageNumber="44">Fig. 32</figureCitation>
|
||
) with internoapical area granulostriate. Pretarsus with 2 similar bifid claws; empodium short, with 2 setae.
|
||
</paragraph>
|
||
<paragraph id="8179B45AFF98547DFF75FA80FB48ECED" blockId="1.[151,1437,581,2013]" pageId="1" pageNumber="44">
|
||
Mesothorax (
|
||
<figureCitation id="19FDA8DFFF98547DFED3FA80FE0AEC5D" box="[353,495,1337,1361]" captionStart-0="FIGURES 40 – 45" captionStart-1="FIGURES 46 – 51" captionStartId-0="9.[151,264,1707,1730]" captionStartId-1="10.[151,264,1865,1888]" captionTargetBox-0="[222,1342,193,1665]" captionTargetBox-1="[202,1405,201,1824]" captionTargetId-0="figure@9.[222,1343,193,1667]" captionTargetId-1="figure@10.[166,1410,193,1828]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="FIGURES 40 – 45. Compsodactylus martinezi pterothorax; 40, anterior, right without mesoalinotum, limit of meso-metathorax in blue, limit of mesoalinotum-postnotum in green; 41, posterior, right without mesoscutelum; 42, metanotum internal, left with mesopostnotum complex; 43, internodorsal, left without metepisternum-epimeron and associated region; 44, lateral; 45, internolateral. afm, anterior flange of metendosternite; alc, alacrista; alm, anterior lobe of metanotum; apn, postnotal apodeme; awp, anterior notal wing process; ba 2 – 3, meso-metabasalare; dm 2 – 3, dorsal mese-metepimeral arm; em 2 – 3, mesemetepimeron; es 2 – 3, mese-metepisternum; fu 2 – 3, meso-metafurca; its, intersegmentar sclerite; klm, metakatepisternal loop; ls 2 – 3, lateral scutellar arm; mfm, medial flange of metendosternite; msb, metendosternite bridge; mwp, medial notal wing process; pe 2 – 3, meso-metaprealare; ph 1 – 3, first, second, and third phragma; pl 2 – 3, lateral of meso-metapostnotum; pla, pleural arm; pn 2 – 3, meso-metapostnotum; po 2 – 3, meso-metapostalare; pwp, posterior notal wing process; sc 2 – 3, mesometascutum; su 3, metasubalare; st 2 – 3, meso-metascutellum; vt 2 – 3, meso-metaventrite; wp 2 – 3, pleural wing process of mesothorax and metathorax." captionText-1="FIGURES 46 – 51. Compsodactylus martinezi; 46, pterothorax dorsal, right without mesoalinotum, limit of meso-metathorax in blue; 47, mesepisternum-epimeron and associated (internal); 48, mesoalinotum internal, internoexternal limit in red; 49 – 51, abdomen male (lateral, ventral), female (ventral). aap, anterior abdominal process; im 7 – 8, intersegmentar membrane VII / VIII; lpf, lateral phragma flange; pcp, prealare contact point; ppg, propygidium; pyg, pygidium; smk, scutellar medial keel; us 2 – 8, urosternite II – VIII; ut 1 – 8, urotergite I – VIII; v 1 – 6, ventrite I – VI. Other abbreviations on the last plate." httpUri-0="https://zenodo.org/record/213487/files/figure.png" httpUri-1="https://zenodo.org/record/213488/files/figure.png" pageId="1" pageNumber="44">Figs. 40–48</figureCitation>
|
||
). Scutellum with anterior margin rounded, not bordered. Episternum as wide as epimeron or near so, not reaching the mesocoxal cavity. Elytra (
|
||
<figureCitation id="19FDA8DFFF98547DFC13FAE4FC1DEC79" box="[929,1016,1373,1397]" captionStart="FIGURES 52 – 62. 52 – 60" captionStartId="11.[151,264,1811,1834]" captionTargetBox="[191,1382,200,1761]" captionTargetId="figure@11.[188,1382,193,1762]" captionTargetPageId="11" captionText="FIGURES 52 – 62. 52 – 60, Compsodactylus martinezi; 52 – 53, wing (distended, folded); 54, wing articulation and base; 55 – 56, first axillary (dorsal, ventral); 57, head of first axillary (anterior); 58 – 59, second axillary (dorsal, ventral); 60, third axillary (dorsal); 61, C. scabrosus wing distended; 62, C. martinezi posterior view of elytral medial contact section. Ax 1, 2, 3, first, second, third axillary; HP, humeral plate; med, medial plate (with two pieces). Wing base and venal: prefix B, basivenalia; suffix A, anterior; suffix P, posterior; b, BMP-BCu brace; PC, precosta; C, costa; R, radius; M, media; Cu, cubitus; A anal; J, jugal. Wing fold (red): af, anal fold; jf, jugal fold; lf, longitudinal fold; mf, medial fold; tf, transversal fold. Other abbreviations on the last plate." httpUri="https://zenodo.org/record/213489/files/figure.png" pageId="1" pageNumber="44">Fig. 58</figureCitation>
|
||
) about 1.2 times longer than wide (combined), striatopunctate; anterior margin abruptly rounded, not bordered; external and internal margins bordered; lateral margin without external membrane. Mesoleg (
|
||
<figureCitation id="19FDA8DFFF98547DFCDCFA1CFC20ECB1" box="[878,965,1445,1469]" captionStart="FIGURES 28 – 39. 28 – 35, 37" captionStartId="8.[151,264,1904,1927]" captionTargetBox="[157,1420,196,1882]" captionTargetId="figure@8.[151,1436,193,1883]" captionTargetPageId="8" captionText="FIGURES 28 – 39. 28 – 35, 37, Compsodactylus martinezi; 28 – 30, prothorax (frontal, posterior, ventral); 31, detail of hypomeron posterior arm; 32, male protarsomere I (internal); 33, male protibia and protarsus; 34, male mesotibia and mesotarsus; 35, male metafemur, metatibia, and metatarsus; 36, C. parvulus female metaleg; 37, C. martinezi male tibial apex; 38, C. parvulus female tibial apex; 39, C. scabrosus male tibial apex." httpUri="https://zenodo.org/record/213486/files/figure.png" pageId="1" pageNumber="44">Fig. 34</figureCitation>
|
||
). Femur with internal area longitudinally carinate. Tibia conical, transversal carina with denticle-like setae; 2 continuous spurs present.
|
||
</paragraph>
|
||
<paragraph id="8179B45AFF98547DFF75FA54FA7CEF94" blockId="1.[151,1437,581,2013]" pageId="1" pageNumber="44">
|
||
Metathorax (
|
||
<figureCitation id="19FDA8DFFF98547DFEE5FA54FE04EF09" box="[343,481,1517,1541]" captionStart-0="FIGURES 40 – 45" captionStart-1="FIGURES 46 – 51" captionStartId-0="9.[151,264,1707,1730]" captionStartId-1="10.[151,264,1865,1888]" captionTargetBox-0="[222,1342,193,1665]" captionTargetBox-1="[202,1405,201,1824]" captionTargetId-0="figure@9.[222,1343,193,1667]" captionTargetId-1="figure@10.[166,1410,193,1828]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="FIGURES 40 – 45. Compsodactylus martinezi pterothorax; 40, anterior, right without mesoalinotum, limit of meso-metathorax in blue, limit of mesoalinotum-postnotum in green; 41, posterior, right without mesoscutelum; 42, metanotum internal, left with mesopostnotum complex; 43, internodorsal, left without metepisternum-epimeron and associated region; 44, lateral; 45, internolateral. afm, anterior flange of metendosternite; alc, alacrista; alm, anterior lobe of metanotum; apn, postnotal apodeme; awp, anterior notal wing process; ba 2 – 3, meso-metabasalare; dm 2 – 3, dorsal mese-metepimeral arm; em 2 – 3, mesemetepimeron; es 2 – 3, mese-metepisternum; fu 2 – 3, meso-metafurca; its, intersegmentar sclerite; klm, metakatepisternal loop; ls 2 – 3, lateral scutellar arm; mfm, medial flange of metendosternite; msb, metendosternite bridge; mwp, medial notal wing process; pe 2 – 3, meso-metaprealare; ph 1 – 3, first, second, and third phragma; pl 2 – 3, lateral of meso-metapostnotum; pla, pleural arm; pn 2 – 3, meso-metapostnotum; po 2 – 3, meso-metapostalare; pwp, posterior notal wing process; sc 2 – 3, mesometascutum; su 3, metasubalare; st 2 – 3, meso-metascutellum; vt 2 – 3, meso-metaventrite; wp 2 – 3, pleural wing process of mesothorax and metathorax." captionText-1="FIGURES 46 – 51. Compsodactylus martinezi; 46, pterothorax dorsal, right without mesoalinotum, limit of meso-metathorax in blue; 47, mesepisternum-epimeron and associated (internal); 48, mesoalinotum internal, internoexternal limit in red; 49 – 51, abdomen male (lateral, ventral), female (ventral). aap, anterior abdominal process; im 7 – 8, intersegmentar membrane VII / VIII; lpf, lateral phragma flange; pcp, prealare contact point; ppg, propygidium; pyg, pygidium; smk, scutellar medial keel; us 2 – 8, urosternite II – VIII; ut 1 – 8, urotergite I – VIII; v 1 – 6, ventrite I – VI. Other abbreviations on the last plate." httpUri-0="https://zenodo.org/record/213487/files/figure.png" httpUri-1="https://zenodo.org/record/213488/files/figure.png" pageId="1" pageNumber="44">Figs. 40–48</figureCitation>
|
||
). Episternum not reaching the mesocoxal cavity, extended between mesepimeron and metepimeron. Metendosternite with anterior flange rounded. Wing (
|
||
<figureCitation id="19FDA8DFFF98547DFC29F9A8FBC6EF25" box="[923,1059,1553,1577]" captionStart="FIGURES 52 – 62. 52 – 60" captionStartId="11.[151,264,1811,1834]" captionTargetBox="[191,1382,200,1761]" captionTargetId="figure@11.[188,1382,193,1762]" captionTargetPageId="11" captionText="FIGURES 52 – 62. 52 – 60, Compsodactylus martinezi; 52 – 53, wing (distended, folded); 54, wing articulation and base; 55 – 56, first axillary (dorsal, ventral); 57, head of first axillary (anterior); 58 – 59, second axillary (dorsal, ventral); 60, third axillary (dorsal); 61, C. scabrosus wing distended; 62, C. martinezi posterior view of elytral medial contact section. Ax 1, 2, 3, first, second, third axillary; HP, humeral plate; med, medial plate (with two pieces). Wing base and venal: prefix B, basivenalia; suffix A, anterior; suffix P, posterior; b, BMP-BCu brace; PC, precosta; C, costa; R, radius; M, media; Cu, cubitus; A anal; J, jugal. Wing fold (red): af, anal fold; jf, jugal fold; lf, longitudinal fold; mf, medial fold; tf, transversal fold. Other abbreviations on the last plate." httpUri="https://zenodo.org/record/213489/files/figure.png" pageId="1" pageNumber="44">Figs. 52–61</figureCitation>
|
||
) with anterodistal margin setose, RA3 and RA4+
|
||
<collectingCountry id="F9D1F4CAFF98547DFE8BF98FFEB9EF41" box="[313,348,1590,1613]" name="Poland" pageId="1" pageNumber="44">RP</collectingCountry>
|
||
1 not reaching the apex, MP3 indistinct. Metaleg (
|
||
<figureCitation id="19FDA8DFFF98547DFC32F98CFBEFEF41" box="[896,1034,1589,1613]" captionStart="FIGURES 28 – 39. 28 – 35, 37" captionStartId="8.[151,264,1904,1927]" captionTargetBox="[157,1420,196,1882]" captionTargetId="figure@8.[151,1436,193,1883]" captionTargetPageId="8" captionText="FIGURES 28 – 39. 28 – 35, 37, Compsodactylus martinezi; 28 – 30, prothorax (frontal, posterior, ventral); 31, detail of hypomeron posterior arm; 32, male protarsomere I (internal); 33, male protibia and protarsus; 34, male mesotibia and mesotarsus; 35, male metafemur, metatibia, and metatarsus; 36, C. parvulus female metaleg; 37, C. martinezi male tibial apex; 38, C. parvulus female tibial apex; 39, C. scabrosus male tibial apex." httpUri="https://zenodo.org/record/213486/files/figure.png" pageId="1" pageNumber="44">Figs. 35–39</figureCitation>
|
||
). Coxa medially contiguous, area of ventral articulation with trochanter enlarged. Tibia with reduced transversal carina more developed in female; male tibial apex (
|
||
<figureCitation id="19FDA8DFFF98547DFEAEF939FE47EF94" box="[284,418,1664,1688]" captionStart="FIGURES 28 – 39. 28 – 35, 37" captionStartId="8.[151,264,1904,1927]" captionTargetBox="[157,1420,196,1882]" captionTargetId="figure@8.[151,1436,193,1883]" captionTargetPageId="8" captionText="FIGURES 28 – 39. 28 – 35, 37, Compsodactylus martinezi; 28 – 30, prothorax (frontal, posterior, ventral); 31, detail of hypomeron posterior arm; 32, male protarsomere I (internal); 33, male protibia and protarsus; 34, male mesotibia and mesotarsus; 35, male metafemur, metatibia, and metatarsus; 36, C. parvulus female metaleg; 37, C. martinezi male tibial apex; 38, C. parvulus female tibial apex; 39, C. scabrosus male tibial apex." httpUri="https://zenodo.org/record/213486/files/figure.png" pageId="1" pageNumber="44">Figs. 37, 39</figureCitation>
|
||
) with internal tooth, lacking spurs; female tibial (
|
||
<figureCitation id="19FDA8DFFF98547DFC7CF939FBFAEF94" box="[974,1055,1664,1688]" captionStart="FIGURES 28 – 39. 28 – 35, 37" captionStartId="8.[151,264,1904,1927]" captionTargetBox="[157,1420,196,1882]" captionTargetId="figure@8.[151,1436,193,1883]" captionTargetPageId="8" captionText="FIGURES 28 – 39. 28 – 35, 37, Compsodactylus martinezi; 28 – 30, prothorax (frontal, posterior, ventral); 31, detail of hypomeron posterior arm; 32, male protarsomere I (internal); 33, male protibia and protarsus; 34, male mesotibia and mesotarsus; 35, male metafemur, metatibia, and metatarsus; 36, C. parvulus female metaleg; 37, C. martinezi male tibial apex; 38, C. parvulus female tibial apex; 39, C. scabrosus male tibial apex." httpUri="https://zenodo.org/record/213486/files/figure.png" pageId="1" pageNumber="44">Fig. 38</figureCitation>
|
||
) apex with two contiguous spurs.
|
||
</paragraph>
|
||
<paragraph id="8179B45AFF98547DFF75F91DFAD5EE98" blockId="1.[151,1437,581,2013]" pageId="1" pageNumber="44">
|
||
Abdomen (
|
||
<figureCitation id="19FDA8DFFF98547DFEFDF91DFE06EFB0" box="[335,483,1700,1724]" captionStart="FIGURES 46 – 51" captionStartId="10.[151,264,1865,1888]" captionTargetBox="[202,1405,201,1824]" captionTargetId="figure@10.[166,1410,193,1828]" captionTargetPageId="10" captionText="FIGURES 46 – 51. Compsodactylus martinezi; 46, pterothorax dorsal, right without mesoalinotum, limit of meso-metathorax in blue; 47, mesepisternum-epimeron and associated (internal); 48, mesoalinotum internal, internoexternal limit in red; 49 – 51, abdomen male (lateral, ventral), female (ventral). aap, anterior abdominal process; im 7 – 8, intersegmentar membrane VII / VIII; lpf, lateral phragma flange; pcp, prealare contact point; ppg, propygidium; pyg, pygidium; smk, scutellar medial keel; us 2 – 8, urosternite II – VIII; ut 1 – 8, urotergite I – VIII; v 1 – 6, ventrite I – VI. Other abbreviations on the last plate." httpUri="https://zenodo.org/record/213488/files/figure.png" pageId="1" pageNumber="44">Figs. 49–51</figureCitation>
|
||
). Spiracular area VII (
|
||
<figureCitation id="19FDA8DFFF98547DFCB7F91DFC84EFB0" box="[773,865,1700,1724]" captionStart="FIGURES 46 – 51" captionStartId="10.[151,264,1865,1888]" captionTargetBox="[202,1405,201,1824]" captionTargetId="figure@10.[166,1410,193,1828]" captionTargetPageId="10" captionText="FIGURES 46 – 51. Compsodactylus martinezi; 46, pterothorax dorsal, right without mesoalinotum, limit of meso-metathorax in blue; 47, mesepisternum-epimeron and associated (internal); 48, mesoalinotum internal, internoexternal limit in red; 49 – 51, abdomen male (lateral, ventral), female (ventral). aap, anterior abdominal process; im 7 – 8, intersegmentar membrane VII / VIII; lpf, lateral phragma flange; pcp, prealare contact point; ppg, propygidium; pyg, pygidium; smk, scutellar medial keel; us 2 – 8, urosternite II – VIII; ut 1 – 8, urotergite I – VIII; v 1 – 6, ventrite I – VI. Other abbreviations on the last plate." httpUri="https://zenodo.org/record/213488/files/figure.png" pageId="1" pageNumber="44">Fig. 49</figureCitation>
|
||
) narrowed. Male pygidium longer than wide, downturned; female pygidium as long as wide or near so, backwardly projected. Ventrite V longer than ventrite IV. Intersegmentar membrane VII–VIII (between ventrite V and VI) exposed. Spiculum gastrale (
|
||
<figureCitation id="19FDA8DFFF98547DFB04F955FAE2EE08" box="[1206,1287,1772,1796]" captionStart="FIGURES 63 – 75. 63 – 69" captionStartId="12.[151,264,1813,1836]" captionTargetBox="[188,1396,202,1785]" captionTargetId="figure@12.[186,1399,193,1787]" captionTargetPageId="12" captionText="FIGURES 63 – 75. 63 – 69, Compsodactylus martinezi; 63, male genital ring; 64, aedeagus (lateral); 65, parameres (dorsal); 66, parameres (ventral); 67 – 69, endophallus (lateral), ventral basis detail, ornamentation detail; 70 – 71, C. scabrosus; 70, aedeagus (lateral); 71, parameres (dorsal); 72 – 73, C. martinezi; 72, female terminalia and internal associated region (ventral); 73, female external genitalia; 74, C. parvulus female external genitalia; 75, C. scabrosus female external genitalia. acg, accessory gland; bcx, bursa copulatrix; dgx, gonocoxite distal piece; gst, gonostylus; ovd, oviduct; pgx, gonocoxite proximal piece; ppt, paraproct; sga, spiculum gastrale caudal piece; sgc, spiculum gastrale caudal piece; spt, spermatecae; stg, spermatecae gland; ut 9, urosternite; tem, temones." httpUri="https://zenodo.org/record/213490/files/figure.png" pageId="1" pageNumber="44">Fig. 63</figureCitation>
|
||
) with cranial piece longer than wide and 0.66 times shorter than caudal arms. Sternite IX (
|
||
<figureCitation id="19FDA8DFFF98547DFC49F8A9FBA8EE24" box="[1019,1101,1808,1832]" captionStart="FIGURES 63 – 75. 63 – 69" captionStartId="12.[151,264,1813,1836]" captionTargetBox="[188,1396,202,1785]" captionTargetId="figure@12.[186,1399,193,1787]" captionTargetPageId="12" captionText="FIGURES 63 – 75. 63 – 69, Compsodactylus martinezi; 63, male genital ring; 64, aedeagus (lateral); 65, parameres (dorsal); 66, parameres (ventral); 67 – 69, endophallus (lateral), ventral basis detail, ornamentation detail; 70 – 71, C. scabrosus; 70, aedeagus (lateral); 71, parameres (dorsal); 72 – 73, C. martinezi; 72, female terminalia and internal associated region (ventral); 73, female external genitalia; 74, C. parvulus female external genitalia; 75, C. scabrosus female external genitalia. acg, accessory gland; bcx, bursa copulatrix; dgx, gonocoxite distal piece; gst, gonostylus; ovd, oviduct; pgx, gonocoxite proximal piece; ppt, paraproct; sga, spiculum gastrale caudal piece; sgc, spiculum gastrale caudal piece; spt, spermatecae; stg, spermatecae gland; ut 9, urosternite; tem, temones." httpUri="https://zenodo.org/record/213490/files/figure.png" pageId="1" pageNumber="44">Fig. 63</figureCitation>
|
||
) triangular, longer than wide, with apical row of short setae. Aedeagus with phallobasic apodeme longer than phallobase. Parameres longer than phallobase (with apodeme), symmetrical (except by the ventral side), glabrous; pieces free, longer than wide; apex acute and ventrally deflected; laterolongitudinal sulcus present. Intestine with 6 posterior caeca (
|
||
<figureCitation id="19FDA8DFFF98547DFB7DF8C5FAC1EE98" box="[1231,1316,1916,1940]" captionStart="FIGURES 63 – 75. 63 – 69" captionStartId="12.[151,264,1813,1836]" captionTargetBox="[188,1396,202,1785]" captionTargetId="figure@12.[186,1399,193,1787]" captionTargetPageId="12" captionText="FIGURES 63 – 75. 63 – 69, Compsodactylus martinezi; 63, male genital ring; 64, aedeagus (lateral); 65, parameres (dorsal); 66, parameres (ventral); 67 – 69, endophallus (lateral), ventral basis detail, ornamentation detail; 70 – 71, C. scabrosus; 70, aedeagus (lateral); 71, parameres (dorsal); 72 – 73, C. martinezi; 72, female terminalia and internal associated region (ventral); 73, female external genitalia; 74, C. parvulus female external genitalia; 75, C. scabrosus female external genitalia. acg, accessory gland; bcx, bursa copulatrix; dgx, gonocoxite distal piece; gst, gonostylus; ovd, oviduct; pgx, gonocoxite proximal piece; ppt, paraproct; sga, spiculum gastrale caudal piece; sgc, spiculum gastrale caudal piece; spt, spermatecae; stg, spermatecae gland; ut 9, urosternite; tem, temones." httpUri="https://zenodo.org/record/213490/files/figure.png" pageId="1" pageNumber="44">Fig. 72</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C9DCE7D1FF98547EFF75F826FAB0EA50" lastPageId="2" lastPageNumber="45" pageId="1" pageNumber="44" type="discussion">
|
||
<paragraph id="8179B45AFF98547EFF75F826FD54EB30" blockId="1.[151,1437,581,2013]" lastBlockId="2.[151,1437,151,860]" lastPageId="2" lastPageNumber="45" pageId="1" pageNumber="44">
|
||
<emphasis id="B3B26848FF98547DFF75F826FEA0EEB4" bold="true" box="[199,325,1951,1976]" pageId="1" pageNumber="44">Discussion</emphasis>
|
||
. Even with some recently published works the differentiation between
|
||
<taxonomicName id="46C6CFD9FF98547DFB0BF819FA81EEB4" box="[1209,1380,1952,1976]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="44" phylum="Arthropoda" rank="tribe" tribe="Macrodactylini">Macrodactylini</taxonomicName>
|
||
and
|
||
<taxonomicName id="46C6CFD9FF98547DFF25F87DFEC3EED0" box="[151,294,1988,2012]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="44" phylum="Arthropoda" rank="tribe" tribe="Tanyproctini">Tanyproctini</taxonomicName>
|
||
is difficult.
|
||
<taxonomicName id="46C6CFD9FF98547DFE1FF87DFDD9EED0" box="[429,572,1988,2012]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="44" phylum="Arthropoda" rank="tribe" tribe="Tanyproctini">Tanyproctini</taxonomicName>
|
||
were recognized by their strong sexual dimorphism, male antennal club with 3–7 antennomeres, reduced mouthparts, epistomal suture well defined or replaced by angular border, intersegmental membrane VII–VIII well developed ventrally.
|
||
<taxonomicName id="46C6CFD9FF9B547EFCD9FF05FBF3E9D8" box="[875,1046,188,212]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="tribe" tribe="Macrodactylini">Macrodactylini</taxonomicName>
|
||
were recognized by their poorly defined epistomal suture, contiguous metatibial spurs not separated by tarsal insertion, propygidium and ventrite V not separated by a suture, and ventrite V evidently longer than preceding ventrites (
|
||
<bibRefCitation id="E557C9ABFF9B547EFBDFFEBDFF37E833" author="Sanmartin & Martin-Piera" firstAuthor="Sanmartin" pageId="2" pageNumber="45" pagination="2 - 46" refId="ref6567" refString="Sanmartin, I. & Martin-Piera, F. (2000) First phylogenetic analysis of the subfamily Pachydeminae (Coleoptera, Scarabaeoidea, Melolonthidae): the Palearctic Pachydeminae. Journal of Zoological Systematics and Evolutionary Research, 41, 2 - 46." type="journal article" year="2000">Sanmartín & Martín-Piera 2000</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="E557C9ABFF9B547EFF6FFE9EFE90E833" author="Lacroix" box="[221,373,295,319]" firstAuthor="Lacroix" pageId="2" pageNumber="45" refId="ref6468" refString="Lacroix, M. (2007) Pachydeminae du Monde (Scarabaeoidea, Melolonthidae). Genera et Catalogue Commente. Marc Lacroix, Paris, 450 pp., 264 plates." type="book" year="2007">Lacroix 2007</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="E557C9ABFF9B547EFE33FE9EFDCEE833" author="Katovich" box="[385,555,295,319]" firstAuthor="Katovich" pageId="2" pageNumber="45" pagination="1 - 78" refId="ref6233" refString="Katovich, K. (2008) A generic-level phylogenetic review of the Macrodactylini (Coleoptera: Scarabaeidae: Melolonthinae). Insecta Mundi, 23, 1 - 78." type="journal article" year="2008">Katovich 2008</bibRefCitation>
|
||
). Neotropical
|
||
<taxonomicName id="46C6CFD9FF9B547EFD7BFE9EFCBDE833" box="[713,856,295,319]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="tribe" tribe="Tanyproctini">Tanyproctini</taxonomicName>
|
||
occur predominantly in temperate regions (
|
||
<collectingCountry id="F9D1F4CAFF9B547EFAF3FE9EFA7DE833" box="[1345,1432,295,319]" name="Bolivia" pageId="2" pageNumber="45">Bolivia</collectingCountry>
|
||
,
|
||
<collectingCountry id="F9D1F4CAFF9B547EFF25FEF4FF28E868" box="[151,205,333,356]" name="Peru" pageId="2" pageNumber="45">Peru</collectingCountry>
|
||
,
|
||
<collectingCountry id="F9D1F4CAFF9B547EFF68FEF4FED8E869" box="[218,317,333,357]" name="Uruguay" pageId="2" pageNumber="45">Uruguay</collectingCountry>
|
||
,
|
||
<collectingCountry id="F9D1F4CAFF9B547EFEF8FEF5FE59E868" box="[330,444,332,356]" name="Argentina" pageId="2" pageNumber="45">Argentina</collectingCountry>
|
||
,
|
||
<collectingCountry id="F9D1F4CAFF9B547EFE7BFEF5FDE2E868" box="[457,519,332,356]" name="Chile" pageId="2" pageNumber="45">Chile</collectingCountry>
|
||
) and
|
||
<taxonomicName id="46C6CFD9FF9B547EFDFAFEF5FD16E868" box="[584,755,332,356]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="tribe" tribe="Macrodactylini">Macrodactylini</taxonomicName>
|
||
species occur predominantly in tropical areas, especially in
|
||
<collectingCountry id="F9D1F4CAFF9B547EFF25FED6FF3EE88B" box="[151,219,367,391]" name="Brazil" pageId="2" pageNumber="45">Brazil</collectingCountry>
|
||
(
|
||
<bibRefCitation id="E557C9ABFF9B547EFF59FED6FDBDE88B" author="Sanmartin & Martin-Piera" box="[235,600,367,391]" firstAuthor="Sanmartin" pageId="2" pageNumber="45" pagination="2 - 46" refId="ref6567" refString="Sanmartin, I. & Martin-Piera, F. (2000) First phylogenetic analysis of the subfamily Pachydeminae (Coleoptera, Scarabaeoidea, Melolonthidae): the Palearctic Pachydeminae. Journal of Zoological Systematics and Evolutionary Research, 41, 2 - 46." type="journal article" year="2000">Sanmartín & Martín-Piera 2000</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="E557C9ABFF9B547EFDD7FED6FCF5E88B" author="Katovich" box="[613,784,367,391]" firstAuthor="Katovich" pageId="2" pageNumber="45" pagination="1 - 78" refId="ref6233" refString="Katovich, K. (2008) A generic-level phylogenetic review of the Macrodactylini (Coleoptera: Scarabaeidae: Melolonthinae). Insecta Mundi, 23, 1 - 78." type="journal article" year="2008">Katovich 2008</bibRefCitation>
|
||
).
|
||
<taxonomicName id="46C6CFD9FF9B547EFC96FEC9FC38E884" box="[804,989,368,392]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B3B26848FF9B547EFC96FEC9FC38E884" box="[804,989,368,392]" italics="true" pageId="2" pageNumber="45">Compsodactylus</emphasis>
|
||
</taxonomicName>
|
||
share some features with
|
||
<taxonomicName id="46C6CFD9FF9B547EFAB4FED6FA7DE88B" box="[1286,1432,367,391]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="tribe" tribe="Tanyproctini">Tanyproctini</taxonomicName>
|
||
, like the presence of intersegmental membrane VII–VIII, the structure of aedeagus with well-developed phallobasic apodeme and long and symmetrical parameres, but these characteristics are also found in some
|
||
<taxonomicName id="46C6CFD9FF9B547EFB58FE0EFA7CE8C3" box="[1258,1433,439,463]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="tribe" tribe="Macrodactylini">Macrodactylini</taxonomicName>
|
||
. The poorly-developed epistomal suture and enlarged ventrite V are features more commonly present in
|
||
<taxonomicName id="46C6CFD9FF9B547EFF25FE46FEA3EB1B" box="[151,326,511,535]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="tribe" tribe="Macrodactylini">Macrodactylini</taxonomicName>
|
||
. Cladistic studies on
|
||
<taxonomicName id="46C6CFD9FF9B547EFD8AFE46FD06EB1B" box="[568,739,511,535]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="tribe" tribe="Macrodactylini">Macrodactylini</taxonomicName>
|
||
and
|
||
<taxonomicName id="46C6CFD9FF9B547EFCAEFE46FC4FEB1B" box="[796,938,511,535]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="tribe" tribe="Tanyproctini">Tanyproctini</taxonomicName>
|
||
are necessary and certainly will change the present scenario and composition of both tribes.
|
||
</paragraph>
|
||
<paragraph id="8179B45AFF9B547EFF75FDFEFAB0EA50" blockId="2.[151,1437,151,860]" pageId="2" pageNumber="45">
|
||
The transfer of
|
||
<taxonomicName id="46C6CFD9FF9B547EFE32FDF1FDEBEB6C" box="[384,526,584,608]" class="Insecta" family="Melolonthidae" genus="Dicrania" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="martinezi">
|
||
<emphasis id="B3B26848FF9B547EFE32FDF1FDEBEB6C" box="[384,526,584,608]" italics="true" pageId="2" pageNumber="45">C. martinezi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="46C6CFD9FF9B547EFDFEFDF1FD37EB6C" box="[588,722,584,608]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="parvulus">
|
||
<emphasis id="B3B26848FF9B547EFDFEFDF1FD37EB6C" box="[588,722,584,608]" italics="true" pageId="2" pageNumber="45">C. parvulus</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="46C6CFD9FF9B547EFCACFDF0FC66EB6C" box="[798,899,585,608]" class="Insecta" family="Melolonthidae" genus="Dicrania" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B3B26848FF9B547EFCACFDF0FC66EB6C" box="[798,899,585,608]" italics="true" pageId="2" pageNumber="45">Dicrania</emphasis>
|
||
</taxonomicName>
|
||
to
|
||
<taxonomicName id="46C6CFD9FF9B547EFC1DFDF1FB8DEB6C" box="[943,1128,584,608]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B3B26848FF9B547EFC1DFDF1FB8DEB6C" box="[943,1128,584,608]" italics="true" pageId="2" pageNumber="45">Compsodactylus</emphasis>
|
||
</taxonomicName>
|
||
is based on the following characters (in opposition to
|
||
<taxonomicName id="46C6CFD9FF9B547EFE7CFDD4FDD6EB88" box="[462,563,621,644]" class="Insecta" family="Melolonthidae" genus="Dicrania" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B3B26848FF9B547EFE7CFDD4FDD6EB88" box="[462,563,621,644]" italics="true" pageId="2" pageNumber="45">Dicrania</emphasis>
|
||
</taxonomicName>
|
||
): frons with denticle-like setae (thin or scale-like setae); pronotum posteriorly bordered (not bordered); protibia with a spur (without spur); scutellar anteriorly rounded (angled); elytron anteriorly rounded (angled); elytron posteriorly bordered (without defined border); external elytral margin without membrane (with membrane); female elytra covering pygidial base (exposing the pygidium); mesofemur internal side longitudinally carinate (without carina); metacoxae weakly enlarged over trochanter basis (without enlargement over trochanter); abdominal spiracular area VII (
|
||
<figureCitation id="19FDA8DFFF9B547EFCF6FCA6FC70EA3B" box="[836,917,799,823]" captionStart="FIGURES 52 – 62. 52 – 60" captionStartId="11.[151,264,1811,1834]" captionTargetBox="[191,1382,200,1761]" captionTargetId="figure@11.[188,1382,193,1762]" captionTargetPageId="11" captionText="FIGURES 52 – 62. 52 – 60, Compsodactylus martinezi; 52 – 53, wing (distended, folded); 54, wing articulation and base; 55 – 56, first axillary (dorsal, ventral); 57, head of first axillary (anterior); 58 – 59, second axillary (dorsal, ventral); 60, third axillary (dorsal); 61, C. scabrosus wing distended; 62, C. martinezi posterior view of elytral medial contact section. Ax 1, 2, 3, first, second, third axillary; HP, humeral plate; med, medial plate (with two pieces). Wing base and venal: prefix B, basivenalia; suffix A, anterior; suffix P, posterior; b, BMP-BCu brace; PC, precosta; C, costa; R, radius; M, media; Cu, cubitus; A anal; J, jugal. Wing fold (red): af, anal fold; jf, jugal fold; lf, longitudinal fold; mf, medial fold; tf, transversal fold. Other abbreviations on the last plate." httpUri="https://zenodo.org/record/213489/files/figure.png" pageId="2" pageNumber="45">Fig. 53</figureCitation>
|
||
) narrowed (unmodified); ventrite IX large and shield-like (reduced a marginal sclerite); lateral area of parameres with a longitudinal line (with depression).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |